New Neotropical species of Chimarra (Trichoptera, Philopotamidae).

Ten new Neotropical species of Chimarra are described in the subgenera Chimarra, Chimarrita, and Otarrha. New species in the subgenus Chimarra include, in the Chimarra ortiziana group: Chimarra calorisp. n. (southeastern Brazil) and Chimarra onchyrhinasp. n. (Venezuela); in the Chimarra picea group: Chimarra inchoatasp. n. (Venezuela), Chimarra nicehuhsp. n. (Venezuela), and Chimarra sunimasp. n. (Colombia); and in the Chimarra poolei group: Chimarra caucasp. n. (Colombia) and Chimarra desiraesp. n. (Bolivia). New species in the subgenus Chimarrita include, in the Chimarra simpliciforma group: Chimarra curvipenissp. n. (SE Brazil) and Chimarra latiforcepssp. n. (SE Brazil). A single new species in the subgenus Otarrha is also described: Chimarra soroasp. n. (Cuba). Males and females for all of the new species are illustrated, except for Chimarra desirae, for which female specimens were unavailable. Additionally, the female of Chimarra (Chimarrita) camella, which was previously unknown, is illustrated.


Introduction
The genus Chimarra Stephens, 1829, with about 700 species worldwide, is the second largest genus in the order Trichoptera (following Rhyacophila Pictet, 1834, with about Each pinned specimen, or lot of specimens in alcohol, examined during the study, was affixed with a barcode label (4 mil polyester, 8 × 14 mm, code 49) with a unique alphanumeric sequence preceded with the prefix UMSP. The prefix is not meant to imply ownership by the University of Minnesota Collection, but only to indicate that the specimen was databased in that collection. Specimen taxonomic and collection data are stored in Biota® (v. 2.0, Sinauer Associates, Inc.) (Colwell 2003). Specimen barcode label information is included for holotypes in the list of material examined. A detailed list of all material examined, including individual barcode numbers, is maintained at UMSP and can be provided on request.
Holotypes are deposited in the collections of the University of Minnesota, St. Paul, Minnesota, USA (UMSP), the National Museum of Natural History, Smithsonian Institution, Washington DC, USA (NMNH), the Museu de Zoologia, Universidade de São Paulo, São Paulo, Brazil (MZUSP), and the Museo de Historia Natural Noel Kempff Mercado, Santa Cruz de la Sierra, Bolivia (UASC), as designated in the species descriptions. Paratypes are deposited in the same institutions, and also in the Museo del Instituto de Zoología Agrícola, Universidad Central de Venezuela, Maracay, Venezuela (MIZA).

species descriptions
Chimarra (Chimarra) ortiziana group Blahnik (1998) recognized 12 species in this group, distributed from Mexico to Ecuador and Venezuela. The new species in this group from southeastern Brazil represents a significant range extension for the group. The group is recognized primarily on the structures of the male genitalia, and particularly the general shape of the inferior appendages, which is very consistent among the described species, relatively short, with a prominent dorsal thumb-like process, which is typically somewhat mesally curved. The group is closely related to several other species groups, including the C. amica, C. dentosa, and C. bidens groups (and possibly also the C. cornuta and C. virgencita groups), based especially on the structure of the phallotremal sclerite complex and general similarity in structure of the inferior appendages. Of these, only the C. bidens group, with 6 described species, has more than 1 or 2 species. The new species described here represent sister species to 2 of the described species in the C. ortiziana group, based both on characters from the original analysis and also overall similarity. Chimarra (Chimarra) calori Blahnik & Holzenthal, sp. n. urn:lsid:zoobank.org:act:06789F2A-D430-442F-9DEB-9AC4DF5889FB http://species-id.net/wiki/Chimarra_calori Figs 1A-F, 8 Description. This species is most closely related to Chimarra gondela Flint, 1974, based on characters of the male and female genitalia. Character synapomorphies include, es- pecially, the general shape of the lateral lobes tergum X, which are narrow overall and each of which has a raised basodorsal protuberance and basally located sensilla; and also the structure of the phallic apparatus, which includes a short, curved, sclerotized apicoventral endothecal spine in both species, in addition to the usual pair of basal endothecal spines. The lateral lobes in C. calori differ in that they are not quite as narrow or strongly sclerotized apically as in C. gondela. The most significant difference is in the dorsal thumb-like projections of the inferior appendages, which in C. calori are bluntly rounded apically and closely resemble those of many other species in the C. ortiziana group, but in C. gondela are acute apically and very narrowed and bent.
