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Catalogue
An updated list of type material of Ephemeroptera Hyatt & Arms, 1890, deposited at the Zoological Museum of Hamburg (ZMH)
expand article infoMichel Sartori‡, Martin Kubiak§, Hossein Rajaei Sh.|
‡ Musée cantonal de zoologie, Lausanne, Switzerland
§ Zoologisches Museum, Hamburg, Germany
| State Museum of Natural History Stuttgart, Stuttgart, Germany

Corresponding author: Michel Sartori ( michel.sartori@vd.ch )

Academic editor: Eduardo Dominguez
© 2016 Michel Sartori, Martin Kubiak, Hossein Rajaei Sh..
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation: Sartori M, Kubiak M, Rajaei H (2016) An updated list of type material of Ephemeroptera Hyatt & Arms, 1890, deposited at the Zoological Museum of Hamburg (ZMH). ZooKeys 607: 49-68. https://doi.org/10.3897/zookeys.607.9391
ZooBank: urn:lsid:zoobank.org:pub:F2D11E9A-5AC0-4309-BB04-B83142605624
Open Access

Abstract

The type specimens of Ephemeroptera (Insecta) housed at the Zoological Museum of Hamburg (ZMH) are compiled in this document. The current nomenclature of all species is given. In total, Ephemeroptera type material of ZMH encompasses 161 species. Fifty-one holotypes and five lectotypes are present. Forty-one species are represented by syntypes, 85 by paratypes and five by paralectotypes. Material of two species (Cinygma asiaticum Ulmer, 1924 and Pseudocloeon klapaleki Müller-Liebenau, 1982) is missing. The present catalogue is an updated version of Weidner (1964a).

Keywords

Ephemeroptera , Hamburg, type specimens, Ulmer, ZMH

Introduction

The Ephemeroptera collection of the Zoological Museum of Hamburg (ZMH) contains 433 species and approximately 4,500 specimens. In total, 161 species are represented by type material. In this sense, this is one of the largest collections with the highest number of type specimens of this insect order in Germany. Furthermore, the mayfly collection together with the caddisfly collection comprise some of the oldest voucher material of the entomological collection at the ZMH which was mainly collected at the end of the 19th century and in the first decades of the 20th century by the Hamburgian entomologist Dr. h.c. Georg Ulmer (1877–1963) (e.g., Weidner 1964b).

Georg Ulmer was born in Hamburg on 5 March 1877. He was the oldest son of seven siblings. After his father died in 1889, he decided to become a school teacher. During his qualification time in the early 1890s, he began collecting insects in the vicinity of Hamburg (Ulmer 1964). Initially, he started to compile a beetle collection but since he observed a mass emergence of mayflies at the river Fulda near Kassel in 1898, he started to focus his collecting efforts and taxonomic studies on primarily aquatic insect groups like mayflies or caddisflies (Ulmer 1964). In 1899, he finished his teacher training and started his career as a board school teacher in Hamburg for 32 years until 1934. Beside his profession, he studied voluntarily the faunistics, biology, systematics and taxonomy of Ephemeroptera (e.g., Ulmer 1904a, 1926, 1939) and Trichoptera (e.g., Ulmer 1905, 1951), and other freshwater invertebrates (e.g., Ulmer 1901, 1902a).

His first publication (Ulmer 1900) was followed by 175 scientific publications until his death on 15 January 1963 (Illies 1964). In the first years of his scientific career, he mainly studied the faunistics and taxonomy of caddisflies in the vicinity of Hamburg (e.g., Ulmer 1902b), or other localities in Germany (e.g. Ulmer 1904b, 1915). From the year 1905 on, he intensively started to study the material from foreign countries (including tropical regions), which has been lent to him by colleagues or other institutions (e.g., Ulmer 1905).

Besides the Trichoptera, Ulmer worked on mayflies (e.g., Ulmer 1908, 1909, 1912). Between 1904 and 1943, he published thirty research papers on the systematics and taxonomy of Ephemeroptera (Kimmins 1963; Illies 1964) with the description of 111 species (Weidner 1964a). His studies on mayflies resulted in the fundamental revision of the Southeast Asian mayfly fauna with the description of numerous species (Ulmer 1939).

During World War II (in 1943), the Zoological Museum Hamburg (that time located near the central railway station) was bombed and nearly all dry preserved material housed in the collections was destroyed and burned in the fire. Only material deposited in ethanol was transferred into the underground networks and survived the war (Weidner 1967). The valuable Ephemeroptera and Trichoptera collection of Georg Ulmer was kept in his private house outside of Hamburg. In 1964, the insect collection of Georg Ulmer was donated to the ZMH (Weidner 1964b) providing the majority of the recent voucher material of the ZMH in these two insect orders.

