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  <front>
    <journal-meta>
      <journal-id journal-id-type="publisher-id">2</journal-id>
      <journal-id journal-id-type="index">urn:lsid:arphahub.com:pub:45048D35-BB1D-5CE8-9668-537E44BD4C7E</journal-id>
      <journal-id journal-id-type="aggregator">urn:lsid:zoobank.org:pub:91BD42D4-90F1-4B45-9350-EEF175B1727A</journal-id>
      <journal-title-group>
        <journal-title xml:lang="en">ZooKeys</journal-title>
        <abbrev-journal-title xml:lang="en">ZK</abbrev-journal-title>
      </journal-title-group>
      <issn pub-type="ppub">1313-2989</issn>
      <issn pub-type="epub">1313-2970</issn>
      <publisher>
        <publisher-name>Pensoft Publishers</publisher-name>
      </publisher>
    </journal-meta>
    <article-meta>
      <article-id pub-id-type="doi">10.3897/zookeys.1071.66750</article-id>
      <article-id pub-id-type="publisher-id">66750</article-id>
      <article-categories>
        <subj-group subj-group-type="heading">
          <subject>Research Article</subject>
        </subj-group>
        <subj-group subj-group-type="biological_taxon">
          <subject>Diptera</subject>
          <subject>Limoniidae</subject>
        </subj-group>
        <subj-group subj-group-type="scientific_subject">
          <subject>Agriculture and Forestry</subject>
          <subject>Biodiversity &amp; Conservation</subject>
        </subj-group>
        <subj-group subj-group-type="geological_era">
          <subject>Cenozoic</subject>
        </subj-group>
        <subj-group subj-group-type="geographical_area">
          <subject>Asia</subject>
        </subj-group>
      </article-categories>
      <title-group>
        <article-title><italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>)</tp:taxon-name></italic> Macquart (<tp:taxon-name><tp:taxon-name-part taxon-name-part-type="order">Diptera</tp:taxon-name-part></tp:taxon-name>, <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="family">Limoniidae</tp:taxon-name-part></tp:taxon-name>) from Xizang, China</article-title>
      </title-group>
      <contrib-group content-type="authors">
        <contrib contrib-type="author" corresp="no">
          <name name-style="western">
            <surname>Zhang</surname>
            <given-names>Bing</given-names>
          </name>
          <xref ref-type="aff" rid="A1">1</xref>
          <role content-type="http://credit.niso.org/contributor-roles/writing-original-draft/">Writing - original draft</role>
        </contrib>
        <contrib contrib-type="author" corresp="no">
          <name name-style="western">
            <surname>Yang</surname>
            <given-names>Qicheng</given-names>
          </name>
          <uri content-type="orcid">https://orcid.org/0000-0003-4378-053X</uri>
          <xref ref-type="aff" rid="A2">2</xref>
          <role content-type="http://credit.niso.org/contributor-roles/writing-review-editing/">Writing - review and editing</role>
        </contrib>
        <contrib contrib-type="author" corresp="no">
          <name name-style="western">
            <surname>Li</surname>
            <given-names>Yan</given-names>
          </name>
          <xref ref-type="aff" rid="A3">3</xref>
          <role content-type="http://credit.niso.org/contributor-roles/writing-review-editing/">Writing - review and editing</role>
        </contrib>
        <contrib contrib-type="author" corresp="yes">
          <name name-style="western">
            <surname>Yang</surname>
            <given-names>Ding</given-names>
          </name>
          <email xlink:type="simple">dyangcau@126.com</email>
          <xref ref-type="aff" rid="A1">1</xref>
          <role content-type="http://credit.niso.org/contributor-roles/writing-review-editing/">Writing - review and editing</role>
        </contrib>
      </contrib-group>
      <aff id="A1">
        <label>1</label>
        <addr-line content-type="verbatim">Department of Entomology, College of Plant Protection, China Agricultural University, Beijing 100193, China</addr-line>
        <institution>China Agricultural University</institution>
        <addr-line content-type="city">Beijing</addr-line>
        <country>China</country>
      </aff>
      <aff id="A2">
        <label>2</label>
        <addr-line content-type="verbatim">Hubei Insect Resources Utilization and Sustainable Pest Management Key Laboratory, College of Plant Science &amp; Technology of Huazhong Agriculture University, Wuhan, 430070, China</addr-line>
        <institution>Huazhong Agriculture University</institution>
        <addr-line content-type="city">Wuhan</addr-line>
        <country>China</country>
      </aff>
      <aff id="A3">
        <label>3</label>
        <addr-line content-type="verbatim">College of Plant Protection, Shenyang Agricultural University, Shenyang, 110866, China</addr-line>
        <institution>Shenyang Agricultural University</institution>
        <addr-line content-type="city">Shenyang</addr-line>
        <country>China</country>
      </aff>
      <author-notes>
        <fn fn-type="corresp">
          <p>Corresponding author: Ding Yang (<email xlink:type="simple">dyangcau@126.com</email>)</p>
        </fn>
        <fn fn-type="edited-by">
          <p>Academic editor: Gunnar Kvifte</p>
        </fn>
      </author-notes>
      <pub-date pub-type="collection">
        <year>2021</year>
      </pub-date>
      <pub-date pub-type="epub">
        <day>18</day>
        <month>11</month>
        <year>2021</year>
      </pub-date>
      <volume>1071</volume>
      <fpage>155</fpage>
      <lpage>173</lpage>
      <uri content-type="arpha" xlink:href="http://openbiodiv.net/75FFBB9E-CC4A-5620-A231-4B054839CE20">75FFBB9E-CC4A-5620-A231-4B054839CE20</uri>
      <uri content-type="zenodo_dep_id" xlink:href="https://zenodo.org/record/5728542">5728542</uri>
      <history>
        <date date-type="received">
          <day>31</day>
          <month>03</month>
          <year>2021</year>
        </date>
        <date date-type="accepted">
          <day>13</day>
          <month>10</month>
          <year>2021</year>
        </date>
      </history>
      <permissions>
        <copyright-statement>Bing Zhang, Qicheng Yang, Yan Li, Ding Yang</copyright-statement>
        <license license-type="creative-commons-attribution" xlink:href="http://creativecommons.org/licenses/by/4.0/" xlink:type="simple">
          <license-p>This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.</license-p>
        </license>
      </permissions>
      <abstract>
        <label>Abstract</label>
        <p>One new species of the subgenus <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part></tp:taxon-name> Macquart,1838, <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="xizangensis">xizangensis</tp:taxon-name-part></tp:taxon-name><bold>sp. nov.</bold> is described and illustrated from Xizang, China. The following four species are re-described and reported from Xizang for the first time: <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">H.</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">E.</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="latigrisea">latigrisea</tp:taxon-name-part></tp:taxon-name> Alexander, 1971, <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">H.</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">E.</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="nepalensis">nepalensis</tp:taxon-name-part></tp:taxon-name> (Westwood, 1836), <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">H.</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">E.</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="paragnava">paragnava</tp:taxon-name-part></tp:taxon-name> Alexander, 1973 and <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">H.</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">E.</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="perhirsuta">perhirsuta</tp:taxon-name-part></tp:taxon-name> Alexander, 1973. A key to the species of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Eriocera">Eriocera</tp:taxon-name-part></tp:taxon-name></italic> from Xizang is presented.</p>
      </abstract>
      <kwd-group>
        <label>Keywords</label>
        <kwd>Biodiversity</kwd>
        <kwd>craneflies</kwd>
        <kwd>
          <tp:taxon-name>
            <tp:taxon-name-part taxon-name-part-type="subfamily">Limnophilinae</tp:taxon-name-part>
          </tp:taxon-name>
        </kwd>
        <kwd>systematics</kwd>
        <kwd>taxonomy</kwd>
        <kwd>Tibet</kwd>
      </kwd-group>
    </article-meta>
    <notes>
      <sec sec-type="Citation" id="SECID0E5H">
        <title>Citation</title>
        <p>Zhang B, Yang Q, Li Y, Yang D (2021) <italic>Hexatoma (Eriocera)</italic> Macquart (Diptera, Limoniidae) from Xizang, China. ZooKeys 1071: 155–173. <ext-link xlink:type="simple" ext-link-type="doi" xlink:href="10.3897/zookeys.1071.66750">https://doi.org/10.3897/zookeys.1071.66750</ext-link></p>
      </sec>
    </notes>
  </front>
  <body>
    <sec sec-type="Introduction" id="SECID0E6AAC">
      <title>Introduction</title>