Male genitalia. Abdominal segment IX, in lateral view, with pronounced sinuous extension of anteroventral margin and with distinct apodemes from the anterodorsal margin; posteroventral process moderately elongate, subtriangular, relatively wide basally, subacute apically. Tergum X with mesal lobe membranous; lateral lobes sclerotized, each bearing short conical lateral projection basally, with 2 sensilla, lobes relatively narrow, as viewed laterally, sclerotized along dorsal margins; apices of paired lobes diverging, as viewed dorsally, forming small, angular projections. Preanal appendage short, knob-like. Inferior appendage, in lateral view, with subquadrate base, dorsally with flattened, thumb-like extension, flexed caudally as viewed laterally, and mesally as viewed caudally, apex of thumb-like projection rounded, mesal surface with sclerotized ridge. Phallotheca with apicoventral extension; endotheca with minute spines and sclerous region formed into short curved spine, endothecal spines 2, subequal in length, relatively short. Phallotremal sclerite complex composed of elongate rod and ring structure, with associated pair of wishbone-like sclerites from anteroventral margin.
Holotype, male ( Description. This species is very similar to C. platyrhina Flint, 1981, especially in the general shape of the lateral lobes of tergum X, as viewed dorsally. In both species, the lateral lobes of tergum X each have a broadly rounded apicolateral projection. The difference in shape of this projection, however, is distinctive and diagnostic, that of C. platyrhina being broad and uniformly rounded, and that of C. onchyrhina being somewhat angular laterally. Both species also have a linear cluster of small spines located near the ventral apex of the endotheca, also present in some other species of the C. ortiziana group. The latter character is, however, absent in C. ortiziana, which otherwise resembles C. onchyrhina in the general shape of the apex of the lateral lobes of tergum X. An additional difference separating them is that Chimarra ortiziana has the lateral sensilla of tergum X placed on a concavely developed lateral projection. The lateral projections of C. onchyrhina and C. platyrhina are simple in structure. The female genitalia of C. onchyrhina also generally resemble C. platyrhina, especially in the shape of apical sclerites of the vaginal apparatus. However the vaginal apparatus has distinctive lateral projections, not present in C. platyrhina. The overall similarities of these species, both in male and female genitalia, are indicative of their sister relationship. Adult. Forewing length (male) 3.9-5.0 mm, (female) 4.8-5.5 mm. Cuticle of head and thorax dark brown, setae of anteromesal and frontal setal warts whitish, setae of other setal warts dark brown, grizzled (grayish in part or intermixed), femora brown, otherwise color nearly uniformly brownish-black (fuscous), including appendages and antennae. Postocular parietal sclerite very short (not extended behind eye). Second segment of maxillary palp much shorter than segment 3 (about 2/3 length). Male protarsal claws enlarged, asymmetrical in size and shape, outer claw much larger, twisted, nearly linear apically.
Male genitalia. Abdominal segment IX, in lateral view, with pronounced sinuous extension of anteroventral margin and with distinct apodemes from anterodorsal margin; posteroventral process moderately elongate, subtriangular, relatively broad basally, subacute apically. Tergum X membranous mesally, with 2 sclerotized lateral lobes, each bearing short rounded projection laterally in basal portion, with 2 sensilla on tiny projections; terminus of lateral lobe inflated, broadly rounded, but with distinct, obtuse angle on lateral margin. Preanal appendage short, knob-like. Inferior appendage, in lateral view, with rounded to subquadrate base, subacute apically; dorsally with flattened, thumb-like extension, flexed caudally as viewed laterally, and mesally as viewed caudally; apex of thumb-like projection rounded. Phallotheca with apicoventral margin projecting; endotheca forming collar of minute spines in apical 1/2, slightly ballooned laterally, apicoventrally with sclerotized region possessing array of short spines, endothecal spines 2, very short, subequal in length. Phallotremal sclerite complex composed of elongate rod and ring structure with associated pair of wishbone-like sclerites from anteroventral margin.
