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ZooKeys 716: 29-41
doi: 10.3897/zookeys.716.21150
Received: 21 Sep 2017 | Approved: 01 Nov 2017 | Published: 23 Nov 2017
This article is part of:
ZooKeys 716
Authors
   
   
   
Michael S. Engel - Corresponding author
University of Kansas, Lawrence, United States of America
American Museum of Natural History, New York, United States of America
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Abdulaziz S. Alqarni
King Saud University, Riyadh, Saudi Arabia
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Mohamed A. Shebl
King Saud University, Riyadh, Saudi Arabia
Suez Canal University, Ismailia, Egypt
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Javaid Iqbal
King Saud University, Riyadh, Saudi Arabia
Muhammad Nawaz Shareef University of Agriculture, Multan, Pakistan
https://faculty.ksu.edu.sa/en/jiqbal
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Ismael A. Hinojosa-Díaz - Corresponding author
Universidad Nacional Autónoma de México, Mexico City, Mexico
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Academic editor
Michael Ohl
Museum für Naturkunde, Berlin, Berlin, Germany
http://www.naturkundemuseum-berlin.de
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Author contributions
Engel, Alqarni, Shebl, and Iqbal participated in field work and specimen preparation in Saudi Arabia; Hinojosa-Diaz undertook dissections, prepared the photographs and figure plates, and worked on the key; Engel, Shebl, and Hinojosa worked on detailed comparisons with African and Asiatic Koptortosoma and tracking types; Engel composed the description; all authors participated in writing the manuscript.
Conflict of interest
The authors have declared that no competing interests exist.
Supporting agencies
KSU -  King Saud University
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
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