Latest Articles from ZooKeys Latest 100 Articles from ZooKeys https://zookeys.pensoft.net/ Fri, 29 Mar 2024 11:58:53 +0200 Pensoft FeedCreator https://zookeys.pensoft.net/i/logo.jpg Latest Articles from ZooKeys https://zookeys.pensoft.net/ A crane fly of the genus Gynoplistia Macquart (Diptera, Limoniidae) from the early Miocene of New Zealand https://zookeys.pensoft.net/article/115536/ ZooKeys 1192: 103-110

DOI: 10.3897/zookeys.1192.115536

Authors: André Nel, Uwe Kaulfuss

Abstract: The first fossil limoniid fly from the Miocene Fossil-Lagerstätte of Foulden Maar in New Zealand is described on the basis of an isolated well-preserved wing. The specimen is tentatively attributed to a new species Gynoplistia fouldensensis sp. nov. in the large extant genus Gynoplistia, which is well diversified in the country. It is the second fossil record of this genus, the first one being an isolated wing from the Cretaceous Weald Clay Formation in the United Kingdom.

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Research Article Tue, 20 Feb 2024 18:08:27 +0200
A revised, annotated checklist of Mexican non-biting midges (Diptera, Chironomidae) https://zookeys.pensoft.net/article/117223/ ZooKeys 1191: 237-286

DOI: 10.3897/zookeys.1191.117223

Authors: Orestes C. Bello-González, Trond Andersen, Norman Mercado-Silva

Abstract: An updated checklist of Mexican non-biting midges (Chironomidae) is presented. A total of 110 species of Chironomidae are known for Mexico: 52 species in 25 genera belong to the subfamily Chironominae, 30 species in 13 genera to Orthocladiinae, 21 species in nine genera to Tanypodinae, five species in two genera to Telmatogetoninae, and two species in one genus to Diamesinae. In addition, 41 genera without identified species are listed. The highest number of species (29) is recorded from the state of Campeche, while 19 species have been found in Veracruz and 15 in Nuevo León. Few or no records exist for states in Central and Northern Mexico, or those on the Pacific coast. The type localities for 34 species are in Mexico; of these, 27 species (25% of the total number of species recorded in the country) are endemic. Twenty-nine species recorded in Mexico have a Neotropical distribution, 15 a Nearctic distribution, and 39 species are distributed in both the Neotropical and Nearctic regions or more widely. It has been suggested that as many as 1000 species might occur in Mexico; so only a little more than 10% of the expected diversity has so far been recorded.

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Checklist Wed, 14 Feb 2024 10:14:12 +0200
Three new species and two newly recorded species of Tachininae from Tibet, China (Arthropoda, Insecta, Diptera, Tachinidae) https://zookeys.pensoft.net/article/105549/ ZooKeys 1191: 215-235

DOI: 10.3897/zookeys.1191.105549

Authors: Ruiqing Dong, Junjian Li, Hui Yang, Chuntian Zhang

Abstract: During our studying of the fauna of Tibet, China, many specimens of the subfamily Tachininae (Diptera, Tachinidae) were collected and examined. Three species are described here as new to science, Leskia latisurstyla sp. nov., Trichoformosomyia cuonaensis sp. nov., and Tachina jilongensis sp. nov., and two species, Nemoraea javana (Brauer & Bergenstamm, 1894) and N. echinata Mesnil, 1953, are newly recorded from Tibet. In addition to their descriptions, illustrations, and diagnoses, three identification keys are provided. The specimens in this study are kept in the Insect Collection of Shenyang Normal University, China (SYNU).

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Research Article Tue, 13 Feb 2024 16:39:52 +0200
The genus Neurigona Rondani, 1856 (Diptera, Dolichopodidae) from Yunnan, China, with descriptions of seven new species and a key to the species of China https://zookeys.pensoft.net/article/109222/ ZooKeys 1190: 319-345

DOI: 10.3897/zookeys.1190.109222

Authors: Chen Lin, Mengqing Wang, Ding Yang

Abstract: Previously, only three species of the genus Neurigona Rondani of the subfamily Neurigoninae were known from Yunnan Province. Here, we reviewed the species of Neurigona from Yunnan and added the following seven new species: N. apicilata sp. nov., N. basicurva sp. nov., N. brevidigitata sp. nov., N. convexa sp. nov., N. huanglianshana sp. nov., N. quadrimaculata sp. nov., and N. ventriprocessa sp. nov. All seven new species are sympatric and were collected from below a reservoir in the Huanglianshan Nature Reserve in Yunnan using three Malaise traps in 2019. This suggests a very high species richness in the Yunnan fauna. A key to the species of Neurigona from Chinese mainland is provided.

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Research Article Wed, 31 Jan 2024 14:09:41 +0200
Four new species of Zeugodacus Hendel (Diptera, Tephritidae, Dacinae, Dacini) and new records of dacines from India https://zookeys.pensoft.net/article/114031/ ZooKeys 1188: 1-26

DOI: 10.3897/zookeys.1188.114031

Authors: Karamankodu Jacob David, Venkateshaiah Abhishek, Ningthoujam Kennedy, K. M. Ajaykumara, R. G. Gracy, Cheday Bhutia Hissay

Abstract: Four new species of Zeugodacus Hendel are described from India viz., Zeugodacus momordicae David & Ajaykumara, sp. nov. from Arunachal Pradesh infesting male flower buds of Momordica dioica, Zeugodacus nasivittatus David & Abhishek, sp. nov. from Meghalaya, Zeugodacus (Sinodacus) sinuvittatus David & Abhishek, sp. nov. from Himachal Pradesh and Zeugodacus (Zeugodacus) umiam David & Kennedy, sp. nov. from Meghalaya. An illustrated key to all species of Zeugodacus from India is also included. Bactrocera (Parazeugodacus) abbreviata (Hardy) and Dacus (Mellesis) vijaysegarani Drew & Hancock are recorded for the first time from India.

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Research Article Wed, 3 Jan 2024 10:43:45 +0200
A conspectus of Australian Apotropina (Diptera, Chloropidae) with the description of two new species https://zookeys.pensoft.net/article/108497/ ZooKeys 1187: 261-299

DOI: 10.3897/zookeys.1187.108497

Authors: Yuchen Ang, James Lumbers, Paula R. Riccardi

Abstract: The genus Apotropina (Diptera, Chloropidae) has a global distribution with more than 80 valid described species, of which 22 are known to occur in Australia. The Australian Apotropina fauna is poorly studied, with many species known from single type specimens, more with the morphology of the other sex unknown, and there have been no new species descriptions since 1959. Here, we describe two new species from Australia, A. maculigena Riccardi, sp. nov. and A. popeye Ang, sp. nov., and provide an updated illustrated key. We also provide a conspectus of the known Australian Apotropina with images of types and collate all original descriptions and subsequent taxonomic notes of relevance as supplementary information. Finally, we discuss the validity of two known syntype specimens of A. bispinosa due to incongruencies with the species description.

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Research Article Thu, 21 Dec 2023 19:55:33 +0200
A review of the subgenus Loxocera Meigen, 1803 (Diptera, Brachycera, Psilidae) in China https://zookeys.pensoft.net/article/108876/ ZooKeys 1186: 71-96

DOI: 10.3897/zookeys.1186.108876

Authors: Jiale Zhou, Ding Yang

Abstract: The subgenus Loxocera (Loxocera Meigen, 1803) (Diptera, Brachycera, Psilidae) in China is reviewed. Six species are recognized, including four new species: L. (L.) chikuni sp. nov., L. (L.) lonsdalei sp. nov., L. (L.) maculithorax sp. nov., and L. (L.) obscura sp. nov. Two species originally placed in the subgenus Loxocera, L. anulata Wang & Yang, 1998 and L. tianmuensis Wang & Yang, 1998, are transferred to the subgenus Loxocera (Imantimyia Frey, 1925). A key to the species of the subgenus Loxocera occurring in China is provided.

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Research Article Mon, 11 Dec 2023 20:13:05 +0200
A new species of the genus Conosiphon Becker, 1923 and the first records of this genus for Europe (Diptera, Asilidae) https://zookeys.pensoft.net/article/105663/ ZooKeys 1181: 59-79

DOI: 10.3897/zookeys.1181.105663

Authors: Reinoud van den Broek, Piluca Álvarez Fidalgo, John Smit

Abstract: A new species of Conosiphon Becker, 1923, Conosiphon ianus Álvarez Fidalgo & van den Broek, sp. nov., is described from Spain, representing the first record of this genus for Europe. It is illustrated in high-resolution photographs and the first ecological information is provided, as well as a key to all species tentatively placed in this genus.

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Research Article Mon, 2 Oct 2023 10:59:50 +0300
Exploring the relationship between bats (Mammalia, Chiroptera) and ectoparasitic flies (Diptera, Hippoboscoidea) of the Orinoquia Region in South America https://zookeys.pensoft.net/article/103479/ ZooKeys 1179: 1-34

DOI: 10.3897/zookeys.1179.103479

Authors: Erika M. Ospina-Pérez, Fredy A. Rivera-Páez, Héctor E. Ramírez-Chaves

Abstract: Bat flies (Nycteribiidae and Streblidae) have been used to study co-evolutionary patterns between ectoparasites and bats. In the world, Nycteribiidae and Streblidae are represented by approximately 276 and 237 species, respectively. In regions such as the Orinoquia located in the north of South America (Colombia and Venezuela), the richness of bats is high (more than 100 documented species), but studies on Nycteribiidae and Streblidae are scarce and discontinuous. To contribute to the knowledge of ectoparasitic flies in the Orinoquia, records of flies and their interactions with bats were reviewed, including new records and associations using interaction networks. We documented 124 species of Streblidae and only 12 of Nycteribiidae for the Orinoquia in approximately 102 bat species reported in Colombia and Venezuela. New records for six species of bat flies in Colombia were found (Mastoptera guimaraesi, Noctiliostrebla maai, Paradyschiria parvuloides, Trichobius jubatus, Trichobius parasiticus, and Basilia ferrisi) associated with six species of bats (Cynomops planirostris, Desmodus rotundus, Myotis handleyi, Molossus rufus, Noctilio albiventris, and Phyllostomus hastatus). The bat-ectoparasite interaction networks in the Orinoquia revealed a pattern of antagonistic relationships, with high specialization, modularity, and low connectivity and nesting. The identified networks are between bat fly species belonging to different ecomorphological groups with unique host species. This supports the idea of ecological niche partitioning among ectoparasitic bat flies and hosts. Our study expanded the knowledge of the distribution of some fly species and the associations with bat hosts in Colombia, by presenting morphological descriptions and new observations, which are key to understanding the ecology, diversity, and distribution of these species.

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Research Article Fri, 8 Sep 2023 10:56:34 +0300
Description of the female of Atherimorpha latipennis Stuckenberg (Diptera, Rhagionidae): the first record of brachyptery in Rhagionidae https://zookeys.pensoft.net/article/107357/ ZooKeys 1178: 265-277

DOI: 10.3897/zookeys.1178.107357

Authors: John M. Midgley, Burgert S. Muller

Abstract: The genus Atherimorpha White, 1915 is a Gondwanan relic, occurring in South America, Southern Africa and Australia. Females are rarely collected, and are not described for more than half of the known species. The female of Atherimorpha latipennis Stuckenberg, 1956 was collected for the first time in 2021 and is described here, along with a redescription of the male. We describe the differences from the male, with the reduced wings and poorly defined scutellum the most noteworthy. The female of A. latipennis represents the first recorded case of brachyptery in the family Rhagionidae. Possible drivers of brachyptery in Afrotropical Diptera are briefly discussed.

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Research Article Thu, 7 Sep 2023 12:49:08 +0300
New species and records of the genus Hybos Meigen (Diptera, Empidoidea, Hybotinae) from Wuyishan National Park, China https://zookeys.pensoft.net/article/105952/ ZooKeys 1172: 313-351

DOI: 10.3897/zookeys.1172.105952

Authors: Meilin Li, Ding Yang

Abstract: Wuyishan National Park is well known for its rich biodiversity. Previously, only five species of Hybos Meigen, 1803 were known to occur in this region. In this study, 27 species of the genus Hybos from Wuyishan National Park are reviewed based on comparative morphological characteristics. Among these, nine species are described as new to science: Hybos brevidigitatus sp. nov., Hybos constractus sp. nov., Hybos curvitibia sp. nov., Hybos dazhulanus sp. nov., Hybos fujianensis sp. nov., Hybos leucopogus sp. nov., Hybos longidigitatus sp. nov., Hybos modificatus sp. nov., and Hybos wuyishanus sp. nov. Diagnoses, detailed descriptions, remarks, colored illustrations, distributions, and some of the female terminalia characteristics are provided for nine new species. In addition, 13 species of this genus are reported for the first time in Wuyishan National Park. A key to Hybos species from Wuyishan National Park is also given.

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Research Article Thu, 27 Jul 2023 13:04:05 +0300
Nycteribiid bat flies (Arthropoda, Insecta, Diptera, Nycteribiidae) of Kenya https://zookeys.pensoft.net/article/102800/ ZooKeys 1169: 65-85

DOI: 10.3897/zookeys.1169.102800

Authors: Carl W. Dick, Taylor B. Verrett, Paul W. Webala, Bruce D. Patterson

Abstract: Bat flies (Diptera: Nycteribiidae and Streblidae) are hematophagous ectoparasites of bats characterized by viviparous pupiparity and generally high host specificity. Nycteribiid bat flies are wingless, morphologically constrained, and are most diverse in the Eastern Hemisphere. Africa hosts approximately 22% of global bat biodiversity and nearly one-third of all African bat species occur in Kenya, one of Africa’s most bat-rich countries. However, records of nycteribiid bat fly diversity in Kenya remain sparse and unconsolidated. This paper combines all past species records of nycteribiid bat flies with records from a survey of 4,255 Kenyan bats across 157 localities between 2006 and 2015. A total of seven nycteribiid genera and 17 species are recorded, with seven species from the recent ‘Bats of Kenya’ surveys representing previously undocumented country records. Host associations and geographic distributions based on all available records are also described. This comprehensive species catalog addresses and further emphasizes the need for similar investigations of nycteribiid biodiversity across Africa.

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Checklist Thu, 13 Jul 2023 19:32:54 +0300
A review of the Strongylophthalmyia coarctata subgroup (Diptera, Brachycera, Strongylophthalmyiidae) from China, with the descriptions of three new species https://zookeys.pensoft.net/article/104699/ ZooKeys 1168: 329-353

DOI: 10.3897/zookeys.1168.104699

Authors: Jiale Zhou, Neal L. Evenhuis, Ding Yang

Abstract: The species belonging to the Strongylophthalmyia coarctata subgroup of the S. punctata group (Diptera: Brachycera: Strongylophthalmyiidae) from China are reviewed. Six species are recognized, including three new species: S. corniculata sp. nov., S. flagellicornis sp. nov., and S. tangwangana sp. nov. Strongylophthalmyia narwhal Evenhuis, 2020 and S. raricornis Shatalkin, 1981 are recorded from China for the first time, and S. raricornis is also recorded from South Korea for the first time. An identification key to the Asian species of the S. coarctata subgroup is provided.

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Research Article Tue, 4 Jul 2023 09:49:38 +0300
Two new species of the Phaonia boleticola-group (Diptera, Muscidae, Phaoniinae) from China https://zookeys.pensoft.net/article/97845/ ZooKeys 1168: 219-229

DOI: 10.3897/zookeys.1168.97845

Authors: Zhongyan Zhou, Lianmeng Wei

Abstract: Two new species of Phaonia are described: Phaonia agitata Zhou & Wei, sp. nov. and Phaonia nujiangensis Zhou & Wei, sp. nov., which were collected from Guizhou and Yunnan provinces of southwestern China and are assigned to the boleticola-group. A key to the species of this group is provided. The type specimens are deposited in the Wei Lianmeng Model Worker Innovation Studio, Anshun, Guizhou, China (WLMWISAGC).

