Puigpunyant, 39.6167°N, 2.5167°E, 6.x.1981, H. Malicky leg. 1♂ (RW).

Previously only recorded from the type locality in northern Spain.

Calobra, 39.85°N, 2.8°E, 9.v.1978, 90 m a.s.l., H. Malicky leg. 1♂ (RW);

Deiá, town fountain, 39.748072°N, 2.643385°E, 8.ii.2015, G.Kvifte, M. Stokkan & C. Garcia leg. 1♂ (ZFMK);

Puigpunyent, 39.62°N, 2.85°E, 12.v.1978, 200m a.s.l., H. Malicky leg., 1♂; same but 1.X.1979, 6 ♂♂; same but 6.x.1979, 6♂♂ (all RW);

South slope of Piug Major, north of Soller, 39.783°N, 2.767°E, 7.–9.v.1978, 700 m a.s.l., H. Malicky leg. 2♂♂ (RW).

Previously known from France, mainland Spain, Italy, Morocco and Algeria (Vaillant 1974). The generic and subgeneric classification of Mormiina is unstable and requires revision; therefore the placement in Mormia may change.

Esporles, IMEDEA research center, 39.666438°N, 2.580863°E, 13.iv.2015, M. Stokkan leg. 1♀ (ZMFK);

E of Puigpunyent, 39.616667°N, 2.85°E, 1.x.1979, 200m a.s.l., H. Malicky leg. 1♀; same but 6.x.1979, 4♀♀ (all RW).

A widespread near-cosmopolitan species, first recorded from Spain by Tonnoir (1920) under the synonym Telmatoscopus meridionalis (Eaton, 1894). Its biology is summarised in Boumans et al. (2009) and Ježek et al. (2012); for a review of its taxonomy see Ibañez-Bernal (2008).

SW Pollensa, 39.8833°N, 2.9833°E, 3–5.x.1981, H. Malicky leg. 1♂ (RW).

Placed in Lepiseodina Enderlein, 1937 (type species Psychoda tristis Meigen, 1830) by Ježek (1990) because of its asymmetric genitalia. The placement in Clogmia by Ježek (1984), Vaillant (1989) and Wagner (2004) may be a valid option to consider for the future when further character systems are explored.

Previous records are from Austria, Belgium, Czech Republic, France, Germany, Great Britain, Ireland and Slovakia (Oboňa and Ježek 2012).

Banyalbufar, 39.691635°N, 2.514367°E, 25.x.2012, G.Kvifte leg. 1♂1♀ (ZMUB);

Pond west of Cala Figuera, 39.335635°N, 3.152597°E, 11.ii.2015, G.Kvifte leg, 1♂ (ZFMK).

A widespread species or complex of species occurring in the Holarctic region. Ježek and Yağci (2005) list occurences from the following countries: Afghanistan, Algeria, the Azores, Belgium, Bosnia-Herzegovina, Bulgaria, the Canary Islands, China, Corsica, Czech Republic, Denmark, France, Germany, Great Britain, Greece, Hungary, Iran, Ireland, Israel, Italy, Macedonia, Madeira, Mongolia, Morocco, the Netherlands, Olanda island, Poland, Romania, Sardinia, Serbia, Slovakia, Slovenia, Spain, Sweden, Switzerland, Tunisia, Turkey and the USA.

Holotype male. Deiá, town fountain, 39.748072°N, 2.643385°E, 8.ii.2015, G.Kvifte, M. Stokkan & C. Garcia leg (ZMUB).

Diagnosis. Pericoma unipennata can be separated from all other Pericoma species on the presence of one feather-tipped and four spatulate tenacula on each surstylus, as well as the following combination of characters: parameral sheath with shallow U-shaped apical indentation 1/6th as deep as width of parameral appendage base, gonostyle with narrow distal part 1/4th as long as broad basal part, distiphallic spatula with sclerotized side margins not converging apically, parameral appendage concave at lateral sides with distal 1/8th protruding over distiphallic spatula.

