Latest Articles from ZooKeys Latest 70 Articles from ZooKeys https://zookeys.pensoft.net/ Fri, 29 Mar 2024 09:35:50 +0200 Pensoft FeedCreator https://zookeys.pensoft.net/i/logo.jpg Latest Articles from ZooKeys https://zookeys.pensoft.net/ A new cryptic species of terrestrial breeding frog of the Pristimantis danae Group (Anura, Strabomantidae) from montane forests in Ayacucho, Peru https://zookeys.pensoft.net/article/104536/ ZooKeys 1187: 1-29

DOI: 10.3897/zookeys.1187.104536

Authors: Valia Herrera-Alva, Alessandro Catenazzi, César Aguilar-Puntriano

Abstract: Based on morphological and molecular characters, we describe a new species of terrestrial breeding frog of the Pristimantis danae Group from montane forests of La Mar Province, Ayacucho Department in southern Peru, at elevations from 1200 to 2000 m a.s.l. The phylogenetic analysis, based on concatenated sequences of gene fragments of 16S rRNA, RAG1, COI and TYR suggests that the new species is a sister taxon of a clade that includes one undescribed species of Pristimantis from Cusco, Pristimantis pharangobates and Pristimantis rhabdolaemus. The new species is most similar to P. rhabdolaemus, which differs by lacking scapular tubercules and by its smaller size (17.0–18.6 mm in males [n = 5], 20.8–25.2 mm in females [n = 5] in the new species vs. 22.8–26.3 mm in males [n = 19], 26.0–31.9 mm in females [n = 30] of P. rhabdolaemus). Additionally, we report the prevalence of Batrachochytrium dendrobatidis (Bd) in this species.

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Research Article Wed, 20 Dec 2023 17:21:51 +0200
Capitojoppa, a new genus of Ichneumoninae (Hymenoptera, Ichneumonidae) from Peruvian Amazonia https://zookeys.pensoft.net/article/108929/ ZooKeys 1178: 69-76

DOI: 10.3897/zookeys.1178.108929

Authors: Brandon R. Claridge, Kari M. Kaunisto, Ilari E. Sääksjärvi

Abstract: A new monotypic genus of ichneumonine parasitoid wasps (Hymenoptera, Ichneumonidae, Ichneumoninae) is described from Peru; Capitojoppa gen. nov. is described to accommodate Capitojoppa amazonica sp. nov. The new genus is morphologically very distinctive and can be easily separated from all known ichneumonine genera. By describing Capitojoppa from the lowland rain forests of Peru, we aim to draw attention to the considerable diversity and morphological variation of the Amazonian ichneumonine fauna.

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Review Article Fri, 1 Sep 2023 17:46:07 +0300
Peruvian nudibranchs (Mollusca, Gastropoda, Heterobranchia): an updated literature review-based list of species https://zookeys.pensoft.net/article/103167/ ZooKeys 1176: 117-163

DOI: 10.3897/zookeys.1176.103167

Authors: Alessandra Grández, André Ampuero, Sergio P. Barahona

Abstract: Nudibranchs, as a group, have received limited attention in terms of scientific study along the coastline of Peru. Here, an updated and comprehensive list of nudibranch species found in the Peruvian sea is presented, compiled through an extensive review of relevant literature. This compilation encompasses a total of 31 species, classified into two suborders, 10 superfamilies, 20 families, and 28 genera. With respect to the biogeographic provinces along the Peruvian coast, 23 species inhabit the Warm Temperate Southeastern Pacific province, 18 species occur in the Tropical Eastern Pacific province, and 10 species are found in both provinces, crossing the transitional zone between them. In terms of distribution patterns, two species exhibit a cosmopolitan distribution (Glaucus atlanticus and Fiona pinnata), while two species display a circumtropical distribution (Cephalopyge trematoides and Phylliroe bucephala). One species exhibits a bipolar distribution in the Eastern Pacific and possesses an amphi-South American distribution (Rostanga pulchra). Additionally, six species exhibit an amphi-South American distribution (Rostanga pulchra, Diaulula punctuolata, Doto uva, Tyrinna evelinae, Tyrinna delicata, and Doris fontainii), and two species are endemic to Peru (Corambe mancorensis and Felimare sechurana). This study provides comprehensive information on biogeographical aspects, geographical distributions, and taxonomic updates within the nudibranch species documented in Peru. Furthermore, we discuss the status of species listed in previous literature that have not been confirmed by collections, referring to them as potentially occurring species.

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Review Article Wed, 23 Aug 2023 18:16:24 +0300
Exceptional larval morphology of nine species of the Anastrepha mucronota species group (Diptera, Tephritidae) https://zookeys.pensoft.net/article/84628/ ZooKeys 1127: 155-215

DOI: 10.3897/zookeys.1127.84628

Authors: Erick J. Rodriguez, Gary J. Steck, Matthew R. Moore, Allen L. Norrbom, Jessica Diaz, Louis A. Somma, Raul Ruiz-Arce, Bruce D. Sutton, Norma Nolazco, Alies Muller, Marc A. Branham

Abstract: Anastrepha is the most diverse and economically important genus of Tephritidae in the American tropics and subtropics. The striking morphology of the third instars of Anastrepha caballeroi Norrbom, Anastrepha crebra Stone, Anastrepha haplacantha Norrbom & Korytkowski, Anastrepha korytkowskii Norrbom, Anastrepha nolazcoae Norrbom & Korytkowski, and three newly discovered and as yet formally unnamed species (Anastrepha sp. Peru-82, Anastrepha sp. nr. protuberans, and Anastrepha sp. Sur-16), and the more typical morphology of Anastrepha aphelocentema Stone, are described using light and scanning electron microscopy. To contribute to a better understanding of the interspecific and intraspecific variation among species in the mucronota species group and facilitate phylogenetic studies, we integrate molecular and morphological techniques to confirm the identity and describe third instars. Larva-adult associations and the identification of described larvae were confirmed using DNA barcodes. We provide diagnostic characters to distinguish larvae among these nine species of the mucronota group and separate them from those of the 29 other Anastrepha species previously described. We introduce the vertical comb-like processes on the oral margin as a novel character, and the unusual character states, including position and shape of the preoral lobe, and dentate or fringed posterior margins of the oral ridges and accessory plates. Our comparative morphology concurs with most previously inferred phylogenetic relationships within the mucronota group.

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Research Article Thu, 3 Nov 2022 19:09:01 +0200
The South American moth Rheumaptera mochica (Dognin, 1904) (Lepidoptera, Geometridae, Larentiinae) rediscovered after more than a century of anonymity https://zookeys.pensoft.net/article/76868/ ZooKeys 1085: 129-143

DOI: 10.3897/zookeys.1085.76868

Authors: Héctor A. Vargas, M. Alma Solis, Marcelo Vargas-Ortiz

Abstract: Rheumaptera mochica (Dognin, 1904) (Lepidoptera, Geometridae, Larentiinae) is reported from Chile for the first time. It was described from the western slopes of the Andes of southern Peru more than 100 years ago, and was recently rediscovered in Chile after larvae were collected and reared on the shrub Senna birostris var. arequipensis (Meyen ex Vogel) H.S. Irwin & Barneby (Fabaceae). This discovery expands the known distribution of this moth and provides its first host plant record. The genitalia of R. mochica are described and illustrated for the first time and compared to those of R. affirmata (Guenée, [1858]). A maximum likelihood analysis based on mitochondrial DNA sequences clustered R. mochica as sister to R. affirmata with 3.6–3.8% divergence (K2P). A lectotype is designated for Calocalpe mochica Dognin, 1904.

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Research Article Mon, 7 Feb 2022 17:59:33 +0200
Designation of the neotype of Triatoma dimidiata (Latreille, 1811) (Hemiptera, Reduviidae, Triatominae), with full integrated redescription including mitogenome and nuclear ITS-2 sequences https://zookeys.pensoft.net/article/72835/ ZooKeys 1076: 9-24

DOI: 10.3897/zookeys.1076.72835

Authors: Silvia Andrade Justi, Carolina Dale

Abstract: The taxonomic status of Triatoma dimidiata (Latreille, 1811) is, by far, the most discussed within Triatominae. Molecular studies have recovered at least three independently evolving lineages in T. dimidiata across its range. The original description of T. dimidiata (as Reduvius dimidiatus) included few taxonomic characters, and no types were assigned. To define and describe the cryptic diversity within T. dimidiata sensu lato (s.l.), a neotype must be designated. For this purpose, all 199 specimens identified as T. dimidiata from the collections of the Smithsonian Institution – National Museum of Natural History and the American Museum of Natural History, ranging from Peru to Mexico, were studied. Only one specimen (from Tumbes, Peru) matched the combination of characters as listed in the original description, and it is herein formally designated as the neotype for T. dimidiata. The neotype is morphologically described and DNA sequences of its whole mitochondrial genome and the nuclear second internal transcribed spacer region (ITS2), commonly used in triatomine molecular systematics studies, are presented and compared to other publicly available sequences of T. dimidiata s.l. in GenBank. Our results suggest that T. dimidiata sensu stricto (s.s.) is somewhat rare and, therefore, unlikely to serve as a major vector of Chagas disease.

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Research Article Wed, 8 Dec 2021 11:30:39 +0200
Amphibians of the equatorial seasonally dry forests of Ecuador and Peru https://zookeys.pensoft.net/article/69580/ ZooKeys 1063: 23-48

DOI: 10.3897/zookeys.1063.69580

Authors: Diego Armijos-Ojeda, Diana Székely, Paul Székely, Dan Cogălniceanu, Diego F. Cisneros-Heredia, Leonardo Ordóñez-Delgado, Adrián Escudero, Carlos Iván Espinosa

Abstract: Seasonally dry forests (SDFs) are one of the most challenging ecosystems for amphibians, fueling the diversity of this group of vertebrates. An updated inventory of native amphibians present in the Equatorial SDF is provided, which extends along the Pacific coast of Ecuador and northwestern Peru. The study is based on an extensive field sampling (two thirds of the total records) carried out throughout the Equatorial SDF, along with a compilation of the available information on distribution of amphibians in the region from published scientific papers, museum collections and on-line databases. The final dataset included 2,032 occurrence records for 30 amphibian species, belonging to eight anuran families. Additionally, data regarding conservation status, habitat use, spawn deposition site, reproductive mode, and body size, along with an identification key for all encountered species are provided. The results indicate a strong sampling bias with a deficit in the Peruvian part of the study area, and a need for urgent inventories targeted at under-sampled areas, using modern taxonomic methods. The study emphasizes the conservation priorities in the Equatorial SDF, based on the distribution, conservation status and life-history data. This information should be useful for the local authorities and institutions involved in the management and conservation of biodiversity in SDF.