Etymology. This species is named C. onchyrhina from the Greek words onkos, meaning a hook or angle, and rhinos, a nose, and referring to the slightly angled lateral process on tergum X of this species, which distinguishes it from the more uniformly rounded process in C. platyrhina. Chimarra (Chimarra) picea group Blahnik (1998) recognized 11 species in this group, distributed from lower Central America to Ecuador and Venezuela. The species all have very similar, though somewhat differently developed, inferior appendages (very short with elongate lateral setae and small, sclerotized, strongly mesally curved dorsomesal processes, not generally evident in lateral view), and are especially diagnosed by differences in the shape of the lateral lobes of tergum X and their sensilla-bearing protuberances. The 3 new species recognized here are similar in general characters to other described species and bring the total number of species in the group to 14. Description. This species is most closely related to C. creagra Flint, 1981 andC. paracreagra Blahnik, 1998, based on the similarly developed, sclerotized, dorsolateral margins of the mesal lobe of tergum X. In C. inchoata, however, this structure is not as strongly sclerotized, and while somewhat curved apically, does not form the strongly recurved and acutely hooked projections found in the two described species. Additionally, the sclerotized basodorsal, mesally curved, thumb-like processes of the inferior appendage are distinctly evident even in lateral view, and the lateral lobes of tergum X are shorter and have much more broadly rounded lateral sensilla-bearing processes.
Female genitalia. Sternum IX with ventral lobes truncate apically, laterally with small rounded clasper receptacles. Vaginal apparatus relatively membranous apically, with narrow, transverse preapical sclerite dorsally; ventrally with elongate, narrow, membranous lobe extending from apex to about ½ length, lobe with small sclerite at apex; vaginal apparatus laterally with pair of small sclerites at about midlength; anteriorly with deflexed, cup-like sclerite.
Holotype, male ( Etymology. This species is named C. inchoata from the Latin word incohatus (or inchoatus), meaning only begun, incipient, or incomplete, and referring to the development of the lateral lobes of tergum X in this species, which are only suggestively sclerotized as compared to the strongly sclerotized, hooked processes in C. creagra and C. paracreagra. Description. This is perhaps one of the most distinctive of the species in the C. picea group described to date and unlikely to be confused with any other described species. Especially distinctive is the shape of the lateral lobes of tergum X, which are short, but broad apically, as viewed laterally, each with a small, obliquely oriented, lateral sensilla-bearing processes. Like the previous species, it has the lateral margins of the mesal lobe of tergum X somewhat sclerotized, though not projecting apically. The thumb-like dorsolateral projections of the inferior appendages are especially blunt and strongly curled, not clearly evident in either lateral or dorsal views. Additionally the apicoventral margin of the appendage is recurved, and thus evident as a ridge on the mesal surface, as viewed caudally.
Male genitalia. Abdominal segment IX, in lateral view, with pronounced linear extension of anteroventral margin and with distinct, enlarged apodemes from anterodorsal margin; posteroventral process short, broad basally, rounded apically. Tergum X with mesal lobe membranous and weakly incised mesally, lightly sclerotized laterally; lateral lobes sclerotized, relatively short, very broadly truncate apically as viewed laterally, each bearing short, lateral, obliquely flattened projection with 2 sensilla. Preanal appendage short, knob-like. Inferior appendage very short, apicoventral margin weakly projecting and slightly recurved, apically with elongate marginal setae, dorsally with very short, bluntly rounded, mesally curved process, visible in caudal view, but not fully evident in either lateral or dorsal views. Phallotheca with acute apicoventral projection; endotheca membranous, elongate tubular, with numerous minute spines in apical ½, preapically on ventral margin with very short sclerotized spines, endothecal spines 2, basal one elongate, apical one moderately elongate. Phallotremal sclerite complex composed of rod and ring structure, rod relatively elongate, ring reclinate, complex laterally with pair of short sclerites.
