Protagonista species, holotypes and labels. 44 P. lugubris, male 45 P. pertristis stat. rev., female 46 P. fusca, now synonym of P. pertristis, male 47 P. aterrima, now synonym of P. pertristis, female. Scale bars: 5.0 mm.

  Part of: Wang C, Wang Z, Che Y (2016) Protagonista lugubris, a cockroach species new to China and its contribution to the revision of genus Protagonista, with notes on the taxonomy of Archiblattinae (Blattodea, Blattidae). ZooKeys 574: 57-73.