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Research Article Mon, 3 Jul 2023 09:42:06 +0300
First records of the crane fly genus Helius Lepeletier & Serville (Diptera, Limoniidae) in Guangxi, China with description of one new species https://zookeys.pensoft.net/article/104106/ ZooKeys 1168: 131-149

DOI: 10.3897/zookeys.1168.104106

Authors: Yuanyuan Xu, Shenglin Zhang, Guoquan Wang, Ding Yang, Xiao Zhang

Abstract: The genus Helius Lepeletier & Serville, 1828 is recorded in Guangxi, China for the first time with the following three species belonging to the subgenus H. (Helius): H. (H.) damingshanus sp. nov., H. (H.) nipponensis (Alexander, 1913) and H. (H.) stenorhynchus stenorhynchus Alexander, 1954. Among them, H. (H.) stenorhynchus stenorhynchus is also a new record in China. Their descriptions and illustrations, as well as the first key to Chinese H. (Helius) crane flies, are presented.

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Research Article Tue, 27 Jun 2023 16:13:41 +0300
An updated catalogue of biting midges of the genus Culicoides Latreille, 1809 (Diptera, Ceratopogonidae) of Mexico and their known distribution by state https://zookeys.pensoft.net/article/102858/ ZooKeys 1167: 1-47

DOI: 10.3897/zookeys.1167.102858

Authors: Alejandro Mendez-Andrade, Sergio Ibáñez-Bernal

Abstract: An updated catalogue of Culicoides of Mexico is presented. It includes 86 species with their regional distribution and corresponding record references, known immature stages and associated pathogens. In addition, a taxonomic key for subgenera and species groups for Mexico is presented and an index of species by state is included.

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Catalogue Wed, 14 Jun 2023 10:57:36 +0300
Two new Ptychoptera Meigen, 1803 (Diptera, Ptychopteridae) from the Western Palaearctic https://zookeys.pensoft.net/article/96193/ ZooKeys 1166: 91-102

DOI: 10.3897/zookeys.1166.96193

Authors: Libor Dvořák, Katarína Fogašová, Jozef Oboňa, Edina Török, Peter Manko

Abstract: Ptychoptera xanthopleura Dvořák, Oboňa & Manko, sp. nov. from Azerbaijan and Georgia, and Ptychoptera staryi Dvořák, Oboňa & Manko, sp. nov. from Bulgaria are described. P. xanthopleura sp. nov. differs from the other member of the lacustris group mainly by having almost completely yellow pleurae, and by the shape of the epandrium and gonocoxites. The diagnostics of P. staryi sp. nov. and P. incognita Török, Kolcsár & Keresztes, 2015 based on male genitalia are provided.

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Research Article Tue, 6 Jun 2023 17:35:29 +0300
Six new species of Diostracus Loew (Diptera, Dolichopodidae) from Tibet https://zookeys.pensoft.net/article/101533/ ZooKeys 1163: 1-46

DOI: 10.3897/zookeys.1163.101533

Authors: Yajun Zhu, Chufei Tang, Ding Yang

Abstract: Six species of Diostracus from Tibet are described as new to science: D. concavus sp. nov., D. fasciculatus sp. nov., D. laetus sp. nov., D. polytrichus sp. nov., D. strenus sp. nov., and D. translucidus sp. nov. A key to the species from Tibet of the genus is provided. The distribution of the genus in Tibet is also discussed.

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Research Article Wed, 17 May 2023 08:55:49 +0300
Diversity of flesh flies (Sarcophagidae, Sarcophaginae) of pond habitats in rural areas in the Croatian part of Baranja https://zookeys.pensoft.net/article/100878/ ZooKeys 1159: 17-36

DOI: 10.3897/zookeys.1159.100878

Authors: Stjepan Krčmar

Abstract: The diversity of grey flesh flies (Sarcophagidae: Sarcophaginae) from the Croatian part of Baranja was studied during 2019 to 2021, resulting in 37 species, of which the following are new for the area: Ravinia pernix (Harris, 1780); Sarcophaga (Het.) depressifrons Zetterstedt, 1845; S. (Het.) filia Rondani, 1860; S. (Het.) haemorrhoides Böttcher, 1913; S. (Het.) pumila Meigen, 1826; S. (Het.) vagans Meigen, 1826; S. (Lis.) dux Thomson, 1869; S. (Lis.) tuberosa Pandellé, 1896; S. (Meh.) sexpunctata (Fabricius, 1805); S. (Pan.) protuberans Pandellé, 1896; S. (Sar.) carnaria (Linnaeus, 1758); S. (Sar.) variegata (Scopoli, 1763), and S. (Pse.) spinosa Villeneuve, 1912. New locality records are provided for 25 species. Sarcophaga (Sar.) croatica Baranov, 1941 was the most abundant with 37%, followed by S. (Sar.) lehmanni Müller, 1922 (21%), and S. (Pas.) albiceps Meigen, 1826 (5%), making up 63% of all collected specimens. Most species (35) were collected in locality of Zmajevac, while the fewest (3) were collected in Bilje locality. During this study, S. (Pse.) spinosa was recorded in Croatia for the first time. Combined with previous records, 42 species of flesh flies have been recorded from Croatian Baranja, which comprise 27% of the flesh flies known to occur in Croatia. The total number of species of the family Sarcophagidae currently known in Croatia has increased to 156.

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Research Article Mon, 24 Apr 2023 18:30:18 +0300
Description of two new species of Dicranomyia (Erostrata) crane fly (Diptera, Limoniidae) from Korea, with remarks on DNA barcoding and updated taxonomic key https://zookeys.pensoft.net/article/90792/ ZooKeys 1157: 193-206

DOI: 10.3897/zookeys.1157.90792

Authors: Jisoo Kim, Yeon Jae Bae

Abstract: Two new crane fly species, Dicranomyia (Erostrata) jejuensis sp. nov. and D. (E.) koreana sp. nov., from Korea are described on the basis of morphology and mitochondrial COI sequences. DNA barcode sequences for other four D. (Erostrata) species from Korea are also provided for the first time. The identification key for all known D. (Erostrata) species is presented.

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Research Article Tue, 11 Apr 2023 14:21:18 +0300
New records for the Western Balkans cranefly fauna (Diptera, Tipuloidea) with the description of a new Baeoura Alexander (Diptera, Limoniidae) https://zookeys.pensoft.net/article/98997/ ZooKeys 1157: 1-42

DOI: 10.3897/zookeys.1157.98997

Authors: Levente-Péter Kolcsár, Micha Camiel d’Oliveira, Wolfram Graf, Clovis Quindroit, Kozo Watanabe, Marija Ivković

Abstract: The cranefly (Tipuloidea) fauna of the Western Balkans is still poorly known. In this study, occurrence data of 77 species is reported, of which two species are newly recorded for Albania, eight species for Bosnia and Herzegovina, twelve for Croatia, and seven for Slovenia, respectively. A new species, Baeoura neretvaensis Kolcsár & d’Oliveira, sp. nov. is described from Bosnia and Herzegovina, Montenegro, and Slovenia. Images of the habitus, wing, and male and female terminalia of the new species are provided. Furthermore, images of male terminalia and wings of thirteen additional species are presented.

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Research Article Fri, 31 Mar 2023 18:46:38 +0300
New species and records of the genus Antocha Osten Sacken (Diptera, Limoniidae) from Tibet, China with a key to species in Qinghai-Tibet region https://zookeys.pensoft.net/article/86786/ ZooKeys 1156: 53-69

DOI: 10.3897/zookeys.1156.86786

Authors: Hanhuiying Lv, Juan Sun, Ning Wang, Ding Yang, Xiao Zhang

Abstract: Thirty-four known species and subspecies of the genus Antocha Osten Sacken, 1860 have been recorded from China, of which four occur in Tibet. Herein, two new Antocha species, A. (Antocha) curvativa sp. nov. and A. (A.) tibetana sp. nov., are described and illustrated from Tibet. The new species are distinguished from congeners mainly by their male genitalia. Antocha (A.) spiralis Alexander, 1932 and A. (A.) setigera Alexander, 1933, which are newly recorded in Tibet, are redescribed and illustrated. A key to Antocha species in the Qinghai-Tibet region of China is also presented.

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Research Article Fri, 24 Mar 2023 15:51:34 +0200
New species, a new combination, and DNA barcodes of Parachironomus Lenz, 1921 (Diptera, Chironomidae) https://zookeys.pensoft.net/article/98542/ ZooKeys 1153: 121-140

DOI: 10.3897/zookeys.1153.98542

Authors: Wen-Bin Liu, Ying Wang, Kang-Zhu Zhao, Cheng-Yan Wang, Jun-Yu Zhang, Chun-Cai Yan, Xiao-Long Lin

Abstract: The genus Parachironomus has a cosmopolitan distribution including 85 valid described species worldwide. Species records and studies of the genus in the Tibetan Plateau are scarce. In this study, the genus Parachironomus from China is revised and two new species, Parachironomus wangi Liu & Lin, sp. nov. and Parachironomus nankaiensis Liu & Lin, sp. nov., are described based on adult morphology and molecular data. Paracladopelma demissum Yan, Wang & Bu is placed in the genus Parachironomus as a new combination. A neighbor-joining tree was reconstructed based on all known Parachironomus COI DNA barcodes. A key to adult males of the genus Parachironomus from China is also provided.

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Research Article Thu, 16 Mar 2023 16:46:16 +0200
A revision of Centrioncus Speiser (Diptera, Diopsidae, Centrioncinae) with descriptions of new species from Angola, Burundi, and Kenya https://zookeys.pensoft.net/article/95619/ ZooKeys 1144: 1-93

DOI: 10.3897/zookeys.1144.95619

Authors: Hans R. Feijen, Cobi Feijen

Abstract: A diagnosis is presented for the Centrioncinae, the Afromontane Forest Flies or stalkless Diopsidae, while its taxonomic position within the Diopsidae is discussed. Arguments are presented for an eventual raising of the Centrioncinae to family level. The differential characters for its two genera, Centrioncus Speiser and Teloglabrus Feijen, are tabulated. The diagnosis for Centrioncus is updated and a key to the ten species now recognised (including three new species) is provided. Centrioncus crassifemur sp. nov. is described from a single female from Angola. This greatly extends the distribution range for the genus. Centrioncus bururiensis sp. nov. is described from Burundi, while Centrioncus copelandi sp. nov. originates from the Kasigau Massif of Kenya. Diagnoses, descriptive updates, illustrations and notes are presented for all other Centrioncus. Centrioncus aberrans Feijen, described from Uganda, is now also recorded for western Kenya, Rwanda, and possibly eastern DR Congo. This wide range of C. aberrans is unusual for the Centrioncinae species which have allopatric and usually very restricted distribution ranges. Defining characters of C. aberrans from the various regions were examined in detail, but only minor differences were found. Centrioncus decoronotus Feijen, described from Kenya, is now recorded for several other places in Kenya. A distribution map is given for the Eastern African Centrioncus species. The eastern branch of the Great Rift Valley appears to form a barrier between C. aberrans and C. decoronotus. The type species of the genus, C. prodiopsis Speiser from the Kilimanjaro in Tanzania, was only known from the 1905–1906 type series. After more than 100 years it has now been found again on the Kenya side of Kilimanjaro. Various differential characters of Centrioncus and Diopsidae are discussed, while brief discussions on sex ratio and fungal parasites are given. Centrioncus are known to occur on low shrubs and herbaceous plants in rain forests. Now, the possibility is indicated that they also might occur higher up in the tree canopies.

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Research Article Wed, 1 Feb 2023 19:04:32 +0200
New species and new records of Monohelea Kieffer (Diptera, Ceratopogonidae) from Brazil https://zookeys.pensoft.net/article/86680/ ZooKeys 1136: 175-185

DOI: 10.3897/zookeys.1136.86680

Authors: Maria Clara Alves Santarém, Erick Aragão Cardoso, Israel de Souza Pinto, Maria Luiza Felippe-Bauer

Abstract: Two new Brazilian species of Monohelea Kieffer are described and illustrated based on male specimens, Monohelea capixaba sp. nov. from Espírito Santo and Monohelea coimbrai sp. nov. from Rio de Janeiro. New records for M. aguirrei Tavares & Souza, M. archibaldoi Tavares & Souza and M. maculipennis (Coquillet) are given based on specimens from Espírito Santo (all three species) and Amapá (M. maculipennis only). All specimens are deposited in the Ceratopogonidae Collection of Fundação Oswaldo Cruz, Brazil.

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Research Article Mon, 19 Dec 2022 15:40:41 +0200
A new species of Chrysotus Meigen (Diptera, Dolichopodidae) from soybean fields in South Dakota, USA https://zookeys.pensoft.net/article/95026/ ZooKeys 1135: 171-180

DOI: 10.3897/zookeys.1135.95026

Authors: Justin B. Runyon, Eric Beckendorf, Louis S. Hesler

Abstract: A new long-legged fly species, Chrysotus soya sp. nov. (Diptera: Dolichopodidae), is described and illustrated from specimens collected in soybean fields near Brookings, South Dakota, USA. The abundance of this species in soybeans suggests it plays an important role as a beneficial predator.

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Research Article Wed, 14 Dec 2022 10:09:43 +0200
Diversity underfoot of agromyzids (Agromyzidae, Diptera) mining thalli of liverworts and hornworts https://zookeys.pensoft.net/article/94530/ ZooKeys 1133: 1-164

DOI: 10.3897/zookeys.1133.94530

Authors: Makoto Kato, Luna Yamamori, Yume Imada

Abstract: Agromyzidae is a dipteran family that has diversified as internal plant feeders. Although most agromyzid species feed on herbaceous angiosperms, only a limited number of species has been recorded as miners of bryophytes. Extensive searches and rearing of bryophytivores in the Japanese Archipelago were made, resulting in that thallus-mining agromyzids are overwhelmingly widespread and diverse on thalloid liverworts and hornworts. By examining the morphology of adult flies, it was revealed that the agromyzid fauna comprise 39 species, of which 37 species are newly described. All the species are assigned to the genus Phytoliriomyza Hendel based on some shared morphological character states as follows: costa reaching M1; orbital setulae minute and erect (rarely proclinate); male epandrium with combs of fused tubercle-like setae and/or hypertrophied arms bearing tubercle-like setae; male distiphallus comprising a pair of stout, extended tubules; female cercus with two stout, apical, trichoid sensilla. Of the 39 agromyzid species in Japan, 36 species are associated with liverworts: 5 spp. on Marchantia (Marchantiaceae), 2 spp. on Dumortiera (Dumortieraceae), 3 spp. on Plagiochasma, 1 sp. on Asterella, 6 spp. on Reboulia (Aytoniaceae), 1 sp. on Wiesnerella (Wiesnerellaceae), 15 spp. on Conocephalum (Conocephalaceae), and 3 spp. on Riccia (Ricciaceae). Three species are associated with hornworts: 1 sp. on Folioceros (Anthocerotaceae), 1 sp. on Megaceros (Dendrocerotaceae), and 1 sp. on Notothylas, Phaeoceros (Notothyladaceae), and Anthoceros (Anthocerotaceae). The results suggest that 37 of the 39 species are host-specific at least to plant genus level, and that the inter-specific differences in male genitalia and color patterns of scutum, antenna, and maxillary palpus have contributed to reproductive isolation on the bryophytes that the flies share.

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Monograph Wed, 30 Nov 2022 16:00:13 +0200
Japanese species of Ormosia Rondani (Diptera, Limoniidae): revision of the subgenera Oreophila Lackschewitz and Parormosia Alexander https://zookeys.pensoft.net/article/86022/ ZooKeys 1132: 127-162

DOI: 10.3897/zookeys.1132.86022

Authors: Daichi Kato, Kozo Watanabe, Levente-Péter Kolcsár

Abstract: Japanese species of the subgenera Oreophila Lackschewitz and Parormosia Alexander of the genus Ormosia Rondani (Limoniidae) are revised. Two new species Ormosia (Oreophila) komazawai Kato & Kolcsár, sp. nov. and Ormosia (Parormosia) phalara Kato & Kolcsár, sp. nov. are described. The identities of all Japanese species of the two subgenera are clarified and redescribed with images of habitus and wings, and drawings of male and female terminalia. The first DNA barcode sequences of the species Ormosia (Parormosia) diversipes Alexander and Ormosia (Parormosia) phalara Kato & Kolcsár, sp. nov. are also provided. A key to, and distribution maps of, the Japanese species are provided.