Etymology. The name of this species should be considered an arbitrary combination of letters. It resulted from an observation made by the first author when seeing the genitalia of this distinctive species for the first time, "Nice, huh?" Chimarra (Chimarra) sunima Blahnik & Holzenthal, sp. n. urn:lsid:zoobank.org:act:C41FE4CB-F191-4B84-8C4A-6F83A03894A2 http://species-id.net/wiki/Chimarra_sunima Figs 5A-G, 12 Description. This species most closely resembles C. onima Flint, 1991, especially in the structure of the lateral sensilla bearing processes of tergum X. In both species the processes have their posterior margins detached and project "ear-like", as viewed dorsally, although those of C. sunima are perhaps not quite so elongate. Differences are found in the basal part of the lateral lobes of tergum X, which in C. sunima are more dorsally projecting in lateral view, and in the structure of the inferior appendages. The latter are very distinctive in C. onima, with the dorsal margin projecting, so that the dorsomesal processes are clearly apical, as viewed laterally, and the posterior margin concave, whereas in C. sunima, the inferior appendages are more or less typical of a number of other species in the group (i.e., C. tapanti Blahnik, 1998or C. limon Blahnik, 1998 in that the dorsomesal processes are slightly recessed compared to the elongate, fringing posterolateral setae of the appendage. Adult. Forewing length (male) 3.8-4.5 mm, (female) 4.3-5.2 mm. Cuticle of head and thorax dark brown, setae of anteromesal and frontal setal warts light brown or whitish, setae of other setal warts dark brown, grizzled (grayish in part or intermixed), otherwise color nearly uniformly brownish-black (fuscous), including appendages and antennae. Postocular parietal sclerite elongate (extended behind eye). Second segment of maxillary palp much shorter than segment 3 (less than 2/3 length). Male protarsal claws enlarged, asymmetrical in size and shape, outer claw larger, twisted, nearly linear apically.
Female genitalia. Sternum IX with ventral lobes tapering, subacute, extreme apex nearly membranous, laterally with large rounded clasper receptacles. Vaginal apparatus relatively membranous apically, with very narrow, elongate mesal lobe extending from apex to more than ½ length, lobe very lightly sclerotized; laterally with conspicuous, projecting preapical lobes on either side, these largely membranous, but with conspicuous, rounded, premarginal ventral sclerites; anterolateral margins of vaginal apparatus lightly sclerotized; anteriorly with deflexed, cup-like sclerite.
Holotype, male ( Etymology. The species name should be considered an arbitrary combination of letters, suggested by its similarity in form to that of other species in this species group (i.e., C. emima Ross, 1959;C. onima Flint, 1991;and C. jemima Blahnik & Holzenthal, 1992).
Chimarra (Chimarra) poolei group Blahnik (1998) recognized 10 species in this group, broadly distributed from Costa Rica in Central America through the Brazilian subregion of South America, including southeastern Brazil. Species are readily recognized by the spine-like projections from the posterodorsal margin of segment IX (as distinguished from the spine-like modifications of tergum X in several species of the C. picea group). All of the species have very short inferior appendages, either without or with very small dorsomesal projections, not generally evident in lateral view. The group is closely related to the C. bidentata group. The 2 new species recognized here increase the species group to 12 species. The new species and new records of this group from Bolivia also represent a range extension for the species group. Description. This new species is most closely related to C. zamora Blahnik, 1998. Notable synapomorphies are found in the overall structure of the lateral lobes of tergum X, which in both species have a somewhat concavely developed apical protuberance and also projecting, sensilla-bearing lateral projections. In C. cauca, the sensilla-bearing projections are larger and more broadly rounded. Additionally, the spine-like processes of segment IX are much more prominent in C. cauca. Female genitalia for the 2 species, while having some similarities, are also different, confirming that these are indeed distinct species.
