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Article title
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Abstract
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Key words
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Introduction
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Materials and methods
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Measurements
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Indices
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Nomenclature
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Species concept
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Results
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Protanilla wallacei
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Leptanilla belantan
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Leptanilla acherontia
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Leptanilla bethyloides
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Leptanilla najaphalla
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Leptanillinae
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Opamyrmini
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Opamyrma
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Leptanillini
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Protanilla
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Protanilla rafflesi species group
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Protanilla bicolor species group
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Protanilla taylori species group
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Incertae sedis
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Protanilla rafflesi species group
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Protanilla bicolor species group
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Protanilla taylori species group
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Incertae sedis
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Leptanilla
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Leptanilla thai species group
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Leptanilla havilandi species group
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Leptanilla bethyloides species group
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Leptanilla najaphalla species group
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Leptanilla revelierii species group
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Incertae sedis
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Unplaced to species group
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Leptanilla thai species group
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Leptanilla havilandi species group
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Leptanilla bethyloides species group
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Leptanilla najaphalla species group
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Leptanilla revelierii species group
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Incertae sedis
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Unplaced to species group
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Worker-based keys to the Leptanillinae
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Worker-based key to the Protanilla taylori species group
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Worker-based key to the Protanilla bicolor species group
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Worker-based key to the Protanilla rafflesi species group
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Worker-based key to the Leptanilla thai species group and Leptanilla havilandi species group
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Worker-based key to the Leptanilla revelierii species group
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Male-based key to the major subclades of the Leptanillinae
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Male-based species-level key to the Protanilla rafflesi species group
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Male-based species-level key to the Leptanilla thai species group
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Male-based species-level key to the Leptanilla najaphalla species group
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Male-based species-level key to the Leptanilla havilandi species group
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Male-based species-level key to the Leptanilla bethyloides species group
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Male-based species-level key to the Leptanilla revelierii species group
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Discussion
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Taxonomic history
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Biogeography and ecology
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Acknowledgements
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Additional information
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Conflict of interest
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Ethical statement
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Funding
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Author contributions
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Author ORCIDs
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Data availability
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References
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Supplementary material
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