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Research Article Tue, 29 Nov 2022 15:54:14 +0200
Pullimosina (Pullimosina) turfosa sp. nov. and other Sphaeroceridae (Diptera) from peat bogs in the North Caucasus (Russia) https://zookeys.pensoft.net/article/94579/ ZooKeys 1132: 1-49

DOI: 10.3897/zookeys.1132.94579

Authors: Jindřich Roháček, Andrey A. Przhiboro

Abstract: The first data about Sphaeroceridae occurring on eight montane and foothill peat bogs of various types in the North Caucasus (Russia) are presented. A total of 38 species has been recorded and their affinity to peat-bog habitats is discussed. A single species is classified as a tyrphobiont, viz. the strongly brachypterous Pullimosina (Pullimosina) turfosa sp. nov. being strictly associated with Sphagnum hummocks in peat bogs. This new species is described and illustrated in detail and its relationships, biology, and wing reduction are discussed. Only three species are considered tyrphophilous or probably tyrphophilous, viz. Ischiolepta nitida (Duda, 1920), Phthitia (Kimosina) longisetosa (Dahl, 1909), and Spelobia ibrida Roháček, 1983. The majority of recorded species do not have close affinity to peat bogs and are treated as tyrphoneutral, and Rachispoda hostica (Villeneuve, 1917) is probably tyrphoxenous due to occasional occurrence in a peat bog. Species composition of Sphaeroceridae on Caucasian peat bogs is discussed in comparison to those known from peat bogs in other parts of Europe. Taxonomic notes are given on Minilimosina (Svarciella) species of the M. vitripennis group. Six species (including P. turfosa sp. nov.) are new additions to the fauna of Russia.

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Research Article Fri, 25 Nov 2022 17:15:32 +0200
A new species of Saropogon Loew, 1847 (Diptera, Asilidae) from Arizona, with a review of the Nearctic species north of Mexico https://zookeys.pensoft.net/article/81874/ ZooKeys 1130: 1-63

DOI: 10.3897/zookeys.1130.81874

Authors: Charlotte H. E. Alberts, Eric M. Fisher

Abstract: The Nearctic species of Saropogon Loew, 1847 north of Mexico are reviewed, with 19 species recognized and one described as new: Saropogon pyrodes sp. nov. from Arizona. This previously recognized new species has awaited description since its first collection in 1964. Only after a community scientist posted photographs taken in nature to an online database did its description become a priority. All species of Saropogon occurring in the Nearctic Region north of the Mexican border have been reexamined. Photographs and diagnoses of all species are provided with a distribution map of the included specimens studied. An updated key to the Nearctic species north of Mexico is provided. Finally, the need for a review of the diverse Mexican fauna is expressed.

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Research Article Thu, 17 Nov 2022 17:53:38 +0200
New faunistic records of the families Bolitophilidae and Keroplatidae (Diptera, Sciaroidea) from Morocco https://zookeys.pensoft.net/article/85536/ ZooKeys 1128: 47-52

DOI: 10.3897/zookeys.1128.85536

Authors: Ouafaa Driauach, Peter J. Chandler, Mohamed Amin El Mouden, Boutaïna Belqat

Abstract: The family Bolitophilidae is recorded for the first time from Morocco with one species Bolitophila (Bolitophila) saundersii (Curtis, 1836). Ten new species are added to the Moroccan fauna of Keroplatidae, known until now by only two species, raising the number of species currently known in Morocco to 12.

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Checklist Mon, 7 Nov 2022 17:21:16 +0200
Exceptional larval morphology of nine species of the Anastrepha mucronota species group (Diptera, Tephritidae) https://zookeys.pensoft.net/article/84628/ ZooKeys 1127: 155-215

DOI: 10.3897/zookeys.1127.84628

Authors: Erick J. Rodriguez, Gary J. Steck, Matthew R. Moore, Allen L. Norrbom, Jessica Diaz, Louis A. Somma, Raul Ruiz-Arce, Bruce D. Sutton, Norma Nolazco, Alies Muller, Marc A. Branham

Abstract: Anastrepha is the most diverse and economically important genus of Tephritidae in the American tropics and subtropics. The striking morphology of the third instars of Anastrepha caballeroi Norrbom, Anastrepha crebra Stone, Anastrepha haplacantha Norrbom & Korytkowski, Anastrepha korytkowskii Norrbom, Anastrepha nolazcoae Norrbom & Korytkowski, and three newly discovered and as yet formally unnamed species (Anastrepha sp. Peru-82, Anastrepha sp. nr. protuberans, and Anastrepha sp. Sur-16), and the more typical morphology of Anastrepha aphelocentema Stone, are described using light and scanning electron microscopy. To contribute to a better understanding of the interspecific and intraspecific variation among species in the mucronota species group and facilitate phylogenetic studies, we integrate molecular and morphological techniques to confirm the identity and describe third instars. Larva-adult associations and the identification of described larvae were confirmed using DNA barcodes. We provide diagnostic characters to distinguish larvae among these nine species of the mucronota group and separate them from those of the 29 other Anastrepha species previously described. We introduce the vertical comb-like processes on the oral margin as a novel character, and the unusual character states, including position and shape of the preoral lobe, and dentate or fringed posterior margins of the oral ridges and accessory plates. Our comparative morphology concurs with most previously inferred phylogenetic relationships within the mucronota group.

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Research Article Thu, 3 Nov 2022 19:09:01 +0200
First records of Trichina Meigen, Euthyneura Macquart and Oedalea Meigen (Diptera, Hybotidae) from North Africa, with descriptions of two new species https://zookeys.pensoft.net/article/90077/ ZooKeys 1124: 43-58

DOI: 10.3897/zookeys.1124.90077

Authors: Laila Zouhair, Patrick Grootaert, Kawtar Kettani

Abstract: Trichina Meigen, 1830, Euthyneura Macquart, 1836 and Oedalea Meigen, 1820 are reported for the first time in North Africa from northern Morocco, with seven species including two ones new to science, based on material collected at nine sites located in the Moroccan sector of the Mediterranean Intercontinental Biosphere reserve (MIBR). These new records represent also the first evidence of the occurrence of Trichininae and Oedaleinae subfamilies throughout North Africa and bring the total of Moroccan hybotid fauna to 51 species. The new species are described and illustrated. A key to Moroccan Trichina species is provided.

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Research Article Mon, 10 Oct 2022 10:08:23 +0300
Three new fold-winged crane flies of the genus Ptychoptera Meigen, 1803 (Diptera, Ptychopteridae) from southern China https://zookeys.pensoft.net/article/86483/ ZooKeys 1122: 159-172

DOI: 10.3897/zookeys.1122.86483

Authors: Zehui Kang, Gang Gao, Xiao Zhang, Ding Yang, Junping Wang

Abstract: Three new Ptychoptera Meigen, 1803 species from southern China, P. hekouensis sp. nov., P. longa sp. nov., and P. xiaohuangshana sp. nov., are described and illustrated. These new species are mainly distinguished from congeners by their body colors and male genitalia. The genus Ptychoptera is recorded from Guangdong, China for the first time. An updated key to all Chinese Ptychoptera species is provided.

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Research Article Wed, 28 Sep 2022 17:01:51 +0300
The genus Hercostomus Loew (Diptera, Dolichopodidae, Dolichopodinae) from Inner Mongolia, China, with the description of two new species https://zookeys.pensoft.net/article/84403/ ZooKeys 1118: 119-131

DOI: 10.3897/zookeys.1118.84403

Authors: Xingyang Qian, Ning Wang, Ding Yang

Abstract: Previously, only two species of Hercostomus Loew were known to occur in Inner Mongolia. Here two species from Inner Mongolia are described as new to science, namely Hercostomus chifengensis sp. nov. and Hercostomus triangulatus sp. nov. Three new records of Hercostomus in Inner Mongolia are added. A key to the species of Hercostomus in Inner Mongolia is provided.

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Research Article Wed, 24 Aug 2022 14:23:00 +0300
A checklist of black flies (Diptera, Simuliidae) from India https://zookeys.pensoft.net/article/84686/ ZooKeys 1118: 111-118

DOI: 10.3897/zookeys.1118.84686

Authors: Arka Mukherjee, Atanu Naskar, Oishik Kar, Debdeep Pramanik, Dhriti Banerjee

Abstract: An updated checklist of the family Simuliidae from India is presented. A total of 79 species of Simulium belonging to eight different subgenera are listed. Eleven species that were not reported in the previous checklist are added here. The present list contributes to a better understanding of the diversity of Simuliidae in India, as well as the impact of Simulium species on the public health of this mega-diverse country.

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Checklist Wed, 24 Aug 2022 14:21:57 +0300
Updated taxonomic keys for European Hippoboscidae (Diptera), and expansion in Central Europe of the bird louse fly Ornithomya comosa (Austen, 1930) with the first record from Slovakia https://zookeys.pensoft.net/article/80146/ ZooKeys 1115: 81-101

DOI: 10.3897/zookeys.1115.80146

Authors: Jozef Oboňa, Katarína Fogašová, Miroslav Fulín, Stanislav Greš, Peter Manko, Jakub Repaský, Jindřich Roháček, Oldřich Sychra, Martin Hromada

Abstract: The available keys for European Hippoboscidae are outdated and do not cover all species currently known from Europe. Therefore, identification keys to the eleven genera and 31 species of the European hippoboscids are provided here. Ornithomya comosa (Austen, 1930) (Diptera: Hippoboscidae) is recorded for the first time from the territory of Slovakia based on one female found on a sand martin, Riparia riparia (Linnaeus, 1758). The list of keds and louse flies recorded from the territory of Slovakia is increased to 20 species. New host records for Slovakia are presented.

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Research Article Fri, 29 Jul 2022 14:58:44 +0300
First record of Lepidiella Enderlein, 1937 from the Oriental Region (Diptera, Psychodidae) https://zookeys.pensoft.net/article/81668/ ZooKeys 1115: 73-79

DOI: 10.3897/zookeys.1115.81668

Authors: Santiago Jaume-Schinkel, Gunnar Mikalsen Kvifte

Abstract: We provide the first record of the genus Lepidiella Enderlein, 1937 from the Oriental Region with the description of Lepidiella limicornis sp. nov., based on two male specimens collected in Thailand. Additionally, we provide a list of the world species of Lepidiella and discuss the diagnosis and taxonomic placement of the genus.

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Research Article Fri, 29 Jul 2022 14:58:24 +0300
Three new species of the genus Alluaudomyia Kieffer, 1913 (Diptera, Ceratopogonidae) from the National Park of Hainan Tropical Rainforest, China https://zookeys.pensoft.net/article/83021/ ZooKeys 1112: 199-218

DOI: 10.3897/zookeys.1112.83021

Authors: Xiaoxiang Wu, Zehua He, Xiaodan Lin, Bin Deng, Qi Zhai, Jiahui Li

Abstract: Three new species of the predaceous midges of genus Alluaudomyia Kieffer, 1913: A. flavinotum Wu & Li, sp. nov. of the maculipennis group, and A. reflexuralis Wu & Li, sp. nov. and A. limu Wu & Li, sp. nov. of the parva group, are described from the National Park of Hainan Tropical Rainforest, Hainan Island, China. Illustrations and COI barcodes (a fragment of the mitochondrial cytochrome c oxidase subunit 1) of the three new species are also provided. Associations of male and female specimens of two species (A. reflexuralis Wu & Li, sp. nov. and A. limu Wu & Li, sp. nov.) are supported by DNA barcodes. The parva group is reported from China for the first time.

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Research Article Thu, 14 Jul 2022 18:10:27 +0300
Taxonomic and nomenclatural notes on Chinese species of Sarcophaga Meigen, 1824 (Diptera, Sarcophagidae) https://zookeys.pensoft.net/article/83267/ ZooKeys 1108: 141-159

DOI: 10.3897/zookeys.1108.83267

Authors: Chao Wang, Haoran Sun, Weibing Zhu, Thomas Pape, Qiyong Liu, Dong Zhang

Abstract: New taxonomic and nomenclatural data are provided for Chinese species of Sarcophaga Meigen, 1824. Eight new synonyms are proposed: two at the genus level, Magnicauda Wei, 2005 syn. nov. = Sarcophaga Meigen, 1824 and Leigongshanophaga Lehrer & Wei, 2010 syn. nov. = Sarcophaga Meigen, 1824, two at the subgenus level, Magnicauda Wei, 2005 syn. nov. = Pterosarcophaga Ye, 1981 and Leigongshanophaga Lehrer & Wei, 2010 syn. nov. = Cornexcisia Fan & Kano, 2000, and four at the species level, Sarcophaga catoptosa Wei & Yang, 2007 syn. nov. = Sarcophaga suthep Pape & Bänziger, 2003, Pierretia daozhenensis Wei, 2005 syn. nov. = Sarcophaga sichotealini (Rohdendorf, 1938), Pierretia autochthona Wei & Yang, 2007 syn. nov. = Sarcophaga (Liosarcophaga) kanoi Park, 1962, and Parasarcophaga simultaneousa Wei & Yang, 2007 syn. nov. = Sarcophaga huangshanensis (Fan, 1964). Sarcophaga (Liosarcophaga) aegyptica Salem, 1935 is considered a senior synonym of Sarcophaga (Liosarcophaga) parkeri (Rohdendorf, 1937). Correct original spellings are established, by First Reviser action, for the genus-group names Magnicauda Wei, 2005 and Pterosarcophaga Ye, 1981 and for the species-group name Magnicauda linjiangensis Wei, 2005. Chinese material of Sarcophaga (Bellieriomima) genuforceps, S. (Robineauella) huangshanensis (holotype and paratype), S. (Liosarcophaga) kanoi, and S. (L.) aegyptica is photographed for the first time.

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Research Article Fri, 24 Jun 2022 16:57:56 +0300
Phlebotomine sand flies (Diptera, Psychodidae) from Spain: an updated checklist and extended distributions https://zookeys.pensoft.net/article/81432/ ZooKeys 1106: 81-99

DOI: 10.3897/zookeys.1106.81432

Authors: Daniel Bravo-Barriga, Ignacio Ruiz-Arrondo, Rosa Estrada Peña, Javier Lucientes, Sarah Delacour-Estrella

Abstract: Phlebotomine sand flies (Diptera: Psychodidae) are the natural vectors of Leishmania spp. (Kinetoplastida: Trypanosomatidae) and phleboviruses (Bunyavirales: Phenuiviridae). In Spain, these vectors appear to be increasing their geographical distribution and have serious repercussions on public and veterinary health, encouraging studies of sand flies and their associated pathogens. An up-to-date and easily accessible compendium of current and historical data on their presence and detailed distribution is a crucial step towards the development and implementation of appropriate preventive strategies. A checklist on the presence and distribution of sand flies in Spain is compiled from data extracted from a comprehensive review of scientific literature published between 1909 and 2021 and our new records on the presence of sand flies specimens collected under the entomological surveillance of bluetongue vectors from the Spanish Ministry of Agriculture, Fishery and Food (MAPA) during the period 2004–2021. In total, 13 Spanish species of sand flies (two of them with controversial status) belonging to two genera and six subgenera are presented in this updated checklist, including new distribution data for seven species, among which several stand out as confirmed or suspected vectors of Leishmania infantum: Phlebotomus ariasi, Ph. langeroni, Ph. mascittii, and Ph. perniciosus.