Male genitalia. Abdominal segment IX, in lateral view, with pronounced linear extension of anteroventral margin and with distinct small apodemes from anterodorsal margin; posterodorsal margin with prominent pair of acute, spine-like projections, ex- tending about 2/3 length of tergum X; posteroventral process prominent, subtriangular, broad basally, acute apically, length slightly greater than width at base. Tergum X with mesal lobe membranous; lateral lobes sclerotized, moderately elongate, each with distinct, preapical, lateral projection, more or less concave on anterior margin (acute as viewed dorsally), lobe laterally with rounded projection bearing 2 sensilla. Preanal appendage short, knob-like. Inferior appendage, in lateral view, very short and strongly cupped basally; apical margin truncate, dorsally with very short, mesally curved process, process very angularly bent. Phallotheca with acute apicoventral projection; endotheca membranous, apparently with numerous minute spines and cluster of short sclerotized spines, endothecal spines 2, basal one moderately elongate, apical one short. Phallotremal sclerite complex composed of rod and ring structure, rod relatively short and ring with small, but distinct apicodorsal extension; complex also with small lateral sclerites.
Holotype, male ( Etymology. This species is named C. cauca, used as noun in apposition, for the Department (regional area) in Colombia where the type specimens were collected. Chimarra (Chimarra) desirae Blahnik & Holzenthal, sp. n. urn:lsid:zoobank.org:act:AD05D344-F0D0-40FA-B7A8-76EA9E98E4C6 http://species-id.net/wiki/Chimarra_desirae Fig. 7A-E Description. This is a distinctive species, most closely related to Chimarra adamsae Blahnik, 1998. The most distinctive diagnostic feature, as in most species of the C. poolei group, is found in the structure of the lateral lobes of tergum X. In C. desirae these are distinctly sclerotized and have an upright basal projection, and a dorsoventrally flattened lateral crease, so that the apices of the lobes, in dorsal view, appear broadly rounded. Chimarra adamsae has these same general features, but the upright processes are subquadrate and more apical, and the lateral creases are also more apical, resulting in the apices being narrow, as viewed either laterally or dorsally. Additionally, the sensilla of the lateral lobes in C. desirae are located along the lateral crease, whereas those of C. adamsae are basal to the crease.
Adult. Forewing length (male) 4.2-5.0 mm. Cuticle of head and thorax very dark, nearly black, setae of anteromesal and frontal setal warts whitish, setae of other setal warts brownish-black, grizzled (grayish in part or intermixed), otherwise color nearly uniformly brownish-black (fuscous), including appendages and antennae. Postocular parietal sclerite short (not greatly extended behind eye). Second segment of maxillary palp much shorter than segment 3 (about 2/3 length). Male protarsal claws enlarged, asymmetrical in size and shape, outer claw much larger, twisted, nearly linear apically.
Male genitalia. Abdominal segment IX, in lateral view, with anteroventral margin sinuously extended from dorsal margin; anterodorsal margin with short apodemes; posterodorsal margin with pair of acute, spine-like projections, extending about 1/3 length of tergum X; posteroventral process subtriangular, relatively broad basally, subacute apically. Tergum X with very short membranous mesal lobe and 2 sclerotized lateral lobes, each bearing large, upright, broadly rounded basal lobe; lateral lobe also with strongly dorsoventrally flattened lateral crease, extending from near base to apex, base of crease with 2 slightly raised sensilla; terminus of lateral lobe narrowed and slightly declivous as viewed laterally, broadly rounded as viewed dorsally. Preanal appendage short, knob-like. Inferior appendage, in lateral view, very short and strongly cupped basally; dorsomesally with small, barely suggested, angular process. Phallotheca with distinct, elongate, apicoventral extension; ventral surface of phallotheca with lightly sclerotized projection; dorsal surface of endotheca with preapical tract of minute spines, basal ones slightly longer, endothecal spines 2, basal one moderately elongate, apical one short and curved. Phallotremal sclerite complex composed of rod and ring structure, rod relatively short and ring with small, but distinct, apicodorsal extension; rod preapically with 2 small, curved, symmetric sclerites.