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Checklist Mon, 18 Oct 2021 11:46:24 +0300
Biodiversity in tropical rainforests: Calleida Dejean, 1825 at the BIOLAT Biological Station, Rio Manu, Peru, with descriptions of seven new species (Coleoptera Carabidae, Lebiini). Part 1 https://zookeys.pensoft.net/article/64082/ ZooKeys 1044: 479-510

DOI: 10.3897/zookeys.1044.64082

Authors: Achille Casale

Abstract: A monographic contribution is presented on the species of the genus Calleida Dejean, 1825 at the BIOLAT Biological Station, Rio Manu, Pakitza, Peru, sampled by Terry Erwin and his co-workers. The following seven new species are described: C. solitaria sp. nov., C. manuensis sp. nov., C. anomala sp. nov., C. demathani sp. nov. (type locality: Peru, Tarapoto, but sampled also at Rio Manu), C. erwini sp. nov., C. marginithorax sp. nov., and C. maxima sp. nov. Relationships of each species are discussed, and a preliminary survey is presented of the genus Calleida in Peru.

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Research Article Wed, 16 Jun 2021 19:00:14 +0300
Phyllocnistis furcata sp. nov.: a new species of leaf-miner associated with Baccharis (Asteraceae) from Southern Peru (Lepidoptera, Gracillariidae) https://zookeys.pensoft.net/article/53958/ ZooKeys 996: 121-145

DOI: 10.3897/zookeys.996.53958

Authors: José Cerdeña, Jackie Farfán, Héctor A. Vargas, Rosângela Brito, Gislene L. Gonçalves, Ana Lazo, Gilson R. P. Moreira

Abstract: The southwestern Andes of Peru harbors a hidden taxonomic diversity of Lepidoptera. Here a new leaf-mining species of Gracillariidae (Lepidoptera) is described, Phyllocnistis furcata Vargas & Cerdeña, sp. nov., from a dry Andean valley of southern Peru, at 2400 m above sea level. The morphological aspects of adults (male and female) and the immature stages associated with Baccharis alnifolia Meyen & Walp. (Asteraceae) are given, under optical microscopy and scanning electron microscopy. DNA barcodes show that its nearest neighbor is the Atlantic Forest species Phyllocnistis ourea Brito & Moreira, 2017 that feeds on Baccharis anomala DC. The importance of morphological characters from immature stages for diagnosis among congeneric species is also discussed. Phyllocnistis furcata represents the fourth species of Phyllocnistis Zeller for Peru, and first record from the south of Peru for the genus.

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Research Article Tue, 24 Nov 2020 00:00:06 +0200
Seven new species of spider-attacking Hymenoepimecis Viereck (Hymenoptera, Ichneumonidae, Pimplinae) from Ecuador, French Guiana, and Peru, with an identification key to the world species https://zookeys.pensoft.net/article/50492/ ZooKeys 935: 57-92

DOI: 10.3897/zookeys.935.50492

Authors: Diego Galvão de Pádua, Ilari Eerikki Sääksjärvi, Ricardo Ferreira Monteiro, Marcio Luiz de Oliveira

Abstract: Seven new species of Hymenoepimecis Viereck are described from Peruvian Andes and Amazonia, French Guiana and Ecuador: H. andina Pádua & Sääksjärvi, sp. nov., H. castilloi Pádua & Sääksjärvi, sp. nov., H. dolichocarinata Pádua & Sääksjärvi, sp. nov., H. ecuatoriana Pádua & Sääksjärvi, sp. nov., H. longilobus Pádua & Sääksjärvi, sp. nov., H. pucallpina Pádua & Sääksjärvi, sp. nov., and H. rafaelmartinezi Pádua & Sääksjärvi, sp. nov. In addition, the male of the Hymenoepimecis kleini Pádua & Sobczak, 2015 is described, new faunistic records for the genus provided, as well as an illustrated identification key to all known species of the genus.

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Research Article Thu, 21 May 2020 00:01:33 +0300
A new Peruvian species of Scybalocanthon Martínez, 1948 (Coleoptera, Scarabaeidae, Scarabaeinae, Deltochilini) and some remarkable intrapopulational variation in the endophallus of S. pinopterus (Kirsch, 1873) https://zookeys.pensoft.net/article/39322/ ZooKeys 884: 69-80

DOI: 10.3897/zookeys.884.39322

Authors: Fernando A. B. Silva, François Génier

Abstract: Scybalocanthon ashei sp. nov. from Madre de Dios, Peru, is described based on differences in external and male genital morphology. Its diagnostic characters and an updated identification key to the species of the genus are provided as well as new distributional data for the following species: S. acrianus Silva & Valois, 2019, S. aereus (Schmidt, 1922), S. kaestneri (Balthasar, 1939) and S. pinopterus (Kirsch, 1873).

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Research Article Wed, 30 Oct 2019 17:10:41 +0200
Systematics of Huicundomantis, a new subgenus of Pristimantis (Anura, Strabomantidae) with extraordinary cryptic diversity and eleven new species https://zookeys.pensoft.net/article/26766/ ZooKeys 868: 1-112

DOI: 10.3897/zookeys.868.26766

Authors: Nadia B. Páez, Santiago R. Ron

Abstract: Pristimantis is the most diverse genus of tetrapods comprising 532 described species. It contains a large number of morphologically cryptic species that are being discovered with the assistance of genetic evidence. We use molecular, morphological, bioacoustic, and environmental data to assess the phylogenetic relationships and determine the species within an Andean clade of Pristimantis, which is distributed from central Ecuador to northern Peru. We assign to this clade the name Huicundomantis and propose it as a subgenus. Our results show that Huicundomantis is composed of two large clades which we name as the P. phoxocephalus species group and the P. cryptomelas species group. Huicundomantis is composed of 28 species of which 12 have been described and 16 are new. We describe 11 of these undescribed species. The most effective characters to discriminate among species are DNA sequences, qualitative morphology, and advertisement calls. Morphometric and environmental characters are not very useful to define species limits. We clarify the identity of P. riveti and show that populations from southern Ecuador traditionally ascribed to P. riveti are a new species, P. lutzae sp. nov. We also show that P. prometeii is a junior synonym of P. hampatusami. The current diversity and geographic distribution of Huicundomantis are consistent with a model of allopatric speciation. All species have a restricted distribution range (less than 4330 km2) and are assigned to the Red List categories Data Deficient or threatened with extinction. We provide new reasons to increase conservation efforts for these species and their habitat. Taking our results into account, Pristimantis species richness in Ecuador increases from 211 to 221 species, and the number of species endemic to Ecuador from 119 to 129.

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Research Article Thu, 1 Aug 2019 11:32:57 +0300
Three new species and a new genus of majoid crabs from the eastern Pacific (Decapoda, Brachyura) https://zookeys.pensoft.net/article/32271/ ZooKeys 825: 1-24

DOI: 10.3897/zookeys.825.32271

Authors: Jessica Colavite, Amanda Windsor, William Santana

Abstract: Three new species and a new genus of majoid crabs from deep waters in the eastern Pacific are described and illustrated using morphological and molecular data. A new species of inachoidid, Collodes anartius sp. n. is described from Peru, which resembles C. tenuirostris Rathbun, 1893, in the general appearance of the carapace, but is distinguished by the details of tubercles on the carapace and thoracic sternum, proportions of the pereopod articles, and bathymetric distribution. A new epialtid, Nibilia machala sp. n., is described from Ecuador; Nibilia A Milne-Edwards, 1878 has, until now, been considered to be monotypic, occurring only in the western Atlantic. This new species, from the eastern Pacific, closely resembles N. antilocapra (Stimpson, 1871) in the general morphology, but can be distinguished by the number of spines on the carapace and pereopods. Another epialtid, Solinca aulix gen. n. et sp. n, is establish for material collected from Ecuador and Peru, and can be easily identified from other taxa by the presence of a deep furrow between the very inflated branchial regions.

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Research Article Mon, 18 Feb 2019 10:56:22 +0200
A new genus and two new species of South American Myodochini (Hemiptera, Heteroptera, Rhyparochromidae) https://zookeys.pensoft.net/article/21431/ ZooKeys 796: 325-333

DOI: 10.3897/zookeys.796.21431

Authors: Pablo M. Dellapé, María Cecilia Melo, Sara I. Montemayor

Abstract: The new Neotropical genus Henryaria (Heteroptera, Rhyparochromidae, Myodochini) is established to accommodate two new species from Bolivia and Peru. Photographs of the types and their male genitalia are provided. Similarities with other genera of the tribe are discussed, as well as the characters to distinguish the two new species.

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Research Article Thu, 15 Nov 2018 04:08:36 +0200
Three new genera of acidocerine water scavenger beetles from tropical South America (Coleoptera, Hydrophilidae, Acidocerinae) https://zookeys.pensoft.net/article/24423/ ZooKeys 768: 113-158

DOI: 10.3897/zookeys.768.24423

Authors: Jennifer C. Girón, Andrew Edward Z. Short

Abstract: Recent collecting efforts in the Neotropics have led to the discovery of numerous new species and lineages of aquatic beetles. Here, three new genera are described to accommodate fifteen new species of water scavenger beetles of the subfamily Acidocerinae from northern South America: Crucisternum gen. n. for C. escalera sp. n. (Venezuela), C. ouboteri sp. n. (Guyana, French Guiana, Suriname, Venezuela), C. queneyi sp. n. (French Guiana), C. sinuatus sp. n. (Brazil), C. toboganensis sp. n. (Venezuela), C. vanessae sp. n. (Suriname), and C. xingu sp. n. (Brazil); Katasophistes gen. n. for K. charynae sp. n. (Peru), K. cuzco sp. n. (Peru), K. merida sp. n. (Venezuela) and K. superficialis sp. n. (Ecuador); and Nanosaphes gen. n. for N. castaneus sp. n. (Brazil), N. hesperus sp. n. (Suriname), N. punctatus sp. n. (Guyana), and N. tricolor sp. n. (Guyana, Suriname). It was also found that the monotypic Neotropical endemic genus Dieroxenus Spangler, 1979, syn. n. is congeneric with Chasmogenus Sharp, 1882 resulting in the single new combination Chasmogenus cremnobates (Spangler, 1979), comb. n.. Katasophistes merida sp. n. is known exclusively from seepage habitats, while the remaining taxa described herein are primarily associated with the margins of densely forested streams. Diagnoses, illustrations, distribution maps, and habitat summaries are provided for all new genera and species. A key to the genera of Acidocerinae of the New World is provided.

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Research Article Tue, 19 Jun 2018 23:44:35 +0300
Echinotermes biriba, a new genus and species of soldierless termite from the Colombian and Peruvian Amazon (Termitidae, Apicotermitinae) https://zookeys.pensoft.net/article/24253/ ZooKeys 748: 21-30

DOI: 10.3897/zookeys.748.24253

Authors: Daniel Castro, Rudolf H. Scheffrahn, Tiago F. Carrijo

Abstract: A new Apicotermitinae genus and species Echinotermes biriba is described from workers collected on the Andean-Amazon Piedmont in Colombia and Peru. The enteric valve armature of Echinotermes biriba Castro & Scheffrahn, gen. et sp. n. is a remarkably diagnostic character. A Bayesian phylogenetic analysis using the COI gene and including all other Neotropical Apicotermitinae genera, supports the new genus as a distinct terminal.