      <p><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Eriocera">Eriocera</tp:taxon-name-part></tp:taxon-name> Macquart, 1838 is a subgenus of the genus <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part></tp:taxon-name> Latreille, 1809 in the family <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="family">Limoniidae</tp:taxon-name-part></tp:taxon-name>. It is distributed worldwide with 563 known species and subspecies, of which 69 taxa are from the Palaearctic Realm, 34 taxa from the Nearctic Realm, 143 taxa from the Neotropical Realm, 30 taxa from the Afrotropical Realm, 290 taxa from the Oriental region, and five taxa from the Australasian/Oceania realms (<xref ref-type="bibr" rid="B16">Oosterbroek 2021</xref>). The subgenus is thus large and morphologically diverse, and was confirmed to be non-monophyletic by <xref ref-type="bibr" rid="B20">Ribeiro (2008)</xref>. It is characterized by the following characters: body medium to large sized; antenna with four to ten flagellomeres; wings often unpatterned through variously darkened, or with a conspicuous hyaline and yellow cross banded pattern, or abundantly spotted and dotted with brown; cell <italic>dm</italic> present; two or three branches of M reaching margin; clasper of gonostylus narrowed apically into a long curved spine; lobe of gonostylus short and stout with setae; gonocoxite moderately stubby or elongate cylindrical; interbase usually cylindrical, or triangular with a sharp spine at base, or two-layered membranous structure with spine at apex; aedeagus usually short and relatively inconspicuous, or directed ventrally (<xref ref-type="bibr" rid="B8">Alexander and Lloyd 1914</xref>; <xref ref-type="bibr" rid="B11">Edwards 1921</xref>; <xref ref-type="bibr" rid="B2">Alexander 1948</xref>; <xref ref-type="bibr" rid="B7">Alexander and Byers 1981</xref>; <xref ref-type="bibr" rid="B21">Savchenko 1986</xref>; <xref ref-type="bibr" rid="B17">Podenas et al. 2006</xref>; <xref ref-type="bibr" rid="B20">Ribeiro 2008</xref>; <xref ref-type="bibr" rid="B18">Podeniene and Gelhaus 2015</xref>).</p>
      <p>So far, only the subgenus <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part></tp:taxon-name> were known to occur in Xizang (<xref ref-type="bibr" rid="B14">Men and Yu 2015</xref>; <xref ref-type="bibr" rid="B16">Oosterbroek 2021</xref>): <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">H.</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">E.</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="lanigera">lanigera</tp:taxon-name-part></tp:taxon-name> Alexander, 1933, <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">H.</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">E.</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="mediofila">mediofila</tp:taxon-name-part></tp:taxon-name> Alexander, 1933, <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">H.</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">E.</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="nudivena">nudivena</tp:taxon-name-part></tp:taxon-name> Alexander, 1933 and <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">H.</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">E.</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="tibetana">tibetana</tp:taxon-name-part></tp:taxon-name> Alexander, 1933. To enrich the knowledge of the species composition of craneflies in Xizang, we conducted a scientific survey of craneflies in Xizang from 1978 to 2019. Presently, five species including one new species of the subgenus <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part></tp:taxon-name> are added to the fauna of Xizang. The following three species are reported from China for the first time: <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">H.</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">E.</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="latigrisea">latigrisea</tp:taxon-name-part></tp:taxon-name> Alexander, 1971, <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">H.</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">E.</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="paragnava">paragnava</tp:taxon-name-part></tp:taxon-name> Alexander, 1973 and <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">H.</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">E.</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="perhirsuta">perhirsuta</tp:taxon-name-part></tp:taxon-name> Alexander, 1973. <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">E.</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="nepalensis">nepalensis</tp:taxon-name-part></tp:taxon-name> (Westwood, 1836) is for the first time reported from Xizang. <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="xizangensis">xizangensis</tp:taxon-name-part></tp:taxon-name> sp. nov. is described and illustrated from Xizang. A key to the species of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Eriocera">Eriocera</tp:taxon-name-part></tp:taxon-name></italic> from Xizang is presented.</p>
    </sec>
    <sec sec-type="materials|methods" id="SECID0E2HAC">
      <title>Materials and methods</title>
      <p>The specimens were studied and illustrated with a ZEISS Stemi 2000-c stereo microscope. Details of coloration were checked in specimens immersed in 75% ethyl alcohol. Male genitalia were prepared by macerating the apical portion of the abdomen in cold 10% NaOH for 12–15 hours. After examination, it was transferred to fresh glycerine (C<sub>3</sub>H<sub>8</sub>O<sub>3</sub>) and stored in a microvial pinned below the specimen. The specimens studied, which were collected in Xizang are deposited in the Entomological Museum of <named-content xlink:type="simple" content-type="institution" xlink:href="http://grbio.org/institution/china-agricultural-university" id="NCID0E1HAC">China Agricultural University</named-content> (<named-content content-type="dwc:institutional_code" xlink:title="China Agricultural University" xlink:href="http://grbio.org/institution/china-agricultural-university">CAU</named-content>), Beijing, China.</p>
      <p>Holotype material of <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="latigrisea">latigrisea</tp:taxon-name-part></tp:taxon-name> Alexander, 1971 used in this paper was borrowed from the National Museum of Natural History, <named-content xlink:type="simple" content-type="institution" xlink:href="http://grbio.org/institution/smithsonian-institution" id="NCID0EBJAC">Smithsonian Institution</named-content>, Washington, DC, USA (<named-content content-type="dwc:institutional_code" xlink:title="Smithsonian Institution, National Museum of Natural History" xlink:href="http://grbio.org/institution/smithsonian-institution-national-museum-natural-history-0">USNM</named-content>) and holotype material of <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="nepalensis">nepalensis</tp:taxon-name-part></tp:taxon-name> (Westwood, 1836) was borrowed from the <named-content xlink:type="simple" content-type="institution" xlink:href="http://grbio.org/institution/natural-history-museum-london" id="NCID0EBKAC">Natural History Museum, London, UK</named-content> (<abbrev content-type="institution" xlink:title="Natural History Museum" id="ABBRID0ENJAC">BMNH</abbrev>). The terminology applied to the wing veins follows the interpretation of <xref ref-type="bibr" rid="B10">de Jong (2017)</xref>. Terminology of the male terminalia follows <xref ref-type="bibr" rid="B19">Ribeiro (2006</xref>, <xref ref-type="bibr" rid="B20">2008</xref>). The following abbreviations in figures are used: 9s = ninth sternite, 9t = ninth tergite, goncx = gonocoxite, cgonst = clasper of gonostylus, lgonst = lobe of gonostylus, aed = aedeagus, ce = cercus, hy = hypogynial valve.</p>
    </sec>
    <sec sec-type="Taxonomy" id="SECID0E4JAC">
      <title>Taxonomy</title>
      <sec sec-type="Key to species of subgenus Eriocera from Xizang, China (adult)" id="SECID0EBKAC">
        <title>Key to species of subgenus <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part></tp:taxon-name> from Xizang, China (adult)</title>
        <table-wrap content-type="key" position="anchor" orientation="portrait">
          <table id="TID0EWEAC" rules="all">
            <tbody>
              <tr>
                <td rowspan="1" colspan="1">1</td>
                <td rowspan="1" colspan="1">Wing with cell <italic>M<sub>1</sub></italic> (Figs 34, 40)</td>
                <td rowspan="1" colspan="1">
                  <bold>2</bold>
                </td>
              </tr>
              <tr>
                <td rowspan="1" colspan="1">–</td>
                <td rowspan="1" colspan="1">Wing without cell <italic>M<sub>1</sub></italic> (Figs 1, 3, 9, 10, 12, 16, 22, 24, 28, 46, 48)</td>
                <td rowspan="1" colspan="1">
                  <bold>4</bold>
                </td>
              </tr>
              <tr>
                <td rowspan="1" colspan="1">2</td>
                <td rowspan="1" colspan="1">Antenna of male approximately as long as body (Alexander, 1933: 150)</td>
                <td rowspan="1" colspan="1">
                  <bold><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">H.</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">E.</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="tibetana">tibetana</tp:taxon-name-part></tp:taxon-name> Alexander, 1933</bold>
                </td>
              </tr>
              <tr>
                <td rowspan="1" colspan="1">– Antenna of male approximately three times as long as body (Figs 34; Alexander, 1973: 8; Alexander, 1933: 149)</td>
                <td rowspan="1" colspan="1">
                  <bold>3</bold>
                </td>
              </tr>
              <tr>
                <td rowspan="1" colspan="1">3</td>
                <td rowspan="1" colspan="1">Femora yellow (Figs 34; Alexander, 1973: 9); <italic>R<sub>2+3</sub></italic> two times longer than <italic>R<sub>2+3+4</sub></italic> (Figs 34, 40; Alexander, 1973: 4, fig. 6)</td>
                <td rowspan="1" colspan="1">
                  <bold><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">H.</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">E.</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="perhirsuta">perhirsuta</tp:taxon-name-part></tp:taxon-name> Alexander, 1973</bold>
                </td>
              </tr>
              <tr>
                <td rowspan="1" colspan="1">–</td>
