Latest Articles from ZooKeys Latest 37 Articles from ZooKeys https://zookeys.pensoft.net/ Fri, 29 Mar 2024 10:34:27 +0200 Pensoft FeedCreator https://zookeys.pensoft.net/i/logo.jpg Latest Articles from ZooKeys https://zookeys.pensoft.net/ New species of redbait from the Philippines (Teleostei, Emmelichthyidae, Emmelichthys) https://zookeys.pensoft.net/article/111161/ ZooKeys 1196: 95-109

DOI: 10.3897/zookeys.1196.111161

Authors: Matthew G. Girard, Mudjekeewis D. Santos, Katherine E. Bemis

Abstract: We describe a new species of redbait in the genus Emmelichthys collected from fish markets on Panay and Cebu islands in the Visayas region of the Philippines. The species is externally similar to E. struhsakeri but is diagnosable by two prominent fleshy papillae associated with the cleithrum and fewer pectoral-fin rays (18–19 vs. 19–21) and gill rakers (30–33 vs. 34–41). Additionally, mitochondrial DNA differentiates this taxon from other species of Emmelichthys. We generate mitochondrial genomes for two of the three type specimens and several other emmelichthyids to place the new taxon in a phylogenetic context. Analysis of the protein-coding mitochondrial loci calls into question the monophyly of two emmelichthyid genera (Emmelichthys and Erythrocles) and highlights the need for subsequent analyses targeting the intrarelationships of the Emmelichthyidae.

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Research Article Fri, 22 Mar 2024 20:00:04 +0200
Systematic revision of the ant subfamily Leptanillinae (Hymenoptera, Formicidae) https://zookeys.pensoft.net/article/107506/ ZooKeys 1189: 83-184

DOI: 10.3897/zookeys.1189.107506

Authors: Zachary Griebenow

Abstract: The genus-level taxonomy of the ant subfamily Leptanillinae (Hymenoptera: Formicidae) is here revised, with the aim of delimiting genus-level taxa that are reciprocally monophyletic and readily diagnosable based upon all adult forms. This new classification reflects molecular phylogenetics and is informed by joint consideration of both male and worker morphology. Three valid genera are recognized in the Leptanillinae: Opamyrma, Leptanilla (= Scyphodon syn. nov., Phaulomyrma, Leptomesites, Noonilla syn. nov., Yavnella syn. nov.), and Protanilla (= Anomalomyrma syn. nov., Furcotanilla). Leptanilla and Protanilla are further divided into informal, monophyletic species groups. Synoptic diagnoses are provided for all genera and informal supraspecific groupings. In addition, worker-based keys to all described species within the Leptanillinae for which the worker caste is known are provided; and male-based keys to all species for which males are known, plus undescribed male morphospecies for which molecular data are published. The following species are described as new: Protanilla wallacei sp. nov., Leptanilla acherontia sp. nov., Leptanilla belantan sp. nov., Leptanilla bethyloides sp. nov., and Leptanilla najaphalla sp. nov.

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Monograph Tue, 16 Jan 2024 10:44:13 +0200
New species of Nigrobaetis from Southeast Asia (Ephemeroptera, Baetidae) https://zookeys.pensoft.net/article/102941/ ZooKeys 1166: 175-234

DOI: 10.3897/zookeys.1166.102941

Authors: Thomas Kaltenbach, Jean-Luc Gattolliat

Abstract: Based on larvae collected in the Philippines, Borneo (Kalimantan), Sumba, and Sumatra, the presence of the genus Nigrobaetis in the Philippines and Indonesia is reported for the first time. Six new species are described and illustrated, two from the Philippines and four from Indonesia. A larval key to all Nigrobaetis species of the Philippines, Indonesia and neighbouring continental Southeast Asia is provided, morphological differences to the species of Taiwan are discussed as well. The eggs of three new species are described, and the morphology of the eggs of Nigrobaetis is briefly discussed.

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Research Article Fri, 9 Jun 2023 16:44:13 +0300
Molecular data from the holotype of the enigmatic Bornean Black Shrew, Suncus ater Medway, 1965 (Soricidae, Crocidurinae), place it in the genus Palawanosorex https://zookeys.pensoft.net/article/94217/ ZooKeys 1137: 17-31

DOI: 10.3897/zookeys.1137.94217

Authors: Jonathan A. Nations, Thomas C. Giarla, Muhd Amsyari Morni, Julius William Dee, Mark T. Swanson, Anna E. Hiller, Faisal Ali Anwarali Khan, Jacob A. Esselstyn

Abstract: Although Borneo has received more attention from biologists than most other islands in the Malay Archipelago, many questions regarding the systematic relationships of Bornean mammals remain. Using next-generation sequencing technology, we obtained mitochondrial DNA sequences from the holotype of Suncus ater, the only known specimen of this shrew. Several shrews collected recently in Sarawak are closely aligned, both morphologically and mitochondrially, with the holotype of S. ater. Phylogenetic analyses of mitochondrial sequences indicate that the S. ater holotype and new Sarawak specimens do not belong to the genus Suncus, but instead are most closely related to Palawanosorex muscorum. Until now Palawanosorex has been known only from the neighboring Philippine island of Palawan. Additional sequences from nuclear ultra-conserved elements from the new Sarawak specimens strongly support a sister relationship to P. muscorum. We therefore transfer ater to Palawanosorex. The new specimens demonstrate that P. ater is more widespread in northern Borneo than previously recorded. Continued sampling of Bornean mammal diversity and reexamination of type material are critical in understanding the evolutionary history of the biologically rich Malay Archipelago.

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Short Communication Wed, 21 Dec 2022 10:40:17 +0200
A widespread new genus of Baetidae (Baetidae, Ephemeroptera) from Southeast Asia https://zookeys.pensoft.net/article/93800/ ZooKeys 1135: 1-59

DOI: 10.3897/zookeys.1135.93800

Authors: Thomas Kaltenbach, Nikita J. Kluge, Jean-Luc Gattolliat

Abstract: A reinvestigation of type and other material of Baetis javanicus Ulmer, 1913 and Baetis sabahensis Müller-Liebenau, 1984, together with new material from Southeast Asia revealed a new genus, Branchiobaetis gen. nov. The above species are formally assigned to the new genus Branchiobaetis gen. nov. It is characterized by the presence of accessory gills ventrally near fore coxa and at the base of maxillae, a peculiar folding of the gonostyli developing under the cuticle of last instar male larvae, together with a unique combination of other larval characters. Besides the two formerly described species, five new species are identified using a combination of morphology and molecular characters (COI, Kimura 2-parameter distances), four species from Sumatra and one from the Philippines. They are described and illustrated at the larval stage. Additionally, a complementary description of larva and adult stages of the generic type species B. javanicus comb. nov. as well as the first description of the eggs are provided. Furthermore, new reports of B. javanicus comb. nov. and B. sabahensis comb. nov. are indicated. The distribution of Branchiobaetis gen. nov. includes the Indonesian Sunda Islands, Borneo, and the Philippines. A key to the larval stage of all species is provided.

