Two new species and new records of the genus Spinolyprops Pic, 1917 from the Oriental Region (Coleoptera, Tenebrionidae, Lupropini) *

Abstract Two new species of the genus Spinolyprops Pic, 1917 (Tenebrionidae, subfamily Lagriinae Latreille, 1825) are described from Thailand and China (Spinolyprops cribricollis sp. n., Spinolyprops thailandicus sp. n.). The species characters within the genus are discussed, photographs of all seven Oriental species are added, a species key for the Oriental species is compiled, and a map with the distributional patterns is provided.


Introduction
The genus Spinolyprops Pic, 1917(Tenebrionidae, subfamily Lagriinae Latreille, 1825, tribe Lupropini Ardoin, 1958 was based on the type species Spinolyprops rufithorax Pic, 1917 from Zanzibar (Pic 1917). Kulzer (1954) published the first species from the Oriental Region (Sri Lanka). Later Kaszab (1965) and Schawaller (1994Schawaller ( , 1996 described additional Oriental species. The purpose of the present paper is the description of two further species from the Oriental region (Thailand, China), to discuss the species characters, to present for the first time photographs of all seven Oriental species , to provide a key for all Oriental species, and finally to add new faunistic data including a map with the distributional patterns (Fig. 1).
The separation of the genera Pseudolyprops Fairmaire, 1882, Sphingocorse Gebien, 1921, and Spinolyprops Pic, 1917 within the tribe Lupropini is still in a preliminary state and not yet based on discriminating characters. At present, the species with spinelike posterior corners of the pronotum, and with elytral colour pattern, are assigned to Spinolyprops. Congeners of all three genera live in Africa or in the Oriental/Papuan regions, thus zoogeographical aspects should also be considered during a future revision.
All species of this group are soil dwellers and are adapted also to extreme dry conditions (personal observations). All known species have fully developed wings and thus possess a high ability for dispersal. Specimens are usually collected by sifting litter and similar substrates, and are also attracted by light.

Depositories BMNH
The Diagnosis. S. cribricollis sp. n. shares with S. lateralis the rough dorsal punctation of pronotum and elytra, the shape of the pronotum with lateral parts broadly separated from disc and bent up, the frons between eyes smaller than dorsal eye diameter, and the antennomeres 8-10 longer than broad. Both can be separated mainly by the anterior margin of the pronotum with feeble (S. cribricollis sp. n.) or deep excavation (S. lateralis), and by different shape of the aedeagus (in S. lateralis the apicale is pentagonal, compare Figs 20-21). Additionally, S. lateralis is somewhat larger in the average (5.0-6.0 mm), and the elytra are slightly more rounded. S. maculatus has a similar shape of the pronotum with feebly excavated anterior margin, but the aedeagus has the apicale of the aedeagus different triangular with straight sides. S. trautneri has also a different aedeagus with broad pentagonal apicale (Fig. 24). See also the species key above.
Description. Body length 4.5-5.0 mm. Dorsal and ventral surfaces and all appendages brown without metallic shine, head and pronotum slightly darker, elytra bicoloured with darker and lighter parts (see Fig. 2); dorsal surface roughly punctate, punc-tures with long erect setae, surface between punctures shining. Head with punctation similar as on pronotum; genae distinctly swollen, clypeal suture somewhat indistinct by rough punctation, clypeus with punctation as on frons, anterior margin of clypeus straight; eyes reniform, frons between eyes smaller than dorsal eye diameter, temples impunctate; maxillary palps with large securiform terminal palpomere; shape of antennomeres see Fig. 2, antennomere 3 not elongate, terminal three antennomeres not forming club. Pronotum widest in middle, anterior and posterior margins unbordered, Figures 2-7. Dorsal view of Spinolyprops species from the Oriental Region. 2 S. cribricollis sp. n., paratype Thailand CRGT 3 S. himalayicus, non-type Thailand SMNS 4 S. himalayicus, non-type Bali SMNS 5 S. himalayicus, non-type W India SMNS 6 S. himalayicus, non-type Thailand SMNS 7 S. lateralis, non-type Sabah SMNS lateral margins unbordered but distinctly crenulate, anterior corner rounded, posterior corners acute, surface flat with irregular rough and partly confluent punctation, lateral parts broadly separated from disc and bent up; propleura with sparser and smaller punctation and shorter setation as on pronotum, prosternal process not prominent; metaventrite with punctation as on propleura. Scutellum visible, shining, without punctation. Hind wings present. Elytra elongate oval, widest in middle, lateral margin distinctly dentate in humeral region, margin completely visible from above; surface with rough punctation as on pronotum, but not confluent, punctation irregular and not arranged in rows or striae; epipleura with sparser and smaller punctation as on elytral disc, similar as on propleura. Ventrites with fine and widely separate punctation, terminal ventrite unbordered, intersegmental membranes exposed between ventrites 3-5. Legs without particular modifications, tibiae without external keels, tibial spurs short. Aedeagus with triangular apicale with acute tip (Figs 14-15). No distinct external sexual dimorphism.
Remarks. I hope not to fail in assigning the (so far disjunct) Chinese specimens from Yunnan and Guangxi to the same species. Shape and punctation of the pronotum, elytral colour and shape of aedeagus are not distinctly different from the specimens from Thailand. The type locality lies in a lowland habitat (Island Ko Chang), and the paratypes from Thailand were collected in higher altitudes (Doi Pui and Doi Inthanon). Obviously, this species has a wide ecological range.
Etymology. The name refers to the rough punctation of the pronotum.  Distribution. India (type locality Peshok/Darjeeling), Nepal, Bhutan, N Thailand, Vietnam (Schawaller 1996); Andaman Islands, Burma, Laos, Java, Bali (new records). Remarks. The specimen from Tenasserim was already published by Kaszab (1965). This specimen clearly belongs to S. maculatus and shares with the specimens from Sri Lanka the elytral colour pattern apically with an arrow-shaped dark element (Fig. 8).