                <td rowspan="1" colspan="1">Basal 1/3 of femora yellow, outer 2/3 black (Alexander, 1933: 149); <italic>R<sub>2+3</sub></italic> as long as <italic>R<sub>2+3+4</sub></italic> (Alexander, 1933: plate I; Fig. 11)</td>
                <td rowspan="1" colspan="1">
                  <bold><italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">H.</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">E.</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="lanigera">lanigera</tp:taxon-name-part></tp:taxon-name> Alexander</italic> , 1933</bold>
                </td>
              </tr>
              <tr>
                <td rowspan="1" colspan="1">4</td>
                <td rowspan="1" colspan="1">Wing without markings (Figs 22, 24, 28)</td>
                <td rowspan="1" colspan="1">
                  <bold>5</bold>
                </td>
              </tr>
              <tr>
                <td rowspan="1" colspan="1">–</td>
                <td rowspan="1" colspan="1">Wing bicolorous, with markings (Figs 1, 3, 9, 10, 12, 16, 46, 48)</td>
                <td rowspan="1" colspan="1">
                  <bold>7</bold>
                </td>
              </tr>
              <tr>
                <td rowspan="1" colspan="1">5 Wing without stigma (Alexander, 1933: 158); <italic>R<sub>2</sub></italic> longer than <italic>R<sub>2+3</sub></italic> (Alexander, 1933: plate I. Fig. 19)</td>
                <td rowspan="1" colspan="1">
                  <bold><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">H.</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">E.</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="nudivena">nudivena</tp:taxon-name-part></tp:taxon-name> Alexander, 1933</bold>
                </td>
              </tr>
              <tr>
                <td rowspan="1" colspan="1">– Wing with stigma; <italic>R<sub>2</sub></italic> shorter than <italic>R<sub>2+3</sub></italic> (Figs 22, 24, 28)</td>
                <td rowspan="1" colspan="1">
                  <bold>6</bold>
                </td>
              </tr>
              <tr>
                <td rowspan="1" colspan="1">6</td>
                <td rowspan="1" colspan="1">Antenna of male approximately as long as body (Alexander, 1933: 151); <italic>R<sub>2+3</sub></italic> longer than <italic>R<sub>2+3+4</sub></italic> (Alexander, 1933: plate I; Fig. 13)</td>
                <td rowspan="1" colspan="1">
                  <bold><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">H.</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">E.</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="mediofila">mediofila</tp:taxon-name-part></tp:taxon-name> Alexander, 1933</bold>
                </td>
              </tr>
              <tr>
                <td rowspan="1" colspan="1">–</td>
                <td rowspan="1" colspan="1">Antenna of male approximately three times longer than body (Figs 22; Alexander, 1973: 8); <italic>R<sub>2+3</sub></italic> shorter than <italic>R<sub>2+3+4</sub></italic> (Figs 22, 24, 28; Alexander, 1973: 4, fig. 5)</td>
                <td rowspan="1" colspan="1">
                  <bold><italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">H.</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">E.</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="paragnava">paragnava</tp:taxon-name-part></tp:taxon-name> Alexander</italic> , 1973</bold>
                </td>
              </tr>
              <tr>
                <td rowspan="1" colspan="1">7</td>
                <td rowspan="1" colspan="1">Male terminalia yellow (Fig. 46); wing with many hyaline markings; <italic>R<sub>2</sub></italic> contacts vein <italic>R<sub>2+3+4</sub></italic> (Figs 46, 48)</td>
                <td rowspan="1" colspan="1">
                  <bold><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">H.</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">E.</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="xizangensis">xizangensis</tp:taxon-name-part></tp:taxon-name> sp. nov.</bold>
                </td>
              </tr>
              <tr>
                <td rowspan="1" colspan="1">–</td>
                <td rowspan="1" colspan="1">Male terminalia black or dark brown (Figs 1, 9, 10); wing with a whitened or hyaline marking before discal area; <italic>R<sub>2</sub></italic> contacts vein <italic>R<sub>2+3</sub></italic> (Figs 1, 3, 9, 12, 16)</td>
                <td rowspan="1" colspan="1">
                  <bold>8</bold>
                </td>
              </tr>
              <tr>
                <td rowspan="1" colspan="1">8</td>
                <td rowspan="1" colspan="1">Antenna blackish brown; femora blackish brown; abdomen of male with four pale white gray markings (Figs 9, 10) ...... <bold><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">H.</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">E.</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="nepalensis">nepalensis</tp:taxon-name-part></tp:taxon-name> (Westwood, 1836)</bold></td>
              </tr>
              <tr>
                <td rowspan="1" colspan="1">–</td>
                <td rowspan="1" colspan="1">Scape and pedicel blackish brown, flagellum reddish yellow except outer two segments dark brown; basal 5/6 of femora yellow, outer 1/6 dull black; abdomen glossy black or dull black (Fig. 1; Alexander, 1971: 116)</td>
                <td rowspan="1" colspan="1">
                  <bold><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">H.</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">E.</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="latigrisea">latigrisea</tp:taxon-name-part></tp:taxon-name> Alexander, 1971</bold>
                </td>
              </tr>
              <tr>
                <td rowspan="1" colspan="1"/>
              </tr>
            </tbody>
          </table>
        </table-wrap>
        <tp:taxon-treatment>
          <tp:treatment-meta>
            <kwd-group>
              <label>Taxon classification</label>
              <kwd>
                <named-content content-type="kingdom" xlink:type="simple">Animalia</named-content>
              </kwd>
              <kwd>
                <named-content content-type="order" xlink:type="simple">Diptera</named-content>
              </kwd>
              <kwd>
                <named-content content-type="family" xlink:type="simple">Limoniidae</named-content>
              </kwd>
            </kwd-group>
          </tp:treatment-meta>
          <tp:nomenclature>
            <tp:taxon-name><object-id content-type="arpha">A0260828-ED4A-502C-9790-A6B7A4713D4F</object-id>
              <tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="latigrisea">latigrisea</tp:taxon-name-part></tp:taxon-name>
            <tp:taxon-authority>Alexander, 1971</tp:taxon-authority>
            <tp:nomenclature-citation-list>
              <tp:nomenclature-citation>
                <tp:taxon-name>
                  <tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="latigrisea">latigrisea</tp:taxon-name-part></tp:taxon-name>
                <comment>Alexander, 1971: 116. Type locality: India, Assam</comment>
              </tp:nomenclature-citation>
            </tp:nomenclature-citation-list>
          </tp:nomenclature>
          <tp:treatment-sec sec-type="material" id="SECID0EQJAE">
            <title>Specimens examined.</title>
            <p>2 males (<named-content content-type="dwc:institutional_code" xlink:title="China Agricultural University" xlink:href="http://grbio.org/institution/china-agricultural-university">CAU</named-content>), China: Xizang, Motuo, 80K, 2014.VII.31, Yan Li (light trap). <bold><italic>Holotype</italic></bold>: male, Kujjalong, Kameng, North East Frontier Agency, Assam, 4500 feet, June 28–30, 1961 (Schmid).</p>
          </tp:treatment-sec>
          <tp:treatment-sec sec-type="diagnosis" id="SECID0E4JAE">
            <title>Diagnosis</title>
            <p>. Antenna has 8 segments. Wing is brown, but narrowly yellow at base; anal cells are much paler; cells <italic>C</italic> and <italic>Sc</italic> are dark yellow; a narrow white discal area is before cord, including cell <italic>R<sub>1</sub></italic> to cell <italic>M</italic>; <italic>m-cu</italic> is nearly at middle of cell <italic>dm</italic>. Posterior margin of male ninth tergite has a V-shaped notch, both sides of notch have a process; interbase is triangular, swollen at base.</p>
          </tp:treatment-sec>
          <tp:treatment-sec sec-type="redescription" id="SECID0ERKAE">
            <title>Redescription</title>
            <p>. Male (<italic>N</italic> = 2): Body length 18.0–19.8 mm, wing length 14.0–16.7 mm, antenna length 3.6–4.1 mm.</p>
            <p>Thorax (Figs 1, 2) dull black with blackish brown setae. Legs with blackish brown setae; coxae and trochanters blackish brown; basal 5/6 of femora yellow, outer 1/6 dull black; tibiae obscure yellow, black apically; tarsi dark brown. Wing (Figs 1, 3) brown, narrowly yellowed at base; anal cells much paler; cells <italic>C</italic> and <italic>Sc</italic> dark yellow; a narrow white discal area before cord, including cell <italic>R<sub>1</sub></italic> to cell <italic>M</italic>; veins brown, more yellow in brightened areas. Venation: <italic>R<sub>2</sub></italic> moderately oblique, <italic>R<sub>2+3</sub></italic> relatively short, shorter than <italic>R<sub>2</sub></italic>; cell <italic>M<sub>1</sub></italic> lacking; <italic>m-cu</italic> nearly at middle of cell <italic>dm</italic>. Halter (Figs 1, 2) length approximately 2.2 mm, halter stem pale brown with brown setae; knob brown with blackish brown setae.</p>
            <fig id="F1" position="float" orientation="portrait">
