Taxonomy of Fissocantharis Pic (Coleoptera, Cantharidae) from Guangxi, China, with descriptions of six new species

Abstract A total of 17 species of Fissocantharis Pic is recorded from Guangxi, China. Six species are described new to science, Fissocantharis sinensomima sp. n., Fissocantharis sexcostata sp. n., Fissocantharis basilaris sp. n., Fissocantharis eschara sp. n., Fissocantharis latipalpa sp. n. and Fissocantharis biprojicientis sp. n., and two previously known species are redescribed, Fissocantharis gracilipes (Pic, 1927) and Fissocantharis sinensis (Wittmer, 1988). These species are presented with habitus of males, abdominal sternites VIII of females and genitalia of both sexes. Fissocantharis flavofacialis (Pic, 1926) is synonymized with Fissocantharis angusta (Fairmaire, 1900); both were originally described in the genus Podabrus Westwood. Additionally, a key and a checklist of all the species of Fissocantharis from Guangxi are provided.


Introduction
The species of Fissocantharis Pic, 1921(redefined by Yang et al. 2009  Antennomeres XI about as twice long as X ... F. tachulanensis (Wittmer, 1988)  Redescription. Male (Fig. 1A). Head black, mouthparts blackish brown, light brown at bases of mandibles and labium, antennae black, yellow at ventral sides of antennomeres I-II, prothorax orange, pronotum sometimes with a large black marking in middle of disc, which extending from anterior to posterior margin, scultellum black, elytra dark purple, with weak metallic shine, legs black, yellow at pro-coxae, trochanters and basal parts of femora, meso-and metasterna and abdomen black. Body densely  (Wittmer, 1988) B F. gracilipes (Pic, 1927) C F. sinensomima sp. n. D F. sexcostata sp. n. Scale bars: 2.0 mm. covered with short decumbent light brown pubescence, also mixed with slightly long semierect pubescence along anterior margin of labrum and on disc of elytra.
Head subquadrate, temples evenly narrowed posteriad, surface semilustrous, finely and densely punctate; eyes strongly protruding, head breadth across eyes distinctly wider than anterior margin of pronotum; maxillary palpomeres IV longer than wide, widest at apical one-third, arcuate and sharp at apical parts of inner margins; antennae almost extending to apical one-third length of elytra, antennomeres II slightly longer than wide at apices, III-X slightly flattened and obliquely widened apically, III about twice as long as II, IV slightly longer than VIII, V-VIII each with a longitudinal smooth impression at apical part of outer margin, XI parallel-sided, slightly longer than X and pointed at apices.
Pronotum about 1.10 times longer than wide, widest near base, anterior margin rounded, anterior angle rounded, lateral margins sinuate, slightly diverging posteriad, posterior angle nearly rectangular, posterior margin arcuate and slightly bordered, disc distinctly convex on postero-lateral parts, surface semilustrous, punctate like that on head.
All tarsal claws bifid, upper claws nearly as long as lower claws. Aedeagus ( Fig. 3A-C): conjoint dorsal plate of parameres well-developed, about half length of ventral processes, nearly parallel-sided, with apical margin tapered apically; ventral process of each paramere evenly narrowed apically, largely hooked at apex.
Female. Similar to male, but eyes not so protruding; antennae shorter, extending to elytral mid-length, antennomeres III-X nearly parallel-sided, V-VIII without impressions; pronotum slightly wider, about 1.05 times longer than wide, moderately convex at posterolateral parts of disc; legs black at profemora. Abdominal sternite VIII (Fig. 8A) slightly emarginated on both sides of posterior margin, middle part between lateral emarginations slightly acute at apex, latero-apical angles widely rounded. Internal organ of reproductive system ( Fig. 9A): vagina stout and abruptly narrowed and extended into a long duct above median oviduct; diverticulum and spermathecal duct arising from the end of the long duct of vagina; diverticulum moderately long, thin and spiral; spermathecal duct distinctly thicker and shorter than diverticulum; spermatheca composed of a spiral tube which is distinctly longer than diverticulum, provided with a very long and thin accessory gland which is much longer than the spiral tube of spermatheca; median oviduct situated in middle of vagina.

Remarks.