Male (n=1). Head (Fig. 1A) longer than wide; vertex rounded with posterior lobe pointed, about half length of head; eyebridge comprising four rows of facets, separated by five facet diameters; interocular suture broadly V-shaped with median swelling; 7 supraocular setae present, 3 on ventral side and 4 on dorsal side; frontal scar patch inversely T-shaped, reaching level of uppermost facet row of eyebridge; clypeus subrectangular, weakly concave at anterior margin, densely setose, not protruding in front of level of eyes; palp with 4th segment fleshy, corrugated, length of palpus segments 93 : 106 : 129 : 231; labellum fleshy, micropilose with 11 larger setae; antennae with 14 fusiform flagellomeres; scape cylindrical, pedicel elongate globular to barrell-shaped; ascoids present on flagellomeres 4-8; flagellomere 14 with apiculus about as long as base of segment; length of antennal segments 69 : 63 : 48 : 45 : 45 : 42 : 42 : 42 : 42 : 42 : 36 : 36 : 33 : 27 : 24 : 36.

Figure 1. 

Pericoma unipennata sp. n., holotype male. A Head B Wing C Aedeagus D Male genitalia E Tenacula. The following abbreviations are used: bp – basiphallus, c r – cercal region, dp lm – distiphallic lateral margin, dp s – distiphallic spatula, f t – feathery tenacula, gcx – gonocoxite, gst – gonostyle, hyp – hypandrium, pm – parameral sclerite, pm j – parameral joint, pms – parameral sheath, s t – spatuliform tenacula.

Thorax without accessory organs; mesonotum and scutellum covered in setae alveoli except on lateral margins; anepisternum and laterotergite covered with setae alveoli; coxae and trochanters with dorsal and ventral stripes of setae alveoli, mid coxa with apicoanterior setose projection; legs without special features;

Wing (Fig. 1B) 2.3 mm long, subovate, membrane without setation or infuscation; radial fork very slightly distad of medial fork, both slightly distad of CuA₂; C with two breaks; Sc weakly curved towards C; apical section of R₂ curved anteriorly; wing apex between R₄ and R₅; medial fork incomplete; apical section of CuA₂ curved posteriorly; jugum obtuse;

Genitalia (Fig. 1C, D) with hypandrium of even width; aedeagus (Fig. 1D) with basiphallus compressed laterally; distiphallus consisting of two phallomeres forming a spatula with sclerotized lateral margins, lateral margins not converging distally; parameral sheath fused with gonocoxites basally, distally narrowing to nearly half its width at base; distally with narrow median incision; parameral sclerite meeting distal section of basiphallus, jointed to parameral sheath with slightly curved transverse band-like sclerite; parameral sclerite half length of distiphallus, consisting of two fused sclerites which are concave laterally, median suture complete; gonocoxites (Fig. 1C) stout, reniform, parabasal process absent but setose field present near connection with parameral sheath; gonocoxal condyles apparently fused with parameral sheath, gonostyle (Fig. 1C) with basal four 5ths liver-shaped, stout, distal 5th pointed, weakly sinous; epandrium slightly wider than long, expanding distally, number of apertures not discernable in specimen, carrying sparse hairs on ventral surface; surstyli about as long as epandrium, with lateral inner third more weakly sclerotized and densely micropilose; apex of surstylus with five aseriate tenacula of which one apical is feathery at the apex, four subapical tenacula spatulate; hypoproct obtusely isosceles trapezoid, micropilose; epiproct oval, densely pilose.

unipennata = with one feather; refers to the presence of a single feather-tipped and four spatulate tenacula on its surstyli.

The specimen was collected at a spring with many bryophytes growing in a seepage stream. Bryophyte material was collected and extracted, however, no psychodid larvae or pupae were found.

Calobra, 39.85°N, 2.8°E, 9.v.1978, 90m a.s.l., H. Malicky leg. 1 ♂ (RW);

Deiá, town fountain, 39.748072°N, 2.643385°E, 8.ii.2015, G.Kvifte, M. Stokkan & C. Garcia leg. 3♂♂ (ZMUB);

Esporles, Torrent de San Vic, 39.670459°N, 2.569193°E, 11.ii.2015, G.Kvifte leg. 1♂ (ZMFK);

Puigpunyent, 39.616667°N, 2.85°E, 15.v.1978, 200m a.s.l., H. Malicky leg. 12 ♂♂; same but 1.x.1979, 5 ♂♂; same but 6.x.1979, 2 ♂♂ (all RW);

South slope of Piug Major, north of Soller, 39.783333°N, 2.766667°E, 7.–9.v.1978, 700 m a.s.l., H. Malicky leg. 11♂♂ (RW).