Georg Ulmer was an outstanding specialist and his comprehensive studies on the faunistics and taxonomy of various groups of freshwater invertebrates formed the foundation of the recent taxonomy and systematics of those groups, especially Ephemeroptera and Trichoptera.

In addition to Ulmer’s collection, the ZMH also stores primary type material described by other authors, such as Klapalek (1905a,b), Lestage (1930, 1935), Malzacher (1976, 2015, 2016), Müller-Liebenau (1971, 1974, 1980a, b, 1981), and Sartori (2008, 2014e). An interesting finding was the discovery of several paratypes of species described by Esben-Petersen (e.g. 1909, 1912a, b, 1913), which were not labelled as such but come from the type series, judging from the dates and places on labels. More information on these mayfly specialists can be found at http://www.ephemeroptera.de/inhaltsverz_deutsch/Galerie____/galerie____.html, a webpage maintained by Udo Jacob and Arne Haybach.

The only attempt, prior to this type catalogue, has been made by Weidner (1962, 1964a, 1977), who published a list of Ephemeroptera type material deposited in ZMH, without any taxonomic remarks. At that time, 121 species were mentioned. This catalogue is accessible online (https://www.cenak.uni-hamburg.de/sammlungen/zoologie/entomologie/typenkatalog.html).

Multiple subsequent taxonomic surveys utilizing a variety of different, high-resolution methods (morphological and molecular) led not only to the description of many new taxa, but also several supraspecific changes (e.g. Peters and Edmunds 1972; Puthz 1977; Braasch and Soldán 1986; Campbell and Suter 1988; Gillies 1990; Grant and Peters 1993; McCafferty and Waltz 1995; Lugo-Ortiz and McCafferty 1996, 1999; Sartori et al. 2008; Sartori 2014a, b, c.).

The catalogue that follows is arranged alphabetically by species name, followed by the original combination, author(s) name and year of publication. For each entry, an actual nomenclature (valid names or synonymy) is given. The kind of type and the metamorphic stage of specimens are abbreviated as indicated in the table header. The present study is a completely independent and new work; data reported by Weidner were not copied as a base for this catalogue. This study is entirely original work, based on the first author’s specimen-by-specimen examination of the ZMH collections from December 2013 to June 2014. Recent publications by Malzacher (2015, 2016) on Ulmer’s collection were also added.

Table 1.
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List of type material of Ephemeroptera Hyatt & Arms, 1890, deposited at the Zoological Museum of Hamburg, with the state and metamorphic stage of materials, current nomenclature, family assignment, related literature and complementary information. Abbreviations in this list are as follows: Type: HT= Holotype; LT= Lectotype; PT= Paratype(s); PLT= Paralectotype(s); ST = Syntypes. (state): (1) Specimen conserved in alcohol; (2) Pinned dry specimen; (3) Specimen or part of it mounted on microscopic slide. Stage: F= Female imago; M= Male imago; N= Nymph; SF= Female subimago; SM= Male subimago. Complementary information: a. Holotype wrongly designated by Weidner (1962, 1964a); b. Holotype designated by Ulmer on the label, not in the publication; c. Holotype apparently not in Stockholm (see http://www.nrm.se/download/18.1cb760b014a762ca801342e9/1421068753427/EPHEMEROPTERA_list.pdf); d. Specimen not labelled, but comes from the type series; e. Not type material! The types are the nymphs and the adults were only described in 1925; f. Only fore- and hindlegs in ZMH; g. Only fore wing in ZMH and remains in Naturkunde Museum, Berlin; h. Only foreleg; i. Only hind wing in ZMH and remains in Naturkunde Museum, Berlin; j. Only one forewing in ZMH and remains apparently not in Stockholm (http://www.nrm.se/download/18.1cb760b014a762ca801342e9/1421068753427/EPHEMEROPTERA_list.pdf); k. Wrongly considered by Ulmer, as a syn. of E. bellieri (Hagen).

