Pselaphus bulbifer Reichenbach, 1816.

(N = 11, 6♂♂, 5♀♀) . Holotype male. “Korea: Jeonnam Prov. Dangsan-ri, Gyegok-myeon, Haenam-gun, 18 May 2019, 34°40'53.0"N, 126°38'56.3"E, 211 m, sifting leaf litter & deadwood debris, J.-S. Park, M.-H. Song” (NIBR). Paratypes. 2♂♂, 2♀♀ (CBNUIC, 1♂, 1♀ slide mounted, 1♂, 1♀ dried). “Korea: Jeonbuk Prov. Sinsi island. Sinsido-gil, Okdo-myeon, Gunsan-si, 4 Jul 2022, 35°49'12.2"N, 126°27'35.1"E, 36 m, sifting leaf & soil litter, M.-H. Song, U.-J. Byeon, J.-W. Kang, T.-Y. Jang”. 2♂♂ (CBNUIC, dried) “Korea: Jeonbuk Prov. Seonyu island. Seonyubuk-gil, Okdo-myeon, Gunsan-si, 16 Jun 2021, 35°48'36.5"N, 126°24'57.4"E, 25 m, sifting leaf, soil litter & fungi, J.-W. Kang, J.-I. Shin”. 1♂, 3♀♀ (CBNUIC, DNA grade). “Korea: Jeonbuk Prov. Seonyu island. 5-1, Seonyunam-gil, Okdo-myeon, Gunsan-si, 4 Jul 2022, 35°48'24.7"N, 126°24'40.3"E, 9 m, sifting leaf & soil litter, M.-H. Song, U.-J. Byeon, J.-W. Kang, T.-Y. Jang”.

Antennal scapes robust, with bowl-like glandular nodule on inner margin (Figs 1E, F, 2A, arrows), 2.45 times as long as pedicels; endophallus of male genitalia with three bifid struts, joined at base (Fig. 1I).

Figure 1. 

Bryaxis grandinodus Choi, Park, Lee & Park, sp. nov., male (A, B, E–I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A–D); 0.1 mm (E, I); 0.5 mm (F–H).

Figure 2. 

Diagnostic characters of Bryaxis grandinodus sp. nov. (A, C) and B. koltzei (Reitter) (B, D). A, B antennal scapes and pedicels C, D protibiae. Scale bars: 0.1 mm.

Body reddish brown, antennae, maxillary palpi, and tarsi slightly lighter, length 1.2–1.32 mm, maximum width 0.51–0.58 mm (Fig. 1A–D). Setae on body yellowish, long. Head 0.92 times as long as wide (Fig. 1E). Frons with U-shaped impression between antennal tubercles; frontal foveae absent; frontal rostrum distinct anteriorly. Vertex slightly convex; longitudinal carina weak; vertexal foveae large. Eyes as long as tempora, with 23–26 facets. Maxillary palpi moderately developed; palpomeres II–III with tubercles; palpomere IV 0.23 mm long and about 3.15 times as long as wide, subcylindrical pseudosegment at apex. Antennae about 0.54 mm long; pedicels subglobose with long setae, 0.89 times as long as wide; antennomere III 1.12 times as long as wide; IV–VIII subequal in length; IX–X transverse, IX 0.64 times as long as wide and X 0.63 times as long as wide; XI largest, pointed at apex, 1.67 times as long as wide (Fig. 1F). Pronotum 0.84 times as long as wide and widest at basal 2/3, lateral antebasal foveae connected by antebasal sulcus. Elytra slightly convex, 0.89 times as long as wide and widest at basal 1/4, each elytron with two basal foveae and subhumeral fovea. Legs slender; protibiae without spine (Figs 1G, 2C); metatibiae with spine at apex (Fig. 1H, arrow). Aedeagus large, 0.37 mm long and 1.69 times as long as wide; penis bulbous and dorsal diaphragm ovoid; parameres short and symmetrical, apices almost encountered, one robust seta and three fine setae on each apex; endophallus divided into three large struts, left dorsal strut branched at basal 1/3 and bifid at apex, right dorsal strut weakly branched basally and bifid dorso-ventrally at apex, ventral strut robust and bifid at apex (Fig. 1I).

Sexual dimorphism. Female eyes shorter than tempora, composed of 11 facets; antennal scapes subcylindrical, without modification; metatibial spines absent.