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Research Article Mon, 12 Dec 2022 18:46:58 +0200
Rhagophthalmidae Olivier, 1907 (Coleoptera, Elateroidea): described genera and species, current problems, and prospects for the bioluminescent and paedomorphic beetle lineage https://zookeys.pensoft.net/article/90233/ ZooKeys 1126: 55-130

DOI: 10.3897/zookeys.1126.90233

Authors: Robin Kundrata, Johana Hoffmannova, Kevin R. Hinson, Oliver Keller, Gabriela Packova

Abstract: Rhagophthalmidae are a small beetle family known from the eastern Palaearctic and Oriental realms. Rhagophthalmidae are closely related to railroad worms (Phengodidae) and fireflies (Lampyridae) with which they share highly modified paedomorphic females and the ability to emit light. Currently, Rhagophthalmidae include 66 species classified in the following 12 genera: Bicladodrilus Pic, 1921 (two spp.), Bicladum Pic, 1921 (two spp.), Dioptoma Pascoe, 1860 (two spp.), Diplocladon Gorham, 1883 (two spp.), Dodecatoma Westwood, 1849 (eight spp.), Falsophrixothrix Pic, 1937 (six spp.), Haplocladon Gorham, 1883 (two spp.), Menghuoius Kawashima, 2000 (three spp.), Mimoochotyra Pic, 1937 (one sp.), Monodrilus Pic, 1921 (two spp. in two subgenera), Pseudothilmanus Pic, 1918 (two spp.), and Rhagophthalmus Motschulsky, 1854 (34 spp.). The replacement name Haplocladon gorhami Kundrata, nom. nov. is proposed for Diplocladon hasseltii Gorham, 1883b (described in subgenus Haplocladon) which is preoccupied by Diplocladon hasseltii Gorham, 1883a. The genus Reductodrilus Pic, 1943 is tentatively placed in Lampyridae: Ototretinae. Lectotypes are designated for Pseudothilmanus alatus Pic, 1918 and P. marginalis Pic, 1918. Interestingly, in the eastern part of their distribution, Rhagophthalmidae have remained within the boundaries of the Sunda Shelf and the Philippines demarcated by the Wallace Line, which separates the Oriental and Australasian realms. This study is intended to be a first step towards a comprehensive revision of the group on both genus and species levels. Additionally, critical problems and prospects for rhagophthalmid research are briefly discussed.

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Research Article Tue, 1 Nov 2022 11:25:12 +0200
Battle of the bands: systematics and phylogeny of the white Goniobranchus nudibranchs with marginal bands (Nudibranchia, Chromodorididae) https://zookeys.pensoft.net/article/72939/ ZooKeys 1083: 169-210

DOI: 10.3897/zookeys.1083.72939

Authors: Giun Yee Soong, Lynn J. Bonomo, James D. Reimer, Terrence M. Gosliner

Abstract: Species identities of Goniobranchus nudibranchs with white bodies and various marginal bands have long been problematic. In this study, specimens of these Goniobranchus nudibranchs from the Philippines, Peninsular Malaysia, Japan, Papua New Guinea, and Madagascar were analyzed and molecular data were obtained in order to re-examine the relationships between species within this “white Goniobranchus with marginal bands” group. The analyses clearly recovered six species groups corresponding to the described species Goniobranchus albonares, G. preciosus, G. rubrocornutus, G. sinensis, and G. verrieri as well as one new species, G. fabulus Soong & Gosliner, sp. nov. Notably, G. preciosus, G. sinensis, G. rubrocornutus, G. verrieri, and G. fabulus Soong & Gosliner, sp. nov. exhibit color variation and polymorphism, suggesting that some aspects of color patterns (e.g., presence or absence of dorsal spots) may not always be useful in the identification of species in the “white Goniobranchus with marginal bands” group, whereas other features such as gill and rhinophore colors and the arrangement and colors of the mantle marginal bands are more diagnostic for each species.

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Research Article Tue, 25 Jan 2022 18:01:01 +0200
Two new species of Byrrhinus Motschulsky, 1858 (Coleoptera, Limnichidae, Limnichinae) from Negros, Philippines https://zookeys.pensoft.net/article/70531/ ZooKeys 1070: 51-72

DOI: 10.3897/zookeys.1070.70531

Authors: Emmanuel D. Delocado, Hendrik Freitag

Abstract: Two new species of Limnichidae beetles, Byrrhinus negrosensis sp. nov. and Byrrhinus villarini sp. nov., are described from the Island of Negros in the Philippines. The adult specimens of the new species can be differentiated by patterns of body punctation, colour and orientation of elytral pubescence, posterolateral angle of pronotum, tarsomere length ratio and aedeagal form. Two clades, representing the two new species, were retrieved in the Maximum Likelihood gene tree using the 3’-end of the COI gene. Maximum genetic divergence within B. negrosensis sp. nov. and B. villarini sp. nov. were recorded to be 2.3% and 1.3%, respectively, while the mean interspecific divergence between the two new species was 19.7%. Morphological descriptions, digital photographs and COI sequences were provided for the two species. The state of knowledge of Byrrhinus is reviewed and an updated Philippine checklist is provided. By coupling morphological and molecular data, this paper provides the first additional new species of Philippine Byrrhinus in the last 28 years.

