Research Article |
Corresponding author: Jan Klimaszewski ( jklimaszewski@cfl.forestry.ca ) Academic editor: Adam Brunke
© 2016 Jan Klimaszewski, David J. Larson, Myriam Labrecque, Caroline Bourdon.
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation:
Klimaszewski J, Larson DJ, Labrecque M, Bourdon C (2016) Twelve new species and fifty-three new provincial distribution records of Aleocharinae rove beetles of Saskatchewan, Canada (Coleoptera, Staphylinidae). ZooKeys 610: 45-112. https://doi.org/10.3897/zookeys.610.9361
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One hundred twenty species of aleocharine beetles (Staphylinidae) are recognized in the province of Saskatchewan. Sixty-five new provincial records, including twelve new species and one new North American record, are presented. Oligota inflata (Mannerheim), a Palearctic species, is newly recorded for North America. The following twelve species are described as new to science: Acrotona pseudopygmaea Klimaszewski & Larson, sp. n., Agaricomorpha pulchra Klimaszewski & Larson, sp. n. (new genus record for Canadian fauna), Aleochara elisabethae Klimaszewski & Larson, sp. n., Atheta (Dimetrota) larsonae Klimaszewski & Larson, sp. n., Atheta (Microdota) pseudopittionii Klimaszewski & Larson, sp. n., Atheta (Microdota) spermathecorum Klimaszewski & Larson, sp. n., Atheta (sensu lato) richardsoni Klimaszewski & Larson, sp. n., Brachyusa saskatchewanae Klimaszewski & Larson, sp. n., Dochmonota langori Klimaszewski & Larson, sp. n., Dochmonota simulans Klimaszewski & Larson, sp. n., Dochmonota websteri Klimaszewski & Larson, sp. n., and Oxypoda domestica Klimaszewski & Larson, sp. n. Colour images of habitus and black and white images of the median lobe of the aedeagus, spermatheca, and tergite and sternite VIII are presented for all new species, Oligota inflata Mannerheim and Dochmonota rudiventris (Eppelsheim). A new synonymy is established: Tetralina filitarsus Casey, syn. n. = Tetralina helenae Casey, now placed in the genus Brachyusa Mulsant & Rey.
Coleoptera , rove beetles, Staphylinidae , new distribution records, new species, Canada, Saskatchewan
Aleocharinae beetles are the most species-rich subfamily of rove beetles (Staphylinidae) with 515 species recorded in the most recent catalog of the Coleoptera of Canada (
Almost all specimens in this study were dissected to examine the genital structures. Extracted genital structures were dehydrated in absolute alcohol, mounted in Canada balsam on celluloid micro-slides, and pinned with the specimen from which they originated. Images of the entire body and the genital structures were taken using an image processing system (Nikon SMZ 1500 stereoscopic microscope; Nikon Digital Camera DXM 1200F, and Adobe Photoshop software).
Morphological terminology mainly follows that used by
Tribes, genera and species within genera are arranged alphabetically in the text and in the Table
Species of Aleocharinae recorded from Saskatchewan and their provincial and territorial distribution within Canada. Provinces and territories in bold denote new records given in the present publication. Species marked with (†) indicate adventive species and species marked with (*) are Holarctic.
ALEOCHARINI | |
Aleochara assiniboin Klimaszewski | BC, MB, ON, SK, YT |
Aleochara bilineata Gyllenhal† | AB, BC, MB, NB, NF, NS, ON, PE, QC, SK; USA: New England states |
Aleochara bimaculata Gravenhorst | AB, BC, LB, MB, NB, NF, NS, ON, QC, SK, NT; USA: widespread |
Aleochara elisabethae Klimaszewski & Larson, sp. n. | SK |
Aleochara gracilicornis Bernhauer | BC, MB, NB, NS, NT, ON, QC, SK; USA: widespread |
Aleochara inexpectata Klimaszewski | NB, NS, ON, QC, SK; USA: MI, WI |
Aleochara lacertina Sharp | AB, BC, MB, NB, NF, NS, ON, QC, SK; USA: widespread |
Aleochara laramiensis (Casey) | BC, SK; USA: CO, WY |
Aleochara lata Gravenhorst† | BC, MB, ON, QC, SK, YT; USA: widespread |
Aleochara rubricalis (Casey) | BC, ON, SK; USA: CA, AZ |
Aleochara sekanai Klimaszewski | AB, LB, MB, NB, NT, ON, SK, YT; USA: AK |
Aleochara speculicollis Bernhauer | AB, ON, QC, SK: USA: CA, CO, AZ, MI, NV, TX |
Aleochara suffusa (Casey) | AB, BC, MB, QC, SK; USA: AK, AZ, CO, NM, WY |
Aleochara tahoensis Casey | AB, BC, MB, NB, NS, NT, ON, SK, YT; USA: CA, CO, MT, NH, NM, NV, OR, WA |
Aleochara verna Say | AB, BC, LB, MB, NB, NF, NS, ON, PE, QC, SK, YT; USA: widespread including AK |
Aleochara villosa Mannerheim† | AB, BC, NB, QC, SK; USA: AK, CA, OR, WA |
Tinotus morion (Gravenhorst) † [now regarded as Aleochara] | AB, BC, NB, NF, NS, ON, QC, SK; USA: CT, NV |
ATHETINI | |