Etymology. We take great pleasure in naming this species C. desirae for Dr. Desiree Robertson-Thompson, who collected the type specimens, in acknowledgment of her contributions to the study of Neotropical caddisflies. Blahnik (1997) recognized 10 species in this species group, which represents the largest and most broadly distributed of the 3 species groups recognized for the subgenus. Described species are mostly known from very few specimens. Species have been recorded from the Guyana Highlands, Amazon Basin, and southeastern Brazil, thus spanning a very broad area. The apparent disjunct distribution of the group probably reflects a very inadequate knowledge about the actual distribution of known species and also the species diversity of the group. Currently 4 species are known from southeastern Brazil. The 2 new species described here are closely related to 2 of the described species. Species in the group are easily recognized by structure of the phallic apparatus in the males, which has a single and typically elongate spine that emerges from the dorsum of the phallotheca. Females in the subgenus are also distinctive because of their elongate genitalia.

Chimarra (Chimarrita) simpliciforma group
Chimarra (Chimarrita) curvipenis Blahnik & Holzenthal, sp. n. urn:lsid:zoobank.org:act:67D04EC4-7571-4CA9-9513-93D906CD4C20 http://species-id.net/wiki/Chimarra_curvipenis Fig. 14A-F, 16A-B Description. Chimarra curvipenis, sp. n. is similar to C. kontilos Blahnik, 1997 and represents a closely related sister species. Similarities are found in the general structure of the inferior appendages and tergum X of the male. However, there are a number of differences. Tergum X has a small lateral, sensilla-bearing projection at past midlength and the inferior appendages are more elongate and have more acute apices. Additionally, the phallotheca is quite different, distinctly curved, rather than elongate, tubular, and the apex of the dorsal phallic spine lacks the distinctive whip-like extension found in C. kontilos. In the key by Blahnik (1997), C. curvipenis would come out with C. tortuosa Blahnik, from which it differs significantly in the shape of the inferior appendages (longer and more strongly incurved apically), and structure of the phallic apparatus (phallotheca more strongly curved and phallic spine less sinuously curved).
Adult. Forewing length (male 4.8 mm, (female) 4.8 mm. Overall color nearly uniformly light brown, palps slightly darker; thorax ventrally, meso-and metacoxae golden brown. Venational branching of forewing typical for Chimarra; Rs straight s, r and r-m of forewing nearly linearly arranged and unpigmented, as is m-cu and apex of Cu 2 ; 2A of forewing with apparent apical "fork", that to 1A elongate and broadly rounded, that to 2A very short (appearing as cross-vein). Rs of hind wing 4-branched, M 3-branched. Head short (postocular parietal sclerite short). Maxillary palps relatively short, segment 2 longer than 3. Male with protarsi unmodified.
Male genitalia. Segment IX relatively wide; as viewed laterally, with anterior margin concave, posterior margin angularly projecting at level of inferior appendages; anteroventral margin noticeably expanded, apex of expansion acutely rounded (as viewed dorsally or ventrally); ventral process narrow, elongate, acute, somewhat curved. Tergum X moderately elongate, fused to segment IX, apex with deep, U-shaped mesal excision, extending about 1/3 length of tergum, forming narrow, paired lobes apically (as viewed dorsally); tergum laterally with short, rounded projections at just past midlength (basal to apical lobes); apical lobes and lateral projections with numerous sensilla. Preanal appendage very small, rounded, fused near base of tergum X. Inferior appendage relatively elongate, nearly linear, except apex deflexed, narrowed, and me- Figure 14. Chimarra (Chimarrita) curvipenis sp. n. Male genitalia: A lateral B segment IX and tergum X, dorsal C inferior appendage, ventral D inferior appendage, dorsal e phallic apparatus, lateral F phallic apparatus, dorsal. sally curved, apex forming acute projection. Phallotheca with somewhat bulbous base, otherwise elongate, narrow, ventral margin projecting and strongly curved; phallic spine single, stout, curved, very elongate (subequal in length to phallotheca), emerging dorsally near base of phallotheca and with slight sinuous twist. Phallotremal sclerite complex (if present) indistinct.