              <object-id content-type="doi">10.3897/zookeys.1071.66750.figures1-3</object-id>
              <object-id content-type="zenodo_dep_id">5728544</object-id>
              <object-id content-type="arpha">70B2040C-7A63-55B9-9A6C-53652AAC1DAB</object-id>
              <label>Figures 1–3.</label>
              <caption>
                <p><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="latigrisea">latigrisea</tp:taxon-name-part></tp:taxon-name> Alexander, male <bold>1</bold> habitus, lateral view <bold>2</bold> male head and thorax, dorsal view <bold>3</bold> right wing. Scale bars: 1.0 mm (1–3).</p>
              </caption>
              <graphic xlink:href="zookeys-1071-155_article-66750__-g001.jpg" position="float" orientation="portrait" xlink:type="simple" id="oo_612049.jpg">
                <uri content-type="original_file">https://binary.pensoft.net/fig/612049</uri>
              </graphic>
            </fig>
            <p>Abdomen (Fig. 1) with short black setae. Segments 1–6 extensively glossy black, segments 7–8 dull black.</p>
            <p>Male terminalia (Figs 1, 4–8) with 180° rotation, dull black with black setae. Posterior margin of ninth tergite with a V-shaped notch, both sides of notch with a lateral projection; posterior margin of ninth sternite with a deep V-shaped shallow; gonocoxite moderately stubby; clasper of gonostylus with long setae at base, slender, terminal spine decurved; lobe of gonostylus short and stout, terminal margin swollen with long setae; interbase triangular; aedeagus longer, apically directed ventrally.</p>
            <fig id="F2" position="float" orientation="portrait">
              <object-id content-type="doi">10.3897/zookeys.1071.66750.figures4-8</object-id>
              <object-id content-type="zenodo_dep_id">5728550</object-id>
              <object-id content-type="arpha">71C68B68-7699-5AFA-9B6B-DA6281D88ACD</object-id>
              <label>Figures 4–8.</label>
              <caption>
                <p><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="latigrisea">latigrisea</tp:taxon-name-part></tp:taxon-name> Alexander, male <bold>4</bold> terminalia, dorsal view <bold>5</bold> terminalia, ventral view <bold>6</bold> aedeagal complex, dorsal view <bold>7</bold> aedeagal complex, lateral view <bold>8</bold> aedeagal complex, ventral view. Scale bars: 0.5 mm (4–8).</p>
              </caption>
              <graphic xlink:href="zookeys-1071-155_article-66750__-g002.jpg" position="float" orientation="portrait" xlink:type="simple" id="oo_612050.jpg">
                <uri content-type="original_file">https://binary.pensoft.net/fig/612050</uri>
              </graphic>
            </fig>
          </tp:treatment-sec>
          <tp:treatment-sec sec-type="Female" id="SECID0EBOAE">
            <title>Female</title>
            <p>. Unknown.</p>
          </tp:treatment-sec>
          <tp:treatment-sec sec-type="distribution" id="SECID0EGOAE">
            <title>Distribution</title>
            <p>. India (Assam), China (Xizang).</p>
          </tp:treatment-sec>
          <tp:treatment-sec sec-type="remarks" id="SECID0ELOAE">
            <title>Remarks</title>
            <p>. This species was known previously only from India. With the present contribution it is recorded from China for the first time.</p>
          </tp:treatment-sec>
        </tp:taxon-treatment>
        <tp:taxon-treatment>
          <tp:treatment-meta>
            <kwd-group>
              <label>Taxon classification</label>
              <kwd>
                <named-content content-type="kingdom" xlink:type="simple">Animalia</named-content>
              </kwd>
              <kwd>
                <named-content content-type="order" xlink:type="simple">Diptera</named-content>
              </kwd>
              <kwd>
                <named-content content-type="family" xlink:type="simple">Limoniidae</named-content>
              </kwd>
            </kwd-group>
          </tp:treatment-meta>
          <tp:nomenclature>
            <tp:taxon-name><object-id content-type="arpha">EF46BDA6-669B-5CC3-9870-F48283AA5FA1</object-id>
              <tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="nepalensis">nepalensis</tp:taxon-name-part></tp:taxon-name>
            <tp:taxon-authority>(Westwood, 1836)</tp:taxon-authority>
            <tp:nomenclature-citation-list>
              <tp:nomenclature-citation>
                <tp:taxon-name>
                  <tp:taxon-name-part taxon-name-part-type="genus" reg="Caloptera">Caloptera</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="nepalensis">nepalensis</tp:taxon-name-part></tp:taxon-name>
                <comment>Westwood, 1836: 681. Type locality: Nepal.</comment>
              </tp:nomenclature-citation>
              <tp:nomenclature-citation>
                <tp:taxon-name>
                  <tp:taxon-name-part taxon-name-part-type="genus" reg="Pterocosmus">Pterocosmus</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="velutinus">velutinus</tp:taxon-name-part></tp:taxon-name>
                <comment>Walker, 1848: 79. Type locality: Nepal.</comment>
              </tp:nomenclature-citation>
              <tp:nomenclature-citation>
                <tp:taxon-name>
                  <tp:taxon-name-part taxon-name-part-type="genus" reg="Lechria">Lechria</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="nepalensis">nepalensis</tp:taxon-name-part></tp:taxon-name>
                <comment>Brunetti, 1918: 317. Type locality: Nepal (Katmandu).</comment>
              </tp:nomenclature-citation>
              <tp:nomenclature-citation>
                <tp:taxon-name>
                  <tp:taxon-name-part taxon-name-part-type="genus" reg="Eriocera">Eriocera</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="nepalensis">nepalensis</tp:taxon-name-part></tp:taxon-name>
                <comment>Westw. (= <italic>velutina</italic>, Walk.): <xref ref-type="bibr" rid="B11">Edwards 1921</xref>: 76.</comment>
              </tp:nomenclature-citation>
              <tp:nomenclature-citation>
                <tp:taxon-name>
                  <tp:taxon-name-part taxon-name-part-type="genus" reg="Pterocosmus">Pterocosmus</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="velutinus">velutinus</tp:taxon-name-part></tp:taxon-name>
                <comment>(<tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Eriocera">Eriocera</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="nepalensis">nepalensis</tp:taxon-name-part></tp:taxon-name>): <xref ref-type="bibr" rid="B11">Edwards 1921</xref>: 99.</comment>
              </tp:nomenclature-citation>
              <tp:nomenclature-citation>
                <tp:taxon-name>
                  <tp:taxon-name-part taxon-name-part-type="genus" reg="Lechria">Lechria</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="nepalensis">nepalensis</tp:taxon-name-part></tp:taxon-name>
                <comment>: <xref ref-type="bibr" rid="B12">Edwards 1924</xref>: 301.</comment>
              </tp:nomenclature-citation>
              <tp:nomenclature-citation>
                <tp:taxon-name>
                  <tp:taxon-name-part taxon-name-part-type="genus" reg="Trichoneura">Trichoneura</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Xipholimnobia">Xipholimnobia</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="nepalensis">nepalensis</tp:taxon-name-part></tp:taxon-name>
                <comment>: <xref ref-type="bibr" rid="B3">Alexander 1963</xref>: 24; <xref ref-type="bibr" rid="B4">Alexander 1968</xref>: 95.</comment>
              </tp:nomenclature-citation>
              <tp:nomenclature-citation>
                <tp:taxon-name>
                  <tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="nepalensis">nepalensis</tp:taxon-name-part></tp:taxon-name>
                <comment>: <xref ref-type="bibr" rid="B15">Mitra et al. 2006</xref>: 234; <xref ref-type="bibr" rid="B14">Men and Yu 2015</xref>: 165.</comment>
              </tp:nomenclature-citation>
            </tp:nomenclature-citation-list>
          </tp:nomenclature>
          <tp:treatment-sec sec-type="material" id="SECID0ENUAE">
            <title>Specimens examined</title>
            <p>. 1 male (<named-content content-type="dwc:institutional_code" xlink:title="China Agricultural University" xlink:href="http://grbio.org/institution/china-agricultural-university">CAU</named-content>), China: Xizang, Chayu, 1570 m, 1978.VI.25, Fasheng Li. 1 male (<named-content content-type="dwc:institutional_code" xlink:title="China Agricultural University" xlink:href="http://grbio.org/institution/china-agricultural-university">CAU</named-content>), China: Xizang, Chayu, 1700 m, 1978.VI.26, Fasheng Li. 1 male (<named-content content-type="dwc:institutional_code" xlink:title="China Agricultural University" xlink:href="http://grbio.org/institution/china-agricultural-university">CAU</named-content>), China: Xizang, Beibeng, 2014.VII.27, Yan Li. 1 female (<named-content content-type="dwc:institutional_code" xlink:title="China Agricultural University" xlink:href="http://grbio.org/institution/china-agricultural-university">CAU</named-content>), China: Xizang, Chayu, 2014.VIII.3, Yan Li. 1 male, 2 females (<named-content content-type="dwc:institutional_code" xlink:title="China Agricultural University" xlink:href="http://grbio.org/institution/china-agricultural-university">CAU</named-content>), China: Xizang, Linzhi, Gongbujiangda, Ganglangcun. <bold><italic>Holotype</italic></bold>: male, Nepal, Hardwicke Bequest; accession no. NHMUK010397658, <abbrev xlink:title="Natural History Museum" id="ABBRID0EKVAE">BMNH</abbrev>(E)247599 (<abbrev xlink:title="Natural History Museum" id="ABBRID0EOVAE">BMNH</abbrev>).</p>
          </tp:treatment-sec>
          <tp:treatment-sec sec-type="diagnosis" id="SECID0ESVAE">