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Research Article Wed, 4 Apr 2018 18:40:07 +0300
Species limits within the widespread Amazonian treefrog Dendropsophus parviceps with descriptions of two new species (Anura, Hylidae) https://zookeys.pensoft.net/article/13864/ ZooKeys 726: 25-77

DOI: 10.3897/zookeys.726.13864

Authors: C. Daniel Rivadeneira, Pablo J. Venegas, Santiago R. Ron

Abstract: The genus Dendropsophus is one of the most speciose among Neotropical anurans and its number of described species is increasing. Herein, molecular, morphological, and bioacoustic evidence are combined to assess species limits within D. parviceps, a widely distributed species in the Amazon Basin. Phylogenetic relationships were assessed using 3040 bp sequences of mitochondrial DNA, genes 12S, ND1, and CO1. The phylogeny shows three well-supported clades. Bioacoustic and morphological divergence is congruent with those clades demonstrating that Dendropsophus parviceps is a species complex. Dendropsophus parvicepssensu stricto occurs in the Amazon basin of Ecuador, northern Peru, southern Colombia and northwestern Brazil. It is sister to two previously undescribed species, D. kubricki sp. n. from central Peru and D. kamagarini sp. n. from southern Peru, northeastern Bolivia, and northwestern Brazil. Genetic distances (uncorrected p, gene 12S) between D. parviceps and the new species is 3 to 4%. Dendropsophus kamagarini sp. n. can be distinguished from D. parviceps by having a prominent conical tubercle on the distal edge of the upper eyelid (tubercle absent in D. parviceps). Dendropsophus kubricki sp. n. differs from D. parviceps by having scattered low tubercles on the upper eyelids (smooth in D. parviceps). Dendropsophus parviceps and both new species differ from all their congeners by their small size (adult maximum SVL = 28.39 mm in females, 22.73 mm in males) and by having a bright orange blotch on the hidden areas of the shanks and under arms. The advertisement call of the two new species has lower dominant frequency relative to D. parviceps. Probable speciation modes are discussed. Available evidence indicates that ecological speciation along an elevation gradient is unlikely in this species complex.

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Research Article Mon, 8 Jan 2018 15:44:22 +0200
Bryophryne phuyuhampatu sp. n., a new species of Cusco Andes frog from the cloud forest of the eastern slopes of the Peruvian Andes (Amphibia, Anura, Craugastoridae) https://zookeys.pensoft.net/article/12152/ ZooKeys 685: 65-81

DOI: 10.3897/zookeys.685.12152

Authors: Alessandro Catenazzi, Alex Ttito, M. Isabel Diaz, Alexander Shepack

Abstract: A new species of Bryophryne from the humid montane forest of the Department of Cusco, Peru, is described. Specimens were collected at 2795–2850 m a.s.l. in the Área de Conservación Privada Ukumari Llaqta, Quispillomayo valley, in the province of Paucartambo. The new species is readily distinguished from all other species of Bryophryne by having green coloration on dorsum, and blue flecks on flanks and ventral parts. Specimens are characterized by lacking a distinct tympanic annulus, tympanic membrane, and dentigerous processes of vomers, and by having dorsal skin shagreen, discontinuous dorsolateral folds, skin tuberculate on flanks, skin areolate on ventral surfaces of the body, and fingers and toes without lateral fringes or webbing. The new species has a snout–vent length of 14.2–16.9 mm in three males and 22.2–22.6 mm in two females, and is smaller than all other congeneric species except for B. abramalagae. Generic allocation is supported by low genetic distances of the 16S mitochondrial gene and morphological similarity with other species of Bryophryne, and geographic distribution. Bryophryne phuyuhampatu sp. n. is only known from the type locality, a cloud forest along the Quispillomayo River in the upper Nusiniscato watershed.

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Research Article Thu, 13 Jul 2017 02:53:45 +0300
A new species of semiarboreal toad of the Rhinella festae group (Anura, Bufonidae) from the Cordillera Azul National Park, Peru https://zookeys.pensoft.net/article/13050/ ZooKeys 673: 21-47

DOI: 10.3897/zookeys.673.13050

Authors: Juan C. Cusi, Jiří Moravec, Edgar Lehr, Václav Gvoždík

Abstract: A new semiarboreal species of the Rhinella festae group is described from montane forests of the Cordillera Azul National Park between 1245 and 1280 m a.s.l. in the Cordillera Oriental, San Martín region, northern Peru. The new species is morphologically and genetically compared with members of the Rhinella acrolopha group (former genus Rhamphophryne) and members of the R. festae group. The new species is characterized by its large size (female SVL 47.1–58.3 mm, n = 4), eight presacral vertebrae, fusion of the sacrum and coccyx, long protuberant snout, snout directed slightly anteroventral in lateral view, cranial crests moderately developed, absence of occipital crest, presence of tympanic membrane, dorsolateral rows of small conical tubercles extending from parotoid gland to groin, hands and feet with long digits, fingers basally webbed and toes moderately webbed. Phylogenetically it is a member of the R. festae group which is most closely related to R. chavin and R. yanachaga from Peru. Morphologically the new species shares similarities with R. tenrec and R. truebae, members of the R. acrolopha group from Colombia.

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Research Article Fri, 12 May 2017 13:10:31 +0300
Delfinoia, a new South American aphid genus (Hemiptera, Aphididae, Macrosiphini) on Cayaponia (Cucurbitaceae) https://zookeys.pensoft.net/article/12247/ ZooKeys 671: 49-60

DOI: 10.3897/zookeys.671.12247

Authors: Juan M. Nieto Nafría, M. Pilar Mier Durante, Sara I. López Ciruelos

Abstract: The genus Delfinoia Nieto Nafría & Mier Durante gen. n. is established, and Utamphorophora peruviana (Essig), originally Amphorophora peruviana and currently Delfinoia peruviana comb. n., is designated species type of the genus. The synonymy between this species and Wahlgreniella australis Delfino syn. n. is established. Apterous and alate viviparous females of D. peruviana are redescribed; the male is also described. The species is currently known from Peru and Argentina; a plant of the genus Cayaponia (Cucurbitaceae) is the only identified host.

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Research Article Wed, 26 Apr 2017 19:05:55 +0300
A new cryptic species and review of the east-Andean leaf chafer genus Mesomerodon Ohaus, 1905 (Coleoptera, Scarabaeidae, Rutelinae) https://zookeys.pensoft.net/article/11815/ ZooKeys 671: 61-85

DOI: 10.3897/zookeys.671.11815

Authors: Matthias Seidel, Mary L. Jameson, Rachel L. Stone

Abstract: The Neotropical scarab beetle genus Mesomerodon Ohaus (Scarabaeidae: Rutelinae: Rutelini) is distributed in the western (lowland) Amazonian region from Colombia to Bolivia. Based on our research, the genus includes three species including a new cryptic species from Ecuador. We use niche modeling to predict potential suitable habitat and identify environmental factors associated with the distribution of Mesomerodon species. We characterize the genus, provide a key to species, diagnose each species, describe a new species, provide spatial and temporal distributions, and discuss distributions of the species in relation to Amazonian landscape biodiversity.

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Research Article Wed, 26 Apr 2017 18:29:05 +0300
Disjunctitermes insularis, a new soldierless termite genus and species (Isoptera, Termitidae, Apicotermitinae) from Guadeloupe and Peru https://zookeys.pensoft.net/article/11599/ ZooKeys 665: 71-84

DOI: 10.3897/zookeys.665.11599

Authors: Rudolf H. Scheffrahn, Tiago F. Carrijo, Anthony C. Postle, Francesco Tonini

Abstract: Disjunctitermes insularis gen. n. & sp. n. is described from workers collected on Guadeloupe and in Peru and is the first soldierless termite found on a deep-water island. As with many soldierless and soil-feeding termite species, the enteric valve morphology is an essential diagnostic character of D. insularis. The D. insularis sequence cluster, derived from a barcode analysis with twelve other described genera of New World Apicotermitinae, is well resolved. Results of a stochastic dynamic spread model suggest that the occurrence of D. insularis on Guadeloupe may be the result of a pre-Colombian overwater dispersal event from mainland South America.

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Research Article Tue, 4 Apr 2017 02:50:28 +0300
A new species of terrestrial-breeding frog (Amphibia, Craugastoridae, Pristimantis) from high elevations of the Pui Pui Protected Forest in central Peru https://zookeys.pensoft.net/article/11394/ ZooKeys 660: 17-42

DOI: 10.3897/zookeys.660.11394

Authors: Edgar Lehr, Rudolf von May

Abstract: We describe a new species of Pristimantis from upper montane forests and high Andean grasslands of the Pui Pui Protected Forest and its close surroundings, Región Junín, central Peru. The description of the new species is based on 34 specimens found at elevations between 3400 and 3936 m a.s.l. Pristimantis attenboroughi sp. n. is characterized by a snout–vent length of 14.6–19.2 mm in adult males (n = 21), 19.2–23.0 mm in adult females (n = 10), and is compared morphologically and genetically with other taxonomically and biogeographically relevant species of Pristimantis. The new species is characterized by having narrow digits that lack circumferential grooves, irregularly shaped, discontinuous dorsolateral folds, and absence of both tympanic membrane and tympanic annulus. The high similarity in morphology between P. attenboroughi sp. n. and members of the Andean genera Phrynopus and Bryophryne provides an example for convergent evolution, and highlights the importance of using molecular data to justify generic assignment. Pristimantis attenboroughi sp. n. is most similar to Phrynopus chaparroi from the Región Junín, suggesting that the generic placement of this species needs to be revised. Phylogenetically the new species belongs to the Pristimantis danae species Group, a clade that includes several Pristimantis species distributed in the montane forests of central Peru, including P. albertus, P. aniptopalmatus, P. ornatus, and P. stictogaster.

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Research Article Tue, 7 Mar 2017 02:10:20 +0200
A new species of Pristimantis (Amphibia, Anura, Craugastoridae) from a montane forest of the Pui Pui Protected Forest in central Peru (Región Junín) https://zookeys.pensoft.net/article/11221/ ZooKeys 645: 85-102

DOI: 10.3897/zookeys.645.11221

Authors: Edgar Lehr, Jiří Moravec

Abstract: A new species of frog of the genus Pristimantis is described from a montane forest between 1700 and 1800 m a.s.l. of the Pui Pui Protected Forest (Región Junín) in central Peru. Pristimantis ashaninka sp. n. is described based on five adult females (snout–vent length 23.1–26.7 mm) and ten juveniles (snout-vent length 10.6–13.4). It differs from its congeners by having the skin on dorsum shagreen with many conical tubercles giving it a spinose appearance, lacking a tympanum, having groin, anterior and posterior surfaces of thighs uniformly grayish brown, and a pale bronze iris with fine black reticulations, a median reddish hint horizontally across iris, and a black narrow vertical streak from pupil across lower and upper half of iris. Among the Peruvian Pristimantis that lack a tympanum, P. ashaninka sp. n. is morphologically most similar to P. lirellus, P. martiae, and P. rhabdocnemus. However, 16S DNA barcoding revealed clear distinctions between all four species of Pristimantis.