In the original publication (Wittmer 1988), some characteristics of antennae for the male is not indicated, which however is important for diagnosis of Fissocantharis species. Herein it is redescribed and also provided with some supplementary characters for abdominal sternite VIII and genitalia of the female. (Pic, 1927) Figs 1B, 3D-F, 8B, 9B
Head subquadrate, temples evenly narrowed posteriad, surface semilustrous, finely and densely punctate; eyes strongly protruding, head breadth across eyes distinctly wider than anterior margin of pronotum; maxillary palpomeres IV longer than wide, widest at apical one-third, arcuate and sharp at apical parts of inner margins; antennae filiform, almost extending to apical one-fourth length of elytra, antennomeres II slightly longer than wide at apices, III about twice as long as II, IV slightly longer than III, IV-XI each with an oblong smooth scar-like bulge at basal part of outer margin, XI slightly longer than X and pointed at apices.
Pronotum about 1.17 times longer than wide, widest near base, anterior margin rounded, anterior angle rounded, lateral margins sinuate, moderately diverging posteriorly, posterior angle nearly rectangular, posterior margin arcuate and slightly bordered, disc distinctly convex on posterolateral parts, surface semilustrous, sparsely and finely punctate.
All tarsal claws bifid, upper claws nearly as long as lower claws. Aedeagus ( Fig. 3D-F): conjoint dorsal plate of parameres greatly reduced, slightly roundly protuberant in middle of apical margin; ventral process of each paramere abruptly narrowed apically, slightly hooked at apex.
Female. Similar to male, but eyes not so protruding; antennae shorter, extending to elytral mid-length, antennomeres IV-XI without bulges; pronotum slightly wider, about 1.10 times longer than wide, moderately convex at posterolateral parts of disc. Abdominal sternite VIII (Fig. 8B) slightly emarginated on both sides of posterior margin, middle part between lateral emarginations arcuate, latero-apical angles widely rounded. Internal organ of reproductive system (Fig. 9B): vagina stout and abruptly narrowed and extended into a long duct above median oviduct; diverticulum and spermathecal duct arising from the end of the long duct of vagina; diverticulum slightly long, thin and spiral; spermathecal duct distinctly thicker and nearly as long as diverticulum; spermatheca composed of a spiral tube which is distinctly longer than diverticulum, provided with a very long and thin accessory gland which is much longer than the spiral tube of spermatheca; median oviduct situated in middle of vagina.
Remarks. The elytra of the holotype are purple, but the coloration could be variable in cantharid species bearing a metallic shine, not only in Fissocantharis, but also in Themus Motschulsky. By contrast, the characteristics of the aedeagus and antennae of the male are much more stable and reliable, which are the basis of our determination of the additional specimens as this species. Description. Male (Fig. 1C). Head black, mouthparts blackish brown, light brown at bases of mandibles and labium, antennae black, yellow at ventral sides of antennomeres I-II, prothorax yellow, pronotum with a large black marking in middle of disc, which extending from anterior to posterior margin, scultellum black, elytra dark purple, with weak metallic shine, legs black, yellow at coxae, trochanters and basal parts of femora, meso-and metasterna and abdomen black. Body densely covered with short decumbent light brown pubescence, also mixed with slightly long semierect pubescence along anterior margin of labrum and on disc of elytra.
Head subquadrate, temples evenly narrowed posteriorly, surface semilustrous, finely and densely punctate; eyes strongly protruding, head breadth across eyes distinctly wider than anterior margin of pronotum; maxillary palpomeres IV longer than wide, widest at apical one-third, arcuate and sharp at apical parts of inner margins; antennae filiform, almost extending to apical one-third length of elytra, antennomeres II slightly longer than wide at apices, III-XI parallel-sided, III about twice as long as II, IV-XI each with a narrow longitudinal smooth impression at basal part of outer margin, IV about one-third longer than III, XI slightly longer than X and pointed at apices. Pronotum about 1.10 times longer than wide, widest near base, anterior margin rounded, anterior angle rounded, lateral margins sinuate, slightly diverging posteriad, posterior angle nearly rectangular, posterior margin arcuate and slightly bordered, disc distinctly convex on posterolateral parts, surface semilustrous, punctate like that on head.
All tarsal claws bifid, upper claws nearly as long as lower claws. Aedeagus (Figs 4A-C): conjoint dorsal plate of parameres greatly reduced, slightly emarginated in middle of apical margin; ventral process of each paramere evenly narrowed apically, largely hooked at apex.