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Checklist Fri, 17 Jun 2022 12:04:03 +0300
Hexatoma crane flies (Diptera, Limoniidae) of Korea https://zookeys.pensoft.net/article/82495/ ZooKeys 1105: 165-208

DOI: 10.3897/zookeys.1105.82495

Authors: Sigitas Podenas, Sun-Jae Park, Hye-Woo Byun, Virginija Podeniene

Abstract: Hexatoma Latreille, 1809 is a large genus of short-palped crane flies with a worldwide distribution. Accounting for more than 60 percent of global species, 362 extant species occur in Asia. Prior to our study, Hexatoma crane flies on the Korean Peninsula (both the Democratic People’s Republic of Korea and the Republic of Korea) had been studied for 90 years starting in 1930, but only three species had been recorded, with one of them being a misidentification. This study adds six species to the fauna of the Korean Peninsula, all of which belong to the subgenus H. (Eriocera) Macquart, 1838. General information on genus and subgenus morphological characters is presented in this paper, with a redescription of each species based on Korean specimens, as well as illustrations of both sexes, the elevation range, the period of activity, habitat information, general distribution, and a distribution map for the Korean Peninsula. Three species H. (E.) ilwola Podenas, sp. nov., H. (E.) pianigra Podenas, sp. nov. and H. (E.) serenensis Podenas, sp. nov. are described as new to science. Hexatoma (E.) lygropis (Alexander, 1920) is deleted from the Korean species list as a misidentification. This publication is a continuation of our previous work on short-palped crane flies (Limoniidae) from Korea.

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Research Article Wed, 15 Jun 2022 13:52:31 +0300
Taxonomic study on the genus Stenochironomus Kieffer from the Baishanzu Nature Reserve, China (Diptera, Chironomidae) https://zookeys.pensoft.net/article/81403/ ZooKeys 1104: 93-113

DOI: 10.3897/zookeys.1104.81403

Authors: Chao Song, Bin-Qing Zhu, Joel Moubayed-Breil, Teng Lei, Xin Qi

Abstract: During the summer of July to September 2020, a biodiversity survey on Chironomidae of Baishanzu Nature Reserve, China was made. In total, five Stenochironomus taxa/species were discovered, of which two belong to undescribed species and one (S. okialbus Sasa, 1990) is reported for the first time from China. The male adults of two new species are described and illustrated. Stenochironomus annulus Song & Qi sp. nov. is distinguished in having a wing with two dark spots restricted to the fork area of FCu and RM, the mid- and hind-femur each with a brown annulus, and the inferior volsella with two setae and one strong terminal spine. Stenochironomus baishanzuensis Song & Qi sp. nov. is distinguished by a combination of characters: a single dark spot on the middle part of the wing, fore legs brown to dark brown except for the basal 3/4 of femur, and the inferior volsella with four long setae and one stout terminal spine. The neighbour-joining tree based on public COI barcodes formed distinct clades with clear support for the new species. An updated key to known male adults of Stenochironomus from China is also provided.

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Research Article Thu, 9 Jun 2022 17:48:36 +0300
Catalogue of the Diptera (Insecta) of Morocco— an annotated checklist, with distributions and a bibliography https://zookeys.pensoft.net/article/62644/ ZooKeys 1094: 1-466

DOI: 10.3897/zookeys.1094.62644

Authors: Kawtar Kettani, Martin J. Ebejer, David M. Ackland, Gerhard Bächli, David Barraclough, Miroslav Barták, Miguel Carles-Tolrá, Milos Černý, Pierfilippo Cerretti, Peter Chandler, Mohamed Dakki, Christophe Daugeron, Herman De Jong, Josef Dils, Henry Disney, Boris Droz, Neal Evenhuis, Paul Gatt, Gustavo Graciolli, Igor Y. Grichanov, Jean-Paul Haenni, Martin Hauser, Oumnia Himmi, Iain MacGowan, Bruno Mathieu, Mohamed Mouna, Lorenzo Munari, Emilia P. Nartshuk, Oleg P. Negrobov, Pjotr Oosterbroek, Thomas Pape, Adrian C. Pont, Grigory V. Popov, Knut Rognes, Marcela Skuhravá, Vaclav Skuhravý, Martin Speight, Guy Tomasovic, Bouchra Trari, Hans-Peter Tschorsnig, Jean-Claude Vala, Michael von Tschirnhaus, Rüdiger Wagner, Daniel Whitmore, Andrzej J. Woźnica, Tadeusz Zatwarnicki, Peter Zwick

Abstract: The faunistic knowledge of the Diptera of Morocco recorded from 1787 to 2021 is summarized and updated in this first catalogue of Moroccan Diptera species. A total of 3057 species, classified into 948 genera and 93 families (21 Nematocera and 72 Brachycera), are listed. Taxa (superfamily, family, genus and species) have been updated according to current interpretations, based on reviews in the literature, the expertise of authors and contributors, and recently conducted fieldwork. Data to compile this catalogue were primarily gathered from the literature. In total, 1225 references were consulted and some information was also obtained from online databases. Each family was reviewed and the checklist updated by the respective taxon expert(s), including the number of species that can be expected for that family in Morocco. For each valid species, synonyms known to have been used for published records from Morocco are listed under the currently accepted name. Where available, distribution within Morocco is also included. One new combination is proposed: Assuania melanoleuca (Séguy, 1941), comb. nov. (Chloropidae).

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Catalogue Tue, 12 Apr 2022 09:47:10 +0300
Two new species of Platensina Enderlein (Diptera, Tephritidae, Tephritinae, Dithrycini) from India https://zookeys.pensoft.net/article/80645/ ZooKeys 1092: 123-146

DOI: 10.3897/zookeys.1092.80645

Authors: K. J. David, D. L. Hancock, K. Sachin, R. G. Gracy, S. Salini

Abstract: Two new species of Platensina Enderlein, P. rabbanii David & Hancock, sp. nov., and P. flavistigma David & Hancock, sp. nov., are described from Meghalaya and southern India, respectively. Platensina rabbanii can be differentiated from P. alboapicalis Hering by the presence of a single hyaline indentation in cell r1 and the apical hyaline band in cell r2+3 restricted to the apex; P. flavistigma differs from P. quadrula Hardy by the presence of a yellow/fulvous pterostigma and shape of the epandrium. DNA barcode sequences of P. acrostacta (Wiedemann), P. flavistigma and P. platyptera Hendel were obtained and reported. Postabdominal descriptions and illustrations of P. acrostacta, P. platyptera and P. zodiacalis (Bezzi) are also provided along with keys to all 23 species and the 7 known from India.

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Research Article Wed, 6 Apr 2022 15:01:25 +0300
DNA barcoding of the horsefly fauna (Diptera, Tabanidae) of Croatia with notes on the morphology and taxonomy of selected species from Chrysopsinae and Tabaninae https://zookeys.pensoft.net/article/78707/ ZooKeys 1087: 141-161

DOI: 10.3897/zookeys.1087.78707

Authors: Stjepan Krčmar, Mladen Kučinić, Marco Pezzi, Branka Bruvo Mađarić

Abstract: In the Croatian fauna, horseflies (Tabanidae) are represented by 78 species belonging to two subfamilies, five tribes, and 10 genera. Identification of these species is based on morphological characteristics. In this study, 43 species of horseflies were analyzed. The highest number of species (19) belongs to the genus Tabanus, followed by the genera Hybomitra with seven species, Haematopota with six species, Chrysops with four species, Atylotus and Philipomyia with two species each, and the genera Silvius, Dasyrhamphis, and Heptatoma with one species each. The standard DNA barcoding region of the mitochondrial cytochrome c oxidase gene, subunit I (COI), was sequenced and compared to the Barcode of Life Database (BOLD). Our analyses confirmed our morphological identifications and added 16 new Barcode Index Numbers (BINs) for Tabanidae to BOLD. Potential problems in the systematics and taxonomy of this family are highlighted.

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Research Article Wed, 23 Feb 2022 10:19:07 +0200
The genus Blepharicera Macquart, 1843 newly recorded from Sichuan, China with descriptions of three new species (Diptera, Blephariceridae) https://zookeys.pensoft.net/article/75885/ ZooKeys 1085: 51-68

DOI: 10.3897/zookeys.1085.75885

Authors: Xiao Zhang, Zehui Kang

Abstract: The genus Blepharicera Macquart, 1843 is recorded from Sichuan, China for the first time with the following three new species: B. gengdica sp. nov., B. balangshana sp. nov. and B. kongsica sp. nov., increasing the number of Chinese Blepharicera species to eleven. The new species are distinguished from congeners mainly by their male genitalia. Descriptions and illustrations for the new species and an updated key to Chinese Blepharicera species are presented.

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Research Article Fri, 4 Feb 2022 15:28:08 +0200
First records of two genera and thirteen species of Tabanidae (Diptera) from Honduras https://zookeys.pensoft.net/article/77038/ ZooKeys 1084: 27-42

DOI: 10.3897/zookeys.1084.77038

Authors: Katerin Veroy, Jesus Orozco, Augusto L. Henriques

Abstract: This works presents information on the diversity of the Tabanidae of Honduras as a product of the examination of 386 specimens and a literature review. Thirteen species and two genera (Bolbodimyia and Dasychela) are recorded from the country for the first time. Eighty-five species distributed in 22 genera, five tribes, and three subfamilies are now known from Honduras. A key to the subfamilies, tribes, and genera of the known Honduran species is also included. All new records are mapped and illustrated to aid in the identification of the species.

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Research Article Wed, 26 Jan 2022 18:04:56 +0200
DNA barcoding of the leaf-miner flies (Diptera, Agromyzidae) of Mitaraka, French Guiana https://zookeys.pensoft.net/article/76651/ ZooKeys 1083: 147-168

DOI: 10.3897/zookeys.1083.76651

Authors: Stéphanie Boucher, Jade Savage

Abstract: Species level identification of Agromyzidae based on morphology is often challenging due to their small size and morphological homogeneity. DNA barcoding has been used regularly to assist with the identification of economically important species of Agromyzidae, but rarely as a tool for species delineation or identification in biodiversity surveys. The main objective of this study was to investigate whether DNA barcoding and the BIN (Barcoding Index) system could assist with species identification, species delineation, male/ female association, and diversity assessment of Agromyzidae material previously determined to morphospecies from Mitaraka, French Guiana. Amplification success was low, with sequences over 400 bp recovered for only 24 (48%) of the selected specimens. Sequences assigned to 17 morphospecies formed 16 distinct branches or clusters separated by very high (minimum of 10%) sequence divergence. Following the reassessment and subsequent reassignment of one specimen, congruence between morphology and DNA barcodes was high with a single instance of two morphospecies sharing identical sequences. While DNA barcoding did not assist with identification (none of our sequences matched those of named taxa in BOLD or GenBank), it did provide support for most of our morphospecies concepts, including male/female associations. The BIN system also provided access to information about the distribution and habitat preferences of several taxa. We conclude that DNA barcoding was a useful approach to study the species diversity of our samples but that much work remains to be done before it can be used as an identification tool for the Agromyzidae fauna of Mitaraka and the rest of the Neotropical region.

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Research Article Tue, 25 Jan 2022 10:53:50 +0200
Notes on the taxonomic status and distribution of some Cylindrotomidae (Diptera, Tipuloidea), with emphasis on Japanese species https://zookeys.pensoft.net/article/75624/ ZooKeys 1083: 13-88

DOI: 10.3897/zookeys.1083.75624

Authors: Levente-Péter Kolcsár, Nikolai Paramonov, Yume Imada, Daichi Kato, Maribet Gamboa, Dai Shinoka, Makoto Kato, Kozo Watanabe

Abstract: A morphological and molecular study of 17 Cylindrotomidae species revealed that the two subspecies of Cylindrotoma distinctissima, the Nearctic C. americana Osten Sacken, 1865, stat. reval. and the Palearctic C. distinctissima (Meigen, 1818), represent separated lineages and consequently are raised to species level. Cylindrotoma japonica Alexander, 1919, syn. nov. and C. distinctissima alpestris Peus, 1952, syn. nov. are now known to be junior synonyms of C. distinctissima. Triogma kuwanai limbinervis Alexander, 1953, syn. nov. and T. nimbipennis Alexander, 1941, syn. nov. are now placed into synonymy under Triogma kuwanai (Alexander, 1913). The Japanese Cylindrotomidae are all redescribed and all available literature and distribution data are summarised. Supplementary descriptions and illustrations for male and female terminalia of Cylindrotoma nigriventris Loew, 1849, Diogma dmitrii Paramonov, 2005, Liogma nodicornis (Osten Sacken, 1865), Phalacrocera replicata (Linnaeus, 1758), P. tipulina Osten Sacken, 1865, and Triogma trisulcata (Schummel, 1829) are provided. The following new distribution records are outlined; Diogma caudata Takahashi, 1960 from Arkhangelsk Oblast, Russia; D. glabrata (Meigen, 1818) from Belarus, Latvia, and Altai Republic, Amur Oblast, Novgorod Oblast, Magadan Oblast, Samara Oblast, and Kuril Islands (Shikotan I and Paramushir I) in Russia; Liogma serraticornis Alexander, 1919 from Khabarovsk Krai, Russia; Phalacrocera replicata from Khabarovsk Krai, Russia; and the presence of Cylindrotoma nigriventris in Altai Republic, Russia is confirmed.

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Research Article Mon, 24 Jan 2022 10:24:07 +0200
A new species of Simulium (Gomphostilbia) (Diptera, Simuliidae) from Thailand, with a key to identify females of 14 species of the Simulium varicorne species-group https://zookeys.pensoft.net/article/77428/ ZooKeys 1083: 1-12

DOI: 10.3897/zookeys.1083.77428

Authors: Wichai Srisuka, Kittipat Aupalee, Yasushi Otsuka, Masako Fukuda, Hiroyuki Takaoka, Atiporn Saeung

Abstract: Simulium (Gomphostilbia) khelangense sp. nov. is described on the basis of females, collected by a sweeping net in Lampang, Phitsanulok and Chiang Mai Provinces, Thailand. This new species is placed in the S. chumpornense subgroup of the S. varicorne species-group in the subgenus Gomphostilbia Enderlein by having the antenna with eight flagellomeres, pleural membrane bare, and female subcosta lacking hairs. It is similar to S. kuvangkadilokae Pramual & Tangkawanit from Thailand in the same subgroup but is barely distinguished from the latter species by the head width relative to the greatest width of the frons and length of the labrum relative to the clypeus. A genetic analysis using the COI gene sequences similarly shows that S. khelangense sp. nov. is most closely related to S. kuvangkadilokae, with a genetic distance of 1.23–2.81%. A revised key to identify females of 14 species of the S. varicorne species-group is provided.

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Research Article Fri, 21 Jan 2022 16:03:11 +0200
Corynoneura Winnertz species from Hunan Province, Oriental China, delineated with morphological and 16S rDNA data (Diptera, Chironomidae) https://zookeys.pensoft.net/article/73019/ ZooKeys 1082: 87-102

DOI: 10.3897/zookeys.1082.73019

Authors: Yue Fu, Xiang-Liang Fang, Xin-Hua Wang, Mi Shen, Yun-Li Xiao

Abstract: The genus Corynoneura Winnertz, 1846 from Hunan Province in Oriental China is reviewed. Four new species, C. enormis Fu sp. nov., C. gibbera Fu sp. nov., C. incuria Fu sp. nov., and C. longshanensis Fu sp. nov. are described and illustrated based on adult males. Sequence data from the 16S rDNA gene were used to infer relationships between these species and complement morphological delineation. Sequences from the mitochondrial large ribosomal subunit (16S rDNA) from these species are uploaded to the National Center for Biotechnology Information (NCBI). Relationships were inferred using the Neighbor-Joining method based on 16S rDNA.

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Research Article Wed, 19 Jan 2022 16:47:11 +0200
Wing interference patterns are consistent and sexually dimorphic in the four families of crane flies (Diptera, Tipuloidea) https://zookeys.pensoft.net/article/69060/ ZooKeys 1080: 135-163

DOI: 10.3897/zookeys.1080.69060

Authors: Robert T. Conrow, Jon K. Gelhaus

Abstract: Wing interference patterns (WIP) are stable structural colors in insect wings caused by thin-film interference. This study seeks to establish WIP as a stable, sexually dimorphic, species-level character across the four families of Tipuloidea and investigate generic level WIP. Thirteen species of Tipuloidea were selected from museum specimens in the Academy of Natural Sciences of Drexel University collection. One wing from a male and female of each representative species was excised and mounted to a slide with coverslip, placed against a black background, and imaged using an integrated microscope camera. Images were minimally retouched but otherwise unchanged. Descriptions of the WIP for each sex of each species are provided. Twelve of thirteen species imaged had WIP, which were stable and species specific while eight of those twelve had sexually dimorphic WIP. Comparisons of three species of Nephrotoma were inconclusive regarding a generic level WIP. Gnophomyia tristissima had higher intraspecific variation than other species examined. This study confirms stable, species specific WIP in all four families of crane flies for the first time. More research must be done regarding generic-level stability of WIP in crane flies as well as the role sexual and natural selection play in the evolution of wing interference patterns in insects.

