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A revised checklist of the Diplopoda of Georgia, with data on species distributions, both within and beyond the country, their statuses, and the main relevant literature sources. Designations: i – introduced; G – entire Georgia; W – western Georgia; C – central Georgia; E – eastern Georgia; R – Russian Caucasus; T – Turkey; Ar – Armenia; Az – Azerbaijan; Cr – Crimean Peninsula; (+) – present; e – endemic to Georgia; se – subendemic to Georgia; t – presumed troglobiont; sc – subcosmopolitan; EuM – Euro-Mediterranean; M – Mediterranean; EM – eastern Mediterranean; EE – eastern European; Ca – Caucasian.

Fauna G R T Ar Az Cr Elevations (m a.s.l.) and status Distribution pattern Main relevant references
Class Diplopoda
Order Polyxenida
Family Polyxenidae
Genus Polyxenus Latreille, 1803
1. Polyxenus lagurus (Linnaeus, 1758) W + + 20–1700, i sc Issaev 1911, Short et al. 2020
2. Polyxenus lankaranensis Short, Vahtera, Wesener & Golovatch, 2020 E + + 100–800 Ca Short et al. 2020
Genus Propolyxenus Silvestri, 1948 W
3. Propolyxenus argentifer (Verhoeff, 1921) G + + + + + 20–1700 EM Short et al. 2020
Family Lophoproctidae
Genus Lophoproctus Pocock, 1894
4. Lophoproctus coecus Pocock, 1894 G + 20–900 EM Short 2015, Short et al. 2020
Order Polyzoniida
Family Hirudisomatidae
Genus Hirudisoma Fanzago, 1881
5. Hirudisoma roseum (Victor, 1839) G + + + 20–1100, se EM Golovatch et al. 2015
Order Siphonocryptida
Family Siphonocryptidae
Hirudicryptus Enghoff & Golovatch, 1985
6. Hirudicryptus abchasicus Golovatch, Evsyukov & Reip, 2015 W + 600–1500, se Ca Golovatch et al. 2015, Zuev 2017
Order Glomerida
Family Glomeridae
Genus Hyleoglomeris Verhoeff, 1910
7. Hyleoglomeris awchasica (Brandt, 1840) W + 20–2100, se Ca Golovatch 1975, 1976a, 1989b
8. H. specialis Golovatch, 1989 E + 500–1400, se Ca Golovatch 1989b
Genus Trachysphaera Heller, 1858
9. Trachyspaera costata (Waga, 1857) G + + + + + 20–2000 EuM Golovatch 1990, 2008
10. T. fragilis Golovatch, 1976 G + 80–460, t, e Ca Golovatch 1976c,1990, Golovatch and Turbanov 2017
11. T. minuta Golovatch, 1976 G + + + 20–1700, se Ca Golovatch 1976c, 1990
12. T. orientalis Golovatch, 1976 W 800–1100, t, e Ca Golovatch 1976c, 1990
13. T. radiosa (Lignau, 1911) W + 20–1800, se Ca Golovatch 1976a, 1990
14. T. solida Golovatch, 1976 W, C 20–2020, se Ca Golovatch 1976c, 1976c, 1990, 1993
Family Glomeridellidae
Genus Typhloglomeris Verhoeff, 1898
15. Typhloglomeris lohmanderi (Golovatch, 1989) C, E + + 600–1450, se Ca Golovatch 1989a, 2003
16. Typhloglomeris palatovi Golovatch & Turbanov, 2018 W 650, t, e Ca Golovatch and Turbanov 2017
Order Chordeumatida
Family Anthroleucosomatidae
Genus Acanthophorella Antić & Makarov, 2016
17. Acanthophorella barjadzei Antić & Makarov, 2016 W 1120–1200, t, e Ca Antić and Makarov 2016
Genus Adshardicus Golovatch, 1981
18. Adshardicus strasseri Golovatch, 1981 W + 20–530, se Ca Enghoff 2006, Antić and Makarov 2016
Genus Alpinella Antić & Makarov, 2016