            <title>Diagnosis</title>
            <p>. Antenna has 8 segments. Wing is brown, but yellowed at basal 1/5; anal cells are much paler; a white discal area is before cord, including cell <italic>R<sub>1</sub></italic> to cell <italic>CuA<sub>1</sub></italic>; <italic>m-cu</italic> is near 2/3 of cell <italic>dm</italic>. Posterior margin of tergite 9 is produced; interbase is triangular, stubby at base.</p>
          </tp:treatment-sec>
          <tp:treatment-sec sec-type="redescription" id="SECID0EEWAE">
            <title>Redescription</title>
            <p>. Male (<italic>N</italic> = 4): Body length 12.5–22.5 mm, wing length 12.4–15.5 mm, antenna length 3.3–3.8 mm.</p>
            <p>Head (Figs 9–11) velvet black with black setae. Rostrum and palpus blackish brown. Antenna 8 segmented, blackish brown with brown setae.</p>
            <fig id="F3" position="float" orientation="portrait">
              <object-id content-type="doi">10.3897/zookeys.1071.66750.figure9</object-id>
              <object-id content-type="zenodo_dep_id">5728552</object-id>
              <object-id content-type="arpha">D4BE6089-AFF9-5B5B-87D1-882F1DF30380</object-id>
              <label>Figure 9.</label>
              <caption>
                <p><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="nepalensis">nepalensis</tp:taxon-name-part></tp:taxon-name> (Westwood, 1836), male. Photo by Qicheng Yang.</p>
              </caption>
              <graphic xlink:href="zookeys-1071-155_article-66750__-g003.jpg" position="float" orientation="portrait" xlink:type="simple" id="oo_612051.jpg">
                <uri content-type="original_file">https://binary.pensoft.net/fig/612051</uri>
              </graphic>
            </fig>
            <p>Thorax (Figs 9–11) velvet black with blackish brown setae. Legs with blackish brown setae; coxae and trochanters velvet black with long setae; femora, tibiae and tarsi blackish brown. Wing (Figs 9–10, 16) brown, basal 1/5 yellowed, anal cells much paler; a white discal area before cord, including cell <italic>R<sub>1</sub></italic> to cell <italic>CuA<sub>1</sub></italic>; veins brown, more yellowed in brightened areas. Venation: <italic>R<sub>2</sub></italic> oblique, <italic>R<sub>2+3</sub></italic> relatively short, shorter than <italic>R<sub>2</sub></italic>; cell <italic>M<sub>1</sub></italic> lacking; <italic>m-cu</italic> near 2/3 of cell <italic>dm</italic>. Halter (Figs 9–11) length approximately 2.1 mm, grayish brown with blackish brown setae.</p>
            <fig id="F4" position="float" orientation="portrait">
              <object-id content-type="doi">10.3897/zookeys.1071.66750.figures10-16</object-id>
              <object-id content-type="zenodo_dep_id">5728554</object-id>
              <object-id content-type="arpha">06D8BFB1-DF88-54E0-A303-02A58B67C297</object-id>
              <label>Figures 10–16.</label>
              <caption>
                <p><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="nepalensis">nepalensis</tp:taxon-name-part></tp:taxon-name> (Westwood, 1836) <bold>10</bold> male habitus, lateral view <bold>11</bold> male head and thorax, dorsal view <bold>12</bold> female habitus, lateral view <bold>13</bold> ovipositor, dorsal view <bold>14</bold> ovipositor, ventral view <bold>15</bold> ovipositor, lateral view <bold>16</bold> male right wing. Scale bars: 1 mm (10–16).</p>
              </caption>
              <graphic xlink:href="zookeys-1071-155_article-66750__-g004.jpg" position="float" orientation="portrait" xlink:type="simple" id="oo_612052.jpg">
                <uri content-type="original_file">https://binary.pensoft.net/fig/612052</uri>
              </graphic>
            </fig>
            <p>Abdomen (Figs 9–10) with short black setae. Segments 2–5 elongate, shining black at base, pale white gray in middle, velvet black at tip; segments 1 and 6–7 velvet black; segment 8 shining black.</p>
            <p>Male terminalia (Figs 9–10, 17–21) shining black with blackish brown setae. Posterior margin of ninth tergite produced, its margin concave; posterior margin of ninth sternite with a deep U-shaped shallow; gonocoxite moderately stubby; clasper of gonostylus with long setae at base, slender, terminal spine decurved; lobe of gonostylus short and stout, terminal margin swollen with long setae; interbase triangular, stubby at base; aedeagus longer, apically directed ventrally.</p>
            <fig id="F5" position="float" orientation="portrait">
              <object-id content-type="doi">10.3897/zookeys.1071.66750.figures17-21</object-id>
              <object-id content-type="zenodo_dep_id">5728556</object-id>
              <object-id content-type="arpha">97B12D26-C5ED-54C1-A04A-47FAD57F4A51</object-id>
              <label>Figures 17–21.</label>
              <caption>
                <p><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="nepalensis">nepalensis</tp:taxon-name-part></tp:taxon-name> (Westwood, 1836), male <bold>17</bold> terminalia, dorsal view <bold>18</bold> terminalia, ventral view <bold>19</bold> aedeagal complex, dorsal view <bold>20</bold> aedeagal complex, lateral view <bold>21</bold> aedeagal complex, ventral view. Scale bars: 0.5 mm (17–21).</p>
              </caption>
              <graphic xlink:href="zookeys-1071-155_article-66750__-g005.jpg" position="float" orientation="portrait" xlink:type="simple" id="oo_612053.jpg">
                <uri content-type="original_file">https://binary.pensoft.net/fig/612053</uri>
              </graphic>
            </fig>
            <p><bold>Female</bold> (<italic>N</italic>=3): Body length 17.3–19.4 mm, wing length 13.3–15.4 mm, antenna length 3.6–4.4 mm.</p>
            <p>Female (Figs 12–15) resembles male. Abdomen shorter; tergites 2, 4–5 pale white gray at base, velvet black at tip; tergites 1, 3, 6–7 velvet black; sternites 1–7 velvet black; segment 8 velvet orange.</p>
            <p>Ovipositor (Figs 12–15) elongate, velvet orange. Cercus narrowed toward tip. Hypogynial valve shorter, narrowed toward tip.</p>
          </tp:treatment-sec>
          <tp:treatment-sec sec-type="distribution" id="SECID0E51AE">
            <title>Distribution</title>
            <p>. Afghanistan; China (Sichuan, Guangdong, Xizang), India (Assam and/or Arunachal Pradesh), Malaysia (Peninsula), Nepal.</p>
          </tp:treatment-sec>
          <tp:treatment-sec sec-type="remarks" id="SECID0ED2AE">
            <title>Remarks</title>
            <p>. This species is here recorded from Xizang, China for the first time.</p>
          </tp:treatment-sec>
        </tp:taxon-treatment>
        <tp:taxon-treatment>
          <tp:treatment-meta>
            <kwd-group>
              <label>Taxon classification</label>
              <kwd>
                <named-content content-type="kingdom" xlink:type="simple">Animalia</named-content>
              </kwd>
              <kwd>
                <named-content content-type="order" xlink:type="simple">Diptera</named-content>
              </kwd>
              <kwd>
                <named-content content-type="family" xlink:type="simple">Limoniidae</named-content>
              </kwd>
            </kwd-group>
          </tp:treatment-meta>
          <tp:nomenclature>
            <tp:taxon-name><object-id content-type="arpha">7178BB81-9F29-5EBF-A15E-668B11802478</object-id>
              <tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="paragnava">paragnava</tp:taxon-name-part></tp:taxon-name>
            <tp:taxon-authority>Alexander, 1973</tp:taxon-authority>
            <tp:nomenclature-citation-list>
              <tp:nomenclature-citation>
                <tp:taxon-name>
                  <tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="paragnava">paragnava</tp:taxon-name-part></tp:taxon-name>
                <comment>Alexander, 1973: 7. Type locality: India, Assam.</comment>
              </tp:nomenclature-citation>
            </tp:nomenclature-citation-list>
          </tp:nomenclature>
          <tp:treatment-sec sec-type="material" id="SECID0EA4AE">
            <title>Specimens examined</title>
            <p>. 19 males, 2 females (<named-content content-type="dwc:institutional_code" xlink:title="China Agricultural University" xlink:href="http://grbio.org/institution/china-agricultural-university">CAU</named-content>), China: Xizang, Motuo, Beibeng, Jiangxincun, 800 m, 2019.V.29, Qicheng Yang (light trap). 1 female (<named-content content-type="dwc:institutional_code" xlink:title="China Agricultural University" xlink:href="http://grbio.org/institution/china-agricultural-university">CAU</named-content>), China: Xizang, Motuo, Beibeng, Jiangxincun, 800 m, 2019.V.30, Qicheng Yang (light trap). 4 males, 3 females (<named-content content-type="dwc:institutional_code" xlink:title="China Agricultural University" xlink:href="http://grbio.org/institution/china-agricultural-university">CAU</named-content>), China: Xizang, Motuo, 2019.V.31, Qicheng Yang (light trap).</p>
          </tp:treatment-sec>
          <tp:treatment-sec sec-type="diagnosis" id="SECID0ES4AE">
            <title>Diagnosis</title>
            <p>. Front and mouth parts are very reduced; antenna is very long and 6-segmented; vertical tubercle is very large. Wing is pale yellow; <italic>m-cu</italic> is near 1/6 of cell <italic>dm</italic>. Posterior margin of ninth tergite has a U-shaped notch; interbase is two-layered membranous structure with spine-like apex.</p>
          </tp:treatment-sec>
          <tp:treatment-sec sec-type="redescription" id="SECID0E34AE">
            <title>Redescription</title>
            <p>. Male (<italic>N</italic>=23): Body length 8.6–11.0 mm, wing length 10.2–15.0 mm, antenna length 35.5–47.0 mm.</p>