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Research Article Thu, 12 Jan 2017 20:57:29 +0200
New South American species of Lamiinae (Coleoptera, Cerambycidae) https://zookeys.pensoft.net/article/11001/ ZooKeys 637: 81-88

DOI: 10.3897/zookeys.637.11001

Authors: Maria Helena M. Galileo, Antonio Santos-Silva

Abstract: Two new species of cerambycid beetles are described from South America: Ataxia camiriensis (Pteropliini), from Bolivia, and Falsamblesthis uniformis (Forsteriini), from Peru. The new species are included in previous keys.

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Research Article Fri, 2 Dec 2016 10:46:17 +0200
A new species of frog of the genus Pristimantis from Tingo María National Park, Huánuco Department, central Peru (Anura, Craugastoridae) https://zookeys.pensoft.net/article/8507/ ZooKeys 610: 113-130

DOI: 10.3897/zookeys.610.8507

Authors: Germán Chávez, Alessandro Catenazzi

Abstract: A new species of Craugastoridae frog encountered from 1000–1700 m in elevation in the premontane forests of the Peruvian central Andes is described. The new species is similar in appearance to many other species of Pristimantis, but is easily distinguishable from these species by having bright red coloration on the groin, posterior surface of thighs, and shanks. The new species is only known for two localities 27 km apart in the Huánuco Region.

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Research Article Thu, 11 Aug 2016 15:01:57 +0300
A new species of Pristimantis (Amphibia, Anura, Craugastoridae) from the foothills of the Andes in Manu National Park, southeastern Peru https://zookeys.pensoft.net/article/8295/ ZooKeys 594: 143-164

DOI: 10.3897/zookeys.594.8295

Authors: Alexander Shepack, Rudolf von May, Alex Ttito, Alessandro Catenazzi

Abstract: We describe a new species of Pristimantis from the humid sub-montane forest of the Región Cusco in Peru. Pristimantis pluvialis sp. n. was collected in the Kosñipata and Entoro valleys at elevations from 740 to 1110 m a.s.l., near the borders of Manu National Park and within the Huachiperi Haramba Queros Conservation Concession. The new species can be distinguished from other members of the genus Pristimantis by its rostral tubercle, smooth dorsal skin, and by its advertisement call. Pristimantis lacrimosus and P. waoranii superficially most resemble the new species, but P. pluvialis sp. n. differs from both species by having a rostral tubercle (absent in P. waoranii and variable in P. lacrimosus) and larger size, from P. lacrimosus by its call emitted at a lower frequency, and from P. waoranii for its dorsal coloration with dark markings. Two other species have partially overlapping distributions and resemble the new species, P. mendax and P. olivaceus, but they produce advertisement calls with much higher dominant frequencies than the advertisement call of the new species. Furthermore, P. mendax differs from the new species by lacking a rostral tubercle and by having a sigmoid inner tarsal fold, whereas P. olivaceus differs by being smaller and by having dorsal skin shagreen with scattered tubercles. The new species has snout-vent length of 21.8–26.9 mm in males (n = 12) and 28.8 mm in a single female.

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Research Article Mon, 30 May 2016 03:22:40 +0300
Synopsis of Central Andean Orthalicoid land snails (Gastropoda, Stylommatophora), excluding Bulimulidae https://zookeys.pensoft.net/article/7906/ ZooKeys 588: 1-199

DOI: 10.3897/zookeys.588.7906

Authors: Abraham S.H. Breure, Valentín Mogollón Avila

Abstract: A faunal overview is presented of the molluscan families Amphibulimidae, Megaspiridae, Odontostomidae, Orthalicidae, Simpulopsidae in Bolivia, Ecuador, and Peru. These Central Andean countries are known for their biodiverse malacofauna, of which the superfamily Orthalicoidea takes relatively a large share. In this paper the five families containing 103 (sub)species, for which systematic information (original publication, type locality, type depository, summarizing literature) and distributional records are presented. All species are illustrated by photographs of the type material or, if this could not be located, by a reproduction of the original figure. The following new taxon is introduced: Thaumastus (Thaumastus) sumaqwayqusp. n. Junior subjective synonyms are established for: Plekocheilus (Sparnotion) Pilsbry, 1944 = Plekocheilus (Eudolichotis) Pilsbry, 1896; Scholvienia (Thomsenia) Strebel, 1910 = Scholvienia Strebel, 1910; Sultana (Trachyorthalicus) Strebel, 1909 = Sultana (Metorthalicus) Pilsbry, 1899; Plekocheilus (Eurytus) conspicuus Pilsbry, 1932 = Thaumastus (Thaumastus) hartwegi (Pfeiffer in Philippi, 1846); Zebra gruneri Strebel, 1909 = Orthalicus maracaibensis (Pfeiffer, 1856); Scholvienia jaspidea minor Strebel, 1910 = Scholvienia alutacea (Reeve, 1850); Bulimus bifasciatus unicolor Philippi, 1869 = Scholvienia brephoides (d’Orbigny, 1835). A new status is given to Plekocheilus mcgintyi ‘Pilsbry’ H.B. Baker, 1963 (subspecies of Bulinus piperitus Sowerby I, 1837); Strophocheilus superstriatus var. prodeflexus Pilsbry, 1895 (subspecies of Bulinus piperitus Sowerby I, 1837); Thaumastus (Quechua) salteri maximus Weyrauch, 1967 (subspecies of Thaumastus (Quechua) olmosensis Zilch, 1954); Pseudoglandina agitata Weyrauch, 1967 (nomen inquirendum). New combinations are: Clathrorthalicus corydon (Crosse, 1869), and Cyclodontina chuquisacana (Marshall, 1930). Lectotypes are now designated for Bulimus incisus Hupé, 1857 and Bulinus piperitus Sowerby I, 1837.

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Research Article Thu, 12 May 2016 03:21:39 +0300
Uncovering the diversity in the Amazophrynella minuta complex: integrative taxonomy reveals a new species of Amazophrynella (Anura, Bufonidae) from southern Peru https://zookeys.pensoft.net/article/6084/ ZooKeys 563: 43-71

DOI: 10.3897/zookeys.563.6084

Authors: Rommel R. Rojas, Juan Chaparro, Vinicius Carvalho, Robson Avila, Izeni Farias, Tomas Hrbek, Marcelo Gordo

Abstract: A new species of the genus Amazophrynella (Anura, Bufonidae) is described from the departments of Madre de Dios, Cusco and Junin in Peru. An integrative taxonomy approach is used. A morphological diagnosis, morphometrics comparisons, description of the advertisement call, and the phylogenetic relationships of the new species are provided. Amazophrynella javierbustamantei sp. n. differs from other species of Amazophrynella by: intermediate body-size (snout-vent length 14.9 mm in males, n = 26 and 19.6 mm in females, n = 20), tuberculate skin texture of body, greatest hand length of the Amazophrynella spp. (3.6 mm in males, n = 26 and 4.6 mm in females, n = 20), venter coloration yellowish, tiny rounded black points covering the venter, and thirteen molecular autapomorphies in the 16S RNA gene. Its distribution varies from 215 to 708 m a.s.l. This discovery highlights the importance of the remnant forest in preserving the biodiversity in Peru, and increase in seven the species formally described in the genus Amazophrynella.

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Research Article Mon, 15 Feb 2016 13:39:26 +0200
Ashea megacephala Kim & Ahn (Coleoptera, Staphylinidae, Aleocharinae), a new gyrophaenine genus and species from Peru https://zookeys.pensoft.net/article/6110/ ZooKeys 530: 91-99

DOI: 10.3897/zookeys.530.6110

Authors: Yoon-Ho Kim, Kee-Jeong Ahn

Abstract: Ashea megacephala, a new Peruvian genus and species is described. The specimens were collected on mushrooms and mushroom-associated trees. This genus can be readily distinguished from the other genera of the subtribe Gyrophaenina by the large head and the three indistinctly articulated labial palpomeres. A key to the known genera of Gyrophaenina is provided. A habitus photograph and illustrations of diagnostic characters are also presented.

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Research Article Wed, 28 Oct 2015 16:16:27 +0200
Additions to the checklist of Scoliidae, Sphecidae, Pompilidae and Vespidae of Peru, with notes on the endemic status of some species (Hymenoptera, Aculeata) https://zookeys.pensoft.net/article/6051/ ZooKeys 519: 33-48

DOI: 10.3897/zookeys.519.6501

Authors: Eduardo Fernando dos Santos, Yuri Campanholo Grandinete, Fernando Barbosa Noll

Abstract: The first checklist of the Peruvian Hymenoptera listed 1169 species and subspecies of aculeate wasps, including 173 species of Pompilidae, seven of Scoliidae, 39 of Sphecidae and 403 of Vespidae. Herein are reported 32 species as new for Peru based mainly on the collection of the Natural History Museum, London. The loss of the endemic status of two species is also reported: Entypus peruvianus (Rohwer) (Pompilidae: Pepsinae) and Omicron ruficolle schunkei Giordani Soika (Vespidae: Eumeninae).

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Checklist Mon, 31 Aug 2015 18:20:57 +0300
A new species of Noblella (Amphibia, Anura, Craugastoridae) from the humid montane forests of Cusco, Peru https://zookeys.pensoft.net/article/5846/ ZooKeys 516: 71-84

DOI: 10.3897/zookeys.516.9776

Authors: Alessandro Catenazzi, Vanessa Uscapi, Rudolf von May

Abstract: A new species of Noblella is described from the humid montane forest of the Región Cusco in Peru. Specimens were collected at 2330–2370 m elevation in Madre Selva, near Santa Ana, in the province of La Convención. The new species is readily distinguished from all other species of Noblella by having a broad, irregularly shaped, white mark on black background on chest and belly. The new species further differs from known Peruvian species of Noblella by the combination of the following characters: tympanic membrane absent, small tubercles on the upper eyelid and on dorsum, tarsal tubercles or folds absent, tips of digits not expanded, no circumferential grooves on digits, dark brown facial mask and lateral band extending from the tip of the snout to the inguinal region. The new species has a snout-to-vent length of 15.6 mm in one adult male and 17.6 mm in one adult female. Like other recently described species in the genus, this new Noblella inhabits high-elevation forests in the Andes and likely has a restricted geographic distribution.

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Research Article Thu, 6 Aug 2015 00:00:00 +0300
A new species of Agaporomorphus Zimmermann, 1921 from Peru (Coleoptera, Dytiscidae, Copelatinae) https://zookeys.pensoft.net/article/5576/ ZooKeys 512: 63-76

DOI: 10.3897/zookeys.512.9505

Authors: Lars Hendrich, Rico Apenborn, Ernst-Gerhard Burmeister, Michael Balke

Abstract: Agaporomorphus julianeae sp. n. is described from the Biological Field Station Panguana, in Huànuco province of central Peru. The new species belongs to the A. knischi-group sensu Miller 2005. Together with A. knischi Zimmermann, 1921 and A. colberti Miller & Wheeler, 2008 this is the third species of the genus with broadly enlarged male antennomeres. The new species can be separated from A. colberti and A. knischi by the smaller please expanded male antennomere VIII, and the form of the median lobe. Important species characters (median lobe, male antennae, metafemur, colour pattern) of the new species and A. knischi are figured, and the habitat, a temporary blackwater forest pond, and its species rich water beetle coenosis are illustrated and described in detail. The Brazilian A. mecolobus Miller, 2001, only known from the type material from Sao Paulo, is here recorded for Minas Gerais. Habitus photos of four additional Agaporomorphus species and Hydrodytes opalinus (Zimmermann, 1921) are provided. Altogether ten species of Agaporomorphus are now known.