Vaillant (1978).

This species was described from Algeria and subsequently recorded from Mallorca by Vaillant (1978). It also occurs in mainland Spain and in North Africa (Vaillant 1978).

Wagner et al. (2002).

Vaillant (1978) wrote “[Pericoma] trifasciata ist ebenfalls gemein auf der Mallorca-Insel [...]” (“trifasciata is also common on the island of Mallorca”) in the paragraph summarizing the geographical distribution of Pericoma barbarica Vaillant, 1955; this is probably a lapsus as Mallorca is not mentioned in the species account for Pericoma trifasciata. No other records are available in the literature prior to the listing by Wagner et al. (2002), which was based on Vaillant (1978). In Wagner (2004) and the present paper we deem the records from the Baleares to be doubtful and in need of verification through examination of specimens.

Deiá, town fountain, 39.748072°N, 2.643385°E, 8.ii.2015, G.Kvifte leg. 1♂ (ZMFK);

Esporles, town fountain, 39.669519°N, 2.576953°E, 10.ii.2015, G.Kvifte leg. 1♂ (ZMFK);

Palma de Mallorca, 39.570725°N, 2.641432°E, 7.ii.2015, G.Kvifte leg. 3♂1♀ (2♂ ZMFK, 1♂1♀ZMUB).

The males recorded here were identified as Philosepedon pyrenaicus according to the key in Vaillant (1974) because of the eyes separated by three facet diameters, the wing “mittwinkel” 105° and the left phallomere of the aedeagus being longer than the right. However, the specimens differ in that the 1st flagellomere is longer than the combined length of the scape and pedicel, and the eyebridge is two facet rows wide at several points. We deem these differences not to be taxonomically meaningful, as the differences in proportions and eyebridge composition could be interpreted as interspecific variation.

Wagner (2004)

Cala Figuera, 39.332245°N, 3.166440°E, 11.ii.2015, G. Kvifte leg. 2♂♂ (ZMFK);

Deiá, town fountain, 39.748072°N, 2.643385°E, 8.ii.2015, G.Kvifte, M. Stokkan & C. Garcia leg. 1♂ (ZMUB);

Esporles, Torrent de San Vic, 39.670459°N, 2.569193°E, 10.ii.2015, G.Kvifte leg. 1♂ (ZMFK).

Wagner (2004)

Wagner (2004).

Esporles, Torrent de San Vic, 39.670459°N, 2.569193°E, 11.ii.2015, G.Kvifte leg. 1♂.

Recorded from Algeria, Austria, Belgium, Bosnia-Herzegovina, Czech Republic, Denmark, the Faroe Islands, France, Germany, Great Britain, Greece, Hungary, Ireland, Italy, Morocco, the Netherlands, Norway, Slovakia, Slovenia, Sweden, Tunisia and Turkey (Andersen 1999, Ježek 2004, Ježek and Yağci 2005).

Wagner (2004).

Esporles, Torrent de San Vic, 39.670459°N, 2.569193°E, 10.ii.2015, G.Kvifte leg. 1♂.

Wagner (2004).

Esporles, town fountain, 39.669519°N, 2.576953°E, 10.ii.2015, G.Kvifte leg. 1♀ (ZMFK).

Tinearia Schellenberg, 1803 was treated as a genus by Wagner (2004) following Ježek (1977). This species group is undoubtedly monophyletic, however based on morphology and some analyses of molecular data it appears to be deeply nested within Psychoda s.l. (Vaillant 1990, Espindola et al. 2012, Kvifte and Andersen 2012, Kvifte unpubl.). Inclusion of Tinearia as a subgenus within Psychoda would thus allow ease of identification of monophyletic units, whereas recognising Tinearia as a separate taxon would render Psychoda paraphyletic. For a discussion of the monophyly criterion in supraspecific taxonomy, see Komarek and Beutel (2006).

Wagner (2004)

For genus taxonomy, see remarks under Psychoda alternata above.