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  • Article title
  • Abstract
  • Key words
  • Introduction
  • Materials and methods
    • Sampling and preservation
    • Morphology observation
    • DNA extraction, sequencing and phylogenetic analyses
  • Results
    • Systematics
    • Barcoding, phylogenetic relationships and taxonomic implication
  • Discussion
  • Acknowledgements
  • Additional information
    • Conflict of interest
    • Ethical statement
    • Funding
    • Author contributions
    • Author ORCIDs
    • Data availability
  • References
  • Supplementary material
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