Adults of this species are very similar to Bryaxis koltzei (Reitter, 1887) in the general body characters, but can be distinguished by the shape of the antennal scapes and its glandular nodule (Fig. 2A, arrow) and the spineless protibiae in the male (Fig. 2C).

The localities of B. grandinodus sp. nov. probably overlap with those of B. koltzei (Reitter) given that the latter are distributed across the entire country (Fig. 11).

The specific epithet is a combination of the Latin words grandis (“large”, masculine) and nodus (“knob”, masculine) and refers to the shape of the glandular nodules on the male antennal scapes.

The holotype was collected by sifting leaf litter in mixed forest. Paratypes were collected by sifting leaf litter and soil.

Korea (Haenam-gun, Jeollanam-do; Gunsan-si, Jeollabuk-do).

(N = 7, 4♂♂, 3♀♀) . Holotype male. “Korea: Gyeongbuk Prov. Onjeong-myeon, Uljin-gun, 8 Jun 2019, 36°43'23.0"N, 129°20'16.0"E, 180 m, sifting leaf litter near stream, J.-S. Park” (NIBR). Paratypes. 2♂♂ (CBNUIC, 1♂ slide mounted, 1♂ dried). “Korea: Gangwon Prov. Gujeol-ri, Yeoryang-myeon, Jeongseon-gun, 24 Apr 2020, 37°31'08.7"N, 128°46'42.8"E, 591 m, sifting leaf & soil litter, U.-J. Byeon, T.-Y. Jang”. 1♂, 3♀♀ (CBNUIC, dried). “Korea: Gangwon Prov. Gujeol-ri, Yeoryang-myeon, Jeongseon-gun, 24 Apr 2020, 37°30'57.6"N, 128°45'18.8"E, 510 m, sifting ant colony, leaf & soil litter, Y.-J. Choi, U.-J. Byeon”.

Antennal pedicel strongly swollen, subglobose with subcylindrical glandular nodule on basal 1/3 of inner margin (Figs 3F, 4C, arrows); protibiae with spine on internal side of widest (Figs 3G, 4E, arrows); parameres of male genitalia robust and fan-shaped, bearing three setae on each (Fig. 3I).

Figure 3. 

Bryaxis uljinensis Choi, Park, Lee & Park, sp. nov., male (A, B, E–I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A–D); 0.1 mm (E, I); 0.5 mm (F–H).

Figure 4. 

Diagnostic characters of Bryaxis uljinensis sp. nov. (A, C, E) and B. mahunkai Löbl (B, D, F). A, B maxillary palpi C, D antennal scapes and pedicels E, F protibiae. Scale bars: 0.1 mm.

Body reddish brown, antennae, maxillary palpi, legs slightly lighter than body, length 1.25–1.31 mm, maximum width 0.53–0.60 mm (Fig. 3A–D). Setae on body yellowish, and short. Head 0.98 times as long as wide (Fig. 3E). Frons with small U-shaped impression between antennal tubercles; frontal foveae absent; frontal rostrum slightly distinct anteriorly. Vertex convex; longitudinal carina present; vertexal foveae small. Eyes large, longer than twice that of tempora, composed of 32–34 facets. Maxillary palpi moderately developed; palpomeres II–III smooth; palpomere IV 0.22 mm long and about 3.16 times as long as wide, subcylindrical pseudosegment at apex. Antennae about 0.53 mm long; scapes short without modification, 0.92 times as long as pedicels; pedicels long as wide; antennomere III–VIII subequal in length; IX 0.63 times as long as wide; X transverse, 0.61 times as long as wide; XI largest, pointed at apex, 1.61 times as long as wide (Fig. 3F). Pronotum 0.88 times as long as wide and widest at basal 3/5, lateral antebasal foveae connected by antebasal sulcus. Elytra convex, 0.85 times as long as wide and widest at basal 1/3, each elytron with two basal foveae and subhumeral fovea. Legs robust; internal spine on widest of protibiae (Fig. 3G, arrow); metatibiae with spine on apical (Fig. 3H, arrow). Aedeagus small, 0.27 mm long and 1.61 times as long as wide; penis small fusiform and dorsal diaphragm bulbous; parameres symmetrical; endophallus composed with two convergent, slender struts (Fig. 3I).

Sexual dimorphism. Female eyes slightly longer than tempora, composed of 15 facets; antennal pedicels simple; protibial spines and metatibial spines absent.

