Leviea herberti sp. n.

This distinctive genus is named in honour of Herbert Walter Levi, his partner Lorna Rose Levi, and their daughter Frances Levi. Dr. Levi (or, Herb, as he humbly preferred to be known by) was one of the grand arachnologists of the twentieth century, describing over 1200 species of spiders, mentoring many subsequent leaders of the field, and curating one of arachnology’s most important museum collections (Leibensperger 2016). Lorna collaborated in his work in many ways, co-authoring the classic book Spiders and their kin (Levi and Levi 1968), which introduced the first author of this paper to spider diversity. Frances accompanied them in the field and carried on an interest in woven creations. Their contributions, both personal and scientific, will long be remembered (Maddison 2014a, b; Leibensperger 2016). The Levis pronounced the vowels of their name approximately as their IPA equivalents (e as in Ed, i as in eat). The last three letters of Leviea are to be pronounced as three separate vowels (as their IPA equivalents, i-e-a). The name is to be treated grammatically as feminine.

The form of the body is not in the least bit reminiscent of an ant, beetle or wasp, unlike other myrmarachnines. Instead, the body is of standard salticid form (e.g., Icius Simon, 1876, Salticus Latreille, 1804), somewhat glabrous, with chevron markings. Two features possibly retained from antlike ancestors are a female palp that is widened and somewhat dorso-ventrally flattened, and the many long macrosetae on the first tibia in two of the Leviea species. The male embolus is distinctive for ending broadly, not tapering to a point. As in Papuamyr, there is an ectal spur on the paturon (white arrow in Figs 13, 29).

Holotype: male, specimen PNG2008-0360 in UBC–SEM, with data PAPUA NEW GUINEA: Enga Province: Kai-ingri. 5.579 S 143.053 E. 3240 m a.s.l. 7–9 July 2008. W Maddison & Manisé Kulé leg. WPM#08-005. Beating understory of Phyllocladus forest. Paratype: female, specimen PNG2008-0370, with same data as holotype, in UBC–SEM.

Named in honour of Dr Herbert W Levi.

Body somewhat smaller, less elongate, than the other species of Leviea, and with fewer (3 or 3.5 pairs) ventral macrosetae on first tibia. The robust dorsal branch of the RTA ends bluntly like a thumb, unlike the tapering and dorsally pointing tip of the other species. Bulb ca. 90° further rotated than in L. lornae and L. francesae, as indicated by the spermophores (Figs 5, 21, 38).

Male (holotype). Carapace length 1.82; abdomen length 1.96. Carapace (Figs 12, 14, 15): Of typical salticid shape, without constrictions or thoracic hump. Chelicera (Fig. 13): Vertical, simple except for a small ectal spur on the paturon. At least five retromarginal teeth. Palp (Figs 5, 6). With round bulb. Embolus circling 1.8 times around, ending in broad tip. RTA appearing as a finger and thumb (dorsal apophysis and ventral flange, respectively), with the dorsal apophysis pointing somewhat ventrally. Legs unremarkable, first somewhat more robust. First tibia with three pairs of ventral macrosetae, of normal length (right tibia bears an extra small distal anterior macroseta). Colour in life (Figs 9–12): Markings are muted beige to reddish brown, with the digestive diverticula in the ocular area providing two pale stripes that continue as pale areas on the thorax and chevrons on the abdomen. Colour in alcohol (Fig. 13–15): Body and appendages honey coloured, with indistinct dark annuli on the legs, and chevrons on the abdomen. Underside of abdomen with two longitudinal dark stripes.

Female (paratype, specimen PNG2008-0370). Carapace length 1.89; abdomen length 2.16. Carapace, legs, abdomen substantially as in male (Figs 16–18, 20), except for the less robust first legs. First leg tibia with seven ventral macrosetae (three pairs plus one prolateral-distal) (Fig. 19). Epigyne (Figs 7, 8): Of fairly typical myrmarachnine form. RTA pocket more distinct and larger than in L. lornae. Colour (Figs 16–18, 20): As in male, but with body and chelicerae darker, and palps bright yellow in life.

Figures 5–20. 

Leviea herberti sp. n., holotype male and paratype female. 5, 6 Left palp 5 ventral view 6 retrolateral view 7 epigyne, ventral view (paratype female) 8 cleared vulva, dorsal view (same female) 9–15 holotype male 13 face; arrow shows ectal spur on paturon 14 side of carapace 15 habitus dorsal view; two photographs joined 16–20 paratype female 19 prolateral view of first leg 20 habitus dorsal view; two photographs joined. Scale bars: 0.1 mm (on genitalia); 1.0 mm (on bodies).

