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Article title
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Abstract
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Keywords
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Introduction
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Methods
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Taxa examined
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Data analysis
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Results
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Comparison of pronotum/elytron morphology among lucanid subfamilies
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Ancestral reconstruction of the pronotum and elytron
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Species richness and pronotum and elytron morphology
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Discussion
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Evolution of the pronotum and elytron in Lucanidae
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The inconsistency between morphological diversity and species richness
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Conclusion
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Acknowledgements
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References
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Supplementary material
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