19. Alpinella waltheri Antić & Makarov, 2016 E 2860, e Ca Antić and Makarov 2016
Genus Brachychaetosoma Antić & Makarov, 2016
20. Brachychaetosoma turbanovi Antić & Makarov, 2016 W 300, t, e Ca Antić and Makarov 2016
Genus Caucaseuma Strasser, 1970
21. Caucaseuma kelasuri Antić & Makarov, 2016 W 190, e Ca Antić and Makarov 2016
22. C. variabile Antić & Makarov, 2016 C + 100–2500, se Ca Antić and Makarov 2016
Genus Cryptacanthophorella Antić & Makarov, 2016
23. Cryptacanthophorella manubriata Antić & Makarov, 2016 W, C 800–1700, e Ca Antić and Makarov 2016
Genus Dentatosoma Antić & Makarov, 2016
24. Dentatosoma denticulatum Antić & Makarov, 2016 W 400–900, e Ca Antić and Makarov 2016
25. D. magnum Antić & Makarov, 2016 W + 20–2200, se Ca Antić and Makarov 2016
26. D. zeraboseli Antić & Makarov, 2016 W 20–1700, e Ca Antić and Makarov 2016
Genus Georgiosoma Antić & Makarov, 2016
27. Georgiosoma bicornutum Antić & Makarov, 2016 W 2000, t, e Ca Antić and Makarov 2016
Genus Herculina Antić & Makarov, 2016
28. Herculina oligosagittae Antić & Makarov, 2016 W 1500–1700, e Ca Antić and Makarov 2016
29. H. polysagittae Antić & Makarov, 2016 C 1750, e Ca Antić and Makarov 2016
Genus Heterocaucaseuma Antić & Makarov, 2016
30. Heterocaucaseuma deprofundum Antić & Makarov, 2018 W 2000–2100, t, e Ca Antić et al. 2018
31. H. longicorne Antić & Makarov, 2016 W 100–350, t, e Ca Antić and Makarov 2016, Antić et al. 2018
32. H. mauriesi (Golovatch & Makarov, 2011) W 215, t, e Ca Golovatch and Makarov 2011, Antić and Makarov 2016, Antić et al. 2018
Genus Metamastigophorophyllon Ceuca, 1976
33. Metamastigophorophyllon giljarovi (Lang, 1959) W + 20–1850, se Ca Antić and Makarov 2016
34. M. hamatum Antić & Makarov, 2016 W + 150–2200, se Ca Antić and Makarov 2016
35. M. lamellohirsutum Antić & Makarov, 2016 W 700–800, e Ca Antić and Makarov 2016
36. M. torsivum Antić & Makarov, 2016 G + 800–1700, se Ca Antić and Makarov 2016
Genus Paranotosoma Antić & Makarov, 2016
37. Paranotosoma attemsi Antić & Makarov, 2016 W 1500–1800, e Ca Antić and Makarov 2016
38. P. cordatum Antić & Makarov, 2016 W 20–800, e Ca Antić and Makarov 2016
39. P. subrotundatum Antić & Makarov, 2016 W, C + 350–850, se Ca Antić and Makarov 2016
Genus Pseudoflagellophorella Antić & Makarov, 2016
40. Pseudoflagellophorella eskovi Antić & Makarov, 2016 C, E + + 100–2080, se Ca Antić and Makarov 2016
41. P. mirabilis Antić & Makarov, 2016 W 20–130, e Ca Antić and Makarov 2016
42. P. papilioformis Antić & Makarov, 2016 E + 850–2100, se Ca Antić and Makarov 2016
Genus Ratcheuma Golovatch, 1985
43. Ratcheuma excorne Golovatch, 1985 W 1180, t, e Ca Golovatch 1984/85, Antić and Makarov 2016
Order Julida
Family Blaniulidae
Genus Cibiniulus Verhoeff, 1927
44. Cibiniulus phlepsii (Verhoeff, 1897) W + 20–130 EuM Enghoff 1984, 2006
Genus Nopoiulus Menge, 1851
45. Nopoiulus brevipilosus Enghoff, 1984 W 130, t, e Ca Enghoff 1984, Golovatch and Enghoff 1990
46. N. densepilosus Enghoff, 1984 W + + 1500–1700 Ca Enghoff 1984, Golovatch and Enghoff 1990