Taxon Original name Author(s), year Type (state) Stage Current nomenclature Family Literature & Complementary information
alegrettae Massartella alegrettae Ulmer, 1943 HT (2) M Massartella alegrettae Ulmer, 1943 Leptophlebiidae (i.)
ambiguum Pseudocloeon ambiguum Müller-Liebenau, 1982 PT (1) N Liebebiella ambigua (Müller-Liebenau, 1982) Baetidae Waltz and McCafferty 1987
ammophila Oligoneuria ammophila Spieth, 1938 PT (1) N Homoeoneuria ammophila (Spieth, 1938) Oligoneuriidae Edmunds and Allen 1957
anatolica Rhithrogena anatolica Kazanci, 1985 PT (1) M Rhithrogena anatolica Kazanci, 1985 Heptageniidae (d.)
annandalei Polymitarcys annandalei Chopra, 1927 ST (1) F Ephoron annandalei (Chopra, 1927) Polymitarcyidae Hubbard and Srivastava 1985
apicatum Cloeon apicatum Navás, 1933 ST (2) M Cloeon navasi van Bruggen, 1957 Baetidae van Bruggen 1957; (d.)
balcanicus Metretopus balcanicus Ulmer, 1920 HT (3) M Metreletus balcanicus (Ulmer, 1920) Ameletidae Puthz 1977a; (f.)
belgica Torleya belgica Lestage, 1917 (PT) (2) M, F Torleya major (Klapalek, 1905a) Ephemerellidae (e.)
bengalensis Ecdyonurus bengalensis Ulmer, 1920 HT, PT (2, 3) M, F, SM, SF Ecdyonurus bengalensis Ulmer, 1920 Heptageniidae (b.)
berneri Rheobaetis berneri Müller-Liebenau, 1974 HT (1) N Heterocloeon berneri (Müller-Liebenau, 1974) Baetidae McCafferty and Provonsha 1975
bicorne Centroptilum bicorne Ulmer, 1909 PT (1) F Afroptilum bicorne (Ulmer, 1909) Baetidae Gillies, 1990
biobionicum Deleatidium biobionicum Ulmer, 1938 PT (2) F Meridialaris biobionica (Ulmer, 1938) Leptophlebiidae Peters and Edmunds 1972; (g.)
bishopi Platybaetis bishopi Müller-Liebenau, 1980a HT (1) N Platybaetis bishopi Müller-Liebenau, 1980 Baetidae
boettgeri Pseudocloeon boettgeri Ulmer, 1924 ST (1, 2, 3) M, F Pseudocloeon boettgeri Ulmer, 1924 Baetidae
borneonia Epeorella borneonia Ulmer, 1939 LT, PLT (1, 2) M, F, SF Epeorella borneonia Ulmer, 1939 Heptageniidae Sartori 2014c; (a.)
braueri Hagenulodes braueri Ulmer, 1920 PT (1) M Hagenulodes braueri Ulmer, 1920 Leptophlebiidae
brenneriana Rhithrogena brenneriana Klapalek, 1905a PT (1) M Rhithrogena alpestris Eaton, 1885 (syn.) Heptageniidae Puthz 1975
brunneum Cloeon brunneum Esben-Petersen, 1909 ST (1, 2) M, F Americabaetis peterseni (Hubbard, 1974) (syn. obj.) Baetidae Lugo-Ortiz and McCafferty 1999
caenoides Neoephemeropsis caenoides Ulmer, 1939 LT, PLT (1, 3) M, F, N Potamanthellus caenoides (Ulmer, 1939) Neoephemeridae Bae and McCafferty 1998
camerunense Pseudocloeon camerunense Ulmer, 1920 ST (1, 3) M, F Ophelmatostoma camerunense (Ulmer, 1920) Baetidae Gillies et al. 1990
canariensis Baetis canariensis Müller-Liebenau, 1971 HT, PT (1) M, N Baetis canariensis Müller-Liebenau, 1971 Baetidae
cavum Cinygma cavum Ulmer, 1927 PT (1) M Cinygmula cava (Ulmer, 1927) Heptageniidae Levanidova 1972
chinensis Baetis chinensis Ulmer, 1936 ST (1) M, F Baetis chinensis Ulmer, 1936 Baetidae
chinensis Heptagenia chinensis Ulmer, 1920 ST (2) M Heptagenia chinensis Ulmer, 1920 Heptageniidae (d.)
chinensis Siphluriscus chinensis Ulmer, 1920 PT (3) M Siphluriscus chinensis Ulmer, 1920 Siphluriscidae (f.)
corsicus Ecdyonurus corsicus Esben-Petersen, 1912a PT (2) M Ecdyonurus corsicus Esben-Petersen, 1912 Heptageniidae Belfiore 1987; (k.)
