An unusual new species of Hallodapomimus Herczek, 2000 from the Eocene Baltic amber (Hemiptera, Heteroptera, Miridae, Phylinae)

Abstract Hallodapomimus antennatus sp. n. (Hemiptera: Heteroptera, Miridae, Phylinae, Hallodapini) is described from a macropterous female found in Eocene Baltic amber. The new species can be recognized readily from the other species of the genus, mainly due to its unusual second antennal segment. A key for the identification of all known fossil Hallodapini is presented.


Introduction
The present article is а continuation of a series of taxonomic papers on fossil plant bugs (Miridae) from Baltic amber (Prussian Eocene Formation). Miridae represent the largest family among true bugs (Hemiptera: Heteroptera), widespread all over the world, and with approximately 1500 genera and more than 11 000 described species, with potentially thousands more undescribed (Schuh 2002(Schuh -2013Cassis and Schuh 2012;Menard et al. 2013). Most of those included in family Miridae are frequently discovered in the Eocene Baltic amber where mirids are represented mainly by the subfamilies Cylapinae, Isometopinae, Psallopinae, and Mirinae (mainly undescribed) with fewer numbers from the remaining subfamilies (Popov and Herczek 2008). The Phylinae are quite rare among amber inclusions and all species described represent the tribe Hallodapini.
The recent Phylinae is one of the numerous subfamilies of mirids currently divided into six tribes, comprising more than 300 genera among which 50 genera belong to the tribe Hallodapini. Their representatives mainly occur temperate regions but there is also a large fauna in tropical and subtropical Asia (Schuh 1995;Schuh and Menard 2013). Moreover, many phylines have a variable myrmecomorphic habitus (McGiver and Stonedahl 1993), e.g. Hallodapini, Leucopterophorini, Auricillocorini and Pilophorini.

Material and methods
Colour photographs and drawings were made with a Nikon Eclipse E 600 microscope and by the computer program NIS Elements, Ver. 4. 10. Body length was measured from the apex of head to the apex of fore wing; body width, across the maximal width; pronotum length, along midline; pronotum width, across the broadest part at its posterior angles; hemelytron length, from the base to the apex of anterior margin; hemelytron width, at maximal width of the hemelytron. All measurements are in millimeters (mm). Diagnosis. Distinguished from the other extinct hallodapine genus Leptomimoides by a combination of the following characters: smooth, impunctate dorsal surface of body, distinctive coloration (head, pronotum and part of cuneus dark, and clavus partly black), head almost twice as broad as long, pronotum 1.2-1.3 times wider than long; pronotal calli visible.
Diagnosis. Readily recognized among the other species of Hallodapomimus by its unusual flattened and widened second antennal segment, presence of two cavities on the vertex, a small scutellum (except H. succinus), and a large mesoscutum.
Description. Female. Macropterous. Body length up to 7 mm, 2.8 times as long as wide. Dorsal surface almost smooth, impunctate. Ground colour light brown, almost yellow; mesoscutum and scutellum brown, hemelytra with one pale transverse fascia just posterior to scutellum, apical part of cuneus dark; hemelytral membrane dark, hyaline, slightly crumpled (Figs 1, 2). Head more than twice (2.3 times) as broad   2 as long; clypeus distinct and not protruding above frons; genal conus distinct; eyes large, almost globular, distinctly protruding laterally and almost touching pronotal collar; vertex with two slightly concave, polished cavities (Fig. 3), antennae inserted just above the lower margins of eyes; fovea antennalis touching the inner margin of eye; second antennal segment laterally flattened and considerably widened to apex, 2.2 times longer than 3 rd segment, 3 rd almost twice as long as 4 th one; rostrum reaching hind coxae. Pronotum tapering (narrowing) to ca. 1.75 (1.76) its length, 1.37 times wider than long; collar rather broad, flat; calli distinctly developed, quite large, occupying almost half of pronotal disc. Mesoscutum broadly exposed, scutellum quite small, only twice longer than mesoscutum length and ca. one third length of claval commissure, distinctly convex. Hemelytra wholly flattened; cuneus rather short: ca. one third length of corium and one fifth times length of hemelytron; large cell of hemelytral membrane almost rectangular, smaller cell very small, almost 4 times shorter than large cell (Figs 1, 2). All legs rather slender and covered with very short, dense, adpressed setae; hind tibia with two rows of very short spines on dorsal (10-11) and ventral (5-6) surface of its distal part, these clearly shorter than diameter of tibia (Fig. 4); first tarsal segments longest, second shorter than third (Fig. 4); claws short and slightly curved, setiform parempodia easily visible (Fig. 5).
Etymology. The species epithet (Latin "antennatus") refers to the unusual flattened and widened the second antennal segment.