Female. Unknown. Body length: 6.0 mm; width: 1.5 mm. Diagnosis. This species is similar to F. sinensis, but can be distinguished by the antennomeres IV-XI each with a narrow longitudinal smooth impression along basal part of outer margin in male; aedeagus: conjoint dorsal plate of parameres greatly reduced, slightly emarginated in middle of apical margin.
Distribution. China (Guangxi). Etymology. The specific name is derived from Latin mimus (similar, imitating something), referring to its similarity to F. sinensis (Wittmer, 1988). Description. Male (Fig. 1D). Head black, mouthparts blackish brown, light brown at bases of mandibles and labium, antennae black, orange at antennomeres I-II and ventral sides of III, prothorax orange, pronotum with a large inverse-triangular and a slightly small triangular black markings in middle of anterior and posterior parts of disc respectively, two markings almost conjoint, scultellum black, elytra dark purple, with weak metallic shine, legs black, yellow at pro-coxae, trochanters and femora and meso-trochanters and bases of femora, meso-and metasterna and abdomen black. Body densely covered with short decumbent light brown pubescence, also mixed with slightly long semierect pubescence along anterior margin of labrum and on disc of elytra.
Head subquadrate, temples evenly narrowed posteriorly, surface semilustrous, finely and densely punctate; eyes moderately protruding, head breadth across eyes dis-tinctly wider than anterior margin of pronotum; maxillary palpomeres IV longer than wide, widest at apical one-third, arcuate and sharp at apical parts of inner margins; antennae almost extending to apical one-third length of elytra, antennomeres II nearly as long as wide at apices, III-X slightly widened apically, nearly long-triangular, the whole length of III-VII and basal two-thirds length of VIII each with a longitudinal ridge along outer margin, IV slightly longer than III, XI parallel-sided, slightly longer than X and pointed at apices.
Pronotum about 1.10 times longer than wide, widest near base, anterior margin rounded, anterior angle rounded, lateral margins slightly sinuate and diverging posteriad, posterior angle nearly rectangular, posterior margin arcuate and slightly bordered, disc distinctly convex on posterolateral parts, surface semilustrous, punctate like that on head.
All tarsal claws bifid, upper claws nearly as long as lower claws. Aedeagus ( Fig. 4D-F): conjoint dorsal plate of parameres moderately reduced, distinctly shorter than ventral process, with apical margin tapered apically; ventral process of each paramere evenly narrowed apically, largely hooked at apex.
Female. Similar to male, but eyes not so protruding; antennae uniformly black, antennomeres III-X nearly parallel-sided, III-VIII without ridges; pronotum slightly wider, about 1.12 times longer than wide, lateral margins sinuate, moderately diverging posteriad, moderately convex at posterolateral parts of disc, legs orange at procoxae and trochanters. Abdominal sternite VIII (Fig. 8C) slightly emarginated on both sides of posterior margin, middle part between lateral emarginations slightly arcuate, latero-apical angles narrowly rounded. Internal organ of reproductive system (Fig.  9C): vagina stout and abruptly narrowed and extended into a long duct above median oviduct; diverticulum and spermathecal duct arising from the end of the long duct of vagina; diverticulum slightly long, thin and spiral; spermathecal duct distinctly thicker and shorter than diverticulum; spermatheca composed of a spiral tube which is distinctly longer than diverticulum, provided with a long and thin accessory gland which is slightly longer than the spiral tube of spermatheca; median oviduct situated in middle of vagina.
Body length: 6.0-10.0 mm; width: 1.2-2.0 mm. Diagnosis. This species is similar to F. sinensis, but can be easily differentiated by the antennomeres III-VIII with longitudinal ridges along outer margins in male; aedeagus: conjoint dorsal plate of parameres moderately reduced.
Distribution. China (Guangxi). Etymology. The specific name is derived from Latin sex (six) and costatus (ridged), referring to its antnnomeres III-VIII with longitudinal ridges (six ridges in total) along outer margins in male.