Acrotona pseudopygmaea Klimaszewski & Larson, sp. n. | SK |
Acrotona recondita (Erichson) | SK; USA: AR, CA, NH, NV, NY, PA |
Acrotona subpygmaea (Bernhauer) | NB, NS, ON, SK |
Amischa analis (Gravenhorst) † | LB, NB, NF, NS, ON, QC, PE, SK |
Atheta celata (Erichson) * | BC, NB, NF, NS, QC, SK; USA: AK |
Atheta crenuliventris Bernhauer | LB, NB, NF, ON, QC, SK |
Atheta dadopora C.G. Thomson * | AB, BC, LB, NB, NF, NS, ON, PE, SK, YT; USA: AK, NY, PA, RI |
Atheta districta Casey | AB, BC, LB, NB, NF, NS, ON, QC, SK |
Atheta fanatica Casey | AB, BC, LB, NB, NS, QC, SK, YT; USA: AK, NV |
Atheta frosti Bernhauer | BC, LB, NB, NS, ON, QC, SK |
Atheta graminicola (Gravenhorst) * | AB, BC, LB, MB, NB, NF, NT, ON, QC, SK, YT; USA: AK, OR |
Atheta klagesi Bernhauer | AB, BC, NB, NF, NS, ON, PE, QC, SK, YT; USA: IA, ME, MN, NJ, NY, PA |
Atheta larsonae Klimaszewski & Larson, sp. n. | SK |
Atheta longicornis (Gravenhorst) † | BC, NB, NF, NS, QC, SK; USA: CA, MN |
Atheta nigra (Kraatz) † | SK |
Atheta platonoffi Brundin* | AB, BC, LB, NB, NF, NS, ON, SK, YT; USA: AK |
Atheta prudhoensis (Lohse) | BC, LB, NB, NF, NS, ON, QC, SK, YT; USA: AK, VT |
Atheta pseudometlakatlana Klimaszewski & Godin | YT, SK |
Atheta pseudopittionii Klimaszewski & Larson, sp. n. | SK |
Atheta pseudoschistoglossa Klimaszewski & Webster | BC, NB, SK; USA: AK |
Atheta recondita (Erichson) | SK; USA: AR, CA, NH, NV, PA |
Atheta remulsa Casey | AB, BC, LB, NB, NF, NS, ON, QC, SK, YT |
Atheta riparia Klimaszewski & Godin | SK, YT |
Atheta richardsoni Klimaszewski & Larson, sp. n. | SK |
Atheta spermathecorum Klimaszewski & Larson, sp. n. | SK |
Atheta strigosula Casey | BC, LB, NB, NF, ON, QC, SK, YT; USA: NY |
Atheta subsinuata (Erichson) † | YT, SK |
Atheta terranovae Klimaszewski & Langor | LB, NB, NF, ON, SK, YT |
Atheta ventricosa Bernhauer | AB, BC, LB, NB, NF, NS, ON, QC, SK, YT; USA: AK, DC, NC, NJ, NY, PA, VT |
Dinaraea angustula (Gyllenhal) † | AB, LB, NB, NF, NS, ON, PE, QC, SK, YT; USA: CA, NY |
Dinaraea pacei Klimaszewski & Langor | AB, BC, LB, NB, QC, SK, YT; USA: AK |
Dochmonota langori Klimaszewski & Larson, sp. n. | SK |
Dochmonota simulans Klimaszewski & Larson, sp. n. | SK |
Dochmonota websteri Klimaszewski & Larson, sp. n. | SK |
Earota dentata (Bernhauer) | AB, BC, MB, NB, NF, NS, ON, QC, SK, YT; USA: AK |
Lypoglossa franclemonti Hoebeke | AB, MB, NB, NF, NS, NT, ON, QC, SK, YT; USA: NY, VT |
Mocyta breviuscula (Mäklin) | AB, BC, LB, NB, NF, NS, ON, QC, SK, YT; USA: AK |
Mocyta discreta (Casey) | ON, QC, SK; USA: IA, MN |
Mocyta spahgnorum Klimaszewski & Webster | NB, NF, ON, QC, SK |
Nehemitropia lividipennis (Mannerheim) † | NB, NF, NS, ON, PE, QC, SK; USA: CA, LA, MA, MN, NE, NM, NY, PA, VT, TX |
Philhygra botanicarum (Muona) * | BC, LB, NB, NF, NS, ON, SK, YT |
Philhygra falcifera Lohse | MB, SK |
Philhygra jarmilae Klimaszewski & Langor | NB, NF, ON, SK, YT |
Philhygra ripicoloides Lohse | NF, NT, SK, YT |
Philhygra rostrifera Lohse | LB, NT, SK, YT; USA: AK |
Philhygra sinuipennis Klimaszewski & Langor | NB, LB, NF, SK, YT |
Philhygra subpolaris (Fenyes) | AB, SK; USA: AZ |
Philhygra terrestris Klimaszewski & Godin | NB, SK, YT |
Schistoglossa blatchleyi (Bernhauer & Scheerpeltz) | MB, NB, NT, ON, QC, SK, YT; USA: AK, IN |
Seeversiella globicollis (Bernhauer) | AB, BC, NB, NF, NS, ON, QC, SK; USA: AZ, CO, ID, MN, MT, NH, SD, WI |
Strigota ambigua (Erichson) | LB, NB, NS, NF, ON, PE, SK, YT; USA: CA, CO, CT, IA, KS, MO, NC, NJ, NM, NY, TX |
Strigota obscurata Klimaszewski & Brunke | NB, ON, SK |
AUTALIINI | |
Autalia rivularis (Gravenhorst) † | AB, BC, LB, NB, NF, NS, ON, QC, SK; USA: CA, MI, MN, NH, NY, OR |
FALAGRINI | |
Falagria caesa Erichson† | AB, NB, ON, QC, SK; USA: MA to VA, UT |
Falagria dissecta Erichson | AB, BC, MB, NB, NS, ON, QC, SK; USA: widespread |
Myrmecocephalus arizonicus (Casey) | AB, BC, SK |
GYMNUSINI | |
Gymnusa campbelli Klimaszewski | MB, NB, NF, NT, ON, QC, SK, YT; USA: AK |
HOMALOTINI | |
Agaricochara pulchra Klimaszewski & Larson, sp. n. | SK |
Gyrophaena affinis Mannerheim | BC, MB, NB, NF, NS, ON, QC, SK; USA: widespread |
Gyrophaena criddlei Casey | LB, MB, NB, ON, SK, YT |
Gyrophaena insolens Casey | BC, LB, MB, NB, NF, ON, SK; USA: MI |
Gyrophaena keeni Casey | AB, BC, LB, NB, NF, ON, QC, SK, YT; USA: FL, MA, MT, NH, NY, TN, WA |
Gyrophaena lobata Casey | NB, SK; USA: DC, IL, IN, KS, MI, WI |
Gyrophaena uteana Casey | AB, BC, NB, ON, QC, SK; USA: CA, CO, UT |
Gyrophaena subnitens Casey (NCR) | MB, SK; USA: IL, KS, ME, MN, MO, WI |
Leptusa gatineauensis Klimaszewski & Pelletier | AB, BC, NB, NF, NS, ON, QC, SK |
HYPOCYPHTINI | |