Female genitalia-Sternum VII with ventral process; process projecting, subacute, located near middle of segment, as viewed laterally. Segment VIII synsclerous, short dorsally, anterolateral margin nearly straight, indented and narrowed dorsally, rounded ventrally; segment sclerously connected ventrally to sternum IX; anteroventral margin of segment, as viewed ventrally, with deep, mesal emargination extending almost entire length of segment, emargination strongly narrowed posteriorly, bordered laterally by distinct U-shaped sclerotization. Sternum IX elongate, with paired, angular projections, projections continuous posteriorly with elongate, narrow ventral sclerites; sternum membranous ventrally between paired sclerites, and membranous also laterally from acute basal projection to apex. Tergum IX elongate, narrow, slightly curved, moderately setose, anteroventrally with short apodemes. Segment X with elongate basal portion, furrowed dorsally, with mesal tract of setae in furrow; apically with pair of small, rounded, setose lobes, each with short apical cercus. Vaginal apparatus largely membranous, with indistinct sclerites, anterior one forming narrow ring.
Etymology. This species is named C. curvipenis for its curved or bent phallotheca, a character that helps to distinguish it from C. kontilos. Description. Chimarra latiforceps is very similar to C. majuscula Blahnik, 1997, par-ticularly in the general shape of the inferior appendages and tergum X of the male, and would key out to that species in the key by Blahnik (1997). However, it does differ in details of both structures. It is also somewhat darker in overall color. The most diagnostic difference is in the shape of the inferior appendages, which are broader overall and have more truncate apices. Tergum X differs in that the lateral lobes formed by the mesal invagination are narrow, elongate, rather than inflated apically.
Female genitalia. Sternum VII with ventral process; process large, projecting, subacute, emerging near anterior margin of segment, as viewed laterally. Segment VIII synsclerous, short dorsally, anterolateral margin broadly rounded, indented and narrowed dorsally, rounded ventrally; segment sclerously connected ventrally to sternum IX; anteroventral margin of segment, as viewed ventrally, with short, narrow mesal emargination, margins of emargination distinctly sclerotized, sclerotization extending posteriorly as pair of distinct ridges. Sternum IX elongate, with paired, angular projections, projections continuous posteriorly with elongate, narrow ventral sclerites; sternum membranous ventrally between paired sclerites, and membranous also laterally from acute basal projection to apex. Tergum IX elongate, narrow, slightly curved, moderately setose, anteroventrally with short apodemes. Segment X with elongate basal portion, furrowed dorsally, with mesal tract of setae in furrow; apically with pair of small, rounded, setose lobes, each with short apical cercus. Vaginal apparatus largely membranous, with indistinct sclerites, anterior one forming narrow ring.
Holotype, male ( Etymology. This species is named C. latiforceps, from the Latin words latus, wide, and forceps, a pincers or tong, in reference to the inferior appendages, which are shaped something like a pair of broad tipped forceps. Chimarra (Chimarrita) camella Blahnik http://species-id.net/wiki/Chimarra_camella Fig. 18A-B Description. The female of this species was not previously illustrated. It is included here, since it is the only species of the subgenus for which the female is not illustrated. The female can be distinguished from other species by the nearly linear anterior margin of segment VIII and, especially, by the distinctive, subquadrate invagination of the posteroventral margin of the same segment. The mesal margin of the invagination has a lightly sclerotized, tab-like projection.