            <p>Head (Figs 22–23) brown. Front and mouth parts very reduced, brownish yellow, palpus brown. Antenna 6-segmented, very long approximately three or four times as long as wing; scape and pedicel shorter, brownish yellow; flagellum very long, first flagellomere brownish yellow at base, brown at tip; remainder of flagellum brown with short blackish brown setae. Vertical tubercle more brown, very large, bulbous with setae on posterior aspect.</p>
            <fig id="F6" position="float" orientation="portrait">
              <object-id content-type="doi">10.3897/zookeys.1071.66750.figures22-28</object-id>
              <object-id content-type="zenodo_dep_id">5728558</object-id>
              <object-id content-type="arpha">3B6B0FA6-82C3-5381-B625-8625D598372F</object-id>
              <label>Figures 22–28.</label>
              <caption>
                <p><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="paragnava">paragnava</tp:taxon-name-part></tp:taxon-name> Alexander, 1973. <bold>22</bold> male habitus, lateral view <bold>23</bold> male head and thorax, dorsal view <bold>24</bold> female habitus, lateral view <bold>25</bold> ovipositor, dorsal view <bold>26</bold> ovipositor, ventral view <bold>27</bold> ovipositor, lateral view <bold>28</bold> right wing. Scale bars: 1 mm (22–28).</p>
              </caption>
              <graphic xlink:href="zookeys-1071-155_article-66750__-g006.jpg" position="float" orientation="portrait" xlink:type="simple" id="oo_612054.jpg">
                <uri content-type="original_file">https://binary.pensoft.net/fig/612054</uri>
              </graphic>
            </fig>
            <p>Thorax (Figs 22–23) brownish yellow to brown with gray setae. Pronotum brownish yellow; propleuron brown; prescutum dark brown with a broad brown stripe at middle; prescutal suture brown; scutum brownish yellow to brown; scutellum yellow with a slender brownish yellow stripe at middle; mediotergite brownish yellow to brown. Thoracic pleuron mostly brown throughout, except prescutum, anepimeron, katepisternum and metakatepisternum partly brownish yellow. Legs: coxae and trochanters brownish yellow with gray setae; femora and tibiae brownish yellow with short brown setae; tarsi brown with brown setae. Wing (Figs 22, 28) pale yellow; stigma and veins brownish yellow. Venation: <italic>R<sub>2</sub></italic> moderately straight; <italic>R<sub>2+3</sub></italic> near half of <italic>R<sub>2+3+4</sub></italic>; cell <italic>M<sub>1</sub></italic> lacking; <italic>m-cu</italic> near 1/6 of cell <italic>dm</italic>. Halter (Figs 22, 23) length approximately 1.9 mm, whitened gray.</p>
            <p>Abdomen (Fig. 22) with brownish yellow setae. First two segments more yellowed at lateral margin; segments 3–8 dark brown.</p>
            <p>Male terminalia (Figs 22, 29–33) brown with brownish yellow setae. Posterior margin of ninth tergite with a U-shaped notch; gonocoxite large, elongate cylindrical, gently curved; clasper of gonostylus slender, terminal spine decurved; lobe of gonostylus short and stout, swollen with setae at middle; interbase two-layered membranous structure with spine-like apex; aedeagus smaller.</p>
            <fig id="F7" position="float" orientation="portrait">
              <object-id content-type="doi">10.3897/zookeys.1071.66750.figures29-33</object-id>
              <object-id content-type="zenodo_dep_id">5728560</object-id>
              <object-id content-type="arpha">ADA3F1F7-1C4F-51AC-A3D4-47C577722BA3</object-id>
              <label>Figures 29–33.</label>
              <caption>
                <p><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="paragnava">paragnava</tp:taxon-name-part></tp:taxon-name> Alexander, 1973, male <bold>29</bold> terminalia, dorsal view <bold>30</bold> terminalia, ventral view <bold>31</bold> aedeagal complex, dorsal view <bold>32</bold> aedeagal complex, lateral view <bold>33</bold> aedeagal complex, ventral view. Scale bars: 0.5 mm (29–33).</p>
              </caption>
              <graphic xlink:href="zookeys-1071-155_article-66750__-g007.jpg" position="float" orientation="portrait" xlink:type="simple" id="oo_612055.jpg">
                <uri content-type="original_file">https://binary.pensoft.net/fig/612055</uri>
              </graphic>
            </fig>
            <p><bold>Female</bold> (<italic>N</italic>=6): Body length 9.3–13.2 mm, wing length 10.8–12.3 mm, antenna length 1.2–1.4 mm.</p>
            <p>Female (Figs 24–27) resembles male. Thoracic pleuron more brownish yellow. Abdomen plump, dark brown.</p>
            <p>Ovipositor (Figs 24–27) short and fleshy, brownish yellow; cercus oval; Hypogynial valve longer.</p>
          </tp:treatment-sec>
          <tp:treatment-sec sec-type="distribution" id="SECID0EUBAG">
            <title>Distribution</title>
            <p>. India (Assam); China (Xizang).</p>
          </tp:treatment-sec>
          <tp:treatment-sec sec-type="remarks" id="SECID0EZBAG">
            <title>Remarks</title>
            <p>. This species was known previously only from India. With the present contribution it is recorded from China for the first time.</p>
          </tp:treatment-sec>
        </tp:taxon-treatment>
        <tp:taxon-treatment>
          <tp:treatment-meta>
            <kwd-group>
              <label>Taxon classification</label>
              <kwd>
                <named-content content-type="kingdom" xlink:type="simple">Animalia</named-content>
              </kwd>
              <kwd>
                <named-content content-type="order" xlink:type="simple">Diptera</named-content>
              </kwd>
              <kwd>
                <named-content content-type="family" xlink:type="simple">Limoniidae</named-content>
              </kwd>
            </kwd-group>
          </tp:treatment-meta>
          <tp:nomenclature>
            <tp:taxon-name><object-id content-type="arpha">9AFF41CC-1698-5502-95D7-DE471C813413</object-id>
              <tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="perhirsuta">perhirsuta</tp:taxon-name-part></tp:taxon-name>
            <tp:taxon-authority>Alexander, 1973</tp:taxon-authority>
            <tp:nomenclature-citation-list>
              <tp:nomenclature-citation>
                <tp:taxon-name>
                  <tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="perhirsuta">perhirsuta</tp:taxon-name-part></tp:taxon-name>
                <comment>Alexander, 1973: 8. Type locality: India, Assam.</comment>
              </tp:nomenclature-citation>
            </tp:nomenclature-citation-list>
          </tp:nomenclature>
          <tp:treatment-sec sec-type="material" id="SECID0EWDAG">
            <title>Specimens examined</title>
            <p>. 1 male (<named-content content-type="dwc:institutional_code" xlink:title="China Agricultural University" xlink:href="http://grbio.org/institution/china-agricultural-university">CAU</named-content>), China: Xizang, Sejilashan, 3200 m, 2013.VIII.2. 2 females (<named-content content-type="dwc:institutional_code" xlink:title="China Agricultural University" xlink:href="http://grbio.org/institution/china-agricultural-university">CAU</named-content>), China: Xizang, Lulang, 2013.VII.28. 1 male (<named-content content-type="dwc:institutional_code" xlink:title="China Agricultural University" xlink:href="http://grbio.org/institution/china-agricultural-university">CAU</named-content>), China: Xizang, Motuo, 80K, 2014.VII.23, Yan Li (light trap). 1 male (<named-content content-type="dwc:institutional_code" xlink:title="China Agricultural University" xlink:href="http://grbio.org/institution/china-agricultural-university">CAU</named-content>), China: Xizang, Motuo, 80K, 2014.VII.31, Yan Li (light trap). 1 male (<named-content content-type="dwc:institutional_code" xlink:title="China Agricultural University" xlink:href="http://grbio.org/institution/china-agricultural-university">CAU</named-content>), China: Xizang, Yigong, 2017.VIII.8, Qicheng Yang.</p>
          </tp:treatment-sec>
          <tp:treatment-sec sec-type="diagnosis" id="SECID0EQEAG">
            <title>Diagnosis</title>
            <p>. Rostrum is very short; antenna has 6 segments, very long; vertical tubercle is large bulbous. Wing is brownish yellow; <italic>R<sub>2+3</sub></italic> is nearly three times length of <italic>R<sub>2+3+4</sub></italic>; cell <italic>M<sub>1</sub></italic> is present; <italic>m-cu</italic> is near 1/6 of cell <italic>dm</italic>. Posterior margin of ninth tergite has a deep U-shaped notch; interbase is well-developed, a two-layered membranous structure with spine apex.</p>
          </tp:treatment-sec>
          <tp:treatment-sec sec-type="redescription" id="SECID0EGFAG">
            <title>Redescription</title>
            <p>. Male (<italic>N</italic> = 4): Body length 16.0–18.3 mm, wing length 18.2–21.3 mm, antenna length 46.0–61.2 mm.</p>
            <p>Head (Figs 34–35) dark brown with very long abundant brown setae. Rostrum very short, dark brown; palpi brown. Antenna 6-segmented, very long, more than three times as long as wing; scape and pedicel blackish brown with long brown setae; flagellum dark brown with short brown setae. Vertical tubercle large bulbous, dark brown with abundant very long brown setae.</p>
            <fig id="F8" position="float" orientation="portrait">
              <object-id content-type="doi">10.3897/zookeys.1071.66750.figures34-40</object-id>
              <object-id content-type="zenodo_dep_id">5728562</object-id>
              <object-id content-type="arpha">6175E1FB-FFBC-5304-8724-E477F87CE1B6</object-id>
              <label>Figures 34–40.</label>
              <caption>
