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Article title
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Abstract
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Keywords
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Introduction
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Materials and methods
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Study areas and forest features
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Beetle sampling
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Site characteristics
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Data analysis
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Results
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Catch, species diversity, and beetle assemblage patterns
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Regional scale assemblage structure and β-diversity
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Local scale assemblage structure and β-diversity at Lac la Biche
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Interannual variability
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Discussion
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Spatial variation in carabid and staphylinid assemblages
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Regional differences in beetle catch and diversity
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Regional variation in beetle assemblage structure
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Environmental correlates of beetle assemblage structure
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Regional β-diversity
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Local patterns of assemblage structure and β-diversity at Lac la Biche
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Conclusions
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Acknowledgements
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References
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Supplementary material
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