costaricanus Leptohyphes costaricanus Ulmer, 1920 HT (2, 3) F Tricorythodes costaricanus (Ulmer, 1920) Leptohyphidae Bamgardner 2008
crassinervis Polyplocia crassinervis Ulmer, 1939 PT (2) SM Polyplocia campylociella Ulmer, 1939 (syn.) Euthyplociidae Demoulin 1953
croaticus Siphlonurus croaticus Ulmer, 1920 PT (1, 3) M Siphlonurus croaticus Ulmer, 1920 Siphlonuridae
curvata Paraleptophlebia curvata Ulmer, 1927 PT (1, 3) M Paraleptophlebia strandii (Eaton, 1901) (syn.) Leptophlebiidae Kluge 2009
deigma Liebebiella deigma Waltz & McCafferty, 1987 HT, PT (1, 3) N Liebebiella vera (Müller-Liebenau, 1981) (syn.) Baetidae Kluge and Novikova 2011
difficilum Pseudocloeon difficilum Müller-Liebenau, 1982 PT (1, 3) N Liebebiella difficila (Müller-Liebenau, 1982) Baetidae Waltz and McCafferty 1987
diptera Hagenulopsis diptera Ulmer, 1920 ST (1) M, SM, SF Hagenulopsis diptera Ulmer, 1920 Leptophlebiidae
duporti Ephemera duporti Lestage, 1921 PT (2) M Ephemera duporti Lestage, 1921 Ephemeridae
eatoni Rhithrogena eatoni Esben-Petersen, 1912a PT (2) F Rhithrogena eatoni Esben-Petersen, 1912 Heptageniidae (d.)
edmundsi Platybaetis edmundsi Müller-Liebenau, 1980b HT (1) N Platybaetis edmundsi Müller-Liebenau, 1980 Baetidae
ehrhardti Thraulus ehrhardti Ulmer, 1920 ST (1, 2, 3) M, F Needhamella ehrhardti (Ulmer, 1920) Leptophlebiidae Dominguez and Flowers 1989
eloisae Derlethina eloisae Sartori, 2008 PT (1) N Derlethina eloisae Sartori, 2008 Teloganodidae Sartori et al. 2008
feuernborni Pseudoligoneuria feuernborni Ulmer, 1939 ST (1, 3) N Chromarcys magnifica Navás, 1932 (syn.) Oligoneuriidae Kluge 2004
flexus Siphlurus flexus Clemens, 1913 PT (1) M Siphloplecton basale (Walker, 1853) (syn.) Metretopodidae McDunnough 1924
fluviatile Cloeon fluviatile Ulmer, 1920 PT (1, 3) M, F Cloeon fluviatile Ulmer, 1920 Baetidae
formosana Ephemera formosana Ulmer, 1920 PT (2, 3) M, F, SM, SF Ephemera formosana Ulmer, 1920 Ephemeridae
formosanus Chirotonetes formosanus Ulmer, 1912 ST (1, 2, 3) M, SF Isonychia formosana (Ulmer, 1912) Isonychiidae Ueno 1931
fruhstorfferi Massartella fruhstorfferi Ulmer, 1943 PT SF Massartella brieni (Lestage, 1924) (syn.) Leptophlebiidae Pescador and Peters 1990
fuegiensis Ameletus fuegiensis Lestage, 1935 HT (1, 3) N Metamonius anceps (Eaton, 1885) (syn.) Nesameletidae Mercado and Elliott 2004
fulmeki Acentrella fulmeki Ulmer, 1939 HT, PT (1, 3) M, F Labiobaetis fulmeki (Ulmer, 1939) Baetidae McCafferty and Waltz 1995
fusca Atalophlebia fusca Ulmer, 1920 HT (1) M Koorrnonga fusca (Ulmer, 1920) Leptophlebiidae Campbell and Suter 1988; (c.)
gornostajevi Epeorus gornostajevi Tshernova, 1981 PT (1) M Epeorus gornostajevi Tshernova, 1981 Heptageniidae
grandis Chirotonetes grandis Ulmer, 1913 ST (1, 3) M Isonychia grandis (Ulmer, 1913) Isonychiidae Ulmer 1924
guranica Heptagenia guranica Belov, 1981 PT (1) M Heptagenia guranica Belov, 1981 Heptageniidae
haenschi Euthyplocia haenschi Ulmer, 1942 PT (2) M Euthyplocia haenschi Ulmer, 1942 Euthyplociidae
horai Potamanthellus horai Lestage, 1930 HT (2) SM Potamanthellus amabilis (Eaton, 1892) (syn.) Neoephemeridae Bae and McCafferty 1998
hutanis Dudgeodes hutanis Sartori, 2008 PT (1) N Dudgeodes hutanis Sartori, 2008 Teloganodidae Sartori et al. 2008
hyalinus Ecdyonurus hyalinus Ulmer, 1912 ST (1, 2, 3) M, F, SM, SF Afronurus hyalinus (Ulmer, 1912) Heptageniidae Kang and Yang 1994