Holotype: male specimen in RBINS, with data PAPUA NEW GUINEA: Chimbu Province, Mount Wilhelm, Pinde-Yaunde Lake 5.78S, 145.06E. 3700 m a.s.l. 4 October 2013. Sub-alpine forest near the limit tree vegetation. Gewa, Damag, Novotny, Leponce leg. #P3633 Beating understory. Paratypes: 5 males (from collection events #P3609, #P3631, #P3634, #P3637, #P3640) 4 females (from collection events #P3598, #P3632, #3619, #P3633,) in RBINS with same data as the holotype. 3 males and 3 females (events #P3619, #P3637) in UBC–SEM with same data as the holotype. 2 males (events #P3232, #P3241), 1 female (event #P3237) in RBINS with data PAPUA NEW GUINEA: Madang Province: Kombunomambuno 5.80S, 145.07E. 3200 m a.s.l. 4 October 2013. Upper montane forest. Dahl, Kaupa, Novotny, Leponce leg. Beating understory. 4 males 4 females in HNHM with data PAPUA NEW GUINEA: Madang Province: Mt Wilhelm, Moss forest at the meteorological station of Kambugomanbuno [sic!]. 3018 m a.s.l. 1968.09.17. János Balogh leg. NGM C26 Beating. (HNHM Araneae-9251, 9252, 9253, 9254).

Named in honour of Lorna Levi.

Larger and more slender than L. herberti, with more (5 pairs) ventral macrosetae on first tibia. In these features it resembles L. francesae, from which it differs in details of the palp: dorsal branch of RTA slender and distal-pointing; bulb rounder than L. francesae, less rotated than L. herberti. Epigyne with openings further posterior than those of L. herberti, and smaller RTA pocket at rear margin.

Male (paratype in UBC–SEM). Carapace length 2.13; abdomen length 2.35. Carapace (Figs 30, 32): Of typical salticid shape, without constrictions or thoracic hump. Narrower than L. herberti. Chelicera (Fig. 29): Vertical, with ectal spur on the paturon. At least five retromarginal teeth. Palp (Figs 21, 22, 25–27): Bulb round. Embolus circles 1.7 times around, ending in broad tip. RTA with dorsal branch slender, pointed, pointing distally. First leg tibia with five pairs of ventral spines (Fig. 28). Colour in alcohol (Figs 29–32): Markings much like those of L. herberti, though body darker. Honey coloured with black around eyes, dark brown on sides of thorax, and extensive black on abdomen. First leg with dark patches, appearing annulate.

Female (paratype in UBC–SEM). Carapace length 1.89; abdomen length 2.45. Carapace (Fig. 33): As in male. First leg tibia with five pairs of ventral spines. Epigyne (Figs 23, 24): With triangular openings pointing to the posterior. Colour in alcohol (Fig. 33): As in male, but with weaker dark patches on first legs.

The range of this species may extend to the east. A female (specimen PNG2008-3321 in UBC–SEM) slightly differing in epigynal structure is tentatively assigned to this species. It is pictured in Figs 34–37, and has data: PAPUA NEW GUINEA: Eastern Highlands Province: Mt Gahavisuka Provincial Park. S 6.015 E 145.412. elev. 2320 m a.s.l. 1–2 August 2008. W Maddison leg. WPM#08-025.

Figures 21–33. 

Leviea lornae sp. n., holotype (25–32) and paratypes. 21, 22 Left palp 21 ventral view 22 retrolateral view 23 female epigyne, ventral view 24 cleared vulva, dorsal view 25 left palp, prolateral view 26 same, ventral view 27 same, retrolateral view 28 first leg, prolateral view 29 male habitus, oblique lateral-frontal view; arrows shows ectal spurs on paturon 30 dorsal view 31 ventral view 32 lateral view 33 female habitus dorsal view. Scale bars: 0.1 mm (on genitalia); 1.0 mm (otherwise).

Holotype: male, DNA voucher d254, in UBC–SEM, with data PAPUA NEW GUINEA: Enga Province: Kai-ingri. 5.574 S 143.048 E. 3315 m a.s.l. 5–8 July 2008. W Maddison leg. WPM#08-004. Paratype: one male, in UBC–SEM, with data Papua New Guinea: Enga Province: Kumul Lodge @ foot of Mt Hagen, 05.47.548 S 143.58.761 E. 2700 m. 5.xii.2006. Balke & Kimbel leg. (PNG124).