Female genitalia. Sternum VII with ventral process; process projecting, subacute, located near middle of segment, as viewed laterally. Segment VIII synsclerous, short dorsally, anterolateral margin nearly straight, indented and narrowed dorsally and ventrally; segment sclerously connected ventrally to sternum IX; anteroventral margin of segment, as viewed ventrally, with subquadrate mesal emargination, extending about ½ length of segment, margins of emargination very distinctly sclerotized, sclerotization extending somewhat posteriorly, mesal margin of emargination with lightly sclerotized tab-like projection . Sternum IX elongate, with paired, angular projections, projections continuous posteriorly with elongate, narrow ventral sclerites; sternum membranous ventrally between paired sclerites, and membranous also laterally from acute basal projection to apex. Tergum IX elongate, narrow, distinctly curved, moderately setose, anteroventrally with apodemes only suggestively developed. Segment X with elongate basal portion, furrowed dorsally, with mesal tract of setae in furrow; apically with pair of small, rounded, setose lobes, each with short apical cercus.

Chimarra (Otarrha)
The subgenus Otarrha was established by Blahnik (2002) to include 31 species, broadly distributed in northern South America and in the Greater and Lesser Antilles, with individual species also found in lower Central America and southeastern Brazil. A clade of 10 described species occurs in the Greater Antilles. All of these were treated in the work by Blahnik (2002), except for C. koki Botosaneanu (1996), described from the Dominican Republic, and a subsequently described subspecies of C. spinulifera Flint, 1968, C. spinulifera baoruco Flint & Sykora, 2004, also from the Dominican Republic.. The new species described here increases the number of species in the subgenus known from Cuba to 5, and the number of species known from the Greater Antilles to 11. Chimarra (Otarrha) soroa Blahnik & Holzenthal, sp. n. urn:lsid:zoobank.org:act:7D1E3A56-5233-43BD-9B2A-D9C49211CC40 http://species-id.net/wiki/Chimarra_soroa Fig. 19A-F, 20A-C Description. This new species closely resembles C. garciai Botosaneanu, 1980, also known from Cuba, and represents a probable sister species. It is also similar in general features to C. jamaicensis Flint, 1968 andC. machaerophora Flint, 1968, both described from Jamaica. All of these species have a tergum X that is divided mesally, with very elongate lateral lobes, and similarly shaped inferior appendages. Like C. garciai, C. soroa, sp. n. has a tergum VIII with a pair of short spine-like projections, but without the mesal projection of C. machaerophora. In the key by Blahnik (2002), C. soroa would key out to the couplet including C. garciai and C. jamaicensis. It differs from C. garciai in lacking a pair of sclerotized spines at the base of the lobes of tergum X, and from either species in having a distinctly projecting, rounded, sensilla-bearing process on the lateral margin of each of the lobes of tergum X. Additionally, the inferior appendages appear more truncate in lateral view than in either of those species.
Female genitalia. Segment VIII short; anterior margin slightly projecting midlaterally at position of apodemes, apodemes short, angular, undeveloped; posterior margin nearly straight, with deep cleft near ventral margin, extending almost to anterior margin, ventral part of segment (below cleft) forming projecting process, narrow as viewed laterally, rounded apically as viewed ventrally; segment with elongate setae on posterolateral margin and shorter setae at apex of ventral projection. Tergum IX very short, with moderately elongate apodemes from anteroventral margin; posteromesally with projecting, short, rounded, setose projection. Sternum IX membranous. Tergum X bilobed, each lobe divided into prominent, rounded, lightly sclerotized basal region and less sclerotized apical region with cercus at apex. Vaginal apparatus moderately elongate, dorsalaterally with elongate paired sclerites, each divided for nearly entire length to form apparent pair of rod-like sclerites; apically with additional pair of short needle-like sclerites on either side of elongate sclerites; vaginal apparatus anteriorly with indistinct ring-like sclerite.
Etymology. This species is named Chimarra soroa, used as a noun in apposition, after the resort in Cuba near which the type specimen was collected.