insularis Rhithrogena insularis Esben-Petersen, 1913 PT (2) M, F Rhithrogena insularis Esben-Petersen, 1913 Heptageniidae (d.)
irina Cinygmula irina Tshernova & Belov, 1982 PT (1) M Cinygmula irina Tshernova & Belov, 1982 Heptageniidae
jacobsoni Tricorythus jacobsoni Ulmer, 1913 ST (1, 3) M, F Sparsorythus jacobsoni (Ulmer, 1913) Tricorythidae Sroka and Soldán 2008; (a.)
japonicus Chirotonetes japonicus Ulmer, 1920 ST (1, 2, 3) M, F Isonychia japonica (Ulmer, 1920) Isonychiidae Ueno 1931; (a.)
javanicus Afronurus javanicus Ulmer, 1939 HT (1) M Afronurus javanicus Ulmer, 1939 Heptageniidae
javanicus Baetis javanicus Ulmer, 1913 ST (1, 2, 3) F Baetis javanicus Ulmer, 1913 Baetidae
jorgenseni Campsurus jorgenseni Esben-Petersen, 1912b ST (1) M, F Campsurus jorgenseni Esben-Petersen, 1912 Polymitarcyidae
jorgenseni Cloeon jorgenseni Esben-Petersen, 1909 ST (1, 3) M, F Americabaetis jorgenseni (Esben-Petersen, 1909) Baetidae Lugo-Ortiz and McCafferty 1999
karnyi Hagenulus karnyi Ulmer, 1939 ST (1, 3) SM, SF Choroterpes (Euthraulus) karnyi (Ulmer, 1939) Leptophlebiidae Peters and Edmunds 1970
kraepelini Pseudocloeon kraepelini Klapalek, 1905b LT, PLT (1, 3) M, SM Pseudocloeon kraepelini Klapaleck, 1905 Baetidae Waltz and McCafferty 1985
krieghoffi Chitonophora krieghoffi Ulmer, 1920 ST (2) M, S Ephemerella mucronata (Bengtsson, 1909) (syn.) Ephemerellidae Jacob 1974
lacuscoerulei Tasmanophlebia lacuscoerulei Tillyard, 1933 PT (1) SF Tasmanophlebia lacuscoerulei Tillyard, 1933 Oniscigastridae (d.)
laminatum Deleatidum laminatum Ulmer, 1920 PT (3) M Meridialaris laminata (Ulmer, 1920) Leptophlebiidae Peters and Edmunds 1972; (h.)
lamuriensis Thalerosphyrus lamuriensis Sartori, 2014e HT, PT (1, 3) N Thalerosphyrus lamuriensis Sartori, 2014 Heptageniidae
latifrons Cinygmula latifrons Tshernova & Belov, 1982 PT (1) M Cinygmula hirasana (Imanishi, 1935) (syn.) Heptageniidae Kluge 1995
latus Tricorythus latus Ulmer, 1916 PT (1, 2, 3) M Tricorythus latus Ulmer, 1916 Tricorythidae
lestagei Atalophlebiodes lestagei Ulmer, 1938 ST (1, 3) N Meridialaris lestagei (Ulmer, 1938) Leptophlebiidae Dominguez et al. 2006
lieftincki Heptagenia lieftincki Ulmer, 1939 HT, PT (1, 3) M, F Compsoneuria lieftincki (Ulmer, 1939) Heptageniidae Sartori 2014a
lobatus Ecdyonurus lobatus Ulmer, 1924 ST (1, 2, 3) M, F, SF Afronurus lobatus (Ulmer, 1924) Heptageniidae Mol 1987
longilobata Leptophlebia longilobata Tshernova, 1928a PT (1, 3) M Paraleptophlebia longilobata (Tshernova, 1928) Leptophlebiidae Tiunova and Kluge 2016
longus Tricorythus longus Ulmer, 1916 PT (1, 2, 3) M, F Tricorythus longus Ulmer, 1916 Tricorythidae
lucida Atalophlebia lucida Ulmer, 1920 HT (1) M Thraulophlebia lucida (Ulmer, 1920) Leptophlebiidae Demoulin 1955; (c.)
macedonicus Rhoenanthus macedonicus Ulmer, 1920 HT (2) M Neoephemera maxima (Joly, 1871) Neoephemeridae Bae and McCafferty 1998
maculipennis Thraulus maculipennis Ulmer, 1920 HT (1) M Hermanella maculipennis (Ulmer, 1920) Leptophlebiidae Dominguez and Flowers 1989
magnificus Rhoenanthus magnificus Ulmer, 1920 PT (1, 2, 3) M Rhoenanthus magnificus Ulmer, 1920 Potamanthidae
major Choroterpides major Ulmer, 1939 HT, PT (1, 3) M, N Dilatognathus major (Ulmer, 1939) Leptophlebiidae Kluge 2012
malaisei Cinygma malaisei Ulmer, 1927 PT (1, 3) M Cinygmula malesei (Ulmer, 1927) Heptageniidae Levanidova 1972