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Research Article Wed, 10 Nov 2021 09:47:44 +0200
Cryptophyllium, the hidden leaf insects – descriptions of a new leaf insect genus and thirteen species from the former celebicum species group (Phasmatodea, Phylliidae) https://zookeys.pensoft.net/article/61033/ ZooKeys 1018: 1-179

DOI: 10.3897/zookeys.1018.61033

Authors: Royce T. Cumming, Sarah Bank, Joachim Bresseel, Jérôme Constant, Stéphane Le Tirant, Zhiwei Dong, Gontran Sonet, Sven Bradler

Abstract: While the leaf insects (Phylliidae) are a well-supported group within Phasmatodea, the genus Phyllium Illiger, 1798 has repeatedly been recovered as paraphyletic. Here, the Phyllium (Phyllium) celebicum species group is reviewed and its distinctiveness from the remaining Phylliini genera and subgenera in a phylogenetic context based on morphological review and a phylogenetic analysis of three genes (nuclear gene 28S and mitochondrial genes COI and 16S) from most known and multiple undescribed species is shown. A new genus, Cryptophyllium gen. nov., is erected to partially accommodate the former members of the celebicum species group. Two species, Phyllium ericoriai Hennemann et al., 2009 and Phyllium bonifacioi Lit & Eusebio, 2014 morphologically and molecularly do not fall within this clade and are therefore left within Phyllium (Phyllium). The transfer of the remaining celebicum group members from Phyllium Illiger, 1798 to this new genus creates the following new combinations; Cryptophyllium athanysus (Westwood, 1859), comb. nov.; Cryptophyllium celebicum (de Haan, 1842), comb. nov.; Cryptophyllium chrisangi (Seow-Choen, 2017), comb. nov.; Cryptophyllium drunganum (Yang, 1995), comb. nov.; Cryptophyllium oyae (Cumming & Le Tirant, 2020), comb. nov.; Cryptophyllium parum (Liu, 1993), comb. nov.; Cryptophyllium rarum (Liu, 1993), comb. nov.; Cryptophyllium tibetense (Liu, 1993), comb. nov.; Cryptophyllium westwoodii (Wood-Mason, 1875), comb. nov.; Cryptophyllium yapicum (Cumming & Teemsma, 2018), comb. nov.; and Cryptophyllium yunnanense (Liu, 1993), comb. nov. The review of specimens belonging to this clade also revealed 13 undescribed species, which are described within as: Cryptophyllium animatum gen. et sp. nov. from Vietnam: Quang Nam Province; Cryptophyllium bankoi gen. et sp. nov. from Vietnam: Quang Ngai, Thua Thien Hue, Da Nang, Gia Lai, Quang Nam, and Dak Nong Provinces; Cryptophyllium bollensi gen. et sp. nov. from Vietnam: Ninh Thuan Province; Cryptophyllium daparo gen. et sp. nov. from China: Yunnan Province; Cryptophyllium echidna gen. et sp. nov. from Indonesia: Wangi-wangi Island; Cryptophyllium faulkneri gen. et sp. nov. from Vietnam: Quang Ngai and Lam Dong Provinces; Cryptophyllium icarus gen. et sp. nov. from Vietnam: Lam Dong and Dak Lak Provinces; Cryptophyllium khmer gen. et sp. nov. from Cambodia: Koh Kong and Siem Reap Provinces; Cryptophyllium limogesi gen. et sp. nov. from Vietnam: Lam Dong, Dak Lak, and Dak Nong Provinces; Cryptophyllium liyananae gen. et sp. nov. from China: Guangxi Province; Cryptophyllium nuichuaense gen. et sp. nov. from Vietnam: Ninh Thuan Province; Cryptophyllium phami gen. et sp. nov. from Vietnam: Dong Nai and Ninh Thuan Provinces; and Cryptophyllium wennae gen. et sp. nov. from China: Yunnan Province. All newly described species are morphologically described, illustrated, and molecularly compared to congenerics. With the molecular results revealing cryptic taxa, it was found necessary for Cryptophyllium westwoodii (Wood-Mason, 1875), comb. nov. to have a neotype specimen designated to allow accurate differentiation from congenerics. To conclude, male and female dichotomous keys to species for the Cryptophyllium gen. nov. are presented.

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Research Article Thu, 18 Feb 2021 15:04:55 +0200
Tiger beetles (Coleoptera, Cicindelidae) of Northern Mindanao region (Philippines): checklist, distributional maps, and habitats https://zookeys.pensoft.net/article/34500/ ZooKeys 1017: 37-75

DOI: 10.3897/zookeys.1017.34500

Authors: Dale Ann P. Acal, Jürgen Wiesner, Olga M. Nuñeza, Radomir Jaskuła

Abstract: The knowledge about tiger beetle fauna of the Northern Mindanao region (Philippines) is summarized based on literature data and new records. Thirty species classified in ten genera (Tricondyla, Neocollyris, Protocollyris, Therates, Prothyma, Heptodonta, Thopeutica, Lophyra, Calomera, and Cylindera) were documented from the area (56% of tiger beetle fauna of Mindanao and 21% of Philippine species). Twelve species were noted from Northern Mindanao region for the first time, including five taxa, Neocollyris speciosa, Calomera angulata, Cylindera minuta, Lophyra striolata tenuiscripta, and Thopeutica virginea, not recorded from Mindanao before. Distribution maps for all recorded species and the first photographs of habitats for some species in Mindanao and/or in the Philippines are provided. Eight species (27% of recorded fauna) were noted from riverine habitats while 18 tiger beetles (60%) were typical forest taxa; in the case of four species, their habitats in Northern Mindanao region are not known.

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Research Article Fri, 12 Feb 2021 13:35:40 +0200
The success story of Labiobaetis Novikova & Kluge in the Philippines (Ephemeroptera, Baetidae), with description of 18 new species https://zookeys.pensoft.net/article/58017/ ZooKeys 1002: 1-114

DOI: 10.3897/zookeys.1002.58017

Authors: Thomas Kaltenbach, Jhoana M. Garces, Jean-Luc Gattolliat

Abstract: Material collected between 1994 and 2020 in the Philippines, covering most main islands like Luzon, Mindoro, Palawan, Negros, Cebu, Leyte, and Mindanao and some smaller islands, substantially increased our knowledge of Labiobaetis Novikova & Kluge in this archipelago. Only three species were previously reported: L. molawinensis (Müller-Liebenau, 1982) and L. sumigarensis (Müller-Liebenau, 1982) from larvae and L. boettgeri (Ulmer, 1924) from adults. Eighteen new species have been identified using a combination of morphology and genetic distance (COI, Kimura 2-parameter). They are described and illustrated based on their larvae and a key to all species in the Philippines is provided. The total number of Labiobaetis in the Philippines has increased to 21 species. Additional diversity of Labiobaetis based on molecular evidence only is presented as Molecular Operational Taxonomic Units (MOTUs) without description. The interspecific K2P distances in the Philippines are between 15% and 27%, the intraspecific distances are usually between 0% and 3%. The total number of Labiobaetis species worldwide is augmented to 144.

