A, B Cladogram found using Piwe weighting with K values of 8-10. This topology is considered to be the best supported in this study. Bootstrap values above 50% found for nodes indicated. A Oniticellini. B Onthophagini. * = no taxon image.

 
 
  Part of: Philips TK (2016) Phylogeny of the Oniticellini and Onthophagini dung beetles (Scarabaeidae, Scarabaeinae) from morphological evidence. ZooKeys 579: 9-57. https://doi.org/10.3897/zookeys.579.6183