ADH, MF and MS performed the field work; EAV and MF accumulated additional museum material; EAV performed the majority of morphological studies, prepared the first draft and photodocumentation; EAV and MS did the molecular work, analysis of the data and prepared the data for submissions to BOLD; VS, MF and EAV prepared the datasets, wrote and tested the scripts, and submitted the data to BOLD, GBIF, GenBank and Flickr; all authors commented drafts of the paper at different stages and helped with completing the manuscript for submission.