marginatus Thraulus marginatus Ulmer, 1913 ST (1, 3) M, F Choroterpes (Euthraulus) marginatus (Ulmer, 1913) Leptophlebiidae Peters and Edmunds 1970
media Ephemera media Ulmer, 1936 ST (2) M, F, SM, SF Ephemera media Ulmer, 1936 Ephemeridae (a.)
media Teloganopsis media Ulmer, 1939 HT, PT (1, 3) M, N Teloganopsis media Ulmer, 1939 Ephemerellidae Ubero-Pascal and Sartori 2009;
Sartori 2014f; (d.)
melli Thalerosphyrus melli Ulmer, 1926 PT (2, 3) M, F Epeorus melli (Ulmer, 1926) Heptageniidae Zhou et al. 2007
mexicana Heptagenia mexicana Ulmer, 1920 PT (3) M Maccaffertium mexicanum (Ulmer, 1920) Heptageniidae Wang and McCafferty 2004; (f.)
mjoebergi Euphyurus mjoebergi Ulmer, 1917 PT (1, 3) M Ulmerophlebia mjoebergi (Ulmer, 1917) Leptophlebiidae Demoulin 1955
montium Thraulus montium Ulmer, 1943 HT (2) M Traverella montium (Ulmer, 1943) Leptophlebiidae Allen 1973
multus Baetis multus Müller-Liebenau, 1984 PT (3) N Labiobaetis multus (Müller-Liebenau, 1984) Baetidae McCafferty and Waltz 1995
nasuta Heptagenia nasuta Ulmer, 1939 ST (1, 2) M, F Trichogenia nasuta (Ulmer, 1939) Heptageniidae Webb et al. 2006; (a)
natans Chankagenesia natans Buldowsky, 1935 PT (1) M, F Chankagenesia natans Buldowsky, 1935 Palingeniidae
necatii Ecdyonurus necatii Kazanci, 1987 PT (1) SM Electrogena necatii (Kazanci, 1987) Heptageniidae Kazanci 2001
necopinatus Baetis necopinatus Müller-Liebenau, 1981 HT (1, 3) M Labiobaetis necopinatus (Müller-Liebenau, 1981) Baetidae McCafferty and Waltz 1995
nigrescens Baetis nigrescens Navás, 1932 ST (2) M Baetis nigrescens Navás, 1932 Baetidae Müller-Liebenau 1971
nigrescens Tasmanophlebia nigrescens Tillyard, 1933 PT (1) F, N Tasmanophlebia nigrescens Tillyard, 1933 Oniscigastridae (d.)
nigropunctata Caenis nigropunctata Klapalek, 1905b ST (2, 3) F Caenis nigropunctata Klapalek, 1905 Caenidae Malzacher 2015
nigropunctatula Caenis nigropunctatula Malzacher, 2015 HT, PT (1, 3) M, F Caenis nigropunctatula Malzacher, 2015 Caenidae
nitidum Centroptilum nitidum Ulmer, 1916 ST (2) F Bugilliesia nitida (Ulmer, 1916) Baetidae Lugo-Ortiz and McCafferty 1996
novatus Baetis novatus Müller-Liebenau, 1981 HT (3) N Baetis novatus Müller-Liebenau, 1981 Baetidae
obscurum Pseudocloeon obscurum Ulmer, 1913 ST (3) M Pseudocloeon obscurum Ulmer, 1913 Baetidae Müller-Liebenau 1981
olivascens Baetis olivascens Ulmer, 1939 HT, PT (1, 3) M, SM, SF Baetis olivascens Ulmer, 1939 Baetidae
operosus Baetis operosus Müller-Liebenau, 1984 PT (3) N Labiobaetis operus (Müller-Liebenau, 1984) Baetidae McCafferty and Waltz 1995
orientale Pseudocloeon orientale Müller-Liebenau, 1982 PT (1) N Liebebiella orientale (Müller-Liebenau, 1982) Baetidae Waltz and McCafferty 1987
orientalis Palingenia orientalis Chopra, 1927 PT (2) M Palingenia orientalis Chopra, 1927 Palingeniidae
orientalis Raptobaetopus orientalis Müller-Liebenau, 1978 HT (3) N Baetopus orientalis (Müller-Liebenau, 1978) Baetidae Jacob 1991
ornata Rhithrogeniella ornata Ulmer, 1939 HT, PT (1, 3) M, F Rhithrogeniella ornata Ulmer, 1939 Heptageniidae Sartori 2014d
ornatum Procloeon ornatum Tshernova, 1928b LT (1, 3) M Procloeon ornatum Tshernova, 1928 Baetidae Sowa 1975
paradoxa Anagenesia paradoxa Buldowsky, 1935 PT (1) M, F Anagenesia paradoxa Buldowsky, 1935 Palingeniidae