Cypha crotchi (Horn) | AB, BC, SK |
Cypha inexpectata Klimaszewski & Godin | ON, YT, SK |
Oligota inflata (Mannerheim)† (NPR, NCR, NAR) | SK |
LOMECHUSINI | |
Xenodusa reflexa (Walker) | AB, BC, MB, NB, NS, QC, ON, SK |
Zyras obliquus (Casey) | AB, BC, MB, NB, NF, NS, ON, QC, SK; USA: MI, MO, NH, NY, OR |
MYLLAENINI | |
Myllaena arcana Casey | AB, LB, NB, NF, NS, ON, QC, SK; USA: AL, FL, IA, IL, MA, NH, NJ |
Mylaena insomnis Casey | AB, BC, LB, MB, NB, NF, NS, NT, ON, QC, SK, YT; USA: AK, ID, MA, MN, WI |
OXYPODINI | |
Cratarea suturalis (Mannerheim) † | BC, LB, NB, NS, ON, SK; USA: IL, MA, MO, PA, SC, VA, VT |
Devia prospera (Erichson) * | AB, BC, LB, MB, NB, NT, ON, SK, YT; USA: AK, CO, MI, MN, NM, OR, SD, UT, WA, WY |
Gnathusa eva Fenyes | AB, BC, SK, YT; USA: CA |
Hylota ochracea Casey | NB, NS, NT, ON, QC, SK; USA: NY |
Ocyusa canadensis Lohse | NB, NF, ON, SK, YT; USA: AK |
Oxypoda canadensis Klimaszewski | AB, MB, LB, NF, NT, ON, QC, SK, YT; USA: AK |
Oxypoda demissa Casey | LB, NB, NF, NS, ON, QC, SK, YT |
Oxypoda domestica Klimaszewski & Larson, sp. n. | SK |
Oxypoda grandipennis (Casey) | AB, BC, LB, NB, NF, NS, ON, QC, SK, YT; USA: AK, NH |
Oxypoda irrasa Mäklin | AB, SK, YT; USA: AK, OR |
Oxypoda lacustris Casey | AB, BC, LB, MB, NB, NF, NS, NT, ON, QC, SK, YT; USA: AK |
Oxypoda manitobae Casey | BC, MB, SK; USA: CO |
Oxypoda orbicollis Casey | AB, LB, NB, NS, ON, QC, SK, YT; USA: WI |
Oxypoda pseudolacustris Klimaszewski | AB, NB, NF, NS, ON, QC, SK |
Parocyusa fuliginosa (Casey) | LB, ON, SK; USA: MA, NC, PA |
Tachyusa obsoleta Casey | BC, NB, SK |
PLACUSINI | |
Placusa incompleta Sjöberg † | AB, BC, NB, NF, NS, ON, QC, SK; USA: WA |
Placusa pseudosuecica Klimaszewski | AB, BC, ON, QC, SK |
Placusa tachyporoides (Waltl) † | AB, BC, NB, NS, ON, QC, SK; USA: CA, MA |
Placusa tacomae Casey | AB, BC, NB, NF, NS, NT, ON, QC, SK, YT; USA: AZ, MA, WA, WI |
Placusa vaga Casey | BC, NB, NS, NT, ON, QC, SK, YT; USA: CA |
SILUSINI | |
Silusa californica Bernhauer | AB, BC, NB, NF, NS, NT, QC, ON, SK, YT; USA: AK, CA, MN |
TACHYUSINI | |
Brachyusa helenae (Casey) | LB, NB, NF, NT, ON, SK, YT; USA: AK, MT |
Brachyusa saskatchewanae Klimaszewski & Larson, sp. n. | SK |
Gnypeta caerula (C.R. Sahlberg) * | AB, BC, LB, MB, NB, NF, NS, NT, ON, PE, QC, SK, YT; USA: AK |
Gnypeta carbonaria (Mannerheim) | AB, MB, NB, NF, NT, ON, QC, SK; USA: AK |
Gnypeta dentata Klimaszewski | AB, NT, SK |
Gnypeta minuta Klimaszewski & Webster | NB, SK |
Gnypeta saccharina Klimaszewski & Webster | NB, SK |
Gnypeta sellmani Brundin | LB, MB, NF, NT, QC, SK, YT; USA: AK |
Almost all collections reported here were made in southwestern Saskatchewan and adjacent Alberta. This area is in the Mixed Grassland and Cypress Upland Ecoregions of the Prairies Ecozone (
Other habitats within the area from which aleocharines have been collected include sand hills and saline ponds of closed drainage basins, both of which occur mainly to the north of the Cypress Hills, stream margins, and springs and fens that are common in the Cypress Hills. Considerable collecting has been done around large reservoirs. Accumulated plant material along the reservoir water lines (wrack) is often rich in beetles but windward shores (the lee shore of mariners) are often rich collecting sites as flying insects that fall into water are blown onto these shores and can sometimes be found in large numbers pulling themselves up onto the beach. Such insects are referred to as occurring in drift. Species found in wrack may in fact be in their normal habitat, but those recorded from drift are probably vagrants, but they do indicate presence and time of year of dispersal.
The low annual precipitation in the region means that a state of drought or near drought occurs frequently. Aleocharines occur mainly in moist environments, thus the majority of Mixed Grassland collections are from sites with moisture such as margins of water bodies or from moist habitats such as carrion and manure (which is very abundant due to the high populations of cattle). Carrion and manure are rich staphylinid habitat but they promote the widespread synanthropic species and a few of our new records come from these habitats. Mushrooms and other fungi, especially as they age and decay, are rich habitats but again irregularity in precipitation means that occurrence and duration of such habitats is very unpredictable over the season and from year to year. Higher levels of precipitation and lower evapotranspiration in the Cypress Uplands produce a wider and more consistent array of moist habitats and this is where we found the richest aleocharine fauna.