Adults of this species are very similar to Bryaxis mahunkai Löbl, 1975 in having strongly swollen antennal pedicels (Fig. 4C, D). However, they can be distinguished by smooth maxillary palpomere II–III (Fig. 4A), apically symmetrical antennal scapes, pedicels less swollen apically and bearing smaller glandular nodules (Fig. 4C, arrow), fore legs with a tibial spine (Fig. 4E, arrow), and parameres wider than the penis (aedeagus in B. mahunkai as wide as penis; Fig. 10C, D).

The localities of B. uljinensis sp. nov. probably overlap with those of B. mahunkai Löbl given that the latter are nationally distributed (Fig. 11).

This species is named after the type locality, Uljin-gun.

The holotype was collected by sifting leaf litter in mixed forest. Paratypes were collected by sifting leaf litter, soil, and an ant colony.

Korea (Uljin-gun, Gyeongsangbuk-do; Jeongseon-gun, Gangwon-do).

(N = 4, 3♂♂, 1♀) . Holotype male. “Korea: Gangwon Prov. Gujeol-ri, Yeoryang-myeon, Jeongseon-gun, 24 Apr 2020, 37°31'08.3"N, 128°46'43.0"E, 552 m, sifting soil & leaf litter, U.-J. Byeon, T.-Y. Jang” (NIBR). Paratypes. (CBNUIC, 1♂ slide mounted, 1♂, 1♀ dried). “Korea: Gangwon Prov. Gujeol-ri, Yeoryang-myeon, Jeongseon-gun, 24 Apr 2020, 37°31'08.3"N, 128°46'43.0"E, 552 m, sifting soil & leaf litter, U.-J. Byeon, T.-Y. Jang”.

Enlarged fabiform antennal pedicels with subcylindrical glandular nodule on inner margin in male (Figs 5E, F, 6A, arrows).

Figure 5. 

Bryaxis fabaiformis Choi, Park, Lee & Park, sp. nov. male (A, B, E–I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A–D); 0.1 mm (E, I); 0.5 mm (F–H).

Figure 6. 

Diagnostic characters of Bryaxis fabaiformis sp. nov. (A, C, E) and B. kimjongkuki Nomura & Lee (B, D, F). A, B antennal scapes and pedicels C, D protibiae E, F heads. Scale bars: 0.1 mm.

Body reddish brown, antennae, maxillary palpi, legs slightly lighter than body, length 1.29–1.41 mm, maximum width 0.60–0.64 mm (Fig. 5A–D). Setae on body golden and long. Head 0.84–1.00 times as long as wide (Fig. 5F). Frons with U-shaped impression between antennal tubercles; frontal foveae absent; frontal rostrum distinct anteriorly. Vertex weakly convex; longitudinal carina distinct; vertexal foveae enlarged. Eyes large with 31–32 facets. Maxillary palpi moderately developed; palpomeres II–III with sparse tubercles; palpomere IV 0.25–0.29 mm long and about 2.92–3.44 times as long as wide, subcylindrical pseudosegment at apex. Antennae about 0.58–0.61 mm long; scapes short, without modification, 0.73–1.11 times as long as pedicels; pedicels 1.02–1.19 times as long as wide; antennomere III 1.18–1.23 times as long as wide; IV–VIII subequal in length; IX–X transverse, IX 0.66–0.69 times as long as wide and X 0.62–0.63 times as long as wide; XI largest, pointed at apex, 1.65–1.75 times as long as wide (Fig. 5F). Pronotum 0.84–0.89 times as long as wide and widest at basal 3/5, lateral antebasal foveae connected by antebasal sulcus. Elytra convex, 0.85–0.94 times as long as wide and widest at basal 1/3, each elytron with two basal foveae and subhumeral fovea. Legs slender; protibiae without spine (Fig. 5G); metatibiae with spine at apex (Fig. 5H, arrow). Aedeagus robust, 0.43 mm long and 1.79 times as long as wide; penis bulbous and dorsal diaphragm circular; parameres small and symmetrical, each with three setae; endophallus comprising two symmetrical struts, each broadened basally and apically, and shortly branched basally (Fig. 5I).

Sexual dimorphism. Female eyes slightly shorter than tempora, comprising 9 facets; antennal pedicels without modification; metatibial spines absent.

Adults of this species are similar to those of Bryaxis kimjongkuki Nomura & Lee, 1993 in having the maxillary palpomere II–III with tubercles and asymmetrical antennal scapes. However, they can be recognized by having a rounded tempora as long as the eyes (Fig. 6E), a glandular nodule situated at the mid-level of the antennal pedicels (Fig. 6A, arrow), and protibiae without a spine (Fig. 6C).

