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Thitarodes shambalaensis sp. nov. A holotype, male, dorsal view B paratype, female, dorsal view C holotype, male, ventral view D paratype, female, dorsal view E resting position in habitat (photograph credit Hua Zhang). Scale bar: 1 cm.
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A Natural habitat of Thitarodes shambalaensis sp. nov. at Yanzigou glacier entrance point, Mt. Gongga, Sichuan, China (photograph credit Meng Li, May 2018) B disturbed habitat of Thitarodes shambalaensis sp. nov. due to excavation of T. shambalaensis pupae, at Haizidang, Yanzigou glacier (photograph credit Wenbin Ju, May 2019).
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Thitarodes shambalaensis sp. nov. A forewing (up) and hindwing (bottom) venation from dorsal view B antenna, front view C labial palp, caudal view, covered under setae. Scale bar: 1 mm.
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Thitarodes shambalaensis sp. nov. A male genitalia from holotype, ventral view B valva, caudal view C pseudotegumen, lateral view D female genitalia from paratype. Scale bar: 1 mm.
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Thitarodes shambalaensis sp. nov. Foreleg (left), midleg (middle), hindleg (right) with each aerolium enlarged in black rectangles. Scale bar: 1 mm.
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Diagnostic character of Thitarodes shambalaensis sp. nov. in forewing venation in comparison to other species of Thitarodes. Scale bar: 1 mm.
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Diagnostic characters of Thitarodes shambalaensis sp. nov. in comparison to other species of Thitarodes A labial palp B valva C pseudotegumen. Scale bar: 1 mm.
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The distribution of Thitarodes species sampled in this study and historical records of Thitarodes species in the Kangding – Mt. Gongga region.
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A Maximum Likelihood tree of CO1 sequences of species sampled in this study (yellow, red and blue dots) and other known species of the genus Thitarodes (grey dots). Outgroups are in black dots. The corresponding position of the genus Ahamus designated by
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![](https://binary.pensoft.net/fig/352700/singlefigmini/file.jpg)
Summary tree for all known species of Thitarodes. Samples in this study are labeled in color bars (blue, red and yellow). Each tip represents a cluster of all samples that are within 2.5% genetic distance from each other, the length of the grey bar represents the number of individual samples falling into the cluster. Numbers after the dots enumerate the number of samples that fall into a particular sequence type. All named individual samples are labeled. The genus Ahamus as designated by