BGC Benoit Godin collection, Whitehorse, Yukon Territory, Canada
LFC Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, R. Martineau Insectarium, Quebec City, Quebec, Canada
DLC David Larson collection, Maple Creek, Saskatchewan, Canada
AB – Alberta
BC – British Columbia
LB – Labrador
MB – Manitoba
NB – New Brunswick
NF – Newfoundland
NS – Nova Scotia
NT – Northwest
ON – Ontario
PE – Prince Edward Island
QC – Quebec
SK – Saskatchewan
Territories YT – Yukon TerritoryNU – Nunavut
USA state abbreviations follow those of the US Postal Service.
Our knowledge of the diversity and distribution of Aleocharinae in Canada has increased rapidly over the last two decades (
In Canada, the Maritime provinces (NB, NS, NF, PE), and the YT are so far the best-studied regions of the country in terms of the aleocharine fauna (
However, the large majority of central, western and northern Canada remains poorly studied. Large numbers of aleocharines (and other staphylinids) have been collected over the last 25 years as a result of numerous trapping studies in forests, native grasslands, agricultural lands, and wetlands, especially in Alberta (
(female). Canada, Saskatchewan, Bowie Ranch, 20 km NW Piapot, sand dunes, 29-V-2008, D. Larson (LFC). Paratype. Canada, Alberta, Empress, Alberta – Saskatchewan border, 5-VIII-1981, Lot 1, B.F. & J.L. Carr (
This species is named for Dr. Élisabeth Gauthier, research director at LFC, for her continuous support of beetle biodiversity research in Canada.
Body compact, narrowly oval (Fig.
Aleochara (Echochara) elisabethae Klimaszewski & Larson, sp. n.: 1 habitus in dorsal view 2 median lobe of aedeagus in lateral view 3 male tergite VIII 4 male sternite VIII 5 female tergite VIII 6 female sternite VIII 7 spermatheca. Scale bar for habitus = 1 mm, and the remaining scale bars = 0.2 mm.
This species is readily distingushed from remaining Nearctic species of subgenus Echochara by its strongly transverse and orange pronotum (dark brown or black in remaining species), arcuate tubus of median lobe of aedeagus with sharp apex (Fig.
This species is known from the type localities in AB and SK.
The female holotype was captured on a dead ground squirrel in sand dunes. The male was collected in August from unspecified habitat. Species of subgenus Echochara are known from caves and animal burrows (
(for diagnosis and illustrations, see Klimaszewski et al. 1984)
Origin | Nearctic |
Distribution | Canada: NB, NS, ON, QC, SK. USA: MI, WI |
New provincial records | CANADA, Saskatchewan: Larson Ranch, Hwy 21, 16 km S Maple Creek, 20-X-2014, in dry polypore fungus, D. Larson (DLC) 1 female |
References |
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In Saskatchewan, one female was captured in dry polypore fungus in October, and this constitutes the westernmost distribution record for this species. In NB, Aleochara inexpectata was collected from fresh moose dung in an eastern white cedar swamp and in decaying sea wrack resting on vegetation on the upper margin of a salt marsh. Adults were collected during May and June (
(for diagnosis and illustrations, see Klimaszewski et al. 1984)
Origin | Nearctic |
Distribution | Canada: BC, ON?, SK. USA: AZ, CA |
New provincial records | CANADA, Saskatchewan: Larson Ranch, Hwy 21, 16 km S Maple Creek: 20-V-2008, D. Larson (LFC) 1 male; 25-VI-2008, carrion trap, D. Larson (DLC) 3 males, 1 female; 8-IV-2005, D. Larson (LFC) 1 female |
References |
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In Saskatchewan, specimens were collected from March through June, several adults were captured from carrion trap. Elsewhere, one specimen was taken from a mouse nest and other specimens were collected from February to October (
(for diagnosis and illustrations, see Klimaszewski et al. 1984)
In Saskatchewan, one female was captured in June from unspecified habitat.
We tentatively associate the SK specimen with this species because it is missing the spermatheca.
(for diagnosis and illustrations, see Klimaszewski et al. 1984)
Origin | Nearctic |
Distribution | Canada: AB, BC, MB, QC, SK. USA: AK, AZ, CO, NM, WY |
New provincial records | CANADA, Saskatchewan: Larson Ranch, Hwy 21, 16 km S Maple Creek: 27-V-2008, D. Larson (DLC) 1 female; 1-VI-2010, D. Larson (DLC, LFC) 2 females; 24-IX-2008, D. Larson (DLC) 1 female; 25-VI-2008, carrion trap, D. Larson (DLC) 1 female; 14-IX-2008, D. Larson (DLC) 1 sex undetermined; Cypress Lake, E dam, wind-drift, 9-V-2012, D. Larson (DLC) 1 female; Harris Res., 10 km S Maple Creek, 12-VI-2013, wind-drift, D. Larson (DLC) 1 male. |
References |
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In Saskatchewan, one female was captured in a carrion trap and one from wind-drift. Elsewhere, specimens were found under rocks in a high altitude meadow and some from AB were reared in laboratory (
The SK specimens are darker and have only the central part of elytra reddish and the rest of the body piceous whereas the typical form of this species has the entire elytra orange or reddish-brown. Pubescence and punctation pattern and the genitalia of SK specimens are identical to the typical form with orange or reddish elytra.