The localities of B. fabaiformis sp. nov. probably overlap with those of B. kimjongkuki Nomura & Lee given that the latter species was abundantly collected near the type localities of the former (Fig. 11).

The specific epithet is a combination of the Latin words faba (“bean”, feminine) and -formis (“having the form of”, masculine/feminine) and refers to the shape of antennal pedicels in the male.

Specimens of this species were collected by sifting soil and leaf litter in mixed forest.

Korea (Jeongseon-gun, Gangwon-do).

(N = 1, 1♂) . Holotype male. “Korea: Gangwon Prov. Bangdong-ri, Girin-myeon, Inje-gun, 23 Jun 2009, sifting flood debris, T.-K. Kim, CNUIC” (NIBR).

Antennal pedicels less enlarged subglobose, with dorsolateral glandular nodule on subapical (Fig. 7C, D, arrows); protibiae with spine on internal side at widest point (Figs 7E, 9C, arrows); parameres of male genitalia with depression on lateral margin and three setae on apical (Fig. 7G).

Figure 7. 

Bryaxis girinensis Choi, Park, Lee & Park, sp. nov. A dorsal habitus B ventral habitus C head D antenna E fore leg F hind leg G aedeagus. Scale bars: 1 mm (A, B); 0.1 mm (C, G); 0.5 mm (D–F).

Body reddish brown, antennae, maxillary palpi, and tarsi slightly lighter, length 1.62 mm, maximum width 0.70 mm (Fig. 7A, B). Setae on body yellowish, long and dense. Head long as wide (Fig. 7C). Frons with U-shaped impression between antennal tubercles; frontal foveae absent; frontal rostrum distinct anteriorly. Vertex slightly convex; longitudinal carina present; vertexal foveae small. Eyes large with 34 facets. Maxillary palpi moderately developed; palpomeres II–III with dense tubercles; palpomere IV 0.25 mm long and about 3.30 times as long as wide, subcylindrical pseudosegment at apex. Antennae about 0.54 mm long; scapes subcylindrical, without modification, 1.83 times as long as pedicels; pedicels long as wide; antennomere III 1.70 times as long as wide; IV–VIII subequal in length; IX 0.89 times as long as wide; X 0.78 times as long as wide; XI largest, pointed at apex, 1.86 times as long as wide (Fig. 7D). Pronotum 0.79 times as long as wide and widest at basal 3/5, lateral antebasal foveae connected by antebasal sulcus. Elytra slightly convex, 0.98 times as long as wide and widest at basal 1/3, each elytron with two basal foveae and subhumeral fovea. Legs robust; metatibiae with spine at apex (Fig. 7E, arrow). Aedeagus large, 0.36 mm long and 2.05 times as long as wide; penis fusiform and dorsal diaphragm transversely ovoid; parameres short and symmetrical, apices truncated; endophallus composed of two fine struts, asymmetrical (Fig. 7G).

Sexual dimorphism. Unknown.

The adult of this species is similar to Bryaxis nemorosus Choi, Park, Lee & Park sp. nov. in the shape of antennomeres IV–XI (Figs 7D, 8F). However, it can be distinguished by the robust setae on the body (Fig. 7A), large eyes as long as the tempora (Fig. 9E), a strongly tuberculate maxillary palpomere II (Fig. 9A), protibiae with a spine at the widest point (Fig. 9C, arrow), and a simple endophallus of the male genitalia (Fig. 7G).

This species is named after the type locality, Girin-myeon, Inje-gun.

The holotype was collected by sifting flood debris in mixed forest.

Korea (Inje-gun, Gangwon-do).