Remarks. Sometimes the pronotum is uniformly orange, without any black markings, and this variation always occurs on the females. Description. Male ( Fig. 2A). Head black, mouthparts blackish brown, light brown at bases of mandibles and labium, antennae black, yellow at ventral sides of antennomeres I-III, pronotum black, scultellum black, elytra blue, with strong metallic shine, legs black, yellow at apical parts of coxae, trochanters and basal parts of femora, presternum dark brown, meso-and metasterna and abdomen black. Body densely covered with short decumbent dark brown pubescence, also mixed with slightly long semierect pubescence along anterior margin of labrum and on disc of elytra.
Head subquadrate, temples evenly narrowed posteriad, surface semilustrous, finely and densely punctate; eyes strongly protruding, head breadth across eyes distinctly wider than anterior margin of pronotum; maxillary palpomeres IV longer than wide, widest at apical one-third, arcuate and sharp at apical parts of inner margins; antennae almost extending to apical one-fourth length of elytra, antennomeres II slightly longer than wide at apices, III-X slightly flattened and widened apically, III about twice as long as II, IV-XI each with a small rounded smooth impression at base of outer margin, IV about one-third longer than III, XI nearly parallel-sided, slightly longer than X and pointed at apices.
Pronotum about 1.26 times longer than wide, widest near base, anterior margin rounded, anterior angle rounded, lateral margins sinuate, moderately diverging posteriorly, posterior angle nearly rectangular, posterior margin arcuate and slightly bordered, disc distinctly convex on posterolateral parts, surface semilustrous, punctate like that on head.
All tarsal claws bifid, upper claws nearly as long as lower claws. Aedeagus ( Fig. 5A-C): conjoint dorsal plate of parameres greatly reduced, rounded at apical margin; ventral process of each paramere evenly narrowed apically, slightly hooked at apex.
Female. Similar to male, but eyes not so protruding; antennae shorter, extending to elytral mid-length, antennomeres III-X nearly parallel-sided, IV-XI without impressions; pronotum slightly wider, about 1.13 times longer than wide, moderately convex at postero-lateral parts of disc. Abdominal sternite VIII (Fig. 8D) roundly protuberant in middle of posterior margin, latero-apical angels subrounded. Internal organ of reproductive system (Fig. 10A): vagina stout and abruptly narrowed and extended into a long duct above median oviduct; diverticulum and spermathecal duct arising from the end of the long duct of vagina; diverticulum slightly long, thin and spiral; spermathecal duct distinctly thicker and slightly shorter than diverticulum; spermatheca composed of a spiral tube which is distinctly longer than diverticulum, provided with a very long and thin accessory gland, which is much longer than the spiral tube of spermatheca; median oviduct situated in middle of vagina.
Body length: 7.0-11.0 mm; width: 1.2-2.0 mm. Diagnosis. This species is similar to F. langaniformis (Wittmer, 1989), but can be distinguished from the latter by the antennomeres IV-XI each with a rounded smooth impression at base of outer margin in male; aedeagus: conjoint dorsal plate of parameres rounded at apical margin.
Distribution. China (Guangxi). Etymology. The specific name is derived from Latin basilaris (basal), referring to its antennomeres IV-XI each with a rounded impression at base of outer margin in male. Description. Male (Fig. 2B). Head black, mouthparts blackish brown, light brown at bases of mandibles and labium, antennae black, yellow at ventral sides of antennomeres I-II, prothorax orange, scultellum black, elytra dark purple, with weak metallic shine, legs black, yellow at coxae, trochanters and basal parts of femora, mesoand metasterna and abdomen black. Body densely covered with short decumbent light brown pubescence, also mixed with slightly long semierect pubescence along anterior margin of labrum and on disc of elytra.
Head subquadrate, temples evenly narrowed posteriorly, surface semilustrous, finely and densely punctate; eyes strongly protruding, head breadth across eyes distinctly wider than anterior margin of pronotum; maxillary palpomeres IV longer than wide, widest at apical one-third, arcuate and sharp at apical parts of inner margins;  (Wittmer, 1988) D-F F. gracilipes (Pic, 1927). Scale bars: 1.0 mm. antennae filiform, nearly extending to elytral apices, antennomeres II slightly longer than wide at apices, III about twice as long as II, IV-XI each with an oblong smooth scar-like bulge at basal part of outer margin, IV slightly longer than III, XI slightly shorter than X and pointed at apices.