Named in honour of Frances Levi.

Larger and more slender than L. herberti, with more (5 pairs) ventral macrosetae on the first tibia. In these features it resembles L. lornae. Differs from both L. herberti and L. lornae in details of the palp: dorsal branch of RTA swollen basally; bulb oval rather than circular, less rotated than L. herberti.

Male (holotype). Carapace length 2.08; abdomen length 2.38. Carapace (Fig. 41): Like that of L. lornae. Chelicera: Ectal spur on paturon small. Five retromarginal teeth. Palp (Figs 38, 39): Bulb compressed slightly laterally to be oval rather than circular. Embolus thick, circling 1.7 times around. First leg tibia with five pairs of ventral spines. Colour in alcohol (Fig. 41): much like that of L. herberti.

Female. Unknown.

Figures 34–37. 

Leviea cf. lornae, female from Mt. Gahavisuka. 34–36 Living specimen 37 epigyne, ventral view. Scale bars: 0.1 mm (on epigyne); 1.0 mm (otherwise).

Figures 38–41. 

Leviea francesae sp. n., holotype male. 38, 39 Left palp 38 ventral view 39 retrolateral view of tibia 40 carapace 41 habitus dorsal view; two photographs joined. Scale bars: 0.1 mm (on palp); 1.0 mm (on carapace).

Agorioides cherubino sp. n.

Named for the spiders’ superficial resemblance to Agorius Thorell, 1877.

Antlike, with concave-sided carapace, swollen first femur, a long ocular quadrangle, long fourth trochanters, and a spinose first tibia. The carapace is sunken inward (concave) between the PME and PLE, leaving the PLE on prominent tubercles, and yielding a constriction that resembles that of hisponine salticids. The femur of the first leg is shaped like a bird’s lower leg (“drumstick”), swollen in the proximal half but thin distally. The length of the ocular quadrangle is distinctly more than half the length of the carapace. The fourth trochanter is unusually long, longer than either the coxa or the fourth tarsus. Unlike Leviea and Papuamyr, the paturon of the chelicera lacks an ectal spur, and the first tibia has many pairs of long macrosetae; both of these features can be found in some species of Myrmarachnina.

The two species described are closely similar, but distinct in the form of the palp, shape of the carapace, and in colour. They were found only seven km apart, but at distinct elevations (570 m vs. ca. 1000 m). In other salticid genera, closely related species have been observed to segregate along such an elevational gradient in the same area (e.g., Cucudeta zabkai Maddison, 2009 vs. Cucudeta uzet Maddison, 2009 at 1170 m vs. 1450 m [Maddison 2009]).

Holotype: male in UBC–SEM, specimen code PNG2008-2854, with data PAPUA NEW GUINEA: Southern Highlands Province: Putuwé, junction of Lagaip & Uruwabwa Rivers. 5.231 S 142.532 E. 570 m a.s.l. 23–26 July 2008. W Maddison & Luc Fimo Tuki leg. WPM#08-019. On leaf litter.

In the Levis’ country home they hosted many animals, domesticated and not. One of these was a dog of multifarious ancestry, Cherubino, named after the character in Mozart’s opera The Marriage of Figaro. The spider resembles the dog in having a hairy, grizzled appearance.

Differs from A. papagena in having the tibia of the male palp distinctly narrower than the cymbium (Figs 42 vs. 51), the cephalic area distinctly higher (Figs 44 vs. 53), and in the black body with a dusting of white setae. The bulb of the palp is rotated slightly more than in A. papagena (see Diagnosis for that species).

Male (holotype). Carapace length 2.16; abdomen length 2.27. Carapace (Figs 44–50): Strangely shaped, as described for the genus, with a constriction between the PME and PLE. Thoracic hump prominent (Fig. 44). Ocular quadrangle occupies more than half of the length of the carapace. Clypeus extremely narrow. Chelicera: Vertical, simple. Teeth not examined for fear of damaging the specimen. Palp (Figs 42, 43): Embolus wrapping around bulb more than once; RTA simple and unbranched. Legs with relatively few short setae, except a greater density yields weak fringes beneath metatarsi and tarsi 2–4 and tibiae 3–4. First tibia with seven pairs of long ventral macrosetae; first metatarsus with two pairs. First femur distinctly swollen in proximal half, shaped like a bird leg’s drumstick (Figs 47, 49). Fourth trochanters distinctly longest, longer than the fourth tarsi (Figs 44, 48). Colour in life (Figs 48–50): Black body with fully or partly erect white setae on the sides and back of the carapace, the abdomen, and the femora of the fourth legs. First and fourth femora black; other segments translucent white to honey with darker lines or patches. Colour in alcohol (Figs 44–47): Carapace brown with black around eyes. Abdomen black except paler around the constriction and anteriorly. Clypeus dark, with only a few setae. Chelicerae honey coloured. Palp black. Legs pale except black on proximal half of first femur, brown to black fourth femur, and black lines on anterior faces of legs 2–4 and posterior faces of legs 3 and 4.