parvus Ecdyonurus parvus Ulmer, 1912 ST (1, 2, 3) M, F Rhithrogena parva (Ulmer, 1912) Heptageniidae Sartori 2014b
patagonica Atalophlebia patagonica Lestage, 1931 HT (1, 3) F Meridialaris patagonica (Lestage, 1931) Leptophlebiidae Peters and Edmunds 1972
pekinensis Caenis pekinensis Malzacher, 2016 HT, PT (1,3) N Caenis pekinensis Malzacher, 2016 Caenidae
pekingensis Baetis pekingensis Ulmer, 1936 HT, PT (1) M, F, SM, SF Alainites pekingensis (Ulmer, 1936) Baetidae Waltz et al. 1994
peruvianus Baetis peruvianus Ulmer, 1920 ST (2) M, SM Andesiops peruvianus (Ulmer, 1920) Baetidae Lugo-Ortiz and McCafferty 1999
pescadori Jubabaetis pescadori Müller-Liebenau, 1980b HT (3) N Jubabaetis pescadori Müller-Liebenau, 1980 Baetidae
peterseni Leptohyphes peterseni Ulmer, 1920 ST (1, 3) M, F, SM, SF Leptohyphes peterseni Ulmer, 1920 Leptohyphidae
petersi Rheobaetis petersi Müller-Liebenau, 1974 HT (1) M, N Heterocloeon petersi (Müller-Liebenau, 1974) Baetidae McCafferty and Provonsha 1975
philippinensis Caenodes philippinensis Ulmer, 1924 ST (1, 2, 3) M Caenis philippinensis (Ulmer, 1924) Caenidae Kluge 1985
pictipennis Ephemera pictipennis Ulmer, 1924 ST (2) M, F Ephemera pictipennis Ulmer, 1924 Ephemeridae
powelli Lachlania powelli Edmunds, 1951 PT (1) M, F, N Lachlania saskatschevanensis Ide, 1941 (syn.) Oligoneuriidae McCafferty 1996
proba Choroterpes proba Ulmer, 1939 ST (1, 3) N Choroterpes proba Ulmer, 1939 Leptophlebiidae
prominens Habrophlebiodes prominens Ulmer, 1939 HT (1, 3) M Habrophlebiodes prominens Ulmer, 1939 Leptophlebiidae
pseudorhodani Baetis pseudorhodani Müller-Liebenau, 1971 HT, PT (1) M, N Baetis pseudorhodani Müller-Liebenau, 1971 Baetidae
puigae Teloganopsis puigae Ubero-Pascal & Sartori, 2009 PT (1) N Teloganopsis puigae Ubero-Pascal & Sartori, 2009 Ephemerellidae
pulcher Afronurus pulcher Ulmer, 1930 PT (1) SM, SF Afronurus collarti (Navás, 1930) (syn.) Heptageniidae Demoulin 1956
purpurata Ephemera purpurata Ulmer, 1920 PT (2) M Ephemera purpurata Ulmer, 1920 Ephemeridae
ranauensis Caenis ranauensis Malzacher, 2015 HT, PT (1, 3) M, F Caenis ranauensis Malzacher, 2015 Caenidae
rhenicola Caenis rhenicola Malzacher, 1976 HT, PT (1) M Caenis pusilla Navás, 1913 (syn.) Caenidae Alba-Tercedor and Malzacher 1986
rossicus Ecdyonurus rossicus Tshernova, 1928b PT (1) M Kageronia fuscogrisea (Retzius, 1783) (syn.) Heptageniidae Puthz 1977b
sauteri Ephemera sauteri Ulmer, 1912 PT (2) M, F Ephemera sauteri Ulmer, 1912 Ephemeridae
scotti Caenis scotti Ulmer, 1930 PLT (1) M, F Caenis scotti Ulmer, 1930 Caenidae
seminiger Simothraulus seminiger Ulmer, 1939 HT (1) M Simothraulus seminiger Ulmer, 1939 Leptophlebiidae Grant and Peters 1993
sexfasciata Atalophlebia sexfasciata Ulmer, 1916 HT (1, 3) M Atalomicria sexfasciata (Ulmer, 1916) Leptophlebiidae Harker 1954; (j.)
sikorai Euthyplocia sikorai Vayssière, 1895 PT (1, 3) N Proboscidoplocia sikorai (Vayssière, 1895) Euthyplociidae Demoulin 1966
sinensis Iron sinensis Ulmer, 1926 PT (2, 3) M Epeorus sinensis (Ulmer, 1926) Heptageniidae Edmunds and Traver 1954
soror Baetis soror Ulmer, 1908 ST (1) M, F Offadens soror (Ulmer, 1908) Baetidae Lugo-Ortiz and McCafferty 1998
sowai Rhithrogena sowai Puthz, 1972 HT (2) M Rhithrogena sowai Puthz, 1972 Heptageniidae
sudanense Centroptilum sudanense Ulmer, 1916 PT (1) M, SM Bugilliesia sudanense (Ulmer, 1916) Baetidae Lugo-Ortiz and McCafferty 1996