(for diagnosis and illustrations, see Klimaszewski et al. 1984)
Origin | Palaearctic, adventive in Canada |
Distribution | Canada: AB, BC, NB, QC, SK. USA: AK, CA, OR, WA |
New provincial records | CANADA, Saskatchewan: Larson Ranch, Hwy 21, 16 km S Maple Creek: 21-III-2007, sheep barn window, D. Larson (DLC) 2 males; 1-IV-2013, D. Larson (LFC) 1 female; 14-IV-2012, D. Larson (LFC) 1 male; 27-VII-2012, D. Larson (DLC) 1 female; 17-IX-2012, compost, D. Larson (DLC) 1 male. |
References |
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In SK, 2 males were captured from a sheep barn window, and one male was found in compost. SK specimens were collected in March, April, July and September. In New Brunswick, A. villosa was collected from the nest contents of a great horned owl, Bubo virginianus (Gmelin) (
Canada, Saskatchewan, Larson Ranch, Hwy 21, 16 km S Maple Creek, 24-VII-2010, sifted from old mouldy alfalfa hay, D. Larson (LFC). Paratypes. 1 male and 1 female, with same label and collection data as the holotype (
The name of this species derives from the Latin participle pygmaea-, meaning small, and the prefix pseudo-, false. The genital structures of this species are similar to those of Palaearctic Acrotona pygmaea (Gravenhorst).
Body narrowly elongate, moderately convex, uniformly dark brown to almost black except for paler legs (Fig.
Acrotona pseudopygmaea Klimaszewski & Larson, sp. n.: 8 habitus in dorsal view 9 median lobe of aedeagus in lateral view, and 10 in dorsal view 11 male tergite VIII 12 male sternite VIII 13 female tergite VIII 14 female sternite VIII 15 spermatheca. Scale bar for habitus = 1 mm, and the remaining scale bars = 0.2 mm.
Distinguished from all other Acrotona by the shape of median lobe of aedeagus with apex turned dorsally in lateral view (Fig.
This species is known only from the type locality in SK.
The type specimens were sifted from old mouldy alfalfa hay.
This species is similar to Palaearctic A. pygmaea (Gravenhorst) from which it differs by subquadrate antennal articles VI-X, by apex of tubus of median lobe of aedeagus more angular, female sternite VIII emarginated apically and spermatheca with much longer and broadly looped stem. It is also genitally similar to Nearctic Acrotona actuella (Casey) and A. egregiella (Casey), from which it differs by straight and not sinuate ventral margin of tubus of median lobe of aedeagus, by straight and not sinuate basal margin of male tergite VIII, and by differently shaped spermatheca with posterior loop of stem sinuate.
(for diagnosis and illustrations, see
Origin | Nearctic |
Distribution | Canada: NB, NS, ON, SK |
New provincial records | CANADA, Saskatchewan: Larson Ranch, Hwy 21, 16 km S Maple Creek, 5-6-VI-2013, maple litter, D. Larson (DLC) 1 female; 20-XI-2014, sifting willow leaf litter, D. Larson (DLC) 1 female. |
References |
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In SK, one female was captured from maple (Acer negundo) litter and one from willow (Salix spp.) leaf litter in June and October, respectively. In NB, Acrotona subpygmaea was found in litter of a variety of forest types and in wetlands including swamps, sphagnum bog, marshes and river margins. Specimens have also been taken from gilled mushroom and under bark (
(for diagnosis and illustrations, see
Origin | Palaearctic, adventive in Canada |
Distribution | Canada: LB, NB, NF, NS, ON, QC, PE, SK |
New provincial records | CANADA, Saskatchewan: Belanger Creek, Frenchman Valley, 18-X-2014, D. Larson (DLC) 1 female. |
References |
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In SK, one female was captured in October by sifting leaf litter along a creek. In NL, adults were collected in pitfall traps in agricultural fields, an urban field and on coastal sand dunes amidst vegetation, and the activity period was June to September (
(for diagnosis and illustrations, see
Origin | Nearctic |
Distribution | Canada: LB, NB, NF, ON, QC, SK |
New provincial records | CANADA, Saskatchewan, Larson Ranch, Hwy 21, 16 km S Maple Creek: 1-IX-2012, compost, D. Larson (DLC) 1 male; 8-IX-2012, compost, D. Larson (DLC) 1 male; Cypress Lake, E end, 31-VII-2012, sifting wrack, D. Larson (DLC) 1 female; Swift Current Cr., 28-VIII-2011, D. Larson (DLC) 1 female; Prince Albert, 53.9804, 106.2800, 532 m, 4-VI-2013, sand beach, sifting debris, B. Godin & D. Horwood (BGC, LFC) 2 males, 1 female. |
References |
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In SK, two males were found in compost in September, one female in wrack on lakeshore in July, and one female from unknown habitat in August. In NF, adults were collected from May to August in carrion-baited pitfall traps and flight intercept traps in conifer-dominated and mixedwood forests, and on the coastal barrens of southeastern LB (
(for diagnosis and illustrations, see
Origin | Nearctic |
Distribution | Canada: AB, BC, LB, NB, NF, NS, ON, QC, SK |
New provincial records | CANADA, Saskatchewan, Cypress Hills Park, Center Block: Lodgepole Trail, 21-VIII-2013, dry and decaying mushrooms, D. Larson (DLC) 1 male; Highland Trail, 13-IX-2012, sifting spruce litter, D. Larson (DLC) 1 male. |
References |
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In SK, one male was captured from dry and decaying mushroom, and another from spruce litter in September. In NF, adults were collected from June to August in carrion-baited pitfall traps and flight intercept traps in conifer-dominated and mixedwood forests, and on coastal barrens (
(for diagnosis and illustrations, see
Origin | Nearctic |
Distribution | Canada: SK, YT |
New provincial records | CANADA, Saskatchewan, Cypress Hills Park, Center Block: Loch Lomond, 19-IX-2014, decaying mushrooms, D. Larson (DLC) 1 male, 1 female; 7-IX-2014, spruce-aspen, D. Larson (DLC) 1 female; fire guard, 10-IX-2013, decaying mushrooms, D. Larson (LFC) 1 male; Sucker Creek, 23-VI-204, aspen woodland, bracket gilled fungi, D. Larson (DLC) 1 male. |
References |
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In SK, specimens were collected from decaying mushrooms, bracket/gilled fungi, in spruce-aspen and aspen woodland forests. In YT adults were captured in June, July, and August at an elevation of 772 m in a white spruce and mixed white spruce-lodgepole pine forests (
Canada, Saskatchewan, Royal Edward Road, 25 km NW Maple Creek, 5-VI-2011, D. Larson (LFC).