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Research Article Thu, 10 Dec 2020 14:52:50 +0200
Moss-inhabiting flea beetles in the Philippines (Coleoptera, Chrysomelidae, Alticinae) https://zookeys.pensoft.net/article/54011/ ZooKeys 960: 125-142

DOI: 10.3897/zookeys.960.54011

Authors: Albert F. Damaška, Dale Joy Mohagan, Martin Fikácek

Abstract: The Philippine islands are one of the key biodiversity hotspots in the Indo-Pacific area. Knowledge of moss-inhabiting flea beetles (Coleoptera: Chrysomelidae: Alticinae), a diverse and ecologically and morphologically enigmatic group in the Philippines is described. Six species from the Philippines are recorded, belonging to three genera: Benedictus luzonicus Sprecher-Uebersax et al., 2009 (recorded from the Philippines previously), Ivalia antennata sp. nov., I. caligulata sp. nov. and I. postfasciata (Chen, 1934), comb. nov. (transferred from Chabria Jacoby, 1887), Cangshanaltica mindanaoensis sp. nov., and C. luzonica sp. nov. Cox1 barcode sequences of Ivalia antennata and Cangshanaltica mindanaoensis are presented. Biogeography and diversity of moss-inhabiting flea beetles in the Philippines are discussed.

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Research Article Mon, 17 Aug 2020 16:07:58 +0300
New and little-known species of the genus Sphecodes Latreille (Hymenoptera, Halictidae) from Southeast Asia https://zookeys.pensoft.net/article/51708/ ZooKeys 937: 31-88

DOI: 10.3897/zookeys.937.51708

Authors: Yulia V. Astafurova, Maxim Yu. Proshchalykin, Maximilian Schwarz

Abstract: The available information about the cleptoparasitic bees of the genus Sphecodes in Southeast Asia is summarized. Thirty-one species are currently known from this area. Four new species are described: Sphecodes discoverlifei Astafurova & Proshchalykin, sp. nov. (Laos), S. engeli Astafurova & Proshchalykin, sp. nov. (Laos, Vietnam), S. ilyadadaria Astafurova, sp. nov. (Indonesia), and S. pseudoredivivus Astafurova & Proshchalykin, sp. nov. (Laos). Nine species are newly recorded from South East Asia: S. chaprensis Blüthgen, 1927 (Laos), S. howardi Cockerell, 1922 (Malaysia, Myanmar, Thailand), S. kershawi Perkins, 1921 (Indonesia, Malaysia, Myanmar, Thailand), S. laticeps Meyer, 1920 (Thailand, Vietnam), S. montanus Smith, 1879 (Laos), S. sauteri Meyer, 1925 (Laos), S. sikkimensis Blüthgen, 1927 (Laos, Myanmar), S. simlaensis Blüthgen, 1924 (Laos), and S. turneri Cockerell, 1916 (Laos). Based on type specimens, new synonymies have been proposed for Sphecodes kershawi Perkins, 1921 = S. javanensis Blüthgen, 1927, syn. nov.; S. simlaensis Blüthgen, 1924 = S. simlaellus Blüthgen, 1927, syn. nov.; S. laticeps Meyer, 1920 = S. biroi mariae Cockerell, 1930, syn. nov. Lectotypes are designated for Sphecodes biroi Friese, 1909, S. simlaellus Blüthgen, 1927, and S. laticeps Meyer, 1920. The female of Sphecodes sauteri Meyer, 1925, and the male of S. turneri Cockerell, 1916 are described for the first time.

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Research Article Mon, 1 Jun 2020 16:33:27 +0300
Three new species of the genus Chilocorellus Miyatake (Coleoptera, Coccinellidae, Sticholotidini) from the Philippines https://zookeys.pensoft.net/article/50139/ ZooKeys 937: 115-127

DOI: 10.3897/zookeys.937.50139

Authors: Xiaoning Zhang, Xinyue Liang, Xiaosheng Chen, Xingmin Wang

Abstract: The genus Chilocorellus Miyatake, 1994 has been revised. Three new species (C. uncinacanthus Zhang & Wang, sp. nov., C. denspinulifer Zhang & Wang, sp. nov., and C. fistulachaetodontus Zhang & Wang, sp. nov.) from the Philippines are described and illustrated in the present paper. An updated key to the species of the genus Chilocorellus is provided. In addition, a list of all known species and their distributions is also provided.

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Research Article Mon, 1 Jun 2020 16:14:06 +0300
Integrative taxonomy of the genus Dudgeodes Sartori, 2008 (Insecta, Ephemeroptera, Teloganodidae) from the Philippines with description of new species and supplementary descriptions of Southeast Asian species https://zookeys.pensoft.net/article/48659/ ZooKeys 910: 93-129

DOI: 10.3897/zookeys.910.48659

Authors: Jhoana M. Garces, Michel Sartori, Hendrik Freitag

Abstract: COI sequences were used as an initial clustering method to delimit putative species of the genus Dudgeodes in the Philippines. An overview of the diagnostic characters of Philippine species and characters with high intraspecific variability are given. Six new species of Dudgeodes are described and illustrated: D. bauernfeindi Garces & Sartori, sp. nov., D. freitagi Garces & Sartori, sp. nov., D. luntian Garces & Sartori, sp. nov., D. pangantihoni Garces & Sartori, sp. nov., D. tabang Garces & Sartori, sp. nov., and D. vonrinteleni Garces & Sartori, sp. nov., all known from the nymphal stage. Supplementary descriptions are provided for D. pescadori Sartori, 2008, D. hutanis Sartori, 2008, D. stephani Sartori, 2008, D. ulmeri Sartori, 2008, and D. celebensis Sartori, 2008. A key to the nymphs of Philippine Dudgeodes species is proposed.

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Research Article Mon, 10 Feb 2020 15:27:10 +0200
A review of Aulacospira Möllendorff, 1890 and Pseudostreptaxis Möllendorff, 1890 in the Philippines (Gastropoda, Pupilloidea, Hypselostomatidae) https://zookeys.pensoft.net/article/33052/ ZooKeys 842: 67-83

DOI: 10.3897/zookeys.842.33052

Authors: Barna Páll-Gergely, Menno Schilthuizen, Aydin Örstan, Kurt Auffenberg

Abstract: The genera Aulacospira and Pseudostreptaxis of the Philippines are revised based on the collections of the Senckenberg Forschungsinstitut und Naturmuseum (Frankfurt am Main, Germany), the Florida Museum of Natural History (USA), and recently collected material. Three new species are described: Aulacospira lens Páll-Gergely & Auffenberg, sp. n., Aulacospira krobyloides Páll-Gergely & Schilthuizen, sp. n., Pseudostreptaxis harli Páll-Gergely & Schilthuizen, sp. n.