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Research Article Mon, 6 Jul 2015 00:00:00 +0300
Records of Coendou ichillus (Rodentia, Erethizontidae) from the Lower Urubamba Region of Peru https://zookeys.pensoft.net/article/5492/ ZooKeys 509: 109-121

DOI: 10.3897/zookeys.509.9821

Authors: Tremaine Gregory, Darrin Lunde, Hugo Tomás Zamora Meza, Farah Carrasco-Rueda

Abstract: Coendou ichillus was first described in 2001 by Voss and da Silva, with a range from Amazonian Ecuador to Iquitos, Peru. Here, we describe an adult female Coendou ichillus specimen collected in a Tomahawk trap in the forest canopy of the Lower Urubamba Region of Peru in October 2013. We also describe pathologies and behaviors observed through 379 camera trapping photo events (2,196 photos) gathered in natural canopy bridges over the course of a year (7,198 trap nights), including information on activity period over the course of the day and over the course of the lunar cycle. We conservatively estimate that 17 individuals were photographed, including one juvenile. Being 900 km away from Iquitos, Peru (the site of the closest record), discovery of this species in the Lower Urubamba constitutes a significant range extension.

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Research Article Wed, 24 Jun 2015 00:00:00 +0300
Three new species of woodlizards (Hoplocercinae, Enyalioides) from northwestern South America https://zookeys.pensoft.net/article/5032/ ZooKeys 494: 107-132

DOI: 10.3897/zookeys.494.8903

Authors: Omar Torres-Carvajal, Pablo J. Venegas, Kevin de Queiroz

Abstract: The discovery of three new species of Enyalioides from the tropical Andes in Ecuador and northern Peru is reported. Enyalioides altotambo sp. n. occurs in northwestern Ecuador and differs from other species of Enyalioides in having dorsal scales that are both smooth and homogeneous in size, a brown iris, and in lacking enlarged, circular and keeled scales on the flanks. Enyalioides anisolepis sp. n. occurs on the Amazonian slopes of the Andes in southern Ecuador and northern Peru and can be distinguished from other species of Enyalioides by its scattered, projecting large scales on the dorsum, flanks, and hind limbs, as well as a well-developed vertebral crest, with the vertebrals on the neck at least three times higher than those between the hind limbs. Enyalioides sophiarothschildae sp. n. is from the Amazonian slopes of the Cordillera Central in northeastern Peru; it differs from other species of Enyalioides in having caudal scales that are relatively homogeneous in size on each caudal segment, a white gular region with a black medial patch and several turquoise scales in males, as well as immaculate white labials and chin. A molecular phylogenetic tree of 18 species of hoplocercines is presented, including the three species described in this paper and E. cofanorum, as well as an updated identification key for species of Hoplocercinae.

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Research Article Mon, 6 Apr 2015 00:00:00 +0300
An illustrated key to Neotropical species of the genus Meteorus Haliday (Hymenoptera, Braconidae, Euphorinae) https://zookeys.pensoft.net/article/4942/ ZooKeys 489: 33-94

DOI: 10.3897/zookeys.489.9258

Authors: Helmuth Aguirre, Luis Felipe Ventura de Almeida, Scott Richard Shaw, Carlos Eduardo Sarmiento Monroy

Abstract: A comprehensive key for 75 species of Meteorus distributed across 15 Neotropical countries is presented. Eleven new species from Bolivia, Costa Rica and Ecuador are described: M. albistigma, M. carolae, M. eurysaccavorus, M. fallacavus, M. flavistigma, M. haimowitzi, M. magnoculus, M. martinezi, M. microcavus, M. noctuivorus and M. orion. Expanded range distributions are recorded for M. andreae, M. farallonensis, M. guineverae, M. jerodi, M. kraussi, M. papiliovorus and M. quimbayensis. The host of M. jerodi is reported for the first time: a noctuid larva feeding on Asteraceae. Meteorus papiliovorus is recorded attacking Papilionidae larvae in Ecuador, therefore displaying a similar host family preference as formerly documented from Costa Rica and Colombia.

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Research Article Mon, 23 Mar 2015 00:00:00 +0200
Peruvian oribatid mites (Acari, Oribatida) from the German Biological Expedition, with description of a new species of the genus Pergalumna https://zookeys.pensoft.net/article/4882/ ZooKeys 487: 87-96

DOI: 10.3897/zookeys.487.9335

Authors: Sergey Ermilov, Darius J. Gwiazdowicz

Abstract: The present study is based on oribatid mite material (Acari, Oribatida) collected during the German Expedition in 2011 in Peru. An annotated checklist of identified oribatid mites, including 16 species from 14 genera and 8 families, is provided. Thirteen species and two genera (Notohermannia, Zetomimus) are recorded for the first time in Peru; the genus Notohermannia and species Notohermannia obtusa are recorded for the first time in the Neotropical region. A new species of the genus Pergalumna (Galumnidae), P. paraboliviana sp. n., is described. The new species is most similar to Pergalumna boliviana Ermilov, 2013 from Bolivia, however, it differs from the latter by the body size, morphology of porose areas A1 and the presence of interlamellar setae.

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Research Article Mon, 16 Mar 2015 00:00:00 +0200
A new species of Telmatobius (Amphibia, Anura, Telmatobiidae) from the Pacific slopes of the Andes, Peru https://zookeys.pensoft.net/article/4631/ ZooKeys 480: 81-95

DOI: 10.3897/zookeys.480.8578

Authors: Alessandro Catenazzi, Victor Vargas, Edgar Lehr

Abstract: We describe a new species of Telmatobius from the Pacific slopes of the Andes in central Peru. Specimens were collected at 3900 m elevation near Huaytará, Huancavelica, in the upper drainage of the Pisco river. The new species has a snout–vent length of 52.5 ± 1.1 mm (49.3–55.7 mm, n = 6) in adult females, and 48.5 mm in the single adult male. The new species has bright yellow and orange coloration ventrally and is readily distinguished from all other central Peruvian Andean species of Telmatobius but T. intermedius by having vomerine teeth but lacking premaxillary and maxillary teeth, and by its slender body shape and long legs. The new species differs from T. intermedius by its larger size, flatter head, and the absence of cutaneous keratinized spicules (present even in immature females of T. intermedius), and in males by the presence of minute, densely packed nuptial spines on dorsal and medial surfaces of thumbs (large, sparsely packed nuptial spines in T. intermedius). The hyper-arid coastal valleys of Peru generally support low species richness, particularly for groups such as aquatic breeding amphibians. The discovery of a new species in this environment, and along a major highway crossing the Andes, shows that much remains to be done to document amphibian diversity in Peru.

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Research Article Mon, 2 Feb 2015 00:00:00 +0200
Annotated type catalogue of the Megaspiridae, Orthalicidae, and Simpulopsidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London https://zookeys.pensoft.net/article/4485/ ZooKeys 470: 17-143

DOI: 10.3897/zookeys.470.8548

Authors: Abraham S.H. Breure, Jonathan D. Ablett

Abstract: The type status is described for 65 taxa of the Orthalicoidea, classified within the families Megaspiridae (14), Orthalicidae (30), and Simpulopsidae (20); one taxon is considered a nomen inquirendum. Lectotypes are designated for the following taxa: Helix brephoides d’Orbigny, 1835; Simpulopsis cumingi Pfeiffer, 1861; Bulimulus (Protoglyptus) dejectus Fulton, 1907; Bulimus iris Pfeiffer, 1853. The type status of Bulimus salteri Sowerby III, 1890, and Strophocheilus (Eurytus) subirroratus da Costa, 1898 is now changed to lectotype according Art. 74.6 ICZN. The taxa Bulimus loxostomus Pfeiffer, 1853, Bulimus marmatensis Pfeiffer, 1855, Bulimus meobambensis Pfeiffer, 1855, and Orthalicus powissianus var. niveusPreston 1909 are now figured for the first time. The following taxa are now considered junior subjective synonyms: Bulimus marmatensis Pfeiffer, 1855 = Helix (Cochlogena) citrinovitrea Moricand, 1836; Vermiculatus Breure, 1978 = Bocourtia Rochebrune, 1882. New combinations are: Kuschelenia (Bocourtia) Rochebrune, 1882; Kuschelenia (Bocourtia) aequatoria (Pfeiffer, 1853); Kuschelenia (Bocourtia) anthisanensis (Pfeiffer, 1853); Kuschelenia (Bocourtia) aquila (Reeve, 1848); Kuschelenia (Bocourtia) badia (Sowerby I, 1835); Kuschelenia (Bocourtia) bicolor (Sowerby I, 1835); Kuschelenia (Bocourtia) caliginosa (Reeve, 1849); Kuschelenia (Bocourtia) coagulata (Reeve, 1849); Kuschelenia (Bocourtia) cotopaxiensis (Pfeiffer, 1853); Kuschelenia (Bocourtia) filaris (Pfeiffer, 1853); Kara indentata (da Costa, 1901); Clathrorthalicus magnificus (Pfeiffer, 1848); Simpulopsis (Eudioptus) marmartensis (Pfeiffer, 1855); Kuschelenia (Bocourtia) nucina (Reeve, 1850); Kuschelenia (Bocourtia) ochracea (Morelet, 1863); Kuschelenia (Bocourtia) peaki (Breure, 1978); Kuschelenia (Bocourtia) petiti (Pfeiffer, 1846); Clathrorthalicus phoebus (Pfeiffer, 1863); Kuschelenia (Bocourtia) polymorpha (d’Orbigny, 1835); Scholvienia porphyria (Pfeiffer, 1847); Kuschelenia (Bocourtia) purpurata (Reeve, 1849); Kuschelenia (Bocourtia) quechuarum Crawford, 1939; Quechua salteri (Sowerby III, 1890); Kuschelenia (Bocourtia) subfasciata Pfeiffer, 1853; Clathrorthalicus victor (Pfeiffer, 1854). In an addedum a lectotype is being designated for Bulimulus (Drymaeus) interruptus var. pallidus Preston, 1909. An index is included to all taxa mentioned in this paper and the preceding ones in this series (Breure and Ablett 2011, 2012, 2014).