Female. Unknown.

One penultimate instar male (specimen PNG2008-2765, in UBC–SEM), with same data as holotype, also from leaf litter. It has a black body and leg markings as in holotype, but is more glabrous, with only a few white setae.

Figures 42–50. 

Agorioides cherubino sp. n., holotype male. 42, 43 Left palp 42 ventral view 43 retrolateral view of tibia 44–46 carapace 44 lateral view 45 dorsal view 46 oblique dorsal-lateral view 47 male habitus, dorsal view 48–50 living specimen. Scale bars: 0.1 mm (on genitalia); 1.0 mm (otherwise).

Holotype: male in UBC–SEM, specimen code PNG2008-1706 and DNA voucher code d253, with data PAPUA NEW GUINEA: Southern Highlands Province: Tualapa, near Wanakipa. 5.283 S 142.498 E. 1000–1100 m a.s.l. 11–22 July 2008. W Maddison & Luc Fimo Tuki leg. WPM#08-008. Forest interior and riverside on leaf litter.

The Levis’ love of opera was reflected in their animals’ names, including their dog Papagena, named after the character in Mozart’s The Magic Flute. The spider does not look like the dog or the opera character. Although the holotype is a male, the species is named for the female that remains to be found.

Differs from A. cherubino in having the tibia of the male palp almost as wide as the cymbium (Fig. 51), flatter profile of the carapace (Fig. 53), and a distinctly orange body and legs. The bulb of the palp is rotated less in A. papagena than in A. cherubino, as seen by the orientation of the spermophores in Fig. 51 versus Fig. 42. Although this could be due to a slight expansion of the palp of the one known specimen of A. papagena, suggested by the offset of the tip of its embolus from the cymbial apical groove, this offset is of lesser angle than the difference in spermophore orientation.

Male (holotype). Carapace length 2.84; abdomen length 3.24. Structure of body, legs as in A. cherubino (Figs 53–57), with first leg having swollen femur (Figs 57, 58). Chelicera (Fig. 54): Vertical, though robust. Four retromarginal teeth. Palp (Figs 51, 52): Embolus wrapping around bulb more than once; RTA simple and unbranched. Tibia distinctly wider than in A. cherubino. Colour in life (Fig. 58): Orange, darkening to brown on the abdomen and with black around the eyes. Swollen femur of the first leg particularly bright orange. Some partially-erect white setae on carapace sides beneath the eyes, on thorax, and on fourth femora, but not as dense or distinct as in A. cherubino. Colour in alcohol (Figs 53, 54, 56, 57): Except for black around the eyes, carapace, legs and palpi are orange, darkest on the thorax and palest at the leg tarsi. Abdomen a muted orangish gray.

Female. Unknown.

One juvenile (specimen PNG2008-1676, in UBC–SEM, Fig. 59), similarly coloured, with same data as holotype, on leaf litter.

Figures 51–59. 

Agorioides papagena sp. n., holotype male except for 59 (juvenile). 51, 52 Left palp: 51 ventral view 52 retrolateral view of tibia 53 Carapace, lateral view. 54 Frontal-lateral view 55, 56 dorsal habitus 57 first leg, prolateral view (two photographs joined at broken patella-tibia joint) 58 living holotype 59 juvenile from type locality. Scale bars: 0.1 mm (on genitalia); 1.0 mm (otherwise).

A single large female from the Muller Range is clearly an Agorioides by carapace shape and long fourth trochanters (the first legs are missing), but is not formally described here because the specimen is missing most of its legs. It seems likely to represent a distinct species, as it has a carapace that is flatter (Fig. 65) than that of the two described above. It is notably larger than A. cherubino (carapace 3.00, abdomen 3.43). Its epigyne is typical myrmarachnine, with the RTA pocket displaced anteriorly (Figs 62, 63). Its data are: PAPUA NEW GUINEA: Western Province: Muller Range, Camp 1, Gugusu. 05.7292S, 142.2633E. 515m elev. 4–11 September 2009. I Agnarsson leg.