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Research Article Wed, 5 Jan 2022 11:11:24 +0200
Taxonomic revision of Romaleosyrphus Bigot (Diptera, Syrphidae), including descriptions of seven new species https://zookeys.pensoft.net/article/55862/ ZooKeys 1075: 1-32

DOI: 10.3897/zookeys.1075.55862

Authors: Kevin M. Moran, Jeffrey H. Skevington

Abstract: The genus Romaleosyrphus Bigot is reviewed, including the description of seven new species (R. argosi Moran, sp. nov., R. bigoti Moran, sp. nov., R. drysus Moran, sp. nov., R. nephelaeus Moran & Thompson, sp. nov., R. soletluna Moran & Thompson, sp. nov., R. vockerothi Moran & Thompson, sp. nov. and R. woodi Moran, sp. nov.). Romaleosyrphus arctophiloides (Giglio-Tos), comb. nov. is transferred to Romaleosyrphus. Romaleosyrphus stat. rev. is redefined to represent the monophyletic unit of species within Criorhinina which possess holoptic males, a proximal ventral half of vein C with setae, a broad intersection of vein R1 with vein C, the distal part of R4+5 beyond M1 longer than cross-vein h and appressed pile on the abdomen. Descriptions, habitus and genitalia photographs, distributions, and an illustrated key for all nine Romaleosyrphus are presented. DNA barcode data are provided for eight of the species with a cytochrome c oxidase subunit I gene tree presented and discussed.

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Research Article Tue, 7 Dec 2021 11:08:10 +0200
Four new species of the genus Luzonomyza Malloch (Diptera, Lauxaniidae) from China https://zookeys.pensoft.net/article/68392/ ZooKeys 1074: 43-59

DOI: 10.3897/zookeys.1074.68392

Authors: Wenliang Li, Xulong Chen, Keli Feng, Shengjuan Zhao, Ding Yang

Abstract: Four species of the genus Luzonomyza Malloch, 1929 from southwest China are described as new to science: Luzonomyza vittifacies Li & Yang, sp. nov., L. serrata Li & Yang, sp. nov., L. honghensis Li & Yang, sp. nov., and L. brevis Li & Yang, sp. nov. A key to Luzonomyza species is also presented.

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Research Article Wed, 1 Dec 2021 17:20:14 +0200
Hexatoma (Eriocera) Macquart (Diptera, Limoniidae) from Xizang, China https://zookeys.pensoft.net/article/66750/ ZooKeys 1071: 155-173

DOI: 10.3897/zookeys.1071.66750

Authors: Bing Zhang, Qicheng Yang, Yan Li, Ding Yang

Abstract: One new species of the subgenus Eriocera Macquart,1838, Hexatoma (Eriocera) xizangensis sp. nov. is described and illustrated from Xizang, China. The following four species are re-described and reported from Xizang for the first time: H. (E.) latigrisea Alexander, 1971, H. (E.) nepalensis (Westwood, 1836), H. (E.) paragnava Alexander, 1973 and H. (E.) perhirsuta Alexander, 1973. A key to the species of Eriocera from Xizang is presented.

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Research Article Thu, 18 Nov 2021 08:03:33 +0200
Helen’s twins in the Balkans: discovery of two new Paraptychoptera Tonnoir, 1919 species closely related to P. helena Peus, 1958, with systematic revision of the “lacustris” group (Diptera, Ptychopteridae) https://zookeys.pensoft.net/article/58598/ ZooKeys 1071: 63-81

DOI: 10.3897/zookeys.1071.58598

Authors: Lujza Keresztes, Jürgen Kappert, Mária Henning, Edina Török

Abstract: Ptychoptera castor Keresztes & Kappert, sp. nov. and P. pollux Keresztes & Török, sp. nov. both belong to the subgenus Ptychoptera (Paraptychoptera) Tonnoir (1919) and are described from boggy headwaters in the south Balkan area. These new species are closely related to the range-restricted P. helena Peus, 1958, which is known only from Oiti village, Mount Oeta, Phthioitis region, Greece and, together with P. lacustris, forms a morphologically well-defined unit in the subgenus Paraptychoptera. Based on cladistic analyses of 53 different morphological characters using the male antenna, wing, and genital structures, a general revision of the “lacustris” group is proposed with a dichotomous key of Paraptychoptera species.

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Research Article Wed, 17 Nov 2021 17:20:26 +0200
A review of tangle-veined flies (Nemestrinidae, Diptera) in Egypt https://zookeys.pensoft.net/article/70743/ ZooKeys 1071: 11-42

DOI: 10.3897/zookeys.1071.70743

Authors: Arafa Elsayed El-Hashash, Haitham Badrawy Mousa Badrawy, Ayman Mohyie-Eldin Ibrahim

Abstract: The Egyptian fauna of the genus Nemestrinus Latreille, 1802 is revised. In 1967, Steyskal and El-Bialy listed 12 species from the region, but only six species are now recognized. The primary type specimens of the species N. aegyptiacus (Wiedemann, 1828), N. rufipes (Olivier, 1810), and N. lateralis Wiedemann, 1828 (N. lateralis being a synonym of N. rufipes) deposited in the Museum für Naturkunde, Berlin, Germany were examined. Two species (N. abdominalis Olivier, 1811 and N. fascifrons (Bigot, 1888) are placed as new synonyms of N. ater (Olivier, 1811), and N. ruficornis Macquart, 1840 is synonymized with N. rufipes (Olivier, 1811). Nemestrinus jullieni (Efflatoun, 1925) is confirmed as a synonym of N. aegyptiacus. Furthermore, three species (N. caucasicus Fischer, 1806, N. pallipes (Olivier, 1811), and N. persicus Lichtwardt, 1909) have been removed and are doubtful records from Egypt. A key to the species, lists of specimens examined, and Illustrations and distributions for each species are provided. The status of species of doubtful occurrence in Egypt is discussed.

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Research Article Tue, 16 Nov 2021 08:14:56 +0200
New species of the genus Ptychoptera Meigen, 1803 (Diptera, Ptychopteridae) from Zhejiang, China with an updated key to Chinese species https://zookeys.pensoft.net/article/67779/ ZooKeys 1070: 87-99

DOI: 10.3897/zookeys.1070.67779

Authors: Jiaqi Shao, Zehui Kang

Abstract: We revise the taxonomy of the genus Ptychoptera Meigen, 1803 from Zhejiang, East China. One new species from Zhejiang, P. tianmushana sp. nov., is described and illustrated. Morphologically, the new species is most similar to P. emeica Kang, Xue & Zhang, 2019 and P. formosensis Alexander, 1924, but it can be distinguished by the coloration of the abdomen and the details of the male genitalia. In addition, P. bellula Alexander, 1937 is recorded from Zhejiang for the first time. Two known species from Zhejiang, P. longwangshana Yang & Chen, 1998 and P. gutianshana Yang & Chen, 1995, are redescribed and illustrated. A key to Chinese species of Ptychoptera is provided.

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Research Article Thu, 11 Nov 2021 09:06:35 +0200
Two new species of the genus Ptychoptera Meigen, 1803 (Diptera, Ptychopteridae) from Yunnan, China with remarks on the distribution of Chinese species https://zookeys.pensoft.net/article/58859/ ZooKeys 1070: 73-86

DOI: 10.3897/zookeys.1070.58859

Authors: Xiao Zhang, Zehui Kang

Abstract: Sixteen known species of the genus Ptychoptera Meigen, 1803 have been recorded from China, of which three occur in Yunnan Province. Herein, two new species from Yunnan, P. cordata sp. nov. and P. yunnanica sp. nov., are described from China. An updated key to Ptychoptera is presented for all Chinese species.

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Research Article Thu, 11 Nov 2021 09:05:48 +0200
On the genus Polypedilum, subgenus Collartomyia, with description of P. (Col.) baishanzuensis sp. nov. from Baishanzu Nature Reserve, China (Diptera, Chironomidae) https://zookeys.pensoft.net/article/69870/ ZooKeys 1065: 1-12

DOI: 10.3897/zookeys.1065.69870

Authors: Chao Song, Binqing Zhu, Wei Liu, Xin Qi

Abstract: A new species of the genus Polypedilum Kieffer, 1912 is described from Baishanzu Nature Reserve, China, based on molecular and morphological data. Molecular phylogenetic analysis based on standard barcode sequences confirmed a new clade of Polypedilum (Collartomyia) species. The new species is easily distinguished from its congeners by a combination of the following morphological characters: membrane of wing with a large spot occupying 70% of the proximal area; tergite without dark brown band pigmentation; tarsi I–V dark brown; superior volsella with three outer lateral setae and six long setae along inner base; inferior volsella with setose tubercules. An updated key to adult males of the subgenus Collartomyia is also provided.

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Research Article Fri, 22 Oct 2021 13:57:03 +0300
Annotated catalogue of the Tachinidae (Insecta, Diptera) of Chile https://zookeys.pensoft.net/article/62972/ ZooKeys 1064: 1-200

DOI: 10.3897/zookeys.1064.62972

Authors: James E. O'Hara, D. Monty Wood, Christian R. González

Abstract: The Tachinidae (Diptera) of Chile are catalogued and information is given on distributions, name-bearing types, synonyms, nomenclatural issues, and pertinent literature. The history of tachinid collectors in Chile and authors who have contributed to the systematic knowledge of Chilean tachinids is extensively reviewed. The classification has been updated and 122 genera and 264 species are recognised in Chile. There is a significant amount of endemism with 28 genera and 100 species known only from Chile. There are also 113 species with distributions shared only between Chile and Argentina, particularly in the southern portions of these countries comprising Patagonia. The catalogue is based on examination of the original descriptions of all nominal species and all other references known to us containing relevant taxonomic and distributional information, for a total of approximately 450 references. Many of the name-bearing types and other Chilean specimens housed in collections were examined. Taxa are arranged hierarchically and alphabetically under the categories of subfamily, tribe, genus, subgenus (where recognised), and species. Nomenclatural information is provided for genus-group and species-group names, including lists of synonyms (mostly restricted to Neotropical taxa) and name-bearing type data. Species distributions are recorded by country within the New World and by larger geographical divisions in the Old World. Additional information is given in the form of notes and references under valid names at the level of tribe, genus, and species. Two genera are newly recorded from Chile: Chaetoepalpus Vimmer & Soukup, 1940 (Tachinini) (also newly recorded from Argentina) and Patelloa Townsend, 1916 (Goniini). Four species are newly recorded from Chile or other countries: Lypha ornata Aldrich, 1934 (Chile); Chaetoepalpus coquilleti Vimmer & Soukup, 1940 (Argentina and Chile); Phytomyptera evanescens (Cortés, 1967) (Argentina); and Xanthobasis unicolor Aldrich, 1934 (Chile). Eight species previously recorded from Chile are deemed to have been misidentified or misrecorded from Chile (known distributions in parentheses): Archytas incertus (Macquart, 1851) (Argentina, Brazil, Paraguay, Uruguay); Archytas seminiger (Wiedemann, 1830) (Brazil, Colombia); Gonia crassicornis (Fabricius, 1794) (Brazil, Peru, Venezuela, Middle America, West Indies, Nearctic); Lespesia andina (Bigot, 1888) (Cuba); Lespesia archippivora (Riley, 1871) (widespread Nearctic and most of Neotropical); Neoethilla ignobilis (van der Wulp, 1890) (Mexico, United States); Siphona (Siphona) geniculata (De Geer, 1776) (Palaearctic, Nearctic [introduced]); and Winthemia quadripustulata (Fabricius, 1794) (Palaearctic, Nearctic, Oriental]. As First Reviser we fix Paratheresia rufiventris Townsend, 1929 as the senior homonym and Sarcoprosena rufiventris Townsend, 1929 as the junior homonym when the two are placed together in Billaea Robineau-Desvoidy, 1830; and we fix Mayophorinia angusta Townsend, 1927 as the senior homonym and Metarrhinomyia angusta Townsend, 1927 as the junior homonym when the two are placed together in Myiopharus Brauer & Bergenstamm, 1889. New replacement names are proposed for eight preoccupied names of Neotropical species (country of type locality in parentheses): Billaea rufescens O’Hara & Wood for Sarcoprosena rufiventris Townsend, 1929, preoccupied in the genus Billaea Robineau-Desvoidy, 1830 by Paratheresia rufiventris Townsend, 1929 (Peru), nom. nov.; Billaea triquetrus O’Hara & Wood for Sarcoprosena triangulifera Townsend, 1927, preoccupied in the genus Billaea Robineau-Desvoidy, 1830 by Dexia triangulifera Zetterstedt, 1844 (Peru), nom. nov.; Eucelatoria nudioculata O’Hara & Wood for Eucelatorioidea nigripalpis Thompson, 1968, preoccupied in the genus Eucelatoria Townsend, 1909 by Chetolyga nigripalpis Bigot, 1889 (Trinidad), nom. nov.; Eucelatoria oblonga O’Hara & Wood for Urodexodes elongatum Cortés & Campos, 1974, preoccupied in the genus Eucelatoria Townsend, 1909 by Exorista elongata van der Wulp, 1890 (Chile), nom. nov.; Lespesia thompsoni O’Hara & Wood for Sturmiopsoidea obscura Thompson, 1966, preoccupied in the genus Lespesia Robineau-Desvoidy, 1863 by Eurigaster obscurus Bigot, 1857 (Cuba), nom. nov.; Myiopharus charapensis O’Hara & Wood for Metarrhinomyia angusta Townsend, 1927, preoccupied in the genus Myiopharus Brauer & Bergenstamm, 1889 by Mayophorinia angusta Townsend, 1927 (Peru), nom. nov.; Myiopharus incognitus O’Hara & Wood for Stenochaeta claripalpis Thompson, 1968, preoccupied in the genus Myiopharus Brauer & Bergenstamm, 1889 by Neoxynopsoidea claripalpis Thompson, 1968 (Trinidad), nom. nov.; and Myiopharus rufopalpus O’Hara & Wood for Paralispe palpalis Townsend, 1929, preoccupied in the genus Myiopharus Brauer & Bergenstamm, 1889 by Myioxynops palpalis Townsend, 1927 (Peru), nom. nov. New type species fixations are made under the provisions of Article 70.3.2 of the ICZN Code for three genus-group names: Parafabricia Brauer & Bergenstamm, 1894 (synonym of Archytas Jaennicke, 1867), type species newly fixed as Parafabricia perplexa Townsend, 1931; Tachinodes Brauer & Bergenstamm, 1889 (synonym of Archytas Jaennicke, 1867), type species newly fixed as Jurinia metallica Robineau-Desvoidy, 1830; and Willistonia Brauer & Bergenstamm, 1889 (synonym of Belvosia Robineau-Desvoidy, 1830), type species newly fixed as Willistonia aldrichi Townsend, 1931. Lectotypes are designated for the following four nominal species, all described or possibly described from Chile: Echinomyia pygmaea Macquart, 1851 (a valid name in the genus Peleteria Robineau-Desvoidy, 1830); Gonia chilensis Macquart, 1844 (a junior synonym of Gonia pallens Wiedemann, 1830); Masicera auriceps Macquart, 1844 (a valid name in the genus Lespesia Robineau-Desvoidy, 1863); and Prosopochoeta nitidiventris Macquart, 1851 (a valid name in the genus Prosopochaeta Macquart, 1851). The following 27 new or revived combinations are proposed (distributions in parentheses): Blepharipeza andina Bigot, 1888 is moved to Lespesia Robineau-Desvoidy, 1863 as L. andina, nomen dubium (Cuba), comb. nov.; Camposodes evanescens Cortés, 1967 is moved to Phytomyptera Rondani, 1845 as P. evanescens (Argentina, Chile), comb. nov.; Ectophasiopsis ypiranga Dios & Nihei, 2017 is moved to Trichopoda Berthold, 1827 and assigned to subgenus Galactomyia Townsend, 1908 as T. (G.) ypiranga (Argentina, Brazil), comb. nov.; Embiomyia australis Aldrich, 1934 is moved to Steleoneura Stein, 1924 as S. australis (Argentina, Chile), comb. nov.; Eurigaster modestus Bigot, 1857 is moved to Lespesia as L. modesta (Cuba), comb. nov.; Eurigaster obscurus Bigot, 1857 is moved to Lespesia as L. obscura (Cuba), comb. nov.; Macropatelloa tanumeana Townsend, 1931 is moved to Patelloa Townsend, 1916 as P. tanumeana (Argentina, Chile), comb. nov.; Masicera insignis van der Wulp, 1882 is moved to Drino Robineau-Desvoidy, 1863 as D. insignis (Argentina, Chile), comb. nov.; Parasetigena hichinsi Cortés, 1967 is moved to Chetogena Rondani, 1856 as C. hichinsi (Chile), comb. nov.; Parasetigena porteri Brèthes, 1920 and junior synonym Stomatotachina splendida Townsend, 1931 are moved to Chetogena as C. porteri (Chile), both comb. nov.; Phorocera calyptrata Aldrich, 1934 is moved to Admontia Brauer & Bergenstamm, 1889 as A. calyptrata (Argentina, Chile), comb. nov.; Poliops auratus Campos, 1953 is moved to Admontia Brauer & Bergenstamm, 1889 as A. aurata (Chile), comb. nov.; Poliops striatus Aldrich, 1934 is moved to Admontia as A. striata (Argentina, Chile), comb. nov.; Ruiziella frontosa Cortés, 1951 is moved to Chaetoepalpus Vimmer & Soukup, 1940 and placed in synonymy with C. coquilleti Vimmer & Soukup, 1940 (Argentina, Chile, Peru), comb. nov.; Ruiziella luctuosa Cortés, 1951 is moved to Chaetoepalpus as C. luctuosus (Argentina, Chile), comb. nov.; Sarcoprosena luteola Cortés & Campos, 1974 is moved to Billaea Robineau-Desvoidy, 1830 as B. luteola (Chile), comb. nov.; Sarcoprosena rufiventris Townsend, 1929 is moved to Billaea where it is a junior secondary homonym and is renamed B. rufescens O’Hara & Wood (Peru), comb. nov.; Sarcoprosena triangulifera Townsend, 1927 is moved to Billaea where it is a junior secondary homonym and is renamed B. triquetrus O’Hara & Wood (Peru),comb. nov.; Saundersia aurea Giglio-Tos, 1893 is moved to “Unplaced species of Tachinini” (Mexico), comb. nov.; Schistostephana aurifrons Townsend, 1919 is moved to Billaea as B. aurifrons (Peru), comb. nov.; Siphoactia charapensis Townsend, 1927 is moved to Clausicella Rondani, 1856 as C. charapensis (Peru), comb. nov.; Siphoactia peregrina Cortés & Campos, 1971 is moved to Clausicella as C. peregrina (Chile), comb. nov.; Sturmia festiva Cortés, 1944 is moved to Drino as D. festiva (Argentina, Chile), comb. nov.; Sturmiopsoidea obscura Thompson, 1966 is moved to Lespesia Robineau-Desvoidy, 1863, where it is a junior secondary homonym and is renamed L. thompsoni O’Hara & Wood (Trinidad), comb. nov.; Trichopoda arcuata Bigot, 1876 is returned to Trichopoda from Ectophasiopsis Townsend, 1915 and assigned to subgenus Galactomyia (Argentina, Chile), comb. revived; and Trichopoda gradata Wiedemann, 1830 is returned to Trichopoda from Ectophasiopsis and assigned to subgenus Galactomyia (Argentina, Brazil, Uruguay), comb. revived. New or revived generic and specific synonymies are proposed for the following 14 names: Camposodes Cortés, 1967 with Phytomyptera Rondani, 1845, syn. nov.; Ectophasiopsis Townsend, 1915 with Trichopoda Berthold, 1827, subgenus Galactomyia Townsend, 1908, syn. nov.; Embiomyia Aldrich, 1934 with Steleoneura Stein, 1924, syn. nov.; Fabricia andicola Bigot, 1888 with Peleteria robusta (Wiedemann, 1830), syn. revived; Macropatelloa Townsend, 1931 with Patelloa Townsend, 1916, syn. nov.; Peleteria inca Curran, 1925 with Peleteria robusta (Wiedemann, 1830), syn. revived; Poliops Aldrich, 1934 with Admontia Brauer & Bergenstamm, 1889, syn. nov.; Ruiziella Cortés, 1951 with Chaetoepalpus Vimmer & Soukup, 1940, syn. nov.; Ruiziella frontosa Cortés, 1951 with Chaetoepalpus coquilleti Vimmer & Soukup, 1940, syn. nov.; Sarcoprosena Townsend, 1927 with Billaea Robineau-Desvoidy, 1830, syn. nov.; Schistostephana Townsend, 1919 with Billaea, syn. nov.; Siphoactia Townsend, 1927 with Clausicella Rondani, 1856, syn. nov.; Stomatotachina Townsend, 1931 with Chetogena Rondani, 1856, syn. nov.; and Sturmiopsoidea Thompson, 1966 with Lespesia Robineau-Desvoidy, 1863, syn. nov.

