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Article title
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Abstract
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Keywords
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Introduction
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Systematics
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Clade: CAENOGASTROPODA Cox
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Group 1. Radial sculpture predominant
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Group 2. Spiral sculpture predominant, or radial and spiral sculpture about equally strong, or virtually no sculpture present
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Group 1. Teleoconch whorl entirely detached
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Group 2. Teleoconch whorls (almost) entirely attached. Supraperipheral usually cord present
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Group 3. Teleoconch whorls (almost) entirely attached. Supraperipheral cord absent (check also D. tyloacron)
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Unranked clade: NERITOMORPHA
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Clade: PULMONATA Cuvier
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Group 1. Shell white
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Group 2. Shell pale corneous, to yellowish green or brown
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Group 1. Last whorl with 2 or more distinct spiral threads (rare specimens of P. tambunanensis and P. kusana have 1 spiral thread)
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Group 2. Last whorl with 1 spiral thread, or without a spiral thread
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Group 1. Protoconch with spaced spiral threads which are predominant over the radial sculpture
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Group 2. Protoconch with fine spiral grooves, usually about as strong as the radial sculpture or weaker, or protoconch without spiral sculpture
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Group 1a. Suture between the whorls coinciding with the periphery. Shell 13.5–24.5 mm wide in adult specimens
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Geotrochus labuanensis-group (G. labuanensis, G. kinabaluensis, G. oedobasis)
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Group 1b. Suture between the whorls coinciding with the periphery. Shell 10–12.5 mm wide in adult specimens (check also G. oedobasis)
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Group 2. Suture slightly below the periphery in the last 4–5 whorls of adult specimens
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Acknowledgements
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References
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