The name of this species is dedicated to R.I. Larson. Ruby I. Larson was a geneticist at the Agriculture Canada Research Station, Lethbridge, who worked on wheat genetics. She was very active in promoting science and from 1958 to 1973 ran a Science Club for Junior High and High School age children. Members of this club went on to a variety of professional careers, including three (DJL included) who became professional entomologists. Her love of learning and science was infectious and her support and encouragement were major factors in our career choices. She taught us the joy and personal rewards of following one’s curiosity.
Body narrowly elongate, slightly flattened (particularly on elytra), uniformly dark brown, almost black except for paler, light brown sutural section of elytra and legs (Fig.
Distinguished from all other Atheta (Dimetrota) by narrow head and pronotum, broad and short elytra, strongly glossy integument, and the shape of median lobe of aedeagus with apex produced ventrally in lateral view (Fig.
This species is known only from the type locality in SK.
The holotype was captured in June from unspecified habitat.
This species is superficially similar to Nearctic Atheta (D.) peticapensis Klimaszewski & Webster, with which it shares similar body proportions and enlarged bulbus of median lobe of aedeagus. However, these differences may not necessarily indicate a close relationship between these species.
(for diagnosis and illustrations, see
Origin | Nearctic |
Distribution | Canada: BC, LB, NB, NF, ON, QC, SK, YT; USA: AK, NY |
New provincial records | CANADA, Saskatchewan: Cypress Hills Park, Center Block, fire guard, 8-VIII-2013, gilled mushrooms, D. Larson (DLC) 1 male, 2 females; Lodgepole Trail, 21-VIII-2013, dry and decaying mushrooms, D. Larson (DLC) 1 female. |
References | Casey 2010, |
In SK, several females were found in dry and decaying mushrooms in August. In NF, adults were collected from June to October in carrion-baited and unbaited pitfall traps and in flight intercept traps in many forest types (coniferous, mixedwood and deciduous), and some adults were found in rotting mushrooms in forests (
(for diagnosis and illustrations, see
Origin | Nearctic |
Distribution | Canada: LB, NB, NF, ON, QC, SK, YT |
New provincial records | CANADA, Saskatchewan: Cypress Hills Park, Center Block: fire guard, 18-VIII-2014, old polypore fungus on dead lodgepole pine stump, D. Larson (DLC) 2 males; 7-IX-2014, spruce-aspen, D. Larson (DLC) 1 male, 1 female. |
References |
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This species is frequently associated with forest mushrooms. In SK, specimens were captured from an old polypore fungus on dead lodgepole pine stump, and in spruce-aspen forest, in August and September. In NF, adults were collected from June to August in carrion-baited and unbaited pitfall traps and in flight intercept traps in many forest types (coniferous, mixedwood and deciduous), and some adults were found in rotting mushrooms in forests (
Canada, Saskatchewan, Larson Ranch, Hwy 21, 16 km S Maple Creek, 7-IX-2010, ex Lepiota rhacodes, D. Larson (LFC). Paratypes. Canada, Saskatchewan, Larson Ranch, Hwy 21, 16 km S Maple Creek: 25-VI-2008, carrion trap, D. Larson (
The species name pseudopittionii derived from the prefix pseudo- (false) and the specific name of European species Atheta pittionii Scheerpeltz, to which it is similar externally and has similar genitalia.
Body narrowly subparallel (Fig.
Atheta (Microdota) pseudopittionii Klimaszewski & Larson, sp. n.: 21 habitus in dorsal view 22 median lobe of aedeagus in lateral view, and 23 in dorsal view 24 male tergite VIII 25 male sternite VIII 26 female tergite VIII 27 female sternite VIII 28 spermatheca in lateral view;. Scale bar for habitus = 1 mm, and the remaining scale bars = 0.2 mm.
This species is very similar to European A. pittionii Scheerpeltz, from which it differs by broader and more elongate elytra, larger bulbus of median lobe of aedeagus in dorsal view (Fig.
Adults are known only from SK.
Most adults of this species were collected from Shaggy parasol mushrooms, Chlorophyllum rhacodes (=Lepiota rhacodes), from unspecified mushrooms, and from carrion.
(for details and body image, see
Origin | Nearctic |
Distribution | Canada: SK, YT |
New provincial records | CANADA, Saskatchewan: Cypress Hills Park, Center Block: fire guard, Sucker Creek, 23-VI-2014, aspen woodland bracket/gilled fungi, D. Larson (DLC) 1 male; 7-IX-2014, spruce-aspen, D. Larson (DLC) 1 male. |
References |
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One SK male was captured in bracket/gilled fungi in aspen woodland in June, and the other from spruce-aspen woodland in September. In YT, two males were captured by sifting litter in mixed aspen and white spruce forest in September, and one female was found on a mushroom in August (
(female). Canada, Saskatchewan, Larson Ranch, Hwy 21, 16 km S Maple Creek, 8-VI-2014, D. Larson (LFC). Paratypes. Canada, Saskatchewan, Larson Ranch, Hwy 21, 16 km S Maple Creek: 29-V-2012 (LFC) 1 female; 30-V-2014, D. Larson (
The species name spermathecorum is derived from the name of spermatheca in reference to unusually shaped capsule of the spermatheca of this species.
Body narrowly subparallel (Fig.
It is distinguished from all other Nearctic species of Atheta, subgenus Microdota, by the unique shape of spermatheca bearing bulbus apical projection on top of capsule (Fig.
Adults are known from SK and AB.
Females were captured in March (Alberta), May and July (Saskatchewan): one was found in a decaying polypore mushroom and one was found in lake drift in May.
