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  • Article title
  • Abstract
  • Key words
  • Introduction
  • Materials and methods
    • Taxonomic procedures and terminology
    • Photography and measurements
    • Dissections of genitalia and proventricules
    • Scanning electron microscopy
    • Molecular analysis
    • Assembly of data on bionomics
    • Assembly of data on distribution
  • Results and discussion
    • Morphology
    • Molecular analysis
    • Neotypes
    • Bionomics and distribution
    • Key to the European Cryphalus species
    • Descriptions
  • Acknowledgements
  • Additional information
    • Conflict of interest
    • Ethical statement
    • Funding
    • Author contributions
    • Author ORCIDs
    • Data availability
  • References
  • Supplementary material
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