Pronotum about 1.29 times longer than wide, widest near base, anterior margin rounded, anterior angle rounded, lateral margins slightly sinuate and diverging posteriad, posterior angle nearly rectangular, posterior margin arcuate and slightly bordered, disc distinctly convex on posterolateral parts, surface semilustrous, punctate like that on head. Elytra about 4.0 times longer than pronotum, 3.0 times longer than humeral width, lateral margins nearly parallel, disc surface semilustrous, rugulose-lacunose and finely punctate.
All tarsal claws bifid, upper claws nearly as long as lower claws. Aedeagus (Fig. 5D-F): conjoint dorsal plate of parameres moderately reduced, distinctly shorter than ventral processes, with apical margin tapered apically; ventral process of each paramere evenly narrowed apically, moderately hooked at apex.
Female. Similar to male, but eyes not so protruding; antennae uniformly black, antennomeres IV-XI without scar-like bulges; pronotum slightly wider, about 1.13 times longer than wide, lateral margins sinuate, moderately diverging posteriorly, moderately convex at posterolateral parts of disc, legs orange at pro-coxae and trochanters. Abdominal sternite VIII (Fig. 8E) slightly protuberant on both sides of posterior margin, lateroapical angles subrounded. Internal organ of reproductive system (Fig. 10B): vagina stout and abruptly narrowed and extended into a long duct above median oviduct; diverticulum and spermathecal duct arising from the end of the long duct of vagina; diverticulum slightly long, thin and spiral; spermathecal duct distinctly thicker and slightly shorter than diverticulum; spermatheca composed of a spiral tube which is distinctly longer than diverticulum, provided with a very long and thin accessory gland (surrounded with a slightly sclerotized sheath, which is hard to be stripped) which is slightly longer than the spiral tube of spermatheca; median oviduct situated in middle of vagina.
Diagnosis. This species is similar to F. gracilipes (Pic, 1927), but differs in the smaller body; aedeagus: conjoint dorsal plate of parameres moderately reduced, ventral process of each paramere evenly narrowed apically, moderately hooked at apex.
Distribution. China (Guangxi). Etymology. The specific name is derived from Latin eschara (scar), referring its antennomeres IV-XI with scar-like bulges along the outer margins in male.
Remarks. Sometimes the pronotum presents with a large inverse-triangular and a slightly small triangular black marking in middle of anterior and posterior parts of disc respectively, which are almost conjoint. Description. Male (Fig. 2C). Head yellow, mouthparts yellow, dark brown at apices of mandibles, antennae yellow, slightly darkened at antennomeres XI, pronotum, scultellum and elytra black, legs yellow, slightly darkened at tarsomeres IV-V, presternum yellow, meso-and metasterna black, abdomen black, light yellow at posterior margins of all visible abdominal sternites and apical half of IX. Body densely covered with short decumbent light yellow pubescence, also mixed with slightly long semierect pubescence along anterior margin of labrum and on disc of elytra.
Head subquadrate, temples evenly narrowed posteriad, surface semilustrous, finely and sparsely punctate; eyes slightly protruding, head breadth across eyes distinctly wider than anterior margin of pronotum; maxillary palpomeres II-IV distinctly flattened and widened, II mountain-shapely convex at outer parts of dorsal sides, III wider than long, slightly widened apically, IV longer than wide, distinctly narrowed apically, with outer margin arcuate and sharp at apical part; antennae extending to elytral mid-length, antennomeres II nearly as long as wide at apices, III-X slightly widened apically, III about 1.5 times as long as wide, IV about onethird longer than III, VI longest, XI slightly longer than X, nearly parallel-sided and pointed at apex.
Pronotum about 1.13 times longer than wide, anterior margin rounded, anterior angle rounded, lateral margins slightly diverging posteriorly, posteriad angle nearly rectangular, posterior margin arcuate and slightly bordered, disc moderately convex on postero-lateral parts, surface semilustrous, sparsely and finely punctate.
All tarsal claws bifid, upper claws nearly as long as lower claws. Aedeagus ( Fig. 6A-C): conjoint dorsal plate of parameres greatly reduced, slightly roundly emarginated in middle of apical margin; ventral process of each paramere evenly narrowed apically, largely hooked at apex.