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Research Article Mon, 12 Jan 2015 00:00:00 +0200
A new species and synonymy of the Neotropical Eucelatoria Townsend and redescription of Myiodoriops Townsend https://zookeys.pensoft.net/article/4340/ ZooKeys 464: 63-97

DOI: 10.3897/zookeys.464.8155

Authors: Diego J. Inclán, John O. Stireman

Abstract: The New World tropics represents the most diverse region for tachinid parasitoids (Diptera: Tachinidae), but it also contains the most narrowly defined, and possibly the most confusing, tachinid genera of any biogeographic region. This over-splitting of genera and taxonomic confusion has limited progress toward our understanding the family in this region and much work is needed to revise, redefine, and make sense of the profusion of finely split taxa. In a recent analysis of the Neotropical genus Erythromelana Townsend, two species previously assigned to this genus, Euptilodegeeria obumbrata (Wulp) and Myiodoriops marginalis Townsend were reinstated as monotypic genera. In the present study, we demonstrate that Euptilodegeeria obumbrata (Wulp), previously assigned to three different genera, represents in fact a species of the large New World genus Eucelatoria Townsend, in which females possess a sharp piercer for oviposition. We also show that the species Eucelatoria carinata (Townsend) belongs to the same species group as Eucelatoria obumbrata, which we here define and characterize as the E. obumbrata species group. Additionally, we describe Eucelatoria flava sp. n. as a new species within the E. obumbrata species group. Finally, we redescribe the genus Myiodoriops Townsend and the single species M. marginalis Townsend.

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Research Article Tue, 16 Dec 2014 00:00:00 +0200
The genus Macroteleia Westwood in Middle Miocene amber from Peru (Hymenoptera, Platygastridae s.l., Scelioninae) https://zookeys.pensoft.net/article/3962/ ZooKeys 426: 119-127

DOI: 10.3897/zookeys.426.7822

Authors: Vincent Perrichot, Rodolfo Salas-Gismondi, John Flynn, Pierre-Olivier Antoine, Michael Engel

Abstract: A new species of the scelionine genus Macroteleia Westwood (Platygastridae s.l., Scelioninae) is described and figured from a female beautifully preserved in middle Miocene amber from Peru. Macroteleia yaguarum Perrichot & Engel, sp. n., shows a unique combination of characters otherwise seen independently within its congeners. It is most similar to the modern M. surfacei Brues, but differs from it by the non-foveolate notauli, the contiguous punctures of the vertex, and the continuous propodeum. The new species is the first New World fossil of the genus, suggesting a Cretaceous origin for the group and a relatively old age of the South American, tropical African, and Australian faunas, and a younger age of the modern Holarctic faunas.

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Research Article Thu, 17 Jul 2014 00:00:00 +0300
A geographic distribution database of Mononychellus mites (Acari, Tetranychidae) on cassava (Manihot esculenta) https://zookeys.pensoft.net/article/3793/ ZooKeys 407: 1-8

DOI: 10.3897/zookeys.407.7564

Authors: Aymer Vásquez-Ordóñez, Soroush Parsa

Abstract: The genus Mononychellus is represented by 28 herbivorous mites. Some of them are notorious pests of cassava (Manihot esculenta Crantz), a primary food crop in the tropics. With the exception of Mononychellus tanajoa (Bondar), their geographic distribution is not widely known. This article therefore reports observational and specimen-based occurrence data of Mononychellus species associated with cassava. The dataset consists of 1,513 distribution records documented by the International Center for Tropical Agriculture (CIAT) between 1975 and 2012. The specimens are held at CIAT’s Arthropod Reference Collection (CIATARC). Most of the records are from the genus’ native range in South America and were documented between 1980 and 2000. Approximately 61% of the records belong to M. tanajoa, 25% to M. caribbeanae (McGregor), 10% to M. mcgregori (Flechtmann and Baker) and 2% to M. planki (McGregor). The complete dataset is available in Darwin Core Archive format via the Global Biodiversity Information Facility (GBIF).

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Data Paper Thu, 8 May 2014 00:00:00 +0300
A new species of the Rhinella margaritifera species group (Anura, Bufonidae) from the montane forest of the Selva Central, Peru https://zookeys.pensoft.net/article/3219/ ZooKeys 371: 35-56

DOI: 10.3897/zookeys.371.6580

Authors: Jiri Moravec, Edgar Lehr, Juan Carlos Cusi, Jesus Cordova, Vaclav Gvozdik

Abstract: We describe a new species of the bufonid toad genus Rhinella from transition montane forest of the buffer zones of the Yanachaga-Chemillén National Park and the Pui Pui Protected Forest (eastern slopes of Andes, Selva Central, Peru). The new species belongs to the Rhinella margaritifera species group (confirmed by mtDNA data) and differs from all its members by the absence of tympanic membrane and tympanic annulus. It is characterized by medium size (SVL 57.5–65.5 mm, n = 5), moderately developed cranial crests, absence of neural crest of vertebrae, absence of bone protrusion at angle of jaw, presence of lateral rows of enlarged tubercles, and absence of subgular vocal sac and vocal slits in males. In addition, based on the molecular phylogenetic analyses of selected Rhinella species we propose the monophylum containing R. chavin, R. festae, R. macrorhina, R. manu, R. nesiotes, R. rostrata, and R. yanachaga as a new species group under the name Rhinella festae species group.

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Research Article Fri, 17 Jan 2014 00:00:00 +0200
Systematics of treefrogs of the Hypsiboas calcaratus and Hypsiboas fasciatus species complex (Anura, Hylidae) with the description of four new species https://zookeys.pensoft.net/article/3213/ ZooKeys 370: 1-68

DOI: 10.3897/zookeys.370.6291

Authors: Marcel Caminer, Santiago Ron

Abstract: We review the systematics of the Hypsiboas calcaratus species complex, a group of widely distributed Amazonian hylid frogs. A comprehensive analysis of genetic, morphological, and bioacoustic datasets uncovered the existence of eleven candidate species, six of which are confirmed. Two of them correspond to Hypsiboas fasciatus and Hypsiboas calcaratus and the remaining four are new species that we describe here. Hypsiboas fasciatus sensu stricto has a geographic range restricted to the eastern Andean foothills of southern Ecuador while Hypsiboas calcaratus sensu stricto has a wide distribution in the Amazon basin. Hypsiboas almendarizae sp. n. occurs at elevations between 500 and 1950 m in central and northern Ecuador; the other new species (H. maculateralis sp. n., H. alfaroi sp. n., and H. tetete sp. n.) occur at elevations below 500 m in Amazonian Ecuador and Peru. The new species differ from H. calcaratus and H. fasciatus in morphology, advertisement calls, and mitochondrial and nuclear DNA sequences. Five candidate species from the Guianan region, Peru, and Bolivia are left as unconfirmed. Examination of the type material of Hyla steinbachi, from Bolivia, shows that it is not conspecific with H. fasciatus and thus is removed from its synonymy.

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Research Article Wed, 15 Jan 2014 00:00:00 +0200
Ophioderma peruana, a new species of brittlestar (Echinodermata, Ophiuroidea, Ophiodermatidae) from the Peruvian coast https://zookeys.pensoft.net/article/3595/ ZooKeys 357: 53-65

DOI: 10.3897/zookeys.357.6176

Authors: Tania Pineda-Enríquez, Francisco Solís-Marín, Yuri Hooker, Alfredo Laguarda-Figueras

Abstract: Ophioderma peruana sp. n. is a new species of Ophiodermatidae, extending the distribution of the genus Ophioderma to Lobos de Afuera Island, Peru, easily distinguishable from its congeners by its peculiarly fragmented dorsal arm plates. Dense granules, rounded or polygonal cover the disc, the radial shields may be naked or completely covered by granules. A good character for recognizing this species in the field is the dorsal side of the disc which is brown with disc granules lighter cream and brown, the arms are mottled with whitish spots and the ventral part of the disc on the interradial part is brown and the radial part bright yellow.

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Research Article Mon, 2 Dec 2013 00:00:00 +0200
Treerunners, cryptic lizards of the Plica plica group (Squamata, Sauria, Tropiduridae) of northern South America https://zookeys.pensoft.net/article/3574/ ZooKeys 355: 49-77

DOI: 10.3897/zookeys.355.5868

Authors: John Murphy, Michael Jowers

Abstract: The arboreal, Neotropical lizard Plica plica (Linnaeus, 1758) has been long considered a widespread species with a distribution east of the Andes. A preliminary examination of 101specimens from about 28 locations mostly north of the Amazon suggests Plica plica is a cryptic species complex with taxa that can be distinguished on the basis of the number of scale rows at mid-body; the arrangement, shape and ornamentation of scales on the snout; the number of lamellae on the fourth toe; the number of subocular plates; as well as other commonly used external morphological traits. The allopatric species discussed here are concordant with northern South American geography. Plica plica (Linnaeus,1758) is associated with the Guiana Shield (Suriname, Guyana and Venezuela). A second species, P. caribeana sp. n. is associated with the Caribbean Coastal Range of Venezuela including Trinidad and Tobago. A third, distinctive species, P. rayi sp. n. is associated with the Middle Orinoco at the eastern edge of the Guiana Shield. Two other species, P. kathleenae sp. n. and P. medemi sp. n., each based upon a single specimen, one from the Sierra Acarai Mountains of Guyana, and the other from southern Meta, Colombia are described. In addition to morphological analyses, we sequenced 12S and 16S rDNA gene fragments from one Plica plica from Trinidad to assess its relationship and taxonomy to other mainland Plica cf plica. The results suggest Plica caribeana sp. n. likely diverged prior to the separation of Trinidad from Northern Venezuela. Isolation in the Caribbean Coastal Range during its rapid uplift in the late Miocene, combined with a marine incursion into northern Venezuela may have contributed to their genetic divergence from other populations.

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Research Article Mon, 25 Nov 2013 00:00:00 +0200
Four new species of Oidardis Hermann, 1912 (Diptera, Asilidae, Laphriinae, Atomosiini) from two major faunistic surveys in the Atlantic Rainforest https://zookeys.pensoft.net/article/3505/ ZooKeys 350: 47-74

DOI: 10.3897/zookeys.350.6096

Authors: Lucas Cezar, Eric Fisher, Carlos Lamas

Abstract: Two recent faunistic surveys in the Brazilian Atlantic Forests region, the PROFAUPAR and the Biota/FAPESP Program, have provided important material for the discovery of new taxa from Brazil. We describe herein four new species of robber-flies of the genus Oidardis (O. falcimystax sp. n., O. fontenellei sp.n., O. maculiseta sp. n. and O. marinonii sp. n.), including illustrations and details on male hypopygia and female genitalia. A distribution map and a key to the species of Oidardis from the Brazilian Atlantic Forests region, including O. triangularis (Hermann), 1912, are also provided.

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Research Article Thu, 14 Nov 2013 00:00:00 +0200
The remarkable scorpion diversity in the Ecuadorian Andes and description of a new species of Tityus C. L. Koch, 1836 (Scorpiones, Buthidae) https://zookeys.pensoft.net/article/3208/ ZooKeys 307: 1-13

DOI: 10.3897/zookeys.307.5334

Authors: Wilson Lourenco, Eric Ythier

Abstract: A new species of Tityus, subgenus Atreus (Scorpiones: Buthidae) is described from the Province of Pichincha in the Ecuadorian Andes. Ecuadorian scorpion fauna remains one of the less well studied among those of South America. Nevertheless, some comments are addressed about its remarkable diversity and high level of endemic elements.