47. N. golovatchi Enghoff, 1984 W + 20–130, se Ca Enghoff 1984, 1990
48. N. kochii (Gervais, 1847) G + + + + 10–2200, i? sc Enghoff 1984, Golovatch and Enghoff 1990
Family Nemasomatidae
Genus Nemasoma C.L. Koch, 1847
49. Nemasoma caucasicum (Lohmander, 1932) G + + + + 20–2000, se Ca Kobakhidze 1965, Enghoff 1985
Family Julidae
Genus Archileucogeorgia Lohmander, 1936
50. Archileucogeorgia abchasica Lohmander, 1936 W 130, t, e Ca Lohmander 1936
51. A. satunini Lohmander, 1936 W 130, e Ca Lohmander 1936
Genus Brachyiulus Berlese, 1884
52. Brachyiulus lusitanus Verhoeff, 1898` C + 100, i M Lohmander 1936
Genus Byzantorhopalum Verhoeff, 1930
53. Byzantorhopalum rossicum (Timotheew, 1897) W? + + + 30–1500 EE Lohmander 1936, Vagalinski and Lazányi 2018
Genus Catamicrophyllum Verhoeff, 1900
54. Catamicrophyllum caucasicum (Attems, 1901) G + + + 700–2000, se Ca Lohmander 1936, Enghoff 1995
Genus Calyptophyllum Brolemann, 1922
55. Calyptophyllum sp. W 100? ? Lohmander 1936, Enghoff 1995
Genus Chaetoleptophyllum Verhoeff, 1898
56. Chaetoleptophyllum flexum Golovatch, 1979 G + 15–2200, se Ca Golovatch 1979, Evsyukov et al. (2020)
Genus Cylindroiulus Verhoeff, 1894
57. Cylindroiulus bellus (Lignau, 1903) W? + + 100 EM Lignau 1903, Read 1992, Chumachenko 2016
58. C. crassiphylacum Read, 1992 W, C + 600–1700, se Ca Read 1992
59. C. kacheticus Lohmander, 1936 E + 500–1250, se Ca Lohmander 1936, Read 1992
60. C. olgainna Read, 1992 W 300–1100, e Ca Read 1992
61. C. parvus Lohmander, 1928 C, E + 500–2100, se Ca Lohmander 1936, Read 1992
62. C. placidus (Lignau, 1903) W, C + 20–2200, se Ca Lignau 1903, Read 1992
63. C. pterophylacum Read, 1992 W, C + 20–1600, se Ca Read 1992, Zuev 2014
64. C. quadrus Read, 1992 W, C 700–1000, e Ca Read 1992
65. C. ruber (Lignau, 1903) W + 100–2000, se Ca Lignau 1903, 1915, Read 1992
66. C. schestoperovi Lohmander, 1936 W + 400–1800, se Ca Lohmander 1936, Read 1992
67. C. truncorum (Silvestri, 1896) W + + 130, i sc Read 1992
Genus Grusiniulus Lohmander, 1936
68. Grusiniulus redikorzevi Lohmander, 1936 C 800–900, e Ca Lohmander 1936, Vagalinski and Lazányi 2018
Genus Julus Linnaeus, 1758
69. Julus colchicus Lohmander, 1936 G + + 20–2850, se Ca Lohmander 1936, Enghoff 2006, Evsyukov et al. 2018
70. J. kubanus Lohmander, 1936 W, E + 300–2100, se Ca Lohmander 1936, Kobakhidze 1965, Evsyukov et al. 2018
71. J. lignaui Verhoeff, 1910 W + 1500–2800, se Ca Evsyukov et al. 2018
72. J. lindholmi Lohmander, 1936 W + 450–2200, se Ca Lohmander 1936, Evsyukov et al. 2018
Genus Kubaniulus Lohmander, 1936
73. Kubaniulus gracilis Lohmander, 1936 W + 20–700, se Ca Lohmander 1936, Evsyukov et al. 2020
Genus Leptoiulus Verhoeff, 1894
74. Leptoiulus hastatus Lohmander, 1932 C + 800–1530, se Ca Lohmander 1936, Enghoff 2006, Evsyukov et al. 2020
75. L. tanymorphus (Attems, 1901) C, E + + + 80–1800, se Ca Lohmander 1936, Evsyukov et al. 2020
Genus Leucogeorgia Verhoeff, 1930
76. Leucogeorgia longipes Verhoeff, 1930 W 170, t, e Ca Verhoeff 1930, Barjadze et al. 2019