sumatrana Baetis sumatrana Ulmer, 1939 ST (1, 3) M, F Baetis sumatrana Ulmer, 1939 Baetidae Müller-Liebenau 1981
sumatranus Ecdyonurus sumatranus Ulmer, 1939 HT (1, 3) F Rhithrogena sumatrana (Ulmer, 1939) Heptageniidae Sartori 2014b
thienemanni Compsoneuriella thienemanni Ulmer, 1939 LT, PLT (1, 2, 3) M, F, N Compsoneuriella thienemanni Ulmer, 1939 Heptageniidae Sartori 2014a
tibiale Cinygma tibiale Ulmer, 1920 PT (1, 3) M Rhithrogena tibiale (Ulmer, 1920) Heptageniidae Tshernova and Belov 1982
tibialis Atopopus tibialis Ulmer, 1920 PT (2, 3) M, F Atopopus tibialis Ulmer, 1920 Heptageniidae
traverae Rheobaetis traverae Müller-Liebenau, 1974 HT (1) N Heterocloeon curiosum (McDunnough, 1923) (syn.) Baetidae McCafferty and Provonsha 1975
triangularis Siphlonurus triangularis Clemens, 1915 PT (1) M, F, N Siphlonurus quebecensis (Provancher, 1878) (syn.) Siphlonuridae McDunnough 1925
truncatus Campsurus truncatus Ulmer, 1920 PT (2) M, F Campsurus truncatus Ulmer, 1920 Polymitarcyidae
ulmeri Asionurus ulmeri Braasch & Soldán, 1986 HT, PT (1, 3) N Asionurus ulmeri Braasch & Soldán, 1986 Heptageniidae
ulmeri Baetis ulmeri Müller-Liebenau, 1981 HT (1, 3) M Labiobaetis ulmeri (Müller-Liebenau, 1981) Baetidae McCafferty and Waltz 1995
ulmeri Behningia ulmeri Lestage, 1930 HT (1) N Behningia ulmeri Lestage, 1930 Behningiidae
ulmeri Dudgeodes ulmeri Sartori, 2008 HT, PT (1, 3) M, SM, F, N Dudgeodes ulmeri Sartori, 2008 Teloganodidae Sartori et al. 2008
Sartori 2014f
ulmeri Trichogenia ulmeri Braasch & Webb, 2006 HT, PT (1, 3) N Trichogenia ulmeri Braasch & Webb, 2006 Heptageniidae
ulmeriana Caenis ulmeriana Malzacher, 2015 HT, PT (1, 3) M, F Caenis ulmeriana Malzacher, 2015 Caenidae
umbrata Teloganella umbrata Ulmer, 1939 HT (1) SF Teloganella umbrata Ulmer, 1939 Teloganellidae
unguicularis Euphyurus unguicularis Ulmer, 1917 PT (1) M Austrophlebiodes unguicularis (Ulmer, 1917) Leptophlebiidae Campbell and Suter 1988
unguiculatus Campsurus unguiculatus Ulmer, 1920 ST (2) M Tortopsis unguiculatus (Ulmer, 1920) Polymitarcyidae Molineri 2010
ussingi Rhithrogena ussingi Esben-Petersen, 1910 PT (2) M Rhithrogena germanica Eaton, 1885 (syn.) Heptageniidae Sowa 1971
verum Pseudocloeon verum Müller-Liebenau, 1982 PT (1, 3) N Liebebiella vera (Müller-Liebenau, 1981) Baetidae Waltz and McCafferty 1987
virens Cloeon virens Klapalek, 1905b ST (2, 3) M, F Cloeon virens Klapalek, 1905 Baetidae
vitellinum Centroptilum vitellinum Ulmer, 1939 HT (2) M Centroptilum vitellinum Ulmer, 1939 Baetidae
winckleri Isonychia winckleri Ulmer, 1939 ST (2) M, F, SF Isonychia winckleri Ulmer, 1939 Isonychiidae (a.)
wui Leptophlebia wui Ulmer, 1936 ST (1) M, F Leptophlebia wui Ulmer, 1936 Leptophlebiidae
wui Caenis wui Malzacher, 2016 HT, PT (1,3) N Caenis wui Malzacher, 2016 Caenidae
Missing material
asiaticum Cinygma asiaticum Ulmer, 1924 ST (2) M Incertae sedis Heptageniidae Tshernova and Belov 1982
klapaleki Pseudocloeon klapaleki Müller-Liebenau, 1982 PT (1) N Liebebiella klapaleki (Müller-Liebenau, 1982) Baetidae Waltz and McCafferty 1987

Acknowledgements

The authors thank Kai Schütte (Hamburg) for his kind support and encouragement for this project. Comments by Luke Jacobus (Columbus, Indiana, USA) helped to improve the manuscript.

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