                <p><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="perhirsuta">perhirsuta</tp:taxon-name-part></tp:taxon-name> Alexander, 1973 <bold>34</bold> male habitus, lateral view <bold>35</bold> male head and thorax, dorsal view <bold>36</bold> female habitus, lateral view <bold>37</bold> ovipositor, dorsal view <bold>38</bold> ovipositor, ventral view <bold>39</bold> ovipositor, lateral view <bold>40</bold> right wing. Scale bars: 1 mm (34–40).</p>
              </caption>
              <graphic xlink:href="zookeys-1071-155_article-66750__-g008.jpg" position="float" orientation="portrait" xlink:type="simple" id="oo_612056.jpg">
                <uri content-type="original_file">https://binary.pensoft.net/fig/612056</uri>
              </graphic>
            </fig>
            <p>Thorax (Figs 34–35) dark brown with abundant very long brownish yellow setae. Legs: coxae dark brown with abundant very long brownish yellow setae; trochanters brownish yellow with long brownish yellow setae; femora yellow with short brownish yellow setae; tibiae and tarsi brown with short brown setae. Wing (Figs 34, 40) brownish yellow, stigma slightly brown; veins slightly brown, very inconspicuous against the ground. Venation: <italic>R<sub>2</sub></italic> moderately straight; <italic>R<sub>2+3</sub></italic> nearly three times as long as <italic>R<sub>2+3+4</sub></italic>; cell <italic>M<sub>1</sub></italic> slightly longer than its petiole; <italic>m-cu</italic> near 1/6 of cell <italic>dm</italic>. Halter (Figs 34–35) length approximately 3.2 mm, halter stem grayish brown with brown setae; knob brown.</p>
            <p>Abdomen (Fig. 34) with abundant very long brownish yellow setae. First three segments brown; segments 4–8 dark brown.</p>
            <p>Male terminalia (Figs 34, 41–45) brown with long brownish yellow setae. Posterior margin of ninth tergite with a deep U-shaped notch; posterior margin of ninth sternite with a deep V-shaped shallow; gonocoxite large, elongate cylindrical; clasper of gonostylus long and slender, terminal spine decurved; lobe of gonostylus short and stout, swollen with setae at middle; interbase well-developed, a two-layered membranous structure with spine-like apex; aedeagus smaller.</p>
            <fig id="F9" position="float" orientation="portrait">
              <object-id content-type="doi">10.3897/zookeys.1071.66750.figures41-45</object-id>
              <object-id content-type="zenodo_dep_id">5728564</object-id>
              <object-id content-type="arpha">C2DB8226-7024-50C9-B922-3091FE63E70D</object-id>
              <label>Figures 41–45.</label>
              <caption>
                <p><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="perhirsuta">perhirsuta</tp:taxon-name-part></tp:taxon-name> Alexander, 1973, male <bold>41</bold> terminalia, dorsal view <bold>42</bold> terminalia, ventral view <bold>43</bold> aedeagal complex, dorsal view <bold>44</bold> aedeagal complex, lateral view <bold>45</bold> aedeagal complex, ventral view. Scale bars: 0.5 mm (41–45).</p>
              </caption>
              <graphic xlink:href="zookeys-1071-155_article-66750__-g009.jpg" position="float" orientation="portrait" xlink:type="simple" id="oo_612057.jpg">
                <uri content-type="original_file">https://binary.pensoft.net/fig/612057</uri>
              </graphic>
            </fig>
            <p><bold>Female</bold> (<italic>N</italic>=2): Body length 21.3–24.7 mm, wing length 19.6–22.5 mm, antenna length 3.1–3.5 mm.</p>
            <p>Female (Figs 36–39) resembles male. Thorax more brownish yellow. Abdomen longer, venter more brownish yellow.</p>
            <p>Ovipositor (Figs 36–39) elongate, reddish yellow. Cercus narrowed toward tip. Hypogynial valve shorter, narrowed toward tip.</p>
          </tp:treatment-sec>
          <tp:treatment-sec sec-type="distribution" id="SECID0E4IAG">
            <title>Distribution</title>
            <p>. India (Assam); China (Xizang).</p>
          </tp:treatment-sec>
          <tp:treatment-sec sec-type="remarks" id="SECID0ECJAG">
            <title>Remarks</title>
            <p>. This species was known previously only from India. This is the first record from China.</p>
          </tp:treatment-sec>
        </tp:taxon-treatment>
        <tp:taxon-treatment>
          <tp:treatment-meta>
            <kwd-group>
              <label>Taxon classification</label>
              <kwd>
                <named-content content-type="kingdom" xlink:type="simple">Animalia</named-content>
              </kwd>
              <kwd>
                <named-content content-type="order" xlink:type="simple">Diptera</named-content>
              </kwd>
              <kwd>
                <named-content content-type="family" xlink:type="simple">Limoniidae</named-content>
              </kwd>
            </kwd-group>
          </tp:treatment-meta>
          <tp:nomenclature>
            <tp:taxon-name><object-id content-type="arpha">C3912DE1-E135-5C01-9E41-CCE1EC224A3E</object-id>
              <tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="xizangensis">xizangensis</tp:taxon-name-part><object-id content-type="zoobank" xlink:type="simple">http://zoobank.org/72D7BB50-F2AB-40AC-AA95-C86E200960B1</object-id></tp:taxon-name>
            <tp:taxon-status>sp. nov.</tp:taxon-status>
          </tp:nomenclature>
          <tp:treatment-sec sec-type="type material" id="SECID0ERKAG">
            <title>Type material</title>
            <p><bold>. <italic>Holotype</italic></bold>: male (<named-content content-type="dwc:institutional_code" xlink:title="China Agricultural University" xlink:href="http://grbio.org/institution/china-agricultural-university">CAU</named-content>), China: Xizang, Beibeng, Jiangxincun, 2019.V.30, Qicheng Yang (light trap).</p>
          </tp:treatment-sec>
          <tp:treatment-sec sec-type="diagnosis" id="SECID0E6KAG">
            <title>Diagnosis</title>
            <p>. Femora are yellow. Wing is brownish yellow with the following markings: an oblique transverse hyaline marking from <italic>R</italic> extended up to <italic>CuA</italic> in base of wing; origin of <italic>Rs</italic> with a small hyaline marking; a longer oblique transverse hyaline marking from <italic>R</italic> extended to wing margin before cord; both sides of <italic>R<sub>2</sub></italic> with a hyaline marking; tip of <italic>R<sub>1</sub></italic> from <italic>Sc</italic> to <italic>R<sub>3</sub></italic> with an oblique transverse hyaline marking. <italic>R<sub>2</sub></italic> is moderately oblique, approximately as long as <italic>R<sub>3+4</sub></italic>, placed before fork of <italic>R<sub>3+4</sub></italic>; cell <italic>M<sub>1</sub></italic> is lacking; <italic>m-cu</italic> is near 2/3 of cell <italic>dm</italic>. Abdomen is brown to darker brown except segments 8–9 yellow. Posterior margin of ninth tergite has two small triangular processes; interbase is cylindrical, but stubby at base.</p>
          </tp:treatment-sec>
          <tp:treatment-sec sec-type="description" id="SECID0EPMAG">
            <title>Description</title>
            <p>. Male (<italic>N</italic> = 1): Body length 8.8 mm, wing length 7.2 mm.</p>
            <p>Head (Figs 46) brown with long brown setae. Rostrum very short, brownish yellow; palpi brown. Antennal scape brown with brown setae; pedicel brownish yellow; flagellum is broken.</p>
            <fig id="F10" position="float" orientation="portrait">
              <object-id content-type="doi">10.3897/zookeys.1071.66750.figures46-4946-48</object-id>
              <object-id content-type="zenodo_dep_id">5728546</object-id>
              <object-id content-type="arpha">7B543728-FDBF-5209-A5BC-C4A96CF73738</object-id>
              <label>Figures 46–49. 46–48</label>
              <caption>
                <p><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="xizangensis">xizangensis</tp:taxon-name-part></tp:taxon-name> sp. nov., male <bold>46</bold> habitus, lateral view <bold>47</bold> male thorax, dorsal view <bold>48</bold> right wing <bold>49</bold> wing of <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="decorata">decorata</tp:taxon-name-part></tp:taxon-name> (Brunetti, 1918: plate VII, fig. 12). Scale bars: 1.0 mm (46–48).</p>
              </caption>
              <graphic xlink:href="zookeys-1071-155_article-66750__-g010.jpg" position="float" orientation="portrait" xlink:type="simple" id="oo_612058.jpg">
                <uri content-type="original_file">https://binary.pensoft.net/fig/612058</uri>
              </graphic>
            </fig>
            <p>Thorax (Figs 46, 47) brown with long brown setae. Legs: coxae and trochanters brown with long brown setae; femora and tibiae yellow with brown setae; basal two segments of tarsi yellow with brown setae, reminder segments brownish yellow to brown with brown setae. Wing (Figs 46, 48) brownish yellow, anal cells more yellow; anal cells with hyaline markings at base; an oblique transverse hyaline marking from <italic>R</italic> extended up to <italic>CuA</italic> in base of wing; origin of <italic>Rs</italic> with a small hyaline marking; a longer oblique transverse hyaline marking from <italic>R</italic> extended to wing margin before cord; both sides of <italic>R<sub>2</sub></italic> with a hyaline marking; tip of <italic>R<sub>1</sub></italic> from <italic>Sc</italic> to <italic>R<sub>3</sub></italic> with an oblique transverse hyaline marking; veins slightly brown, very inconspicuous against ground. Venation: <italic>R<sub>2</sub></italic> oblique, approximately as long as <italic>R<sub>3+4</sub></italic>, placed before fork of <italic>R<sub>3</sub></italic><sub>+</sub><italic><sub>4</sub></italic>; cell <italic>M<sub>1</sub></italic> lacking; <italic>m-cu</italic> near 2/3 of cell <italic>dm</italic>. Halter (Figs 46, 47) length approximately 1.2 mm, pale brown.</p>
            <p>Abdomen (Fig. 46) with brown setae. Segments 1–5 brown, segments 6–7 dark brown, segment 8 yellow.</p>