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Research Article Tue, 7 May 2019 14:58:41 +0300
A new species of sardine, Sardinella pacifica from the Philippines (Teleostei, Clupeiformes, Clupeidae) https://zookeys.pensoft.net/article/30688/ ZooKeys 829: 75-83

DOI: 10.3897/zookeys.829.30688

Authors: Harutaka Hata, Hiroyuki Motomura

Abstract: A new sardine, Sardinella pacifica sp. n., is described on the basis of 21 specimens collected from the Philippines. The new species closely resembles Sardinella fimbriata (Valenciennes, 1847), both species having lateral scales with centrally discontinuous striae, a dark spot on the dorsal-fin origin, more than 70 lower gill rakers on the first gill arch, the pelvic fin with eight rays, and 17 or 18 prepelvic and 12 or 13 postpelvic scutes. However, the new species is distinguished from the latter by lower counts of lateral scales, pseudobranchial filaments, and postpelvic scutes (38–41, 14–19 and 12–13, respectively vs. 44–46, 19–22 and 13–14), and a shorter lower jaw (10.4–11.6% of standard length vs. 11.1–12.2%). Sardinella pacifica sp. n. is known only from the Philippines, whereas S. fimbriata is restricted to the Indian Ocean, although previously considered to be an Indo-West Pacific species, distributed from India to the Philippines.

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Research Article Mon, 11 Mar 2019 11:15:21 +0200
Henryhalticus philippinensis gen. et sp. n., a minute halticine from the Philippines (Insecta, Heteroptera, Miridae, Orthotylinae) https://zookeys.pensoft.net/article/21240/ ZooKeys 796: 187-195

DOI: 10.3897/zookeys.796.21240

Authors: Gerasimos Cassis

Abstract: Henryhalticus philippinensis gen. et sp. n. is described from a single location in the Negros Oriental Province of the Philippines. The male and female genitalia are described and illustrated. On the basis of the genitalic characters, external morphology, and size and color, the new species is erected as a monotypic genus.

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Research Article Thu, 15 Nov 2018 18:16:39 +0200
A new diplommatinid genus and two new species from the Philippines (Gastropoda, Caenogastropoda, Cyclophoroidea) https://zookeys.pensoft.net/article/13059/ ZooKeys 678: 1-10

DOI: 10.3897/zookeys.678.13059

Authors: Barna Páll-Gergely, András Hunyadi, Takahiro Asami

Abstract: A new diplommatinid genus, Luzonocoptis Páll-Gergely & Hunyadi, gen. n. is erected for two new species, Luzonocoptis antenna Páll-Gergely & Hunyadi, sp. n. and L. angulata Páll-Gergely & Hunyadi, sp. n. Both species inhabit the northeastern part of Luzon Island, Philippines. The genus Luzonocoptis gen. n. is mostly characterized by a very slender shell with 14–18 whorls, a strongly expanded peristome, an interrupted, weak columellar lamella, the absence of any additional plicae or lamellae, and a rachidian tooth having five cusps.

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Research Article Tue, 6 Jun 2017 00:37:06 +0300
Taxonomic study on specimens of the genus Micrencaustes deposited in the Bernice P. Bishop Museum (Coleoptera, Erotylidae) https://zookeys.pensoft.net/article/11003/ ZooKeys 645: 27-35

DOI: 10.3897/zookeys.645.11003

Authors: Jing Li, Yan-Chen Zhao, Guo-Dong Ren, Zhiqiang Cheng

Abstract: Specimens of the genus Micrencaustes deposited in Bernice P. Bishop Museum were studied. Two new species Micrencaustes (Mimencaustes) rotundimaculatasp. n. and Micrencaustes (Mimencaustes) serratimaculatasp. n. are described and illustrated. A key to worldwide species of the subgenus Mimencaustes is provided.

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Research Article Thu, 12 Jan 2017 04:17:16 +0200
The amphibians and reptiles of Mindanao Island, southern Philippines, II: the herpetofauna of northeast Mindanao and adjacent islands https://zookeys.pensoft.net/article/9814/ ZooKeys 624: 1-132

DOI: 10.3897/zookeys.624.9814

Authors: Marites B. Sanguila, Kerry A. Cobb, Cameron D. Siler, Arvin C. Diesmos, Angel C. Alcala, Rafe M. Brown

Abstract: We summarize all available amphibian and reptile species distribution data from the northeast Mindanao faunal region, including small islands associated with this subcenter of endemic vertebrate biodiversity. Together with all publicly available historical information from biodiversity repositories, we present new data from several major herpetological surveys, including recently conducted inventories on four major mountains of northeast Mindanao, and adjacent islands of Camiguin Sur, Dinagat, and Siargao. We present species accounts for all taxa, comment on unresolved taxonomic problems, and provide revisions to outdated IUCN conservation status assessments in cases where our new data significantly alter earlier classification status summaries. Together, our comprehensive analysis of this fauna suggests that the greater Mindanao faunal region possesses distinct subcenters of amphibian and reptile species diversity, and that until this area is revisited and its fauna and actually studied, with on-the-ground field work including targeted surveys of species distributions coupled to the study their natural history, our understanding of the diversity and conservation status of southern Philippine herpetological fauna will remain incomplete. Nevertheless, the northeast Mindanao geographical area (Caraga Region) appears to have the highest herpetological species diversity (at least 126 species) of any comparably-sized Philippine faunal subregion.

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Checklist Mon, 17 Oct 2016 18:45:22 +0300
A black-and-red stick insect from the Philippines – observations on the external anatomy and natural history of a new species of Orthomeria https://zookeys.pensoft.net/article/6281/ ZooKeys 559: 35-57

DOI: 10.3897/zookeys.559.6281

Authors: Davide Vallotto, Joachim Bresseel, Thierry Heitzmann, Marco Gottardo

Abstract: A new stick insect of the genus Orthomeria Kirby, 1904 (Phasmatodea, Aschiphasmatidae) is described from the Philippines. Orthomeria (O.) kangisp. n. is readily distinguished from all other congeners by the distinctive blood red colouration of the costal region of the hind wings. Major features of the external morphology of adults, eggs, and first-instar nymphs are illustrated. Locomotory attachment pads are of the smooth type with irregular microgrooves on the contact surface. An unusual condition of male terminalia is the absence of tergal thorn pads on segment 10. The male clasping organs are represented by an elongated vomer terminating in a prominent spine, and by incurved cerci featuring a bilobed apex equipped with a sharp blade-like ridge. Intraspecific variation in body colouration and hind wing length occurs in females. The new species lives at 400-650 m elevation in the surroundings of the Sablang and Tuba regions, in the Benguet Province of Luzon island. Host plants include Ficus spp. (Moraceae), and Pipturus spp. and Leucosyke spp. (Urticaceae). Observations on the mating and defensive behaviour are presented. Orthomeria (O.) catadromus (Westwood, 1859) is recognised as a junior synonym of O. (O.) pandora (Westwood, 1859), syn. n. A lectotype is designated for both species. Finally, an updated identification key to the species of the subgenus Orthomeria is provided.

