A survey of East Palaearctic Lycosidae (Araneae). 7. A new species of Acantholycosa Dahl, 1908 from the Russian Far East

Abstract Acantholycosa azarkinae sp. n. is described from the Maritime Province of Russia on the basis of both sexes. Acantholycosa norvegica (Thorell, 1872) is reported from the Maritime Province for the first time. A key and illustrations to all six species that occur in Far East Asia are provided.


Introduction
Acantholycosa Dahl, 1908 is a relatively small Holarctic genus with 26 species and one subspecies (Platnick 2011). It is a well delimited genus that can easily be recognized by having 4-6 pairs of ventral tibial spines on legs I and II, and a modifi ed palea. Th e genus was recently revised by Marusik et al. (2004). Acantholycosa has a rather unusual geographical distribution, with two centres of species richness, including an extraordinary degree of endemism in the northern Palaearctic (Marusik et al. 2004). Twenty-one species of this genus are known from the Altai-Sayan mountainous region, of which 17 are local endemics. Four species of Acantholycosa are known from the Maritime Province (A. aborigenica Zyuzin & Marusik, 1998;A. lignaria (Clerck, 1757), A. oligerae Marusik et al., 2004 andA. sundukovi Marusik et al., 2004), two of which are local endemics. No other areas in the Holarctic region have more than two species.
While studying wolf spiders in the Maritime Province of Russia we found two additional species, one of which was new to science. Th e main aim of this paper is to provide a description of the new species. We also review and provide a key to all species known to occur in the whole of the Russian Far East.

Material and methods
Specimens were photographed using an Olympus Camedia E-520 camera attached to an Olympus SZX16 stereomicroscope in the Zoological Museum, University of Turku. Th e images were montaged using "CombineZP" image stacking software. Photographs were taken in dishes of diff erent size with paraffi n at the bottom. Diff erent sized holes were made in the bottom to keep the specimens in the required position. Figures 6-7 , 13-21, 29-40 are reproduced from Marusik et al. (2004) with permission of the coauthors G.N. Azarkina and S. Koponen, in addition to N. Smirnov, the chief editor of Arthropoda Selecta.
Th e standard of description follows that in Marusik et al. (2004). All measurements are in mm.
Th e material treated herein will be deposited in the Zoological Museum of the Moscow State University (ZMMU) and in Gornotayozhnaya Station (GTS).
Etymology. Th e specifi c name is a matronym in honor of our friend and colleague Galina N. Azarkina.
Diagnosis. Th e new species can be easily distinguished from other congeners occurring in the Far East by the shape of the palp, which has a broad embolus tip (Figs 8, 10) (not broad in the other species) and by the shape of the epigyne, which has a broad apical pocket and well developed hoods (Figs 26-28).
Comments. A. azarkinae sp. n. is morphologically close to two other endemic species that occur in the Maritime Province: A. oligerae and A. sundukovi. Th e three species have similar male palps although they diff er from one another by the shape of the tegular apophysis and the embolus.
Epigyne as in Figs 26-28. Apical pocket wide with two distinct hoods, septum distinct, septum with trapezoidal base; spermathecae long, with blind outgrowth in basal third.

Acantholycosa oligerae Marusik, Azarkina & Koponen, 2004
Comments. Th is species was recently described from material found at a single locality in the Lazo Reserve, the Russian Far East.
Comments. Th is species is known from the holotype male only. So far, A. sundukovi is known from a single locality in the Lazo Reserve (Kordon Amerika), the Russian Far East. Tibia-metatarsus I and II with long hairs (Fig. 5); tip of embolus broad and twisted (Figs 1, 3-4, 8-12

Conclusions
Th e number of Acantholycosa species in the Maritime Province of Russia is fewer than that of the Altai-Sayan region only, with 6 and 21 species respectively. Th e same is true for the number of endemic species (3 and 17 respectively). Presently, only the southern region of the Maritime Province has been relatively well studied. Th e huge territories of Sikhote-Alin remain uninvestigated. Given the high level of endemism among petrophilous species of spiders it is reasonable to expect the occurrence of additional new species in the province, especially on isolated screes on mountain tops.