77. L. rediviva Golovatch, 1983 W 330, t, e Ca Golovatch 1983, Barjadze et al. 2019
Genus Megaphyllum Verhoeff, 1894
78. Megaphyllum dioscoriadis (Lignau, 1915) W + 130–1400, se Ca Lignau 1915, Lohmander 1936, Kobakhidze 1965, Chumachenko 2016, Vagalinski and Lazányi 2018
79. M. hercules (Verhoeff, 1901) W + 20, i EM Lazányi and Vagalinski 2013
80. M. spathulatum (Lohmander, 1936) W? ? ? Ca Lohmander 1936, Lazányi and Vagalinski 2013
Genus Omobrachyiulus Lohmander, 1936
81. Omobrachyiulus adsharicus (Lohmander, 1936) W 20–30, e Ca Lohmander 1936, Vagalinski and Lazányi 2018
82. O. brachyurus (Attems, 1899) G + + + + 20–2500 EM Lohmander 1936, Kobakhidze 1965, Enghoff 2006, Vagalinski and Lazányi 2018
83. O. curvocaudatus (Lignau, 1903) W + 30–1700, se Ca Lohmander 1936, Kobakhidze 1965, Vagalinski and Lazányi 2018
84. O. divaricatus (Lohmander, 1936) G + 600 –2000, se Ca Lohmander 1936, Kobakhidze 1965, Vagalinski and Lazányi 2018
85. O. hortensis (Golovatch, 1981) W 150, e Ca Golovatch 1981, Vagalinski and Lazányi 2018
86. O. implicitus Lohmander, 1936 (= O. i. ritsensis (Golovatch, 1981)) W + 400–1800, se Ca Lohmander 1936, Chumachenko 2016, Vagalinski and Lazányi 2018, Vagalinski in litt.
87. O. macrourus (Lohmander, 1928) (= O. m. abchasicus (Lohmander, 1936)) W, C 130–2000, e Ca Lohmander 1936, Kobakhidze 1965, Vagalinski and Lazányi 2018, Vagalinski in litt.
Genus Pachyiulus Berlese, 1883
88. Pachyiulus flavipes (C.L. Koch, 1847) W + 30, i M Lohmander 1936
89. P. krivolutskyi Golovatch, 1977 W + 20–1800, se Ca Golovatch 1977, Evsyukov 2016
Genus Syrioiulus Verhoeff, 1914
90. Syrioiulus adsharicus (Lohmander, 1936) W 120, e Ca Lohmander 1936, Golovatch 2018
91. S. georgicus (Lohmander, 1932) C 800–900, e Ca Lohmander 1932, Golovatch 2018
Order Polydesmida
Family Paradoxosomatidae
Genus Oxidus Cook, 1911
92. Oxidus gracilis (C.L. Koch, 1847) W + 20–100, i Ca Lignau 1915, Lohmander 1936, Chumachenko 2016
Genus Strongylosoma Brandt, 1833
93. Strongylosoma kordylamythrum Attems, 1898 G + + 20–2200 Ca Lohmander 1936, Kobakhidze 1965, Evyukov et al. 2016
94. S. lenkoranum Attems, 1898 C + + + 80–1650 Ca Lohmander 1936, Kobakhidze 1965, Evyukov et al. 2016
Family Polydesmidae
Genus Brachydesmus Heller, 1858
95. Brachydesmus assimilis Lohmander, 1936 C, E + + + 600–2800, se Ca Golovatch et al. 2016
96. B. furcatus Lohmander, 1936 W + 20–1900, se Ca Golovatch et al. 2016
97. B. kalischewskyi Lignau, 1915 G + + + + 50–2400, se Ca Golovatch et al. 2016
98. B. kvavadzei Golovatch, Evsyukov & Reip, 2016 W 70–1520, e Ca Golovatch et al. 2016
99. B. simplex Golovatch, Evsyukov & Reip, 2016 W + 20–1100, se Ca Golovatch et al. 2016
100. B. superus Latzel, 1884 W + 150–450, i sc Golovatch et al. 2016
Genus Polydesmus Latreille, 1803
101. Polydesmus abchasius Attems, 1899 W, C + 10–2230, se Ca Golovatch et al. 2016
102. P. lignaui Lohmander, 1936 W + 100–2200, se Ca Golovatch et al. 2016
103. P. mediterraneus Daday, 1889 W + 100, i EM Golovatch et al. 2016
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