(N = 5, 4♂♂, 1♀) . Holotype male. “Korea: Jeonnam Prov. Mt. Doksil, Gageo island. Gageodo-gil, Heuksan-myeon, Sinan-gun, 13 Jul 2021, 34°05'06.1"N, 125°06'17.4"E, 468 m, sifting leaf & soil litter, J.-W. Seo” (NIBR). Paratype. 1♂ (CBNUIC, slide mounted). “Korea: Jeonnam Prov. Mt. Doksil, Gageo island. Gageodo-gil, Heuksan-myeon, Sinan-gun, 13 Jul 2021, 34°05'06.1"N, 125°06'17.4"E, 468 m, sifting leaf & soil litter, J.-W. Seo”. 1♂ (CBNUIC, dried). “Korea: Jeonnam Prov. Mt. Doksil, Gageo island. Gageodo-gil, Heuksan-myeon, Sinan-gun, 8 Jul 2020, 34°05'35.0"N, 125°06'25.0"E, 590 m, sifting leaf & soil litter, T.-Y. Jang”. 1♂ (CBNUIC, dried). “Korea: Jeonnam Prov. Mt. Doksil, Gageo island. Gageodo-gil, Heuksan-myeon, Sinan-gun, 8 Jul 2020, 34°04'40.0"N, 125°06'23.0"E, 540 m, sifting leaf & soil litter, U.-J. Byeon, T.-Y. Jang”. 1♀ (CBNUIC, dried). “Korea: Jeonnam Prov. Mt. Doksil, Gageo island. Gageodo-gil, Heuksan-myeon, Sinan-gun, 7 Jul 2020, 34°04'34.7"N, 125°06'28.8"E, 534 m, sifting leaf & soil litter, T.-Y. Jang”.

Elongated head with small eyes situated on mid-length of head (Fig. 8E, arrow).

Figure 8. 

Bryaxis nemorosus Choi, Park, Lee & Park, sp. nov. male (A, B, E–I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A–D); 0.1 mm (E, I); 0.5 mm (F–H).

Body reddish brown, antennae, maxillary palpi, legs slightly lighter than body, length 1.57–1.58 mm, maximum width 0.61–0.67 mm (Fig. 8A–D). Setae on body yellowish. Head 1.15 times as long as wide (Fig. 8E). Frons with U-shaped impression between antennal tubercles; frontal foveae absent; frontal rostrum distinct anteriorly. Vertex weakly convex; longitudinal carina absent; vertexal foveae small. Eyes reduced with 9 facets. Maxillary palpi moderately developed; palpomeres II smooth; III with tubercles; palpomere IV 0.24–0.28 mm long and about 3.06–3.24 times as long as wide, subcylindrical pseudosegment at apex. Antennae about 0.61–0.68 mm long; scapes subcylindrical and elongated, 2–2.18 times as long as pedicels, without modification; pedicels subcylindrical, 1.38–1.40 times as long as wide; antennomere III 1.67–1.72 times as long as wide; IV–VII subequal in length; VIII subglobose as long as wide; IX 0.82–0.86 times as long as wide; X transverse, 0.64–0.65; XI largest, pointed at apex, 1.76–1.84 times as long as wide (Fig. 8F). Pronotum 0.85–0.89 times as long as wide and widest at basal 2/3, lateral antebasal foveae connected by antebasal sulcus. Elytra slightly convex, 0.87 times as long as wide and widest at basal 1/3, each elytron with two basal foveae and subhumeral fovea. Legs slender; protibiae without spine (Fig. 8G); metatibiae with spine on apical (Fig. 8H, arrow). Aedeagus robust, 0.53 mm long and 1.79 times as long as wide; penis bulbous and dorsal diaphragm small, transversely ovoid; parameres symmetrical, each bearing two setae; endophallus comprising simple strut basally and two symmetrical struts curved along with parameres, thickened basally (Fig. 8I).

Sexual dimorphism. Female metatibial spines absent.

Adults of this species are similar to that of Bryaxis girinensis Choi, Park, Lee & Park sp. nov. in the shape of antennomeres IV–XI (Figs 7D, 8F). However, they can be distinguished by having angular tempora much longer than the eyes (Fig. 9F), smooth maxillary palpomere II (Fig. 9B), unadorned antennal scapes and pedicels (Fig. 8F), slender protibiae without a spine (Fig. 9D), and an endophallus composed of three long struts (Fig. 8I).

Figure 9. 

Diagnostic characters of Bryaxis girinensis sp. nov. (A, C, E) and B. nemorosus sp. nov. (B, D, F). A, B maxillary palpi C, D protibiae E, F heads. Scale bars: 0.1 mm.

The specific epithet is the Latin word nemorosus (“wooded, shady”, masculine) that refers to habitat where the types collected.

Specimens of this species were collected by sifting leaf and soil litter in wet forest with dense canopy, which is located on an island.

Korea (Gageo island, Sinan-gun, Jeollanam-do).