Spinolyprops lateralis
It is the only specimen of S. maculatus out of Sri Lanka, so probably it was mislabelled (and is not mapped herein).
Description. Body length 4.3-5.3 mm. Dorsal and ventral surfaces and all appendages brown without metallic shine, elytra bicoloured with darker and lighter parts in different variation (Figs 10-12); dorsal surface roughly punctate, punctures with long erect setae, surface between punctures shining. Head with punctation similar as on pronotum; genae distinctly swollen, clypeal suture somewhat indistinct by rough punctation, clypeus with punctation as on frons, anterior margin of clypeus straight; eyes reniform, frons between eyes as broad as dorsal eye diameter, temples impunctate; maxillary palps with large securiform terminal palpomere; shape of antennomeres see Figs 10-12, antennomere 3 not elongate, terminal three antennomeres not forming club. Pronotum widest slightly before middle, anterior and posterior margins unbordered, lateral margins unbordered but distinctly crenulate, anterior corner rounded, posterior corners acute, surface flat with irregular rough and partly confluent punctation, lateral parts broadly separated from disc and bent up; propleura with sparser and smaller punctation and shorter setation as on pronotum, prosternal process slightly prominent; metaventrite with punctation as on propleura. Scutellum visible, shining, without punctation. Hind wings present. Elytra elongate oval, widest in middle, lateral margin distinctly dentate in humeral region, margin completely visible from above; surface with rough punctation as on pronotum, but not confluent, punctation irregular and not arranged in rows or striae; epipleura with sparser and smaller punctation as on elytral disc, similar as on propleura. Ventrites with fine and widely separate punctation, terminal ventrite unbordered, intersegmental membranes exposed between ventrites 3-5. Legs without particular modifications, tibiae without external keels, tibial spurs short. Aedeagus with broad spade-like apicale with rounded tip (Fig. 25). No distinct external sexual dimorphism.
Etymology. Named after the type locality in Thailand. subsequently processed by him with Auto-Montage (Syncroscopy) software. At last but not least I thank the referees Dr. Kiyoshi Ando (Osaka) and Dr. Ottó Merkl (Budapest) for proofreading and valuable improvements.