            <p>Male terminalia (Figs 46, 50–54) yellow with brownish yellow setae. Posterior margin of ninth tergite with two small triangular processes, with abundant brown setae; gonocoxite moderately stubby; clasper of gonostylus slender, terminal spine decurved; lobe of gonostylus short and stout, middle margin swollen with setae; interbase cylindrical, stubby at base; aedeagus longer, apically directed ventrally.</p>
            <fig id="F11" position="float" orientation="portrait">
              <object-id content-type="doi">10.3897/zookeys.1071.66750.figures50-5650-54</object-id>
              <object-id content-type="zenodo_dep_id">5728548</object-id>
              <object-id content-type="arpha">842EFDF4-42B7-51A3-B4AA-7BF8AD13DA31</object-id>
              <label>Figures 50–56. 50–54</label>
              <caption>
                <p><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="xizangensis">xizangensis</tp:taxon-name-part></tp:taxon-name> sp. nov., male <bold>50</bold> terminalia, dorsal view <bold>51</bold> terminalia, ventral view <bold>52</bold> aedeagal complex, dorsal view <bold>53</bold> aedeagal complex, lateral view <bold>54</bold> aedeagal complex, ventral view <bold>55–56</bold><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">Hexatoma</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="decorata">decorata</tp:taxon-name-part></tp:taxon-name> (Brunetti, 1918), male. <bold>55</bold> ninth tergite (Joseph, 1977: 427, fig. 17) <bold>56</bold> aedeagus (Joseph, 1977: 427, fig. 20). Scale bars: 0.5 mm (50–54).</p>
              </caption>
              <graphic xlink:href="zookeys-1071-155_article-66750__-g011.jpg" position="float" orientation="portrait" xlink:type="simple" id="oo_612059.jpg">
                <uri content-type="original_file">https://binary.pensoft.net/fig/612059</uri>
              </graphic>
            </fig>
          </tp:treatment-sec>
          <tp:treatment-sec sec-type="Female" id="SECID0ETRAG">
            <title>Female</title>
            <p>. Unknown.</p>
          </tp:treatment-sec>
          <tp:treatment-sec sec-type="distribution" id="SECID0EYRAG">
            <title>Distribution</title>
            <p>. China (Xizang).</p>
          </tp:treatment-sec>
          <tp:treatment-sec sec-type="etymology" id="SECID0E4RAG">
            <title>Etymology</title>
            <p>. The species is named after Xizang Autonomous Region, where the type locality is located.</p>
          </tp:treatment-sec>
          <tp:treatment-sec sec-type="remarks" id="SECID0ECSAG">
            <title>Remarks</title>
            <p>. This new species is very similar to <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">H.</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">E.</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="decorata">decorata</tp:taxon-name-part></tp:taxon-name> (Brunetti, 1918) from India (W Bengal) in having similar wing markings, but can be separated from it by the wing with a longer oblique transverse hyaline marking from <italic>R</italic> extended to wing margin before cord (Figs 46, 48) and the posterior margin of the ninth tergite with two small triangular processes (Fig. 50). In <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Hexatoma">H.</tp:taxon-name-part> (<tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">E.</tp:taxon-name-part>) <tp:taxon-name-part taxon-name-part-type="species" reg="decorata">decorata</tp:taxon-name-part></tp:taxon-name>, the oblique transverse hyaline marking from <italic>R</italic> extends to the <italic>CuP</italic> before the cord (Fig. 49 i.e., Brunetti, 1918: plate VII. fig. 12; Edwards, 1924: 304; Joseph, 1977: 427, fig. 16) and the posterior margin of the ninth tergite is produced, its margin concave (Fig. 55 i.e., Joseph, 1977: 427, fig. 7). These two species are very special within the subgenus <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="subgenus" reg="Eriocera">Eriocera</tp:taxon-name-part></tp:taxon-name>, be<named-content content-type="dwc:institutional_code" xlink:title="China Agricultural University" xlink:href="http://grbio.org/institution/china-agricultural-university">CAU</named-content>se of their unique position and slope of vein <italic>R<sub>2</sub></italic> (Fig. 46, 48–49): it contacts vein <italic>R<sub>3+4</sub></italic> (and not vein <italic>R<sub>3</sub></italic> as is common in all <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Eriocera">Eriocera</tp:taxon-name-part></tp:taxon-name></italic> with a short vein <italic>R<sub>2+3+4</sub></italic>), and is sloping forwards (also a very uncommon feature in <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Eriocera">Eriocera</tp:taxon-name-part></tp:taxon-name></italic>).</p>
          </tp:treatment-sec>
        </tp:taxon-treatment>
      </sec>
    </sec>
  </body>
  <back>
    <ack>
      <title>Acknowledgements</title>
      <p>We are grateful to Prof. Fasheng Li (Beijing) for collecting the specimens from Xizang. We are also particularly grateful to Prof. Herman de Jong for his valuable suggestions on this paper. The research was funded by the National Science &amp; Technology Fundamental Resources Investigation Program of China (Grant No. 2019FY100400).</p>
    </ack>
    <ref-list>
      <title>References</title>
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        <mixed-citation xlink:type="simple"><person-group><name name-style="western"><surname>Alexander</surname><given-names>CP</given-names></name></person-group> (<year>1933</year>) <article-title>New or little-known <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="family">Tipulidae</tp:taxon-name-part></tp:taxon-name> from eastern Asia (<tp:taxon-name><tp:taxon-name-part taxon-name-part-type="order">Diptera</tp:taxon-name-part></tp:taxon-name>). XV.</article-title><source>Philippine Journal of Science</source><volume>52</volume>: <fpage>131</fpage>–<lpage>166</lpage>.</mixed-citation>
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        <mixed-citation xlink:type="simple"><person-group><name name-style="western"><surname>Alexander</surname><given-names>CP</given-names></name></person-group> (<year>1948</year>) <article-title>Notes on the tropical American species of <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="family">Tipulidae</tp:taxon-name-part></tp:taxon-name> (<tp:taxon-name><tp:taxon-name-part taxon-name-part-type="order">Diptera</tp:taxon-name-part></tp:taxon-name>). V. The specialized <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="tribe">Hexatomini</tp:taxon-name-part></tp:taxon-name>: <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Limnophila</tp:taxon-name-part></tp:taxon-name></italic>, <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Shannonomyia</tp:taxon-name-part></tp:taxon-name></italic>, <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Gynoplistia</tp:taxon-name-part></tp:taxon-name></italic>, <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Hexatoma</tp:taxon-name-part></tp:taxon-name></italic>, <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Atarba</tp:taxon-name-part></tp:taxon-name></italic>, <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Elephantomyia</tp:taxon-name-part></tp:taxon-name></italic>, and allies.</article-title><source>Revista de Entomologia</source><volume>19</volume>: <fpage>509</fpage>–<lpage>556</lpage>.</mixed-citation>
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        <mixed-citation xlink:type="simple"><person-group><name name-style="western"><surname>Alexander</surname><given-names>CP</given-names></name><name name-style="western"><surname>Byers</surname><given-names>GW</given-names></name></person-group> (<year>1981</year>) <article-title><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="family">Tipulidae</tp:taxon-name-part></tp:taxon-name>.</article-title> In: <person-group><name name-style="western"><surname>McAlpine</surname><given-names>JF</given-names></name><name name-style="western"><surname>Peterson</surname><given-names>BV</given-names></name><name name-style="western"><surname>Shewell</surname><given-names>GE</given-names></name><name name-style="western"><surname>Teskey</surname><given-names>HJ</given-names></name><name name-style="western"><surname>Vockeroth</surname><given-names>JR</given-names></name><name name-style="western"><surname>Wood</surname><given-names>DM</given-names></name></person-group> (<role>Eds</role>) <issue-title>Manual of Nearctic Diptera. Vol. 1.</issue-title><source>Biosystematics Research Institute, Ottawa, Ontario, Monograph</source><volume>27</volume>: <fpage>153</fpage>–<lpage>190</lpage>.</mixed-citation>
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        <mixed-citation xlink:type="simple"><person-group><name name-style="western"><surname>Alexander</surname><given-names>CP</given-names></name><name name-style="western"><surname>Lloyd</surname><given-names>JT</given-names></name></person-group> (<year>1914</year>) <article-title>The biology of the North American crane-flies (<tp:taxon-name><tp:taxon-name-part taxon-name-part-type="family">Tipulidae</tp:taxon-name-part></tp:taxon-name>, <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="order">Diptera</tp:taxon-name-part></tp:taxon-name>). I. The genus <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus">Eriocera</tp:taxon-name-part></tp:taxon-name></italic> Macquart.</article-title><source>Journal of Entomology and Zoology</source><volume>6</volume>: <fpage>12</fpage>–<lpage>34</lpage>. <ext-link xlink:type="simple" ext-link-type="doi" xlink:href="10.5962/bhl.part.26522">https://doi.org/10.5962/bhl.part.26522</ext-link></mixed-citation>
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</article>
