New species of oribatid mites of the genera Lepidozetes and Scutozetes (Acari, Oribatida, Tegoribatidae) from Nepal

Abstract Two new species of oribatid mites, Lepidozetes acutirostrum sp. n. and Scutozetes clavatosensillus sp. n., are described from Nepal. The genera Lepidozetes and Scutozetes are recorded for the first time for the Oriental region. The identification keys to the known species of these genera are provided.


Introduction
In the course of taxonomic identification of Nepalese oribatid mites 1 (Acari: Oribatida) we found two new species of the family Tegoribatidae, belonging to the genera Lepidozetes Berlese, 1910 andScutozetes Hammer, 1952. The purpose of this paper is 1 to describe and illustrate these species under the names Lepidozetes acutirostrum sp. n. and Scutozetes clavatosensillus sp. n.
Lepidozetes is a small genus that was proposed by Berlese (1910) with Lepidozetes singularis Berlese, 1910 as the type species. Currently, the genus comprises four 2 species, which distributed in the Holarctic region (Subías 2004(Subías , online version 2013. Hence, the genus Lepidozetes is recorded in the Oriental region for the first time. The main generic characters of the genus were summarized by Bayartogtokh and Aoki (1999) and Weigmann (2006).
Scutozetes is a small genus that was proposed by Hammer (1952) with Scutozetes lanceolatus Hammer, 1952 as the type species. Currently, the genus comprises two species, which distributed in the Holarctic and Neotropical regions (Subías 2004(Subías , online version 2013. Hence, the genus Scutozetes is recorded in the Oriental region for the first time. The main generic characters of the genus were presented by Hammer (1952) and summarized by Bayartogtokh and Aoki (1999).
The identification keys to the known species of Lepidozetes and Scutozetes are provided below.
All specimens were studied in lactic acid, mounted in temporary cavity slides for the duration of the study, and then stored in 70% ethanol in vials. Body measurements are presented in micrometers. The body length was measured in lateral view, from the tip of the rostrum to the posterior edge of the ventral plate. Notogastral width refers to the maximum width in dorsal aspect. Lengths of body setae were measured in lateral aspect. Formulae for leg setation are given in parentheses according to the sequence of trochanter-femur-genu-tibia-tarsus (famulus included). Formulae for leg solenidia are given in square brackets according to the sequence of genu-tibia-tarsus. Terminology used in this paper mostly follows that of Norton and Behan-Pelletier (2009 Subías (2004Subías ( , online version 2013 includes the genus Onazetes Bugrov, 1991 with the type species, Onazetes umbellatus Bugrov, 1991(see Bugrov 1991, in Lepidozetes. However, O. umbellatus has an additional pair of porose areas close to notogastral setae c (it absent in all Lepidozetes species) and five pairs of genital setae (six pairs in all Lepidozetes-species). Further research is needed to establish the taxonomical position of Onazetes.
Integument. Body color brown. Body surface distinctly microfoveolate; foveolae rounded (diameter up to 1) or elongated. Dorsal sides of lamellae with longitudinal striae.
Type deposition. The holotype and one paratype are deposited in the collection of the Senckenberg Institution, Frankfurt, Germany; two paratypes are deposited in the collection of the Siberian Zoological Museum, Novosibirsk, Russia; three paratypes are deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia.
Legs. Similar to Lepidozetes acutirostrum sp. n. Type deposition. The holotype and one paratype are deposited in the collection of the Senckenberg Institution, Frankfurt, Germany; two paratypes are deposited in the collection of the Siberian Zoological Museum, Novosibirsk, Russia; two paratypes are deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia. Etymology. The specific name "clavatosensillus" refers to the clavate sensilli. Comparison. Scutozetes clavatosensillus sp. n. can be distinguished from all known species of the genus Scutozetes by the key, which is presented below.