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Research Article Wed, 5 Jun 2013 00:00:00 +0300
Two sympatric new species of woodlizards (Hoplocercinae, Enyalioides) from Cordillera Azul National Park in northeastern Peru https://zookeys.pensoft.net/article/3737/ ZooKeys 277: 69-90

DOI: 10.3897/zookeys.277.3594

Authors: Pablo Venegas, Omar Torres-Carvajal, Vilma Duran, Kevin de Queiroz

Abstract: We report the discovery of two sympatric new species of Enyalioides from a montane rainforest of the Río Huallaga basin in northeastern Peru. Among other characters, the first new species is distinguishable from others Enyalioides by the combination of the following characters: strongly keeled ventral scales, more than 37 longitudinal rows of dorsals in a transverse line between the dorsolateral crests at midbody, low vertebral crest on the neck with vertebrals on neck similar in size to those between hind limbs, projecting scales on body or limbs absent, 96 mm maximum SVL in both sexes, and caudals increasing in size posteriorly within each autotomic segment. The second new species differs from other species of Enyalioides instrongly keeled ventral scales, scales posterior to the superciliaries forming a longitudinal row of strongly projecting scales across the lateral edge of the skull roof in adults of both sexes, 31 or fewer longitudinal rows of strongly keeled dorsals in a transverse line between the dorsolateral crests at midbody, vertebrals on neck more than five times the size of vertebrals between hind limbs in adult males, projecting scales on body or limbs absent, and caudals increasing in size posteriorly within each autotomic segment. We also present an updated molecular phylogenetic tree of hoplocercines including new samples of E. rudolfarndti, E. rubrigularis, both species described in this paper, as well as an updated identification key for species of Hoplocercinae.

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Research Article Fri, 15 Mar 2013 00:00:00 +0200
Two new species of Phrynopus (Anura, Strabomantidae) from high elevations in the Yanachaga-Chemillén National park in Peru (Departamento de Pasco) https://zookeys.pensoft.net/article/3259/ ZooKeys 235: 51-71

DOI: 10.3897/zookeys.235.3885

Authors: Edgar Lehr, Jiri Moravec, Juan Carlos Cusi

Abstract: We describe two new species of Phrynopus from a cloud forest of the Cordillera Yanachaga, Yanachaga-Chemillén National Park in Peru and describe and document the first clutch and case of parental care for a species of Phrynopus. One of the new species of Phrynopus is described based on two females (SVL 19.1–21.0 mm) which were found in leaf litter and moss layer in a cloud forest at 2900 m elevation. This new species is most similar to Phrynopus bracki. The second new species of Phrynopus was found in the transitional formation between cloud forest and wet puna at 3000 m elevation. Its description is based on a single female (SVL 20.7 mm) that was observed guarding nine eggs under moss. This new species is most similar to Phrynopus nicoleae. The eggs had a diameter of 5.7–5.8 mm (n = 3) and froglets when hatched a SVL of 6.2–6.5 mm (n = 3). Sympatric anurans include Gastrotheca sp., Pristimantis aniptopalmatus, P. bromeliaceus, Pristimantis sp., and Rhinella yanachaga.

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Research Article Wed, 31 Oct 2012 00:00:00 +0200
The status of the genus Bostryx Troschel, 1847, with description of a new subfamily (Mollusca, Gastropoda, Bulimulidae) https://zookeys.pensoft.net/article/2998/ ZooKeys 216: 1-3

DOI: 10.3897/zookeys.216.3646

Authors: Abraham Breure

Abstract: The status of the genus Bostryx is discussed and, based on morphological and molecular data, restricted to a group of species related to B. solutus, for which the new subfamily name Bostrycinae is introduced.

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Short Communication Tue, 21 Aug 2012 00:00:00 +0300
Three new species of Sesioctonus Viereck (Hymenoptera, Braconidae, Agathidinae) from Peru https://zookeys.pensoft.net/article/2810/ ZooKeys 196: 1-10

DOI: 10.3897/zookeys.196.3086

Authors: Lidia Sulca, Michael Sharkey

Abstract: Three new species of Sesioctonus (Braconidae: Agathidinae) are described and illustrated, i.e., Sesioctonus huggerti, S.wayquecha, and S. bina. Two new Peruvian species records for Sesioctonus are reported: S. longinoi and S. diazi. A revised key to all known species of Sesioctonus is presented.

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Research Article Mon, 21 May 2012 00:00:00 +0300
A remarkable new species of Euragallia from Peru (Insecta, Hemiptera, Cicadellidae, Agalliini), including the description of a peculiar structure of the male genitalia https://zookeys.pensoft.net/article/2644/ ZooKeys 178: 51-58

DOI: 10.3897/zookeys.178.3038

Authors: Luiz Rodrigues, Ana Gonçalves, Gabriel Mejdalani

Abstract: A new species of Euragallia Oman, 1938 from Peru (Pasco Department) is described and illustrated. Euragallia batmani sp. n. can be distinguished from the other species of the genus by the very posteriorly pronounced male pygofer, with an apical hook-like projection, and by the well-developed dorsal area of the aedeagal base, resembling the open wings of a bat. With the addition of E. batmani sp. n., the number of Euragallia species is increased to 21. Only one additional species of the genus is recorded from Peru (E. prion Kramer, 1976). A comparison between the new species and E. prion is provided. A conspicuous structure, which connects the subgenital plates to the styles, is described in detail and named.

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Research Article Thu, 29 Mar 2012 00:00:00 +0300
First record of the genus Phradis Förster (Hymenoptera, Ichneumonidae, Tersilochinae) from the Neotropical Region https://zookeys.pensoft.net/article/2444/ ZooKeys 169: 31-38

DOI: 10.3897/zookeys.169.2333

Authors: Andrey Khalaim, Santiago Bordera

Abstract: One new species of the genus Phradis, P. peruvianus sp. n., from the mountainous part of Peruvian Amazonia, is described and illustrated. This is the first record of the genus from South America and the Neotropical region.

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Research Article Fri, 10 Feb 2012 00:00:00 +0200
Discovery of the genus Meggoleus Townes, 1971 (Hymenoptera, Ichneumonidae, Tersilochinae) in Peru, with the description of two new species https://zookeys.pensoft.net/article/2380/ ZooKeys 163: 83-90

DOI: 10.3897/zookeys.163.2291

Authors: Mabel Alvarado

Abstract: The genus Meggoleus Townes, 1971 (Ichneumonidae, Tersilochinae) currently comprises two species, one from Brazil and one from Gabon. The genus is recorded from Peru for the first time, with a range extension of the type species, Meggoleus spirator Townes, 1971, and the discovery of two new species – Meggoleus fuscatus sp. n. and Meggoleus pampahermosensis sp. n. A key to the world’s species is provided.

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Research Article Mon, 9 Jan 2012 00:00:00 +0200
Euglossa williamsi, a new species of orchid bee from the Amazon Basin of Ecuador and Peru, with notes on its taxonomic association and biogeography (Hymenoptera, Apidae) https://zookeys.pensoft.net/article/2337/ ZooKeys 159: 49-63

DOI: 10.3897/zookeys.159.2239

Authors: Ismael Hinojosa-Díaz, Michael Engel

Abstract: Euglossawilliamsi sp. n. is here described from the lowland Amazonianregion in Ecuador and Peru, and as part of a small species assemblage within Euglossa consisting of E. dodsoni Moure and E. obtusa Dressler. An identification key to the males of the group is provided plus detailed figures of the new species and representative illustrations for the others. A brief discussion of the taxonomic and biogeographical implications of the new species is provided. New records in Honduras and Nicaragua are provided for the related E. dodsoni.

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Research Article Fri, 23 Dec 2011 00:00:00 +0200
A new genus and species of termites (Isoptera, Termitidae, Nasutitermitinae) from the rainforest of northern Peru https://zookeys.pensoft.net/article/2343/ ZooKeys 159: 1-9

DOI: 10.3897/zookeys.159.2311

Authors: Carolina Cuezzo, David A. Nickle

Abstract: A new genus and species of nasutitermitine termites are described and illustrated, based on soldier and worker characters. Sinqasapatermes gen. n., can be distinguished from all other nasutitermitine genera by its singular worker gut coiling and enteric valve characters: distal margin of the enteric valve not everted into the paunch but bending towards the ileum, that is, directed against the flow of food; enteric valve armature with one ring of six equal subtriangularly-shaped ridges, each ridge with short spines on the entire surface; enteric valve armature situated on external face of cone, facing the internal ileum wall; enteric valve seating tri-lobed and separated from remaining portion of the paunch; paunch subdivided. Sinqasapatermes sachae sp. n., was collected on a tree in a very narrow flattened tunnel that was well concealed beneath lichens in a northern Peru rainforest (Arcadia, Loreto Province).

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Research Article Fri, 23 Dec 2011 00:00:00 +0200
Carabidae diversity along an altitudinal gradient in a Peruvian cloud forest (Coleoptera) https://zookeys.pensoft.net/article/2918/ ZooKeys 147: 651-666

DOI: 10.3897/zookeys.147.2047

Authors: Sarah Maveety, Robert Browne, Terry Erwin

Abstract: Carabid beetles were sampled at five sites, ranging from 1500 m to 3400 m, along a 15 km transect in the cloud forest of Manu National Park, Perú. Seasonal collections during a one year period yielded 77 morphospecies, of which 60% are projected to be undescribed species. There was a significant negative correlation between species richness and altitude, with the number of carabid species declining at the rate of one species for each 100 m increase in altitude. The majority of species (70.1 %) were restricted to only one altitudinal site and no species was found at more than three of the five altitudinal sites. Only one genus, Pelmatellus (Tribe Harpalini), was found at all five sites. Active (hand) collections yielded approximately twice as many species per individuals collected than passive (pitfall trap) collections. This study is the first systematic sampling of carabid beetles of a high altitude gradient in the cloud forests of southeastern Perú and supports the need to conserve the zone of extremely high biodiversity present on the eastern slopes of the Peruvian Andes.

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Research Article Wed, 16 Nov 2011 00:00:00 +0200
A new bush anole (Iguanidae, Polychrotinae, Polychrus) from the upper Marañon basin, Peru, with a redescription of Polychrus peruvianus (Noble, 1924) and additional information on P. gutturosus Berthold, 1845 https://zookeys.pensoft.net/article/2851/ ZooKeys 141: 79-107

DOI: 10.3897/zookeys.141.1678

Authors: Claudia Koch, Pablo Venegas, Antonio García Bravo, Wolfgang Boehme

Abstract: We herein describe a new colorful species of Polychrus with a conspicuous sexual dimorphism from the dry forest of the northern portion of Región de La Libertad, Peru. The new species differs from all other Polychrus species, in that this species has very small dorsal scales and thus a higher number of scales around midbody and in the middorsal line from behind the occipital scales to the level of the posterior edge of the thigh. Furthermore, we redescribe P. peruvianus whose original description is short and lacks information on intraspecific variation and sexual dimorphism. Also, we add some information on intraspecific variation and ecology of P. gutturosus. Finally, we synonymize P. spurrelli Boulenger with P. gutturosus.

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Research Article Fri, 28 Oct 2011 00:00:00 +0300
A new species of Quexua from southeastern Peru (Hymenoptera, Crabronidae) https://zookeys.pensoft.net/article/2855/ ZooKeys 141: 65-70

DOI: 10.3897/zookeys.141.1965

Authors: Daniel Bennett

Abstract: A distinctive new species of the crabronine wasp genus Quexua Pate is described and figured from a single male collected from lowland Amazonian rain forest in southeastern Peru. Quexua cicra sp. n. is the only species in the genus known with a sessile metasoma.