(for details and body image, see
Origin | Palaearctic, adventive in Canada |
Distribution | Canada: SK , YT |
New provincial records | CANADA, Saskatchewan, Larson Ranch, Hwy 21, 16 km S Maple Creek, 1-VI-2011, D. Larson (DLC) 1 female; 27-VI-2010, old wet alfalfa hay with Coprinus, D. Larson (DLC, LFC) 3 females; 24-VII-2010, sifted from old mouldy alfalfa hay, D. Larson (DLC) 1 female; 1-IX-2012, compost, D. Larson (LFC) 1 female. |
References |
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Like many introduced species, A. subsinuata appears to be synanthropic, as all collections have been made from artificial habitats. The Saskatchewan specimens were sifted from old mouldy alfalfa hay in June and July, and one female was taken in September from compost. In YT, four specimens were captured in a compost pile in September 2005 (
(for details and illustrations, see
Origin | Nearctic |
Distribution | Canada: BC, LB, NB, NS, ON, QC, SK; USA: MA, NC, NH, NY, PA, RI, VT |
New provincial records | CANADA, Saskatchewan, Cypress Hills, Center Block, Lake, Lodgepole Trail, 24-IX-2014, decaying mushrooms, D. Larson (DLC) 1 female. |
References |
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The SK female was captured in decaying mushrooms in September. In LB, adults were abundant in pitfall traps during July and August in an open spruce forest with sandy soil and Cladina lichen cover, and a few adults were captured using pitfall traps in a birch-dominated forest (
The following species have uncertain subgeneric affiliation in the large and diverse genus Atheta. Some of the species belong to a group of species described in Europe by
(for details, genitalia and body image, see
Origin | Nearctic |
Distribution | Canada: NB, SK |
New provincial records | CANADA, Saskatchewan: Cypress Hills Park, Sucker Creek, 21-VIII-2012, aspen-pine litter, D. Larson (DLC) 1 male; Cypress Hills Park, Center Block, Lodgepole Trail, 18-IX-2012, pine-spruce litter near stream, D. Larson (DLC) 1 female; Belanger Creek, Frenchman Valley, 18-X-2014, D. Larson (DLC) 1 male. |
References |
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The SK specimens were captured from aspen/pine litter and pine/spruce litter in August through October. In NB, most adults of A. pseudoschistoglossa were found in or near wetland habitats including among cobblestones, drift material, and flood debris along river margins, moist leaves along vernal pond margin in a silver maple swamp, in leaf litter and moss along brook margins in alder swamps, and in litter at base of red maple, in Carex hummock in Carex marshes, in leaf litter in a red oak forest near seasonally flooded marsh, in a salt marsh, in marsh litter in a Carex-sedge marsh, and in litter and sphagnum at the base of a tree in a marsh (
(for details and illustrations, see
Origin | Nearctic |
Distribution | Canada: AB, BC, LB, NB, NF, NS, ON, QC, SK, YT |
New provincial records | CANADA, Saskatchewan: Cypress Hills Park, Center Block, Lodgepole Trail, 21-VIII-2013, dry and decaying mushrooms, D. Larson (DLC) 1 female. |
References |
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In SK one female was captured from dry and decaying mushrooms. In NL, adults were collected from June to September using unbaited and carrion-baited pitfall traps and flight intercept traps in many forest types (deciduous, mixedwood, coniferous, riparian), and also in rotting mushrooms in forests (
Canada, Saskatchewan, Hwy 21, 20 km N Maple Creek, 25-VI-2010, Gramma-stipa pasture, Richardson ground squirrel burrow, D. Larson (LFC). Paratype. Canada, Saskatchewan, Grassland National Park, W Block Larson’s Prairie Dog colony, 11-VI-2009, D. Larson (LFC) 1 female.
This species name is derived from the surname of Sir John Richardson, the surgeon-naturalist who participated in 19th century British naval expeditions to the arctic coast of “British North America”, now Canada. In 1820 he discovered a new species of ground squirrel along the Saskatchewan River, which was later named after him as Urocitellus richardsonii. The holotype of Atheta richardsoni was found in a Richardson’s ground squirrel burrow.
Body narrowly subparallel (Fig.
Atheta (sensu lato) richardsoni Klimaszewski & Larson, sp. n.: 33 habitus in dorsal view 34 median lobe of aedeagus in lateral view, and 35 in dorsal view 36 tergite VIII 37 sternite VIII 38 female tergite VIII 39 female sternite VIII 40 spermatheca. Scale bar for habitus = 1 mm, and the remaining scale bars = 0.2 mm.
Distinguished from all other species of Nearctic Atheta by its small size, densely and finally punctate and pubescent forebody, nearly all pronotal pubescence directed straight posteriad (Fig.
Adults are known from SK.
The single male from SK was captured in a ground squirrel burrow, and the single female was found in a Prairie Dog colony in June.
This species in body size and general appearance is similar to species of the subgenus Microdota of Atheta. However, it has a different pubescence pattern of pronotum with microsetae along midline of disc directed straight posteriad and elsewhere straight or obliquely posteriad (Fig.
(for details and illustrations, see
Origin | Palaearctic, adventive in Canada |
Distribution | Canada: AB, LB, NB, NF, NS, ON, PE, QC, SK, YT. USA: CA, NY |
New provincial records | CANADA, Saskatchewan: Saskatoon, 16-VI-1976, D. Larson (DLC) 1 female; Larson Ranch, Hwy 21, 16 km S Maple Creek, 5-V-2008, D. Larson (DLC, LFC) 2 males; 22-VI-2014, D. Larson (LFC) 1 male. |
References |
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The SK specimens were captured in May and June from unspecified habitat. Elsewhere, this species is associated with soil and organic debris in agricultural fields and disturbed urban meadows. It is also found in marsh litter, in leaf litter in mixed forests, in compost, under bark of decaying spruce logs, amongst vegetation on a coastal sand dune, in litter in a cattail marsh, in leaf litter along a vernal pond, and in drift material along a lakeshore (
(for details and illustrations, see
Origin | Nearctic |
Distribution | Canada: AB, BC, LB, NB, ON, PE, QC, SK, YT. USA: AK |
New provincial records | CANADA, Saskatchewan: Cypress Hills Park, Center Block: fire, Sucker Creek, 23-VI-2014, aspen woodland bracket/gilled fungi, D. Larson (DLC) 1 female; 1-VI-2004, under aspen bark, Hooper & Larson (DLC) 1 male. |
References |
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The SK specimens were captured from aspen woodland bracket/gilled fungi, and from under aspen bark. Adults in NF and LB were collected from June to August using pitfall traps and flight intercept traps in various coniferous forest types, and one specimen was collected under the bark of a dead red pine (
(for synonymies and discussion, see
Untill now, only one native species, Dochmonota rudiventris (Eppelsheim) (Figs
Dochmonota rudiventris (Eppelsheim): 41 habitus in dorsal view 42 median lobe of aedeagus in lateral view, and 43 in dorsal view 44 male tergite VIII 45 male sternite VIII 46 female tergite VIII 47 female sternite VIII 48 spermatheca. Scale bar for habitus = 1 mm, and the remaining scale bars = 0.2 mm.
