Description of Onthophagus humboldti and Uroxys bonplandi, two new scarab beetles (Coleoptera, Scarabaeidae, Scarabaeinae) from Costa Rica, with notes on tropical mountain brachyptery and endemicity

Bert Kohlmann; Ángel Solís; Guillermo Alvarado

doi:10.3897/zookeys.881.38026

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ZooKeys 881: 23-51
https://doi.org/10.3897/zookeys.881.38026 (17 Oct 2019)

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Roisin A. Stanbrook, W. Edwin Harris, Charles P. Wheater, Martin Jones (2021)
Evidence of phenotypic plasticity along an altitudinal gradient in the dung beetle Onthophagus proteus. PeerJ 9: e10798.

DOI: 10.7717/peerj.10798

BERT KOHLMANN, ÁNGEL SOLÍS (2025)
A review of the species groups of the Western Hemisphere Onthophagus Latreille (Coleoptera: Scarabaeidae: Scarabaeinae) using COI barcoding and gene trees. Zootaxa 5604: 401.

DOI: 10.11646/zootaxa.5604.4.1

VICTOR MOCTEZUMA, ALEJANDRO ESPINOSA DE LOS MONTEROS, GONZALO HALFFTER (2024)
Phylogenetic analyses of the subfamily Scarabaeinae (Coleoptera: Scarabaeidae) provide new insights into the Mexican Transition Zone theory. Zootaxa 5415: 501.

DOI: 10.11646/zootaxa.5415.4.1

Marianela Cambronero Sánchez (2025)
Catálogo de ejemplares tipo del orden Coleoptera (Insecta) en el Museo Nacional de Costa Rica. ACTA ZOOLÓGICA MEXICANA (N.S.) : 1.

DOI: 10.21829/azm.2025.4112742

Mario Cupello, Fernando A. B. Silva, Fernando Z. Vaz-de-Mello (2023)
The Taxonomic Revolution of New World dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae). Frontiers in Ecology and Evolution 11: .

DOI: 10.3389/fevo.2023.1168754

Michele Rossini, François Génier (2024)
A New Classification for the Onthophagus clypeatus Group (Coleoptera: Scarabaeidae: Scarabaeinae) and Description of a New Species from Western Ecuador. The Coleopterists Bulletin 78: .

DOI: 10.1649/0010-065X-78.3.415

Alfonsina Arriaga-Jiménez, Bert Kohlmann, Lorenzo Vázquez-Selem, Yhenner Umaña, Matthias Rös (2020)
Past and future sky-island dynamics of tropical mountains: A model for two Geotrupes (Coleoptera: Geotrupidae) species in Oaxaca, Mexico. The Holocene 30: 1462.

DOI: 10.1177/0959683620932977

Giulio Montanaro, Vasily V Grebennikov, Michele Rossini, Alessandro Grapputo, Enrico Ruzzier, Sergei Tarasov, Julian Dupuis (2024)
Microallopatric speciation in the relict dung beetle genus Grebennikovius (Coleoptera: Scarabaeidae) in the Eastern Arc Mountains. Insect Systematics and Diversity 8: .

DOI: 10.1093/isd/ixae004

Ángel Solís, Bert Kohlmann, Guillermo E. Alvarado (2024)
Landscapes and Landforms of Costa Rica. Chapter 10: 189.

DOI: 10.1007/978-3-031-64940-0_10

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