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Research Article Wed, 3 Feb 2016 01:58:19 +0200
The oribatid mite subgenus Galumna (Galumna) (Acari, Oribatida, Galumnidae) in the Philippines https://zookeys.pensoft.net/article/4216/ ZooKeys 452: 1-13

DOI: 10.3897/zookeys.452.8212

Authors: Sergey Ermilov, Leonila Corpuz-Raros, Andrey Tolstikov

Abstract: Five species of the subgenus Galumna (Galumna) (Acari, Oribatida, Galumnidae) are registered in the Philippine oribatid mite fauna. A new species, G. (G.) makilingensissp. n., is described; it is most similar morphologically to G. (G.) tokyoensis Aoki, 1966, but differs from the latter by the morphology of porose areas Aa and Ap, rostral setae, and length of interlamellar setae. Three species, G. (G.) crenata Deb & Raychaudhuri, 1975, G. (G.) cf. exigua Sellnick, 1925 and G. (G.) khoii Mahunka, 1989, are recorded in the Philippines for the first time. The species G. (G.) crenata is redescribed. An identification key to the Philippine species of Galumna (Galumna) is given.

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Research Article Tue, 4 Nov 2014 00:00:00 +0200
Description of Medwayella independencia (Siphonaptera, Stivaliidae), a new species of flea from Mindanao Island, the Philippines and their phoretic mites, and miscellaneous flea records from the Malay Archipelago https://zookeys.pensoft.net/article/3794/ ZooKeys 408: 107-123

DOI: 10.3897/zookeys.408.7479

Authors: Michael Hastriter, Sarah E. Bush

Abstract: Medwayella independencia, a new species of flea, is described from the tupaiid host Urogale everetti (Thomas) from Mindanao Island, Philippines. Several other species of fleas are also recorded from the Philippines including a single male of Lentistivalius philippinensis Hastriter & Bush, 2013 (previously known only from two males), the bat fleas Thaumapsylla breviceps orientalis Smit and Thaumapsylla longiforceps Traub, a single unidentified female species of Macrostylophora Ewing collected from the murid Bullimus bagobos Mearns, and a pair of Medwayella robinsoni ssp. from Sundasciurus hoogstraali (Sanborn) from Busuanga Island, Philippines. Representatives of Medwayella Traub, 1972 and Macrostylophora have not previously been recorded from the Philippines. A key to the male sex of Medwayella is provided. Phoretic mites of the genus Psylloglyphus (family Winterschmidtiidae) were present under the abdominal sclerites of several male and female specimens of M. independencia. This is the second report of a phoretic mite on a species of Medwayella Traub. The co-evolutionary implications between phoretic mites and fleas are discussed.

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Research Article Tue, 13 May 2014 00:00:00 +0300
The “Fungia patella group” (Scleractinia, Fungiidae) revisited with a description of the mini mushroom coral Cycloseris boschmai sp. n. https://zookeys.pensoft.net/article/3217/ ZooKeys 371: 57-84

DOI: 10.3897/zookeys.371.6677

Authors: Bert Hoeksema

Abstract: The recent taxonomic history of extant free-living Cycloseris species is briefly reviewed, resulting in the description of Cycloseris boschmai sp. n. (Scleractinia, Fungiidae) and a discussion on the validity of two other recently described species. Some Cycloseris species were previously considered to belong to the Fungia patella group, which also concerned misidentified museum specimens that actually belong to the new species. Other specimens of C. boschmai sp. n. were photographed and collected in the course of 30 years of fieldwork. The new mushroom coral is compared with other free-living Cycloseris species by means of an identification key. With a maximum diameter of 50 mm, it is the smallest free-living mushroom coral discovered so far. It can also be distinguished by its large primary order costae and variable colouration. Its distribution range is limited to the Coral Triangle, where it can be observed as an uncommon species on lower reef slopes.

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Research Article Fri, 17 Jan 2014 00:00:00 +0200
Hydraena (Hydraenopsis) ateneo, new species (Coleoptera, Hydraenidae) and other aquatic Polyphaga from a small habitat patch in a highly urbanized landscape of Metro Manila, Philippines https://zookeys.pensoft.net/article/3623/ ZooKeys 329: 9-21

DOI: 10.3897/zookeys.329.5955

Authors: Hendrik Freitag

Abstract: Seven species of Hydraenidae, Hydrophilidae and Elmidae are recorded from temporary freshwater habitats at the Ateneo de Manila University Campus in the metropolitan area of Manila, Philippines. They were identified as Enochrus (Lumetus) fragiloides d’Orchymont, Helochares (Hydrobaticus) lepidus d’Orchymont, Helochares (Helochares) pallens (MacLeay), Hydraena (Hydraenopsis) scabra d’Orchymont, Hydraena (Hydraenopsis) palawanensis Freitag & Jäch (new record for Luzon Island), Stenelmis sp. A further hydraenid species was unknown to science and is newly described: Hydraena (Hydraenopsis) ateneo Freitag, sp. n. Aedeagus, gonocoxite, spermatheca, and female tergite X are illustrated by computer-based line drawings. Habitus images of all three Hydraena Kugelann species recorded and a checklist of the Philippine Hydraena are provided. The presence of these seven species in the Ateneo campus is briefly discussed in regard to the area’s history. Measures to maintain and extend semi-natural islands of biodiversity in urban areas are suggested.

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Research Article Thu, 5 Sep 2013 00:00:00 +0300
Ancyronyx Erichson, 1847 (Coleoptera, Elmidae) from Mindoro, Philippines, with description of the larvae and two new species using DNA sequences for the assignment of the developmental stages https://zookeys.pensoft.net/article/3493/ ZooKeys 321: 35-64

DOI: 10.3897/zookeys.321.5395

Authors: Hendrik Freitag

Abstract: Ancyronyx buhid sp. n. and A. tamaraw sp. n. are described based on adults and larvae, matched using their cox1 or cob DNA sequence data. Additional records of A. schillhammeri Jäch, 1994 and A. minerva Freitag & Jäch, 2007 from Mindoro are listed. The previously unknown larva of A. schillhammeri is also described here, aided by cox1 data. The new species and larval stages are described in detail and illustrated by SEM and stacked microscopic images. Keys to the adult and larval Ancyronyx species of Mindoro and an up­dated checklist of Philippine Ancyronyx species are provided. The usefulness as bioindicators, the phylogenetic relationships and biogeographic aspects affecting the distribution patterns are briefly discussed.