(N = 60, 34♂♂, 26♀♀) . 1♂ (CBNUIC, dried). “Korea: Chungbuk Prov., Mt. Songnisan, Beopjusa-ro, Songnisan-myeon, Boeun-gun, 20 Sep 2019, 36°32'55.6"N, 127°51'19.8"E, 476 m, flood debris, Y.-J. Choi, J.-W. Kang”. 2♂♂, 11♀♀ (CBNUIC, dried). “Korea: Gangwon Prov., Garakjae-ro, Hwachon-myeon, Hongcheon-gun, 5 May 2019, 37°46'26.0"N, 127°54'48.0"E, 240 m, sifting leaf litter near stream, J.-S. Park”. 1♂ (CBNUIC, dried). “Korea: Gangwon Prov., Jangjeon-gil, Jinbu-myeon, Pyeongchang-gun, 11 Mar 2019, 37°27'58.0"N, 128°32'18.4"E, 901 m, sifting leaf litter & dead wood debris & moss, J.-W. Kang”. 7♂♂, 7♀♀ (CBNUIC, dried). “Korea: Gyeongbuk Prov., Gowol-gil, Yeongyang-eup, Yeongyang-gun, 19 Mar 2019, 36°38'48.1"N, 129°09'18.7"E, 265 m, sifting leaf litter, Y.-J. Choi”. 2♂♂ (CBNUIC, dried). “Korea: Gyeongbuk Prov., Yongmunsa-gil, Yongmun-myeon, Yecheon-gun, 1 Jun 2019, 36°43'45.0"N, 128°22'14.0"E, 358 m, sifting leaf litter & soil near stream, U.-J. Byeon”. 7♂♂ (CBNUIC, dried). “Korea: Gyeonggi Prov., Mt. Bukhansan, Daeseomun-gil, Deogyang-gu, Goyang-si, 23 Aug 2019, 37°39'43.7"N, 126°59'11.2"E, 491 m, sifting leaf litter & soil & dead wood debris, Y.-J. Choi, T.-Y. Jang”. 4♂♂ (CBNUIC, dried). “Korea: Gyeonggi Prov., Mt. Yeoninsan, Yongchu-ro, Gapyeong-eup, Gapyeong-gun, 15 Apr 2019, 37°51'29.5"N, 127°28'01.0"E, 193 m, sifting leaf litter & moss near stream, J.-Y. Kang, J.-W. Kang”. 5♂♂, 3♀♀ (CBNUIC, dried). “Korea: Gangwon Prov., Hwanseon-ro, Singi-myeon, Samcheok-si, 23 Aug 2018, 37°20'22.6"N, 129°03'28.5"E, 172 m, sifting leaf litter near mountain stream, Y.-J. Choi”. 5♂♂, 5♀♀ (CBNUIC, dried). “Korea: Jeonnam Prov., Mt. Heukseoksan, Biseuran-gil, Gyegok-myeon, Haenam-gun, 18 May 2019, 34°40'44.9"N, 126°37'10.9"E, 160 m, sifting mushroom & leaf litter & plant root under rock in bamboo forest, S.-H. Choi, U.-J. Byeon”.

Adult males of this species are characterized by the following combination of characters: maxillary palpomeres II–III tubercular ventrally; antennal scapes tubiform, more curved on the internal side; pedicels globularly enlarged and a glandular nodule situated at the basal 1/3 (Fig. 10A, B, arrows); parameres with four setae each; and an endophallus consisting of two slender struts, converging subapically (Fig. 10C, D).

Figure 10. 

Diagnostic characters of Bryaxis mahunkai Löbl. A, B antennal scapes and pedicels C, D aedeagi. Scale bars: 0.1 mm.

Dorsal habitus of the holotype is available in Park and Jeon (2012; https://ecolibrary.me.go.kr/nibr/#/search/detail/5513253). Illustrations of antenna and aedeagus were obtained from Löbl (1975), and compared to those of specimens examined in this study. All specimens collected in Korea were recognized as B. mahunkai Löbl based on the antennal scapes (curved internally), pedicels (swollen and bearing upward glandular nodule), and the aedeagus (structure of endophallus).

Korea (Kaesong-si, Gyeonggi-do; Gapyeong-gun, Gyeonggi-do; Goyang-si, Gyeonggi-do; Boeun-gun, Chungcheongbuk-do; Hongcheon-gun, Gangwon-do; Samcheok-si, Gangwon-do; Pyeongchang-gun, Gangwon-do; Yeongyang-gun, Gyeongsangbuk-do; Yecheon-gun, Gyeongsangbuk-do; Haenam-gun, Jeollanam-do).