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Catalogue Thu, 21 Oct 2021 17:11:24 +0300
Revision of the genus Niphta (Diptera, Thaumaleidae) Theischinger of South America, with descriptions of nine new species and a new immature morphotype https://zookeys.pensoft.net/article/71180/ ZooKeys 1063: 49-104

DOI: 10.3897/zookeys.1063.71180

Authors: Robert J. Pivar, Bradley J. Sinclair, John K. Moulton

Abstract: The Niphta Theischinger fauna of South America is revised to include 11 species, nine of which are described as new to science (N. acus Pivar, sp. nov., N. bifurcata Pivar & Moulton, sp. nov., N. bispinosa Pivar & Sinclair, sp. nov., N. brunnea Pivar, sp. nov., N. courtneyi Pivar, sp. nov., N. daniellae Pivar, sp. nov., N. downesi Pivar, sp. nov., N. eurydactyla Pivar, sp. nov., N. mapuche Pivar, sp. nov.). The genus Niphta is redefined, both previously described Chilean species are redescribed, N. halteralis (Edwards) and N. nudipennis (Edwards), and females are described or redescribed where possible. The first descriptions of the immature stages of South American Niphta are provided, which represent a new larval morphotype in Thaumaleidae, as larvae and pupae possess ventral adhesive structures. Furthermore, these larvae were collected from vegetation rather than rocky substrates. Illustrations and micrographs are provided for all species, and scanning electron microscopy images are included for select immatures. A key to species, distribution maps, and discussions regarding phylogenetic affinities and habitat are also included.

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Research Article Tue, 19 Oct 2021 07:00:01 +0300
The Herniosina story continues in the Mediterranean: H. calabra sp. nov. from Calabria and H. erymantha Roháček, new female from the Peloponnese (Diptera, Sphaeroceridae) https://zookeys.pensoft.net/article/72235/ ZooKeys 1061: 165-190

DOI: 10.3897/zookeys.1061.72235

Authors: Jindřich Roháček

Abstract: A study of recently acquired material of Herniosina Roháček, 1983 (Diptera: Sphaeroceridae: Limosininae) in the Mediterranean subregion revealed a new species, H. calabra sp. nov. (Italy: Calabria: Serre Calabresi Mts) and the first females of H. erymantha Roháček, 2016 (Greece: southern Peloponnese: Taygetos Mts). Herniosina calabra sp. nov. (both sexes) and the female of H. erymantha are described and illustrated in detail including structures of terminalia, their relationships are discussed and new information on their biology (habitat association) is given. An update of a key to all know species of Herniosina species is presented.

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Research Article Mon, 11 Oct 2021 15:48:24 +0300
Tipula (Vestiplex) crane flies (Diptera, Tipulidae) of Korea https://zookeys.pensoft.net/article/49999/ ZooKeys 1061: 23-55

DOI: 10.3897/zookeys.1061.49999

Authors: Pavel Starkevich, Sigitas Podėnas, Virginija Podėnienė, Sun-Jae Park, A-Young Kim

Abstract: The Korean species of Tipula (Vestiplex) Bezzi, 1924 crane flies are taxonomically revised. Five species are recognized. Tipula (V.) coquillettiana Alexander, 1924, T. (V.) kuwayamai Alexander, 1921, T. (V.) tchukchi Alexander, 1934, and T. (V.) verecunda Alexander, 1924 are newly recorded from the Korean Peninsula, and T. (V.) serricauda Alexander, 1914 was previously recorded. The larva of T. (V.) serricauda is described and illustrated, and the larvae of the subgenus T. (Vestiplex) are divided into four groups based on spiracular lobe morphology. An identification key, redescriptions, and illustrations of Korean T. (Vestiplex) adults and grouping of known larvae are presented.

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Research Article Thu, 30 Sep 2021 10:35:06 +0300
Scenopinus jerei, a new species of window fly (Diptera, Scenopinidae) from Finland https://zookeys.pensoft.net/article/70085/ ZooKeys 1059: 135-156

DOI: 10.3897/zookeys.1059.70085

Authors: Jaakko Pohjoismäki, Antti Haarto

Abstract: A new species of window fly (Diptera: Scenopinidae), Scenopinus jerei sp. nov., with characteristic bicoloured legs and completely black halteres, is described from Finland. To exclude potential previously named species, a survey of the relevant type specimens as well as original descriptions of the Palearctic and Nearctic Scenopinus species has been conducted, including old Scenopinus fenestralis (Linnaeus) synonyms. Scenopinus jerei sp. nov. is likely to be an overlooked, boreal forest specialist living in the nests of cavity-nesting birds. An identification key to the European species is provided.

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Research Article Fri, 10 Sep 2021 15:44:25 +0300
Novel lures and COI sequences reveal cryptic new species of Bactrocera fruit flies in the Solomon Islands (Diptera, Tephritidae, Dacini) https://zookeys.pensoft.net/article/68375/ ZooKeys 1057: 49-103

DOI: 10.3897/zookeys.1057.68375

Authors: Luc Leblanc, Francis Tsatsia, Camiel Doorenweerd

Abstract: Results from a snap-shot survey of Dacine fruit flies carried out on three of the Solomon Islands in April 2018 are reported. Using traps baited with the male lures cue-lure, methyl eugenol, and zingerone, 30 of the 48 species previously known to occur in the Solomon Islands were collected. Six species are newly described here: Bactrocera allodistincta sp. nov., B. geminosimulata sp. nov., B. kolombangarae sp. nov., B. quasienochra sp. nov., B. tsatsiai sp. nov., and B. vargasi sp. nov., all authored by Leblanc & Doorenweerd. An illustrated key to the 54 species now known to be present in the country is provided.

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Research Article Fri, 27 Aug 2021 19:34:46 +0300
Synorthocladius federicoi sp. nov., a new species occurring in the middle basin of the Adige River, northern Italy (Diptera, Chironomidae, Orthocladiinae) https://zookeys.pensoft.net/article/68175/ ZooKeys 1057: 105-116

DOI: 10.3897/zookeys.1057.68175

Authors: Valeria Lencioni, Joel Moubayed

Abstract: An adult male Synorthocladius was collected in the middle basin of the Adige River in the city of Verona, northern Italy. A combination of atypical characters for the genus signalled a new species. Synorthocladius federicoi sp. nov. is here diagnosed and described. The new species is known only from its type locality and is presumed to be a local biogeographical representative of the Italian Pre-Alps. An emended generic diagnosis, a key to known Synorthocladius from Europe and comments on the taxonomic position of the new species are given.

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Short Communication Fri, 27 Aug 2021 16:41:12 +0300
A review of Madagopsina Feijen, Feijen & Feijen (Diptera, Diopsidae) with description of a new species, key to the species, and discussion of intrageneric relationships https://zookeys.pensoft.net/article/67433/ ZooKeys 1057: 1-21

DOI: 10.3897/zookeys.1057.67433

Authors: Hans R. Feijen, Frida A. A. Feijen, Cobi Feijen, Benoît Gilles

Abstract: For the recently established genus Madagopsina (Diopsidae, stalk-eyed flies), Madagopsina makayensis Feijen, Feijen & Feijen, sp. nov. is described from Madagascar. A concise catalogue is given for the genus and an identification key is presented for its six species. The differential character states are listed for the two species groups of the genus: the Madagopsina apollo species group and the Madagopsina apographica species group. The intrageneric relations are discussed based on morphology, geometric morphometrics analysis of wing shape, and allometric data for eye span against body length. Each of these three procedures places the new species in the M. apollo species group with Madagopsina parvapollina as its closest relative. New records are presented for M. apographica and M. parvapollina.

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Research Article Tue, 24 Aug 2021 16:05:39 +0300
The genus Dettopsomyia Lamb, 1914 (Diptera, Drosophilidae) from southern China https://zookeys.pensoft.net/article/56996/ ZooKeys 1056: 73-94

DOI: 10.3897/zookeys.1056.56996

Authors: Ya-Lian Wang, Qiao Li, Masanori J. Toda, Jian-Jun Gao

Abstract: The genus Dettopsomyia was established by Lamb in 1914 for a single species, De. formosa described therein. It contains 13 known species recorded from the Old World (the Oriental, Australasian, Palearctic and Afrotropical regions). In the present paper, five new species discovered from southern China are described as members of Dettopsomyia: De. acutipenis Wang & Gao, sp. nov., De. serripenis Wang & Gao, sp. nov., De. discontinua Wang & Gao, sp. nov., De. camelonota Wang, Li & Gao, sp. nov. and De. paranigrovittata Wang, Li & Gao, sp. nov. The new species were delimitated, based on not only morphological characters but also molecular data.

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Research Article Thu, 19 Aug 2021 17:48:53 +0300
Review of the Nordic Gymnocheta Robineau-Desvoidy (Diptera, Tachinidae) with report of two species new to Europe https://zookeys.pensoft.net/article/52761/ ZooKeys 1053: 145-184

DOI: 10.3897/zookeys.1053.52761

Authors: Jaakko Pohjoismäki, Christer Bergström

Abstract: The genus Gymnocheta Robineau-Desvoidy, 1830 (Diptera, Tachinidae) has until now been represented by two species in Europe, G. viridis (Fallén, 1810) and G. magna Zimin, 1958. Two species are newly recorded from Finland and Sweden, Gymnocheta lucida Zimin, 1958 and G. zhelochovtsevi Zimin, 1958, both previously known only from the Russian Far East and Japan. These four European species are redescribed and illustrated, including the first description of the female of G. zhelochovtsevi. A key is provided to seven of the eight described species of Palaearctic Gymnocheta. The holotype of G. viridis was examined and found to differ from the present concept of the species, instead matching the concept of the more recently described G. magna. In the interests of nomenclatural stability, the two names are maintained in their current usage pending a request to the International Commission on Zoological Nomenclature to replace the current holotype of G. viridis with a neotype that corresponds to the long-established concept of that species.

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Research Article Mon, 2 Aug 2021 14:27:09 +0300
Manual of North American Agromyzidae (Diptera, Schizophora), with revision of the fauna of the “Delmarva” states https://zookeys.pensoft.net/article/64603/ ZooKeys 1051: 1-481

DOI: 10.3897/zookeys.1051.64603

Authors: Owen Lonsdale

Abstract: Тhe agromyzid (Diptera: Schizophora: Agromyzidae) fauna of America north of Mexico is described in the first part of this publication, including a genus key and discussions on morphology, life history and classification. The second part is a species-level revision of the family in the “Delmarva” states of the United States of America, that is, of the District of Columbia and the surrounding states of Delaware, Maryland, Virginia and West Virginia. The fauna of this region includes 156 species. This study presents 346 new state and provincial records and 23 new country records, two of which are new continental records (Agromyza abiens Zetterstedt and A. apfelbecki Strobl). Liriomyza endiviae Hering is no longer considered to occur in North America. Fifteen species are newly described: Agromyza echinalis sp. nov., Melanagromyza brunkei sp. nov., M. eoflacensis sp. nov., M. glyptos sp. nov., M. rutella sp. nov., Ophiomyia capitolia sp. nov., O. cuprea sp. nov., O. galiodes sp. nov., O. heleios sp. nov., O. kalia sp. nov., O. laticolis sp. nov., Cerodontha (Poemyza) ungula sp. nov., Phytomyza avicursa sp. nov., P. catenula sp. nov., and P. winkleri sp. nov. Four new species-level synonyms and one genus-level synonym are provided: Agromyza marmorensis Spencer syn. nov. is included as a synonym of A. aristata Malloch; Melanagromyza fastosa Spencer, syn. nov. is included as a junior synonym of Ophiomyia tiliae (Couden); Melanagromyza verbesinae Spencer is considered a synonym of M. vernoniana Steyskal; Phytomyza ranunculoides Spencer, syn. nov. is included as a junior synonym of Phytomyza loewii Hendel; the genus Liomycina Enderlein, syn. nov. is included as a junior synonym of Phytobia Lioy. Ophiomyia ultima (Spencer), comb. nov. is recombined from Melanagromyza. Euhexomyza albicula Spencer, stat. reinst., comb. nov. is resurrected from synonymy with E. winnemanae (Malloch). New host records are given.