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Research Article Tue, 6 Aug 2013 00:00:00 +0300
Two new species of the genus Ancistrocerus Wesmael (Hymenoptera, Vespidae, Eumeninae) from China, with a key to the Oriental species https://zookeys.pensoft.net/article/3144/ ZooKeys 303: 77-86

DOI: 10.3897/zookeys.303.4922

Authors: Ju You, Bin Chen, Tingjing Li

Abstract: Two new species, namely Ancistrocerus transpunctatus You and Li, sp. n. and Ancistrocerus deqinensis You and Li, sp. n. are described and illustrated from Yunnan, China. A key to the Oriental species of the genus Ancistrocerus is provided.

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Research Article Tue, 21 May 2013 00:00:00 +0300
Cardiocondyla pirata sp. n. – a new Philippine ant with enigmatic pigmentation pattern (Hymenoptera, Formicidae) https://zookeys.pensoft.net/article/3127/ ZooKeys 301: 13-24

DOI: 10.3897/zookeys.301.4913

Authors: Bernhard Seifert, Sabine Frohschammer

Abstract: A new species of the ant genus Cardiocondyla Emery, 1869 – Cardiocondyla pirata sp. n. – is described from the Philippines. The species belongs to an Indo-Malayan group of six species that is characterized by workersa strongly bilobate postpetiolar sternite and a thickset mesosoma with strongly convex dorsal profile as well as wingless, ergatoid males with sickle-shaped mandibles. The female castes show a pigmentation pattern not known from any ant worldwide. Ifany adaptive value, a possible function of this structure is supposed to be visual dissolution of body shape in order to irritate predators.

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Research Article Fri, 17 May 2013 00:00:00 +0300
The amphibians and reptiles of Luzon Island, Philippines, VIII: the herpetofauna of Cagayan and Isabela Provinces, northern Sierra Madre Mountain Range https://zookeys.pensoft.net/article/3682/ ZooKeys 266: 1-120

DOI: 10.3897/zookeys.266.3982

Authors: Rafe Brown, Cameron Siler, Carl Oliveros, Luke Welton, Ashley Rock, John Swab, Merlijn Van Weerd, Jonah van Beijnen, Edgar Jose, Dominic Rodriguez, Edmund Jose, Arvin Diesmos

Abstract: We provide the first report on the herpetological biodiversity (amphibians and reptiles) of the northern Sierra Madre Mountain Range (Cagayan and Isabela provinces), northeast Luzon Island, Philippines. New data from extensive previously unpublished surveys in the Municipalities of Gonzaga, Gattaran, Lasam, Santa Ana, and Baggao (Cagayan Province), as well as fieldwork in the Municipalities of Cabagan, San Mariano, and Palanan (Isabela Province), combined with all available historical museum records, suggest this region is quite diverse. Our new data indicate that at least 101 species are present (29 amphibians, 30 lizards, 35 snakes, two freshwater turtles, three marine turtles, and two crocodilians) and now represented with well-documented records and/or voucher specimens, confirmed in institutional biodiversity repositories. A high percentage of Philippine endemic species constitute the local fauna (approximately 70%). The results of this and other recent studies signify that the herpetological diversity of the northern Philippines is far more diverse than previously imagined. Thirty-eight percent of our recorded species are associated with unresolved taxonomic issues (suspected new species or species complexes in need of taxonomic partitioning). This suggests that despite past and present efforts to comprehensively characterize the fauna, the herpetological biodiversity of the northern Philippines is still substantially underestimated and warranting of further study.

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Checklist Thu, 7 Feb 2013 00:00:00 +0200
Description of Lentistivalius philippinensis, a new species of flea (Siphonaptera, Pygiosyllomorpha, Stivalidae), and new records of Ascodipterinae (Streblidae) on bats and other small mammals from Luzon, The Philippines https://zookeys.pensoft.net/article/3576/ ZooKeys 260: 17-30

DOI: 10.3897/zookeys.260.3971

Authors: Michael Hastriter, Sarah Bush

Abstract: During May 2009 and July 2011, we collected 357 mammals and examined each for ectoparasites. Among the ectoparasites collected, a new species of flea was discovered. This new species, Lentistivalius philippinensis, is described from the male sex only. Two males were recovered from two specimens of the soricid Crocidura grayi Dobson in Municipality Maria Aurora, Aurora Province, Luzon, Philippines. Additional fleas included Thaumapsylla breviceps orientalis Smit, Thaumapsylla longiforceps Traub, and Ischnopsyllus indicus Jordan. Although the latter species is common in Japan and documented in Guam (as well as mainland Southeast Asia) also on Pipistrellus javanicus (Gray), I. indicus represents a new record in the Philippine Islands. The ascodipterinae (Streblidae) Maabella stomalata and Ascodipteron speiserianum Muir collected from Rhinolophus inops K. Andersen and Rhinolophus subrufus K. Andersen, respectively, also represent new host records. A key to the species of the flea genus Lentistivalius Traub is provided.

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Research Article Fri, 18 Jan 2013 00:00:00 +0200
Forever in the dark: the cave-dwelling azooxanthellate reef coral Leptoseris troglodyta sp. n. (Scleractinia, Agariciidae) https://zookeys.pensoft.net/article/3193/ ZooKeys 228: 21-37

DOI: 10.3897/zookeys.228.3798

Authors: Bert Hoeksema

Abstract: The coral species Leptoseris troglodyta sp. n. (Scleractinia, Agariciidae) is described as new to science. It is the first known azooxanthellate shallow-water agariciid and is recorded from the ceilings of caves at 5-35 m depth in West Pacific coral reefs. The corals have monocentric cup-shaped calices. They may become colonial through extramural budding from the basal coenosteum, which may cause adjacent calices to fuse. The size, shape and habitat of L. troglodyta are unique compared to other Leptoseris species, many of which have been recorded from mesophotic depths. The absence of zooxanthellae indicates that it may survive well in darkness, but endolithic algae in some corals indicate that they may be able to get some light. The presence of menianes on the septal sides, which may help to absorb light at greater depths in zooxanthellate corals, have no obvious adaptive relevance in the new species and could have been inherited from ancestral species that perhaps were zooxanthellate. The new species may be azooxanthellate as derived through the loss of zooxanthellae, which would be a reversal in Leptoseris phylogeny.

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Research Article Thu, 11 Oct 2012 00:00:00 +0300
A synoptic review of the ant genera (Hymenoptera, Formicidae) of the Philippines https://zookeys.pensoft.net/article/2844/ ZooKeys 200: 1-111

DOI: 10.3897/zookeys.200.2447

Authors: David General, Gary Alpert

Abstract: An overview of the history of myrmecology in the Philippine archipelago is presented. Keys are provided to the 11 ant subfamilies and the 92 ant genera known from the Philippines. Eleven ant genera (12%), including 3 undescribed genera, are recorded for the first time from the Philippines. The biology and ecology of the 92 genera, illustrated by full-face and profile photo-images, of Philippine ants are summarized in the form of brief generic accounts. A bibliography of significant taxonomic and behavioral papers on Philippine ants and a checklist of valid species and subspecies and their island distributions are provided.