Figures 60–65. 

Agorioides sp., female from Gugusu, Muller Range. 60 Habitus, dorsal view 61 carapace, dorsal view 62 epigyne, ventral view 63 cleared vulva, dorsal view 64 carapace, oblique dorsal-lateral view 65 carapace, lateral view. Scale bars: 0.1 mm (on genitalia); 1.0 mm (otherwise).

Papuamyr omhifosga sp. n.

A blend of Papua and the first syllable of the name of the related genus Myrmarachne. To be treated as feminine, as is Myrmarachne.

Small antlike salticids with somewhat swollen male first legs, vertical and excavated male chelicerae with an ectal spur on the paturon, a round bulb on the palp, and a relatively long RTA. In contrast, the Ligonipes group of genera has the bulb of the male palp compressed laterally (Davies and Żabka 1989: figs 7 and 9), while most Myrmarachne males have projecting chelicerae and thin first legs. An apparent synapomorphy uniting the two Papuamyr species described below is a complex fold in the embolus at the point where it crosses over previous loops before terminating (see black arrows in Figs 70 and 78).

Papuamyr lacks the morphological diagnostic characters of the Ligonipedina and Myrmarachnina, but we have not found morphological characters that place it more clearly. The molecular data, however, are clear that Papuamyr is within the Levieina and a close relative of Agorioides (Fig. 1).

Holotype: male, specimen UBC–SEM AR00215 in UBC–SEM, with data PAPUA NEW GUINEA: Southern Highlands Province: Putuwé, junction of Lagaip & Uruwabwa Rivers. 5.231 S 142.532 E. 570 m a.s.l. 23–26 July 2008. W Maddison & Luc Fimo Tuki leg. WPM#08-019. Beating. Paratypes: 5 males, 5 females, 2 juveniles, with same data.

A combination of letters derived from the first letters of words in a statement of Herbert Levi’s from approximately 1985. He had forgotten to do some bureaucratic task, and in dismay, he exclaimed “Och, my head is full of spider genitalia!” Truly, it was, and for that knowledge which he conveyed to us, arachnology is forever enriched. The name is particularly apt for this species: its genitalia are distinctive and elegant, the palp having a long transparent RTA and sharply bent embolus, the epigyne an RTA pocket displaced far to the anterior.

Distinct for its unusual genitalia and its orange and black bicoloured body. The embolus tip extends onto the retrolateral side of the cymbium before looping back to terminate ventrally, much as seen in many amycoid salticids such as Tartamura (Bustamante and Ruiz 2017). Most notable, however, is the long RTA whose posterior branch extends almost to the tip of the cymbium (Fig. 67). Accordingly, the RTA pocket of the epigyne is displaced far anteriorly (Fig. 68).

Male (holotype). Carapace length 1.31; abdomen length 1.24. Carapace (Figs 71–75): Narrow and flat, with a thoracic hump but without a strong constriction (Fig. 72). Carapace glabrous and shiny, without evident microsculpture. Ocular quadrangle less than half the length of the carapace. Clypeus extremely narrow. Chelicera (Fig. 71): Excavated medially, and with an ectal spur on the paturon. Five very small retromarginal teeth. Palp (Figs 66, 67, 70): Bulb round, with embolus with a sharp bend near the terminus (black arrow in Fig. 70), at the point where it passes a previous loop, just before extending to the retrolateral side of the cymbium. Ventral lobe of RTA projects ventrally; dorsal lobe extremely long. Legs with relatively few, short setae. First legs distinctly more robust than others. First tibia with two short anterior ventral macrosetae in the distal third, and two extremely small posterior ventral macrosetae in the proximal two thirds. Colour (Figs 71–75): Orange (vibrant in life, pale in alcohol) except for the dark cephalic region, the black posterior half of the abdomen, and dark lines on legs 3 and 4.

Female (paratype, specimen # PNG2008-2603). Carapace length 1.13; abdomen length 1.46. Carapace (Figs 76, 77): As in male. Chelicera: At least four retromarginal teeth. Legs similar to male except for the smaller first leg. First tibia with ventral macrosetae longer than in male, of normal length. Four macrosetae on first tibia (as in male, from distal to basal one anterior- one pair - one posterior), though five in other specimens (two pairs distally, and one posterior basally). Epigyne (Figs 68, 69): Of typical myrmarachnine form (see description of the tribe), except that RTA pocket is far to the anterior. Colour (Figs 76, 77): As in male.