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Monograph Thu, 29 Jul 2021 10:17:22 +0300
Psychodidae (Diptera) of Azerbaijan and Georgia – faunistics with biodiversity notes https://zookeys.pensoft.net/article/66063/ ZooKeys 1049: 15-42

DOI: 10.3897/zookeys.1049.66063

Authors: Jan Ježek, Peter Manko, Jozef Oboňa

Abstract: Records of 46 Psychodidae (Sycoracinae 1, Trichomyiinae 1, Psychodinae 44) species/subspecies are presented in this paper based on specimens collected by sweep-netting in Azerbaijan and Georgia in 2019. Nine species are recorded for the first time since their original description; 12 species are new for Transcaucasia; 22 species are new for Azerbaijan; and 17 species are new for Georgia. Saraiella ressli montana Ježek, 1990 is proposed as a synonym of S. ressli Wagner, 1983, syn. nov. Knowledge of some aspects of the ecology and biogeography of selected (especially rare) species has been expanded and a clear pattern was found in species richness, rare species, and new records in relation to land use, habitat diversity, and preservation of the environment surrounding the sampling site.

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Research Article Thu, 15 Jul 2021 13:17:59 +0300
A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group https://zookeys.pensoft.net/article/67564/ ZooKeys 1048: 145-175

DOI: 10.3897/zookeys.1048.67564

Authors: Vladimir I. Lantsov, Valentin E. Pilipenko

Abstract: The caucasica species group in the subgenus Lunatipula is redefined and now consists of five species native to the Caucasus. Tipula (L.) eleniya sp. nov. is described as new to science, and variations in the male terminalia in two populations are noted. Two subspecies (quadridentata quadridentata and quadridentata paupera) are elevated to species rank. Detailed photo’s complement the descriptions of all five species (caucasica, eleniya, paupera, quadridentata, talyshensis), and data on ecology and distribution patterns are included as well as identification keys to males and females. Tipula caucasica is recorded from the West Caucasus and Tipula quadridentata is recorded from Dagestan (Russia) for the first time. Parallel evolution is traced in the male terminalia of the new species and in several non caucasica species group of Palaearctic Lunatipula.

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Research Article Tue, 13 Jul 2021 14:19:43 +0300
A new species of Dactylolabis subgenus Dactylolabis Osten Sacken, 1860 from China (Diptera, Limoniidae) https://zookeys.pensoft.net/article/62033/ ZooKeys 1047: 91-100

DOI: 10.3897/zookeys.1047.62033

Authors: Shang Gao, Bing Zhang, Ding Yang

Abstract: Only two species of Dactylolabis subgenus Dactylolabis Osten Sacken, 1860 were previously known from China. Here, a new species, Dactylolabis (Dactylolabis) wudangensis sp. nov., is reported from China. Dactylolabis (D.) gracilistylus Alexander, 1926 is re-described and illustrated. A key to males of species of the subgenus Dactylolabis from China is presented.

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Research Article Mon, 28 Jun 2021 10:05:07 +0300
Parasitoids of Delia planipalpis (Meigen) and Delia platura (Stein) (Diptera, Anthomyiidae) in Mexico https://zookeys.pensoft.net/article/64405/ ZooKeys 1046: 177-187

DOI: 10.3897/zookeys.1046.64405

Authors: Paulina Nava-Ruiz, Ricardo Meraz-Álvarez, Jorge Valdez-Carrasco, Onésimo Chávez-López, Néstor Bautista-Martínez

Abstract: Among the insect pests that affect crucifer crops in Mexico are Delia planipalpis (Meigen) and D. platura (Stein). They are a threat to the production of these vegetables since the damage they cause directly and indirectly affects yield, quality, and commercialization of these crops. Nevertheless, the existence of natural enemies of these dipterans is still unknown. It is fundamental to determine which parasitoids or predators can be considered possible biological control agents in an integrated pest management program. The sampling sites were located in Guanajuato, Puebla, and the State of Mexico, where plants of Brassica oleracea L. var. italica Plenk and capitata L., B. napus L., and Raphanus sativus L. infested with Delia spp. were selected. The symptoms observed were wilting, yellowish, flaccid leaves and individuals less developed than the rest of the crop. These plants were extracted with their root and the surrounding soil. Also, wild crucifers were collected, such as Raphanus raphanistrum L., Brassica campestris L., and Sisymbrium irio L. The first records of Aphaereta pallipes Say (Hymenoptera, Braconidae), Trybliographa rapae (Westwood) (Hymenoptera, Figitidae), and Aleochara bimaculata Gravenhorst (Coleoptera, Staphylinidae) are reported parasitizing the puparia of these anthomyiid flies on cultivated and wild crucifers. This represents only a starting point for the continuous study of these parasitoids, which is needed to consider them useful for the biological control of D. planipalpis and D. platura.

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Research Article Mon, 21 Jun 2021 18:06:21 +0300
Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data https://zookeys.pensoft.net/article/57052/ ZooKeys 1046: 1-141

DOI: 10.3897/zookeys.1046.57052

Authors: Kurt Jordaens, Georg Goergen, Jeffrey H. Skevington, Scott Kelso, Marc De Meyer

Abstract: The Afrotropical representatives of the hover fly genus Mesembrius Rondani, 1857 (Diptera) are divided into two subgenera, namely Mesembrius s.s. and Vadonimyia Séguy, 1951 and, in this present work, the subgenus Mesembrius s.s. is revised. A total of 23 Mesembrius s.s. species are recognised for the Afrotropics. Known species are re-described and six species new to science are described: Mesembrius arcuatus sp. nov., M. copelandi sp. nov., M. longipilosus sp. nov., M. sulcus sp. nov., M. tibialis sp. nov. and M. vockerothi sp. nov. Mesembrius africanus (Verrall, 1898) is considered a junior synonym of M. senegalensis (Macquart, 1842), M. ctenifer Hull, 1941 a junior synonym of M. caffer (Loew, 1858), M. lagopus (Loew, 1869) a junior synonym of M. capensis (Macquart, 1842) and M. platytarsis Curran, 1929 a junior synonym of M. simplicipes Curran, 1929. The females of Mesembrius chapini Curran, 1939, M. rex Curran, 1927 and M. regulus (Hull, 1937) are described for the first time. Lectotypes are designated for Mesembrius caffer, M. capensis, M. cyanipennis (Bezzi, 1915), M. minor (Bezzi, 1915), M. senegalensis, M. strigilatus (Bezzi, 1912) and M. tarsatus (Bigot, 1883). Separate identification keys for males and females are presented. We obtained 236 DNA barcodes for 18 species. The relationships amongst the different Mesembrius species are briefly discussed, based on morphological and DNA barcode data.

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Research Article Mon, 21 Jun 2021 11:03:05 +0300
New species and records of the subgenus Libnotes (Laosa) Edwards (Diptera, Limoniidae) from China with a key to world species https://zookeys.pensoft.net/article/65906/ ZooKeys 1041: 101-112

DOI: 10.3897/zookeys.1041.65906

Authors: Zehui Kang, Xiao Zhang

Abstract: Twenty species of Libnotes (Laosa) Edwards, 1926 are known worldwide and three are known from China so far. Here, two species of Laosa are added to the Chinese fauna, of which L. (L.) baiyunensis sp. nov. is described and illustrated as new to science, and L. (L.) fuscinervis Brunetti, 1912 is newly recorded from China. Morphologically, the new species is most similar to L. (L.) charmosyne (Alexander, 1958) but can be distinguished by the pleura of the thorax, the relative position of the additional cross veins in cell r3 and r5, and the details of the male genitalia. A key to the world species of Laosa is presented.

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Research Article Tue, 1 Jun 2021 17:03:30 +0300
New or little-known Boreoheptagyia (Diptera, Chironomidae) in China inferred from morphology and DNA barcodes https://zookeys.pensoft.net/article/66527/ ZooKeys 1040: 187-200

DOI: 10.3897/zookeys.1040.66527

Authors: Xiao-Long Lin, Hai-Jun Yu, Xin-Hua Wang, Wen-Jun Bu, Chun-Cai Yan, Wen-Bin Liu

Abstract: The male adult of Boreoheptagyia zhengi Lin & Liu, sp. nov. is described and illustrated based on material collected in China. Associated morphological characteristics and reference to its DNA barcode are provided. Boreoheptagyia kurobebrevis (Sasa & Okazawa, 1992) is newly recorded from China based on both a male and female, with additional associated data on the DNA barcode of the male adult. A neighbor-joining tree based on available Boreoheptagyia DNA barcodes and a key to the adults of Boreoheptagyia from China are given.

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Research Article Fri, 28 May 2021 10:58:11 +0300
Two new crane fly species of the subgenus Vestiplex Bezzi, 1924 (Diptera, Tipulidae, Tipula) from Yunnan and Sichuan, China, with a key to species in the immota species group https://zookeys.pensoft.net/article/64376/ ZooKeys 1040: 33-48

DOI: 10.3897/zookeys.1040.64376

Authors: Qiu-Lei Men, Pavel Starkevich, Aidas Saldaitis

Abstract: Two new crane fly species, Tipula (Vestiplex) gongdangensis sp. nov. and T. (V.) dechangensis sp. nov. are described and illustrated based on materials collected in the Yunnan and Sichuan provinces, China. A key is provided to distinguish males of the new species from those of other species in the T. (V.) immota Alexander, 1935 species group which is proposed here for the first time.

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Research Article Wed, 26 May 2021 12:36:11 +0300
High endemicity in aquatic dance flies of Corsica, France (Diptera, Empididae, Clinocerinae and Hemerodromiinae), with the description of a new species of Chelipoda https://zookeys.pensoft.net/article/66493/ ZooKeys 1039: 177-197

DOI: 10.3897/zookeys.1039.66493

Authors: Marija Ivković, Marija Perović, Patrick Grootaert, Marc Pollet

Abstract: All known records of aquatic dance flies (Empididae, Clinocerinae: 21 species; Hemerodromiinae: eight species) from the island of Corsica (France) are summarized, including previously unpublished data and data on the newly described species Chelipoda puschae Ivković, Perović & Grootaert, sp. nov. This species was collected during the “La Planète Revisitée Corsica 2019” survey and represents the first description of a new species in the genus Chelipoda from the European–Mediterranean region in more than 180 years. A key to European species of Chelipoda is provided. Including the new species, five species are recorded from Corsica for the first time: Dolichocephala malickyi Wagner, 1995, Dolichocephala oblongoguttata (Dale, 1878), Dolichocephala ocellata (Costa, 1854), Chelifera subangusta Collin, 1961, and Hemerodromia unilineata Zetterstedt, 1842. The new species is described and illustrated, and new records of aquatic dance flies from Corsica are given, with new data on 17 species in eight different genera. At present, 29 species of aquatic dance flies are known from Corsica, with 10 species endemic to the island.

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Research Article Tue, 25 May 2021 19:35:04 +0300
Revision of the Afrotropical genus Fainia Zumpt, 1958, with notes on the morphology of Rhiniidae subfamilies (Diptera, Oestroidea) https://zookeys.pensoft.net/article/58539/ ZooKeys 1033: 127-157

DOI: 10.3897/zookeys.1033.58539

Authors: Arianna Thomas-Cabianca, Anabel Martínez-Sánchez, Martin H. Villet, Santos Rojo

Abstract: The taxonomy and diversity of Fainia Zumpt, 1958, an exclusive Afrotropical genus, had not been reviewed recently. The genus included six nominal species, but the status of several of them was debated. Identification of most Fainia species depends on characters of the male terminalia; females are poorly known and, in several cases, are not adequately diagnosed. We conducted a taxonomic revision of the genus and generated identification tools. Based on the study of type material and specimens available in entomological collections in Africa and Europe, we recognise here three of the six species as valid (F. albitarsis (Macquart, 1846), F. elongata (Bezzi, 1908) and F. inexpectata Zumpt, 1973). We also provide an identification key to both sexes, redescriptions of the species, updated distribution records and high resolution photographs of males’ and females’ habitus and male terminalia. The description of Fainia kagerana Lehrer, 2007a nom. nud. is an invalid nomenclatural act in terms of ICZN Article 13.1.1. Based on examinations of their holotypes, F. sambura Lehrer, 2008 syn. nov. is proposed as a junior synonym of F. albitarsis; F. kirinyaga Lehrer, 2007b syn. nov. is proposed as a junior synonym of F. inexpectata; and Fainia giriama Lehrer, 2007b is moved from the genus Fainia to the genus Rhinia Robineau-Desvoidy, as Rhinia giriama (Lehrer, 2007b) comb. nov.. We propose two apomorphies that support the status of the subfamily Rhiniinae.

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Research Article Thu, 22 Apr 2021 20:50:03 +0300
Neotropical Nilothauma Kieffer, 1921 (Diptera, Chironomidae): Key, eleven new species, re-descriptions, new combination and new records https://zookeys.pensoft.net/article/60686/ ZooKeys 1033: 81-125

DOI: 10.3897/zookeys.1033.60686

Authors: Luiz Carlos Pinho, Trond Andersen

Abstract: Nine new species of Nilothauma Kieffer, N. hamadae sp. nov., N. jupau sp. nov., N. karitiana sp. nov., N. leccii sp. nov., N. marianoi sp. nov., N. mateusi sp. nov., N. txukuyana sp. nov., N. werekena sp. nov. and N. yekwana sp. nov. are described and figured, based on adult males collected in Brazil and N. maya sp. nov. on an adult male from Mexico; N. terena sp. nov. is described as male, pupa and larva based on a reared specimen from Brazil. Nilothauma aleta Roback, 1960 and N. duena Roback, 1960 are re-described and recorded from Brazil. Nilothauma longissimum Mendes & Andersen, 2009 is transferred to Beardius Reiss & Sublette, 1985 and the diagnosis of Nilothauma is emended. New records of thirteen Neotropical Nilothauma species are given and a key to the males of all known species of Nilothauma is provided.

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Research Article Thu, 22 Apr 2021 16:25:17 +0300
Taxonomic complexity in the genus Merodon Meigen, 1803 (Diptera, Syrphidae) https://zookeys.pensoft.net/article/62125/ ZooKeys 1031: 85-124

DOI: 10.3897/zookeys.1031.62125

Authors: Ante Vujić, Snežana Radenković, Laura Likov, Sanja Veselić

Abstract: The genus Merodon Meigen, 1803 is distributed across the Palaearctic and Afrotropical Regions. The present work summarizes the knowledge from recent taxonomic and systematic revisions and includes an identification key for the five monophyletic lineages (namely albifrons, aureus, avidus-nigritarsis, desuturinus and natans), 24 species groups, two species subgroups and 10 unplaced species, along with diagnosis and illustrations. A list of 234 taxa, including 194 described and 40 undescribed species, is appended. Most of the species are distributed in the Palaearctic (209 taxa, 181 described), while 27 species (14 described) are known from the Afrotropical Region. Three lineages (aureus, desuturinus and natans) are present in the Afrotropical Region, as well as in the Palaearctic. The Afrotropical melanocerus species group of the desuturinus lineage and the bombiformis species group of the aureus lineage are endemic to the Afrotropical Region, and all other species groups belong to the Palaearctic fauna. The albifrons lineage contains six species groups (albifrons, constans, equestris, geniculatus, ruficornis and rufus) and two unplaced taxa. The aureus lineage includes five species groups (aureus, bombiformis, funestus, nanus and spinitarsis). The avidus-nigritarsis lineage is divided into 10 species groups (aberrans, aurifer, avidus, clavipes, fulcratus, italicus, nigritarsis, pruni, serrulatus and tarsatus) and eight unplaced taxa. The desuturinus lineage contains two species groups: the Afrotropical melanocerus group, with the melanocerus and planifacies subgroups plus the species M. cuthbertsoni Curran, 1939, and the Palaearctic murorum species group. The natans lineage consists of the natans species group plus the species M. segetum Fabricius, 1794.