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Research Article Fri, 28 Oct 2011 00:00:00 +0300
Black-tie dress code: two new species of the genus Toxomerus (Diptera, Syrphidae) https://zookeys.pensoft.net/article/2850/ ZooKeys 140: 1-26

DOI: 10.3897/zookeys.140.1930

Authors: Ximo Mengual

Abstract: Toxomerus hauseri Mengual sp. n. and T. picudus Mengual sp. n. are described from Peru and Ecuador respectively. Toxomerus circumcintus (Enderlein, 1938) is treated as a valid species and not considered synonym of T. marginatus, and Toxomerus ovatus (Hull, 1942) is considered junior synonym of Toxomerus nitidus (Schiner, 1868). An identification key for the Toxomerus species with dark abdomens is given along with diagnoses for each studied species.

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Research Article Wed, 26 Oct 2011 00:00:00 +0300
Annotated type catalogue of the Amphibulimidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London https://zookeys.pensoft.net/article/2841/ ZooKeys 138: 1-52

DOI: 10.3897/zookeys.138.1847

Authors: Abraham Breure, Jonathan Ablett

Abstract: The type status is described of 39 taxa classified within the family Amphibulimidae (superfamily Orthalicoidea) and kept in the London museum. One taxon, Bulimus elaeodes Pfeiffer, 1853, is removed to the Strophocheilidae. Lectotypes are designated for Bulimus adoptus Reeve, 1849; Bulimus (Eurytus) eros Angas, 1878; Helix onca d’Orbigny, 1835; Amphibulima pardalina Guppy, 1868. The type status of the following taxon is changed to lectotype in accordance with Art. 74.6 ICZN: Strophocheilus (Dryptus) jubeus Fulton, 1908.As general introduction to this and following papers on Orthalicoid types in the Natural History Museum, a brief history of the London collection is given and several examples of handwriting from different authors are presented.

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Research Article Wed, 19 Oct 2011 00:00:00 +0300
First record of Acaenitinae (Hymenoptera, Ichneumonidae) from South America with description of a new species and a key to the world species of Arotes Gravenhorst https://zookeys.pensoft.net/article/2828/ ZooKeys 137: 77-88

DOI: 10.3897/zookeys.137.1788

Authors: Carol Castillo, llari Saaksjarvi, Andrew Bennett, Gavin Broad

Abstract: A new species of Acaenitinae, Arotes ucumari Castillo & Sääksjärvi sp. n., is described and illustrated representing the first record of the subfamily from South America. The new species was collected from a premontane tropical rain forest in the Peruvian Andes at 1500 m. A key to the world species of Arotes Gravenhorst, 1829 is provided. The subspecies Arotes albicinctus moiwanus (Matsumura, 1912) is raised to species rank, Arotes moiwanus stat. n.

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Research Article Fri, 14 Oct 2011 00:00:00 +0300
Two new species of Andean gymnophthalmid lizards of the genus Euspondylus (Reptilia, Squamata) from central and southern Peru https://zookeys.pensoft.net/article/2568/ ZooKeys 109: 1-17

DOI: 10.3897/zookeys.109.1304

Authors: Germán Chávez, Karen Siu-Ting, Vilma Duran, Pablo Venegas

Abstract: Two new species of lizards assigned to the genus Euspondylus from the montane forests of the Peruvian Andes in the Pasco Department (central Peru) and Ayacucho Department (southern Peru) both at elevations of 2550 and 3450 m, respectively, are described. The new species are distinguishable from all other Peruvian and Ecuadorian species of Euspondylus by a unique combination of morphometric, scalation and color pattern characteristics. Natural history data for the new species and for E. spinalis are also provided.

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Research Article Mon, 20 Jun 2011 00:00:00 +0300
Agra, arboreal beetles of Neotropical forests: pusilla group and piranha group systematics and notes on their ways of life (Coleoptera, Carabidae, Lebiini, Agrina) https://zookeys.pensoft.net/article/2430/ ZooKeys 66: 1-28

DOI: 10.3897/zookeys.66.684

Authors: Terry Erwin

Abstract: Revisions of two new species groups of the genus Agra Fabricius are presented with the following species described as new: pusilla group - Agra cruciaria sp. n. (Brazil), Agra grace sp. n. (Ecuador, Perú), Agra max sp. n. (Brazil), Agra minasianus sp. n. (Brazil), Agra notpusilla sp. n. (Brazil), Agra pseudopusilla sp. n. (Brazil); piranha group - Agra ce sp. n. (Perú), Agra risseri sp. n. (Bolivia, Brazil), Agra maia sp. n. (Bolivia), Agra piranha sp. n. (Ecuador); Agra tiputini sp. n. (Ecuador). Species of these two groups have adults that are the smallest in the entire genus, although this does not indicate they are closely related based on other attributes. All species are Amazonian in distribution.

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Research Article Thu, 4 Nov 2010 00:00:00 +0200
New data on Neotropical Scolytus Geoffroy, 1762 with description of five new species from Peru (Coleoptera, Curculionidae, Scolytinae) https://zookeys.pensoft.net/article/2293/ ZooKeys 56: 65-104

DOI: 10.3897/zookeys.56.519

Authors: Alexander Petrov, Michail Mandelshtam

Abstract: Five new species of Scolytus Geoffroy, 1762 (Coleoptera: Curculionidae: Scolytinae) are described from Peru, namely S. woodi, S. carveli, S. vagabundus, S. lindemani, S. mozolevskae. The following new synonym is established: Scolytus angustatus Browne, 1970 (= Scolytus facialis Schedl, 1973, syn. n.) New records of the Scolytus species in Loreto, Junin ,Cusco and Madre de Dios Provinces are given and the biology of the genus representatives is discussed.

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Research Article Fri, 17 Sep 2010 00:00:00 +0300
Revision of the Bee Genus Chlerogella (Hymenoptera, Halictidae), Part II: South American Species and Generic Diagnosis https://zookeys.pensoft.net/article/2202/ ZooKeys 47: 1-100

DOI: 10.3897/zookeys.47.416

Authors: Michael Engel

Abstract: The South American species of the rare bee genus Chlerogella Michener (Halictinae: Augochlorini) are revised, completing the study of the genus. Chlerogella diversity is significantly expanded beyond the five previously described South American species of Cherlogella azurea (Enderlein), comb. n., C. nasus (Enderlein), C. mourella Engel, C. octogesima (Brooks & Engel), comb. n., and C. buyssoni (Vachal). Twenty-two new species are described – C. agaylei sp. n., C. arhyncha sp. n., C. borysthenis sp. n., C. breviceps sp. n., C. cochabambensis sp. n., C. cooperella sp. n., C. cyranoi sp. n., C. dolichorhina sp. n., C. elysia sp. n., C. eumorpha sp. n., C. euprepia sp. n., C. hauseri sp. n., C. hypermeces sp. n., C. materdonnae sp. n., C. oresbios sp. n., C. picketti sp. n., C. rostrata sp. n., C. silvula sp. n., C. terpsichore sp. n., C. tychoi sp. n., C. vachali sp. n., C. xuthopleura sp. n. – and the distribution of the genus is expanded beyond Perú and Ecuador to include Bolivia, Colombia, and Venezuela. The female of C. azurea is described for the first time while the placement of Halictus buyssoni Vachal in Chlerogella is considered tentative, following the usage of previous authors, as the holotype and sole specimen is untraceable. The genus is newly diagnosed based on a greater understanding of variation in malar length across the species and a dichotomous key is provided. New floral records for species of Chlerogella include Psychotria pongoana Standl. (Rubiaceae) and a putative record on Phragmopedium longifolium (Warsz. & Rchb.f.) Rolfe (Orchidaceae).

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Research Article Fri, 21 May 2010 00:00:00 +0300
A checklist to the wasps of Peru (Hymenoptera, Aculeata) https://zookeys.pensoft.net/article/2007/ ZooKeys 15: 1-78

DOI: 10.3897/zookeys.15.196

Authors: Claus Rasmussen, Angelico Asenjo

Abstract: The first checklist to the 225 genera and 1169 reported species-group taxa of aculeate wasps of Peru is presented. The list is based on a literature survey and examination of Peruvian entomological collections and include locality references for each taxon. Bibliographic references for the identification of families, genera, and species are provided when available. The occurrence data are published in addition as a downloadable file (doi: 10.3897/zookeys.15.196.app.2.ds, doi: 10.3897/zookeys.15.196.app.3.ds, and 10.3897/zookeys.15.196.app.4.ds) and were uploaded onto GBIF infrastructure simultaneously with the publication process. The following new combinations are proposed: Ancistroceroides cirrifer (Zavattari, 1912), Ancistrocerus epicus (Zavattari, 1912), and Stenodynerus corallineipes (Zavattari, 1912).

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Catalogue Thu, 23 Jul 2009 00:00:00 +0300
Two new species of Xanthopimpla (Hymenoptera: Ichneumonidae) from Western Amazonia, with a revised key to the Neotropical species of the genus https://zookeys.pensoft.net/article/2005/ ZooKeys 14: 55-65

DOI: 10.3897/zookeys.14.193

Authors: Isrrael Gomez, llari Saaksjarvi, Anu Veijalainen, Gavin Broad

Abstract: Xanthopimpla Saussure, 1892 is one of the largest genera of the family Ichneumonidae. The genus is very species rich in Indo-Australia but poorly represented in the Neotropical region. In this paper two morphologically very distinctive species of the genus, X. amazonica sp.n. Gómez, Sääksjärvi & Veijalainen, and X. jussilai sp.n. Veijalainen, Sääksjärvi & Broad, are described and illustrated. The new species have been collected in the Peruvian and Ecuadorean Amazonia. A revised key to the Neotropical species of the genus is provided. X. aurita Krieger and X. craspedoptera Krieger are recorded for the first time from Peru.

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Research Article Fri, 10 Jul 2009 00:00:00 +0300
A description of four new species of fleas (Insecta, Siphonaptera) from Angola, Ethiopia, Papua New Guinea, and Peru https://zookeys.pensoft.net/article/1977/ ZooKeys 8: 39-61

DOI: 10.3897/zookeys.8.82

Authors: Michael Hastriter

Abstract: Four new species of fleas are described: Aphropsylla truncata sp. n. (Ethiopia), Ectinorus hirsutus sp. n. (Peru), Rhinolophopsylla traubi sp. n. (Angola), and Thaumapsylla wilsoni sp. n. (Papua New Guinea). Our understanding of the genus Aphropsylla is deficient, therefore a discussion of host/parasite relationships and new records from Uganda are provided. A key is provided for the genus Aphropsylla, while representatives of the other three genera are associated with existing keys to facilitate their identification. The presence of lucodiscs on Apropsylla and other genera is briefly discussed. The occurrence of lucodiscs among representatives of the order Siphonaptera deserves further investigation.

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Research Article Tue, 28 Apr 2009 00:00:00 +0300