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Thelphusula capillodigitus sp. n., colour in life, holotype male (23.9 × 18.4 mm) (ZRC 2017.1294), Sabah. A dorsal view B frontal view (photographs Dennis Sim).


Thelphusula capillodigitus sp. n., holotype male (23.9 × 18.4 mm) (ZRC 2017.1294), Sabah. A overall view B dorsal view of carapace C frontal view of cephalothorax D left third maxilliped E pleon F anterior thoracic sternum and pleon G sternopleonal cavity.


Thelphusula capillodigitus sp. n., holotype male (23.9 × 18.4 mm) (ZRC 2017.1294), Sabah. A outer views of chelae B outer view of right chela C dorsal view of dactylus of right chela D surface of dactylus showing dense short setae E epistome F–I first to fourth ambulatory legs, respectively (all to same scale).


Thelphusula capillodigitus sp. n., holotype male (23.9 × 18.4 mm) (ZRC 2017.1294), Sabah. A left G1 (ventral view) B distal part of left G1 (ventral view) C distal part of left G1 (doral view) D left G2 (ventral view). Scale bars: 1.0 mm (A, D); 0.5 mm (B, C).


Habitat and life colour of Arachnothelphusa terrapes. A water-filled tree hole at base of tree in Danum Valley where crab was found B water filled tree hole where crabs were hiding C, D male (30.8 × 20.5 mm) (ZRC 2017.1205) E, F female (30.1 × 20.5 mm, with juvenile crabs) (ZRC 2017.1205).


A, B Thelphusula dicerophilus, female (26.1 × 21.3 mm) (ZRC 2017.1047) (in situ) C, D Parathelphusa valida, male (27.8 × 22.7 mm) (ZRC 2017.1269) E Terrathelphusa secula, holotype male (29.2 × 20.4 mm) (ZRC 2018.0297) (preserved colour) F Isolapotamon ingeri, male (57.4 × 43.5 mm) (ZRC 1997.0799) (preserved colour).


A Geosesarma danumense, ovigerous female (14.9 × 15.1 mm) (ZRC 2017.1273) B, C G. danumense, male (carapace width ca. 1.5–2.0 cm), on log approx. 30 cm from ground, ca. 30 m from river flowing through Danum Valley Field Centre, 10 pm, 25 July 2013, specimen not collected (photographs: Marcus Ng) D G. sabanum, male, on leaf above ground, Tawau Hills National Park, specimen not collected (photograph Ying Seawei).