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Research Article Wed, 14 Apr 2021 23:38:36 +0300
Three new species of the genus Czernyola Bezzi, 1907 (Diptera, Clusiidae) from China https://zookeys.pensoft.net/article/63696/ ZooKeys 1029: 175-183

DOI: 10.3897/zookeys.1029.63696

Authors: Shuai-Lai Yang, Xin-Ming Yin, Yu-Qiang Xi

Abstract: The following three species assigned to the Czernyola biseta group of the genus Czernyola Bezzi, 1907, from China are described as new to science: C. luteigenis sp. nov., C. planipalpis sp. nov., and C. shanxiensis sp. nov. A key to the known species of Czernyola from China is provided.

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Research Article Thu, 8 Apr 2021 15:22:24 +0300
A contribution towards checklist of fungus gnats (Diptera, Diadocidiidae, Ditomyiidae, Bolitophilidae, Keroplatidae, Mycetophilidae) in Georgia, Transcaucasia https://zookeys.pensoft.net/article/63749/ ZooKeys 1026: 69-142

DOI: 10.3897/zookeys.1026.63749

Authors: Olavi Kurina

Abstract: The fungus gnats of Georgia are studied based on 2682 specimens collected from 57 localities during 2011–2019. Altogether, 245 species are recorded including four species of Bolitophilidae, three species of Diadocidiidae, two species of Ditomyiidae, 34 species of Keroplatidae and 202 species of Mycetophilidae. 230 and 188 species are recorded from Georgia and the whole of Transcaucasia for the first time, respectively. Three new species – Sciophila georgei sp. nov., Leia katae sp. nov. and Anatella metae sp. nov. – are described including detailed illustrations of the male terminalia. Photographs are provided for an additional 38 species to highlight a variability of their general facies. Combined with earlier published data, the number of fungus gnat species in Georgia is set at 246. The estimated diversity of fungus gnats in Georgia is calculated using non-parametric methods and discussed with respect to other Western Palaearctic regions.

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Research Article Fri, 26 Mar 2021 16:24:04 +0200
Review of the North American fauna of Drymeia Meigen (Diptera, Muscidae) and evaluation of DNA barcodes for species-level identification in the genus https://zookeys.pensoft.net/article/60393/ ZooKeys 1024: 31-89

DOI: 10.3897/zookeys.1024.60393

Authors: Jade Savage, Vera S. Sorokina

Abstract: The North American fauna of Drymeia Meigen was studied. Four new species are described (Drymeia hucketti sp. nov., Drymeia ponti sp. nov., Drymeia vockerothi sp. nov., Drymeia woodorum sp. nov.), and three new synonymies are proposed: Drymeia amnicola (Huckett, 1966) (= Drymeia rivalis (Huckett, 1966), syn. nov.); Drymeia glacialis (Rondani, 1866) (= Drymeia alpicola (Rondani, 1871), syn. nov.); and Drymeia spinitarsis (Aldrich, 1918) (= Drymeia longiseta Sorokina & Pont, 2015, syn. nov.). An annotated checklist, DNA barcodes (when available), and keys for each sex of the 24 named species of North American Drymeia are provided. The utility of DNA barcodes for the identification of Drymeia species across a wide geographical range was explored using sequences from five countries. A match between morphology and DNA barcodes was found for 71% (22/31) of species studied (including three unnamed taxa). The remaining nine species clustered into two groups of taxa with very little interspecific variation within clusters (groups of two and seven species). Caution is advised against using DNA barcoding as the only determination tool for Drymeia material without prior knowledge of its limitations for certain species groups.

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Research Article Mon, 15 Mar 2021 20:03:28 +0200
Four new West Palaearctic species and new distributional records of Hybotidae (Diptera) https://zookeys.pensoft.net/article/61496/ ZooKeys 1019: 141-162

DOI: 10.3897/zookeys.1019.61496

Authors: Liliana Kanavalová, Patrick Grootaert, Štěpán Kubík, Miroslav Barták

Abstract: Megagrapha starki Barták & Grootaert, sp. nov. (Poland, Russia, Slovakia), Oedalea portugalica Barták & Grootaert, sp. nov. (Portugal), Hybos conicus Grootaert & Barták, sp. nov. (Greece, Turkey), and Platypalpus obscuroides Barták & Grootaert, sp. nov. (Slovakia) are described and illustrated. Diagnostic characters are discussed. The female of Syndyas merzi Shamshev & Grootaert, 2012 is described for the first time. New distributional records are presented: Megagrapha europaea Papp & Földvári, 2001 is first reported from Slovakia and Syndyas merzi Shamshev & Grootaert, 2012 is first reported from Turkey.

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Research Article Mon, 22 Feb 2021 20:50:54 +0200
A new species of Polypedilum (Cerobregma) (Diptera, Chironomidae) from Oriental China https://zookeys.pensoft.net/article/59554/ ZooKeys 1011: 139-148

DOI: 10.3897/zookeys.1011.59554

Authors: Wen-Bin Liu, Yuan Yao, Chun-Cai Yan, Xin-Hua Wang, Xiao-Long Lin

Abstract: Polypedilum (Cerobregma) huapingensis Liu & Lin, sp. nov. is described and illustrated based on an adult male from Huaping National Nature Reserve, Guangxi, China. A DNA barcode analysis, including the known partial COI sequences of species in the Cerobregma subgenus, was conducted. An updated key to adult males of the subgenus Cerobregma is provided.

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Research Article Fri, 22 Jan 2021 21:07:05 +0200
Records of Parochlus steinenii in the Maritime Antarctic and sub-Antarctic regions https://zookeys.pensoft.net/article/56833/ ZooKeys 1011: 63-71

DOI: 10.3897/zookeys.1011.56833

Authors: Melisa Gañan, Tamara Contador, Javier Rendoll, Felipe Simoes, Carolina Pérez, Gillian Graham, Simón Castillo, James Kennedy, Peter Convey

Abstract: This study provides the summary of the reports of the geographical distribution in the Maritime Antarctic and sub-Antarctic regions of Parochlus steinenii (Gercke, 1889) (Diptera, Chironomidae), the only flying insect occurring naturally in the Antarctic continent. The distribution encompasses the South Shetland Islands (Maritime Antarctic), South Georgia (sub-Antarctic), and parts of the Cape Horn Biosphere Reserve (CHBR, southern Chile). In total 78 occurrence records were identified, 53 from our own records, 19 from the literature, and six from other data present in GBIF. Of the 78 records, 66 are from the South Shetland Islands, eight are from South Georgia, and four from the CHBR. This database was developed as one of the main objectives of two Chilean-funded research projects addressing understanding the effects of climate change on sub-Antarctic and Antarctic insects. It provides dataset documenting the distribution of Parochlus steinenii in the Maritime Antarctic, the sub-Antarctic, and the CHBR in southern South America (Chile). The complete dataset is available in Darwin Core Archive format via the Global Biodiversity Information Facility (GBIF).

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Data Paper Mon, 18 Jan 2021 22:13:35 +0200
New species of Limnophyes Eaton (Diptera, Chironomidae) from China and synonymy of L. fuscipygmus Tokunaga, 1940 https://zookeys.pensoft.net/article/58993/ ZooKeys 1011: 51-61

DOI: 10.3897/zookeys.1011.58993

Authors: Wenbin Liu, Cong Zhao, Fanqing Kong, Chuncai Yan, Xinhua Wang

Abstract: Two new species, L. minerus Liu & Yan, sp. nov. and L. subtilus Liu & Yan, sp. nov. are described and illustrated as adult males. Limnophyes minimus (Meigen, 1818) is assigned as a senior synonym of L. minerus Tokunaga, 1940. A key to males of Limnophyes from China is presented.

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Research Article Mon, 18 Jan 2021 15:32:42 +0200
New Metalimnobia crane flies (Diptera, Limoniidae) from China with an update of species distributions https://zookeys.pensoft.net/article/60704/ ZooKeys 1008: 93-105

DOI: 10.3897/zookeys.1008.60704

Authors: Ye Jiang, Xiao Zhang

Abstract: Two new species of the genus Metalimnobia Matsumura, 1911 from southwest China, M. (M.) bicolor sp. nov. and M. (M.) caudifusca sp. nov., are described and illustrated. The new species can be distinguished from congeners by their wing patterns and male genitalia. New distributional data are given for the following species: M. (M.) bifasciata (Schrank, 1781), M. (M.) quadrinotata (Meigen, 1818) and M. (M.) tenua Savchenko, 1976. An updated key for all Chinese Metalimnobia crane flies is presented.

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Research Article Thu, 31 Dec 2020 13:51:07 +0200
Taxonomic revision of the Afrotropical hover fly genus Senaspis Macquart (Diptera, Syrphidae) https://zookeys.pensoft.net/article/56557/ ZooKeys 1003: 83-160

DOI: 10.3897/zookeys.1003.56557

Authors: Marc De Meyer, Georg Goergen, Kurt Jordaens

Abstract: The representatives of the Afrotropical hover fly genus Senaspis Macquart (Diptera) are revised. In total, ten species are recognized. Senaspis apophysata (Bezzi) is herewith placed as junior synonym of S. flaviceps Macquart, S. livida (Bezzi) is herewith placed as junior synonym of S. dentipes (Macquart) and S. griseifacies (Bezzi) is herewith placed as junior synonym of S. haemorrhoa (Gerstaecker). All species are redescribed and an identification key is provided. DNA barcoding analysis (7 species, 64 barcodes) showed that the technique can be used to unambiguously identify the species. The relationships among the different Senaspis species are discussed based on morphological and DNA data.

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Research Article Mon, 14 Dec 2020 23:20:41 +0200
A new species of the genus Noeetomima Enderlein (Diptera, Lauxaniidae) from Guizhou, China with a key to worldwide species https://zookeys.pensoft.net/article/57577/ ZooKeys 1000: 107-123

DOI: 10.3897/zookeys.1000.57577

Authors: Li Shi, Miao Liu, Zheng-Kun Hu

Abstract: A species from the Fanjingshan National Nature Reserve in Guizhou Province, China is described as new to science: Noeetomima huzhengkuni sp. nov. A key to separate worldwide species of Noeetomima, and a list of all species in the genus together with type information, is presented. The habitat of the new species is discussed.

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Research Article Mon, 7 Dec 2020 14:00:39 +0200
Revision of Japanese species of Nipponomyia Alexander, 1924 (Diptera, Pediciidae) https://zookeys.pensoft.net/article/55021/ ZooKeys 1000: 71-105

DOI: 10.3897/zookeys.1000.55021

Authors: Levente-Péter Kolcsár, Daichi Kato, Maribet Gamboa, Kozo Watanabe

Abstract: Japanese species of the genus Nipponomyia Alexander, 1924 are revised. Two new species, Nipponomyia okinawensis Kolcsár & Kato, sp. nov. and N. yakushimensis Kolcsár & Kato, sp. nov. are described from the Ryukyu Islands. Images of habitus and wings, illustrations of male and female terminalia, and distribution maps are provided for the Japanese species. A key to the world species of Nipponomyia is added. DNA barcodes of three Japanese Nipponomyia are provided, representing the first barcodes from the genus.

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Research Article Thu, 3 Dec 2020 21:25:22 +0200
Synopsis of the genus Ulomorpha Osten Sacken, 1869 (Diptera, Limoniidae) in Japan https://zookeys.pensoft.net/article/52831/ ZooKeys 999: 147-163

DOI: 10.3897/zookeys.999.52831

Authors: Daichi Kato, Kozo Watanabe, Levente-Péter Kolcsár

Abstract: Japanese species of the genus Ulomorpha Osten Sacken, 1869 are revised and U. amamiana Kato & Kolcsár, sp. nov. and U. longipenis Kato & Kolcsár, sp. nov. are described. A key to the four Japanese species of the genus is provided, with images of habitus and wings, and drawings of their male terminalia. Ulomorpha amamiana Kato & Kolcsár, sp. nov. is the first representative of the genus discovered from the Oriental region.

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Research Article Mon, 30 Nov 2020 12:30:03 +0200
A third species of the rarely collected Oriental hoverfly genus Furcantenna Cheng, 2008 (Diptera, Syrphidae, Microdontinae) https://zookeys.pensoft.net/article/57087/ ZooKeys 989: 73-78

DOI: 10.3897/zookeys.989.57087

Authors: Menno Reemer

Abstract: Furcantenna malayana sp. nov. is described from Peninsular Malaysia, based on a single female collected in 1962. The other two known species of this genus are also known from single specimens, from Southeastern China and Nepal. A key to the species is given, and the taxonomy and distribution of the genus are discussed.

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Research Article Mon, 9 Nov 2020 11:22:45 +0200
Four new Neotropical species of Eudicrana Loew (Diptera, Mycetophilidae, Sciophilinae) from the Colombian high Andean ecosystems, with comments on the genus https://zookeys.pensoft.net/article/49627/ ZooKeys 988: 129-150

DOI: 10.3897/zookeys.988.49627

Authors: Carolina Henao-Sepúlveda, Marta Wolff, Dalton de Souza Amorim

Abstract: Four new species of the sciophiline genus Eudicrana Loew are described for the Eastern and Central Andes of Colombia–Eudicrana silvaandina sp. nov., E. chingaza sp. nov., E. maculata sp. nov. and E. merizaldei. These are the first species of Eudicrana described from the extreme northern range of the Andes. The altitudinal distribution of these species in the paramos and high Andean forest ecosystems is restricted to 1750–3660 m a.s.l. and some other information on the environment is briefly discussed. A key for the Colombian species of Eudicrana is provided and a discussion is elaborated on the position of these species within the genus.

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Research Article Fri, 6 Nov 2020 12:28:40 +0200
New stiletto flies from New Caledonia (Therevidae, Agapophytinae) https://zookeys.pensoft.net/article/53587/ ZooKeys 984: 83-132

DOI: 10.3897/zookeys.984.53587

Authors: Michael E. Irwin, Shaun L. Winterton, Mark A. Metz

Abstract: Stiletto-flies (Diptera: Therevidae) are highly diverse and species-rich in Australia and New Zealand, yet relatively few species have been recorded from neighbouring Papua New Guinea, Indonesia and throughout the remainder of Oceania. Indeed, in New Caledonia only a single species of the widely distributed Australasian genus Anabarhynchus Macquart (Therevinae) is previously known. Herein we describe two new agapophytine genera (i.e., Jeanchazeauia gen. nov., Calophytus gen. nov.), together comprising nine charismatic new species; this represents a first record of the subfamily from New Caledonia. The new genera and species are described and figured.

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Research Article Wed, 4 Nov 2020 18:15:33 +0200
New and little-known species of Tipula Linnaeus (Diptera, Tipulidae) from Laos, with a new synonym https://zookeys.pensoft.net/article/49781/ ZooKeys 982: 11-31

DOI: 10.3897/zookeys.982.49781

Authors: Bing Zhang, Jinlong Ren, Yan Li, Ding Yang

Abstract: Only seven species of the genus Tipula Linnaeus were previously known to occur in Laos. Here one new species is added to the fauna of Laos, Tipula (Nippotipula) champasakensis sp. nov. Tipula (unplaced) cladomera Alexander, 1936 is designated as a synonym of T. (unplaced) reposita Walker, 1848, syn. nov. A key to species of the genus Tipula from Laos is presented.

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Research Article Mon, 2 Nov 2020 15:55:56 +0200