First record of Tyrodes Raffray (Coleoptera, Staphylinidae, Pselaphinae) in China, with description of T. jenisi sp. n. from Yunnan Province

Abstract

In this paper, we report a new species of Tyrodes from Yunnan, Southwest China. A diagnosis, a description, and illustrations of male diagnostic features are provided. This also represents the first record of the genus in China.

Material and methods
The holotype is housed in the private collection of Peter Hlaváč (pcPH), and will eventually be deposited in the National Museum of Natural History, Prague (NMPC) The collection data of the referred material are quoted verbatim. A slash (/) is used to separate lines on the same label, and a double slash (//) is used to separate different labels.
All measurements are in millimeters. The following acronyms are applied: ALlength of the abdomen along the midline; AW-maximum width of the abdomen; BLlength of the body (= HL + PL + EL + AL); EL-length of the elytra along the sutural line; EW-maximum width of the elytra; HL-length of the head from the anterior clypeal margin to the occipital constriction; HW-width of the head across eyes; PLlength of the pronotum along the midline; PW-maximum width of the pronotum.
Female. Unknown. Comparative notes. Tyrodes jenisi can be separated from all congeners, except for T. clavatus, by the distinct form of the aedeagus, as well as the consideration of distributional patterns.
Tyrodes championi is subequal in size (1.8 mm), but its aedeagal median lobe forms a distinct process at apex; T. histrio is smaller (1.5 mm), with the aedeagus being split at apex, and the endophallus with a long sclerite on the left side; T. janetscheki is greater in size (1.9 mm), its aedeagus has much more complicated structure of endophallus; T. segrex has a greater size as well (1.9-2.0 mm), and has clear different structure of aedeagal endophallus; T. setosus is subequal in size (1.8 mm), but the apical portion of aedeagal median lobe is curving to right, and has a strong apophysis on the right side. Aedeagus of Tyrodes clavatus has not been illustrated in any reference, but it is much smaller (1.4 mm), and is found in Singapore.
Comments. In his world catalog of the genera of Pselaphidae, Raffray (1908) moved Pselaphodes clavatus Raffray, 1895 to Tyrodes, followed in the later Coleopterorum Catalogus (Raffray 1911). Jeannel (1957: 32) compared the new species Tyrodes setosus Jeannel with T. clavatus when treating the pselaphines collected from Tonkin, Vietnam by Albert de Cooman. Besuchet (1970: 316), Newton & Chandler (1989: 60) and Kurbatov (1990: 145) also suggested or mentioned the placement of clavatus in Tyrodes. In the recent catalog of Tyrini (Hlaváč and Chandler 2005), this placement was probably overlooked, the species was remained in the genus Pselaphodes. According to the original description, it is clear that clavatus belongs to Tyrodes.
Distribution. Southwest China: Yunnan. Etymology. The new species is named after Ivo Jeniš (Náklo, Czech Republic), collector of the holotype.