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The single most parsimonious tree retrieved in the cladistic analysis representing Sadocus relationships (182 steps, L = 182; C.I. = 45; R.I. = 53). Number at the nodes are Goodman-Bremer support. Black circles indicate unique transformations, while white circles indicate homoplastic transformations, the arrow indicates the position of the genus Sadocus. The character number is above each circle, and the character state is below. The characters are optimized in ACCTRAN. The character and character states are given in Table


Sadocus asperatus (Gervais). Male (


Sadocus dilatatus Roewer. Male holotype (


Sadocus funestus Butler. Male (


Sadocus ingens (Mello-Leitão). Male (


Sadocus polyacanthus (Gervais). Male (


Female specimens of Sadocus A, B S. asperatus (


Photographs of living specimens of Sadocus A S. asperatus (ma) B S. asperatus (fe) C S. asperatus (ma) D S. dilatatus (ma) E S. asperatus (fe) F S. funestus (ma). A, B taken by R. Pinto-da-Rocha, C–F taken by Pedro H. Martins.


Photographs of living specimens of Sadocus A S. ingens (ma) B S. ingens (fe) C S. polyacanthus (ma) D S. polyacanthus (fe) E S. polyacanthus (ma) F S. polyacanthus (fe). B, E taken by R. Pinto-da-Rocha; A, C, D, F taken by Pedro H. Martins.


Distal part of penis Sadocus and Discocyrtus, dorsal and ventral views, respectively A, B S. asperatus, macrosetae colorized and numbered C, D S. ingens E, F S. polyacanthus G, H S. dilatatus I, J S. funestus K, L D. catharinensis. Scale bar of drawings: 0.1mm.


Distal part of ovipositor of Sadocus A–C S. asperatus A upper view B villi C setae D–F S. funestus D upper view E lateral view F seta G–I S. ingens G upper view H, I setae J–L S. polyacanthus J upper view K lateral view L setae.