Oniticellini topology with characters and states shown. A Topology above using unweighted data shows the tribe as paraphyletic without the Onthophagini B Topology below using the weighted data with K value of 10 shows the tribe as monophyletic.

 
 
  Part of: Philips TK (2016) Phylogeny of the Oniticellini and Onthophagini dung beetles (Scarabaeidae, Scarabaeinae) from morphological evidence. ZooKeys 579: 9-57. https://doi.org/10.3897/zookeys.579.6183