Female. Similar to male, but maxillary palpi normal; antennae shorter, extending to basal one-third length of elytra, antennomeres II about 1.5 times as long as wide at apices, III-X parallel-sided; pronotum slightly wider, nearly as long as wide, slightly convex at postero-lateral parts of disc; elytra with lateral margins slightly diverging posteriad. Abdominal sternite VIII (Fig. 8F) slightly emarginated on both sides of posterior margin, middle part between lateral emarginations subtruncated, latero-apical angles widely rounded. Internal organ of reproductive system (Fig.  10C): vagina stout and abruptly narrowed and extended into a long duct above median oviduct; diverticulum and spermathecal duct arising from the end of the long duct of vagina; diverticulum moderately long, thin and spiral; spermathecal duct distinctly thicker and shorter than diverticulum; spermatheca composed of a spiral tube which is nearly as long as diverticulum, provided with a moderately long and thin accessory gland, which is nearly as long as the spiral tube of spermatheca; median oviduct situated in middle of vagina.
Body length: 6.5-7.5 mm; width: 1.3-1.5 mm. Diagnosis. This species is similar to F. pallidiceps (Pic, 1911), but can be easily distinguished from the latter by the characteristic maxillary palpi in the male, of which palpomeres II-IV are flattened and widened; aedeagus: conjoint dorsal plate of parameres greatly reduced, slightly emarginated in middle of apical margin.
Head subquadrate, temples evenly narrowed posteriad, surface semilustrous, finely and sparsely punctate; eyes slightly protruding, head breadth across eyes distinctly wider than anterior margin of pronotum; maxillary palpomeres II-III excavated wholly on dorsal sides, IV longer than wide, nearly parallel-sided, arcuate and sharp at apices; antennae (Fig. 7) extending to elytral mid-length, antennomeres II short, about twice wider than long, III strongly widened apically, with outer-apical angle distinctly projecting laterad, IV thickened and excavated at ventral sides, with two long and pointed projections at basal parts, dorsal projections slightly shorter than ventral ones, which are triangularly protuberant at lower margins near apices, V-X slightly widened apically, XI slightly shorter than X and pointed at apices.
Pronotum about 1.22 times longer than wide, anterior margin rounded, anterior angle distinctly rounded, lateral margins nearly parallel, posterior angle nearly rectangular, posterior margin arcuate and slightly bordered, disc moderately convex on postero-lateral parts, surface semilustrous, sparsely and finely punctate.
All tarsal claws bifid, upper claws nearly as long as lower claws. Aedeagus (Figs 6D-F): conjoint dorsal plate of parameres greatly reduced, roundly emarginated in middle of apical margin; ventral process of each paramere evenly narrowed apically at apical part, which distinctly narrower than basal part, moderately hooked at apex.
Female. Similar to male, but maxillary palpi normal; antennae orange at antennomeres I-III and bases of IV, II about twice longer than wide, III-IV normal; pronotum slightly wider, about 1.12 times longer than wide, slightly convex at postero-lateral parts of disc. Abdominal sternite VIII (Fig. 8G) triangularly emarginated on both sides and roundly emarginated in middle of posterior margin, the parts between lateral and middle emarginations subrounded at apices, latero-apical angles widely rounded. Internal organ of reproductive system (Fig. 10D): vagina stout and abruptly narrowed and extended into a long duct above median oviduct; diverticulum and spermathecal duct arising from the end of the long duct of vagina; diverticulum moderately long,  (Wittmer, 1988) B F. gracilipes (Pic, 1927) C F. sexcostata sp. n. D F. basilaris sp. n. E F. eschara sp. n. F F. latipalpa sp. n. G F. biprojicientis sp. n. Scale bars: 1.0 mm.  (Wittmer, 1988) B F. gracilipes (Pic, 1927) C F. sexcostata sp. n. Scale bars: 1.0 mm. thin and spiral; spermathecal duct distinctly thicker and shorter than diverticulum; spermatheca composed of a spiral tube which is slightly shorter than diverticulum, provided with a moderately long and thin accessory gland, which is slightly shorter than the spiral tube of spermatheca; median oviduct situated in middle of vagina.
Diagnosis. This species is similar to F. bidifformis (Wittmer, 1988), but it can be differentiated from the latter by the antennomeres IV with two projections on the basal part in the male; aedeagus: conjoint dorsal plate of parameres greatly reduced, roundly emarginated in middle of apical margin.
Distribution. China (Guangxi). Etymology. The specific name is derived from the suffix bi-(two) and projicientis (projecting), referring to its antennomere IV with two projections om the basal part in the male.