Figures 66–77. 

Papuamyr omhifosga sp. n., holotype (66, 67, 70, 75) and paratypes. 66, 67 Left palp of holotype 66 ventral view 67 retrolateral view 68 epigyne of specimen PNG2008-2603, ventral view 69 cleared vulva of same specimen, dorsal view 70 left palp of holotype, oblique ventral-terminal view; arrow shoes bend in embolus 71 face of male, oblique frontal-lateral-dorsal view; arrows show ectal spurs on paturons 72 carapace, lateral view 73, 74 living male 75 habitus, dorsal view, holotype male 76 habitus, dorsal view, female UBC–SEM AR00216 77 living female. Scale bars: 0.1 mm (on genitalia); 1.0 mm (otherwise).

Holotype: male, specimen in RBINS, with data PAPUA NEW GUINEA: Madang Province, Oromongu, 5.73S, 142.53E. 700 m a.s.l. 26 January 2014. Maurice Leponce leg. #P5097. Mixed lowland forest Mixed evergreen forest of foothills and mountains, beating. Paratypes: 3 males, 3 females, with same data (from collecting events: males: #P5088, #P5103, #P5110, females: #P5082, #P5071, #P5106).

From the Greek “all giving” or “all gifted”, referring primarily to the gifts given us by the Levis, most memorably the 1254 species that Herb described, and the wonderfully abundant Thanksgiving celebration dinners that Lorna put on for their students each year. It is also the name of the dog Lorna had when Herb and Lorna were married.

Distinct in having the thoracic hump higher than the ocular area in both sexes (Figs 84, 87, 89). Males can be recognized by the shape of the RTA, with dorsal branch shorter than in P. omhifosga and twisted, and a very short ventral flange (Fig. 80). Females can be recognized by the lack of anterior pocket, the relatively large spermathecae (Fig. 81), and the long lateral extension of the sclerotized ducts near the posterior margin (Fig. 82).

Male (holotype). Carapace length 1.52; abdomen length 1.41. Carapace (Figs 83–85): Thoracic slope with a large hump which is higher than the flat ocular area (Fig. 84). Ocular quadrangle with rugose integument (Fig. 85), approximately as long as wide, occupying half of the carapace (0.75 long). Clypeus extremely narrow (Fig. 83). Chelicera (Figs 83, 84): Excavated medially, with an ectal spur on the paturon. Palp (Figs 78–80): cymbium with 3–4 macroseta on the apical part, bulb round, embolus looped twice around it, with a twist on the prolateral side of the bulb (black arrow Fig. 78) and a slight bend at the end. Ventral flange of RTA small, a slight hump; dorsal lobe well developed, with an elongated “S” shape. Legs: Leg I robust, each segment is at least twice as wide as other legs. Other legs slender. Abdomen: pear shaped, with a slight constriction at the anterior fourth/third. Colour (Figs 85–87 in alcohol): Body dark brown with a transverse pale band in the middle of the carapace and abdomen. Coxa I, leg II, proximal half of coxa IV, patella III-IV, metatarsus III, and all tarsi pale yellow. Trochanter and femur I dark brown, patella and metatarsus I dark yellow with a dark ventral side. Leg III-IV black except the pale yellow segments mentioned earlier (Figs 85–87).

Female: (paratype, from collecting event #P5013). Carapace length 1.41; abdomen length 1.54. Carapace (Figs 88, 89): As in male. Chelicera: unmodified. Legs similar to male except for the less robust first leg. Epigyne (Figs 81, 82): Spermathecae large, touching each other. Sclerotized ducts of spermathecal complex oriented laterally at back margin before proceeding anterior to the fertilization duct. Colour (Figs 88, 89): As in male.

Figures 78–89. 

Papuamyr pandora sp. n., holotype, except 81, 82 and 88–89 (paratype). 78–80 Left palp 78 prolateral view 79 ventral view 80 retrolateral view 81 female epigyne, ventral view 82 cleared vulva, dorsal view 83 male carapace, frontal view 84 oblique lateral-ventral view showing patural teeth (indicated by arrows) 85 male habitus, dorsal view 86 ventral view 87 lateral view 88 female habitus dorsal view 89 lateral view. Scale bars: 0.1 mm (on genitalia and 83); 1.0 mm (otherwise).

Figure 90. 

Distribution of levieines in Papua New Guinea.