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Research Article Tue, 5 Jun 2012 00:00:00 +0300
Revision of the pseudo-orbweavers of the genus Fecenia Simon, 1887 (Araneae, Psechridae), with emphasis on their pre-epigyne https://zookeys.pensoft.net/article/2084/ ZooKeys 153: 1-56

DOI: 10.3897/zookeys.153.2110

Authors: Steffen Bayer

Abstract: The present paper provides a taxonomic revision of the genus Fecenia with emphasis on the characteristics of the pre-epigynes which are integrated for the first time into an identification key. As a result, one species is revalidated, F. protensa Thorell, 1891, stat. n., and two new junior synonyms for F. protensa are recognised: F. sumatrana Kulczyński, 1908, syn. n. and F. nicobarensis (Tikader, 1977), syn. n. New records are reported: F. ochracea (Doleschall, 1859) from Malaysian Borneo, F. macilenta (Simon, 1885) from Sumatra, Indonesia, F. protensa from Thailand and Malaysia, F. travancoria Pocock, 1899 from Sri Lanka and Thailand, and F. cylindrata Thorell, 1895 from Thailand and Laos. Additional information on the biology of Fecenia is provided and the validity of characters for identifying Fecenia species is discussed.

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Research Article Fri, 9 Dec 2011 00:00:00 +0200
Larvae and a new species of Ancyronyx Erichson, 1847 (Insecta, Coleoptera, Elmidae) from Palawan, Philippines, using DNA sequences for the assignment of the developmental stages https://zookeys.pensoft.net/article/2820/ ZooKeys 136: 47-82

DOI: 10.3897/zookeys.136.1914

Authors: Hendrik Freitag, Michael Balke

Abstract: Ancyronyx montanus sp. n. is described based on adults and larvae, matched using their cox1 DNA sequence data. Larvae of six additional species of Ancyronyx Erichson, 1847 were also described here for the first time, aided by cox1 or cob data: A. helgeschneideri Freitag & Jäch, 2007, A. minerva Freitag & Jäch, 2007, A. patrolus Freitag & Jäch, 2007, A. procerus Jäch, 1994, A. punkti Freitag & Jäch, 2007, A. pseudopatrolus Freitag & Jäch, 2007. Ancyronyx procerus is newly recorded from the Philippines by a larval specimen from Busuanga island. The new species and larval stages are described in detail and illustrated by digital and SEM images. A key to the Ancyronyx larvae of Palawan and an updated checklist of Philippine Ancyronyx is provided.

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Research Article Thu, 13 Oct 2011 00:00:00 +0300
The spider family Selenopidae (Arachnida, Araneae) in Australia and Asia https://zookeys.pensoft.net/article/2353/ ZooKeys 99: 1-103

DOI: 10.3897/zookeys.99.723

Authors: Sarah Crews, Mark Harvey

Abstract: The spider family Selenopidae Simon occurs worldwide in tropical and subtropical regions, currently containing nearly 200 species in five genera. We relimit and revise the family to include four new genera and 27 new species from Australia and Asia. The family Selenopidae is redefined, as are the genera Anyphops Benoit, Garcorops Corronca, Hovops Benoit, Selenops Latreille, and Siamspinops Dankittipakul and Corronca, to accommodate the new genera and to correct previous errors in the definition. The species of Selenops that occur throughout India and China are also reviewed. Three species occur in China: S. bursarius Karsch, also known from Japan, Korea and Taiwan, S. ollarius Zhu, Sha, and Chen, and S. radiatus Latreille, the type of the genus and most widespread selenopid. Selenops cordatus Zhu, Sha, and Chen is recognized as a junior synonym of S. radiatus, syn. n. Amamanganops gen. n. is monotypic, with A. baginawa sp. n., and is known only from the Philippine island of Mindoro. Godumops gen. n. is monotypic, with G. careus sp. n., and is known only from Papua New Guinea. Karaops gen. n. occurs throughout Australia and has 24 species: K. australiensis (L. Koch) comb. n., K. gangarie sp. n., K. monteithi sp. n., K. alanlongbottomi sp. n., K. keithlongbottomi sp. n., K. larryoo sp. n., K. jarrit sp. n., K. marrayagong sp. n., K. raveni sp. n., K. badgeradda sp. n., K. burbidgei sp. n., K. karrawarla sp. n., K. julianneae sp. n., K. martamarta sp. n., K. manaayn sp. n., K. vadlaadambara sp. n., K. pilkingtoni sp. n., K. deserticola sp. n., K. ngarutjaranya sp. n., K. francesae sp. n., K. toolbrunup sp. n., the type species K. ellenae sp. n., K. jenniferae sp. n., and K. dawara sp. n. The genus Makdiops gen. n. contains five species from India and Nepal: M. agumbensis (Tikader), comb. n., the type of the genus M. mahishasura sp. n., M. montigenus (Simon), comb. n., M. nilgirensis (Reimoser) comb. n., and M. shiva sp. n. The genus Pakawops gen. n. is monotypic and contains P. formosanus (Kayashima) comb. n. known only from Taiwan. Finally, Selenops aculeatus Simon is transferred to the genus Siamspinops, forming the new combination S. aculeatus (Simon) comb. n. The distribution and diversity of the Australasian selenopid fauna is discussed. Keys are provided to all of the selenopid genera and to the species of Karaops and Makdiops.

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Monograph Thu, 19 May 2011 00:00:00 +0300
The genus Nelcyndana Stål (Hemiptera, Cicadidae, Taphurini) with description of three new species from Borneo https://zookeys.pensoft.net/article/2368/ ZooKeys 61: 11-31

DOI: 10.3897/zookeys.61.487

Authors: Joannes Duffels

Abstract: The type species of Nelcyndana, Nelcyndana tener (Stål, 1870) from the Philippines, is redescribed and illustrated. The taxonomic position of the genus Nelcyndana in the tribe Taphurini is discussed. Three new species from Borneo are described and illustrated: Nelcyndana borneensis sp. n., Nelcyndana vantoli sp. n., and Nelcyndana mulu sp. n. Distributions maps for the three Borneo species are presented and a key for the identification of the four Asian species of the genus is provided.

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Research Article Mon, 11 Oct 2010 00:00:00 +0300