1 | Head about as broad as pronotum (Fig. |
D. langori Klimaszewski & Larson, sp. n. |
– | Head distinctly narrower than pronotum (Figs |
2 |
2 | Elytra at suture longer than pronotum (Fig. |
D. simulans Klimaszewski & Larson, sp. n. |
– | Elytra at suture about as long as pronotum (Figs |
3 |
3. | Elytral posterior corners with strong lateral emarginations (Fig. |
D. rudiventris (Eppelsheim) |
– | Elytral posterior corners with slight emarginations laterally (Fig. |
D. websteri Klimaszewski & Larson, sp. n. |
Dochmonota langori Klimaszewski & Larson, sp. n.: 49 habitus in dorsal view 50 median lobe of aedeagus in lateral view, and 51 in dorsal view 52 male tergite VIII 53 male sternite VIII 54 female tergite VIII 55 female sternite VIII 56 spermatheca. Scale bar for habitus = 1 mm, and the remaining scale bars = 0.2 mm.
Dochmonota simulans Klimaszewski & Larson, sp. n.: 57 habitus in dorsal view 58 median lobe of aedeagus in lateral view, and 59 in dorsal view 60 male tergite VIII 61 male sternite VIII 62 female tergite VIII 63 female sternite VIII 64 spermatheca. Scale bar for habitus = 1 mm, and the remaining scale bars = 0.2 mm.
Dochmonota websteri Klimaszewski & Larson, sp. n.: 65 habitus in dorsal view 66 median lobe of aedeagus in lateral view, and 67 in dorsal view 68 male tergite VIII 69 male sternite VIII 70 female tergite VIII 71 female sternite VIII 72 spermatheca. Scale bar for habitus = 1 mm, and the remaining scale bars = 0.2 mm.
Canada, Saskatchewan, Cypress Lake, E dam, 9-V-2012, wind-drift, D. Larson (LFC) 1 male. Paratypes. Canada, Saskatchewan, Cypress Lake, E dam, 9-V-2012, wind-drift, D. Larson (
The species is named for our friend and professional colleague Dr. David W. Langor, Canadian Forest Service, collaborator and supporter of many joint entomological projects. He contributed to the discovery and descriptions of many new species of aleocharine beetles in Canada, particularly in Newfoundland and Alberta.
Body narrowly subparallel (Fig.
This species is known only from SK.
Adults of this species were collected by sifting wrack on lakeshore beach, and were found in wind-drift on a lake.
Canada, Saskatchewan, Royal Edward Rd., 25 km NW Maple Creek, 5-VI-2011, D. Larson (LFC) 1 male. PARATYPE (female): Canada, Saskatchewan, Hwy 21, 17 km N Maple Ceek, 26-VI-2010, saline slough, D. Larson (LFC).
The species name is derived from Latin adjective simulans-, meaning imitating, in reference to its similarity to the closely related Dochmonota websteri.
Body narrowly subparallel (Fig.
Adults are known only from SK.
The male of this species was captured in June in unspecified habitat, and one female was taken from saline slough, also in June.
Canada, Saskatchewan, Bigstick Lake, 16 km E Golden Prairie, 1-IX-2011, D. Larson (LFC). Paratypes. Canada, Saskatchewan, Bigstick Lake, 16 km E Golden Prairie, 1-IX-2011, D. Larson (LFC) 1 female; Bear Creek at Crane Lake, NE Piapot, 18-VIII-2011, D. Larson (DLC) 1 female. NON-TYPE: Canada, Saskatchewan, Bigstick Lake, N Maple Creek, 4-IX-2012, organic mud/sedges, rushes, etc. near water, D. Larson (DLC) 1 male.
The species is named for Dr. Reginald R. Webster, close friend of JK, and extraordinary entomologist who “understands aleocharine beetles” and who changed the beetle map of New Brunswick by endless discovery of new species. In memory of our “grappa discussions” and fruitful collaboration.
Body moderately narrow, subparallel (Fig.
Adults are known only from SK.
Most adults of this species were collected from shorelines of eutrophic lakes in June, August and September, and one male was captured in organic mud/sedges, and rushes near water.
A male from Bigstick Lake had slightly distorted median lobe of aedeagus and was excluded from the type series.
(for details and illustrations, see
Origin | Nearctic |
Distribution | Canada: AB, BC, MB, NB, NL, NS, ON, QC, SK. USA: AK, AL, AZ, CO, IA, IL, NC, NJ, NM, OR, VA, WA |
New provincial records | CANADA, Saskatchewan: Larson Ranch, Hwy 21, 16 km S Maple Creek, 16-VI-2011, D. Larson (DLC) 2 females. |
References |
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The SK females were captured in June from unspecified habitat. In NL, adults were captured from June to September in the litter of a riparian forest and along the shore of a pond (
(for details and illustrations, see
Origin | Nearctic |
Distribution | Canada: AB, BC, MB, NB, NF, NL, NS, NT, SK, YT. USA: AK, OR |
New provincial records | CANADA, Saskatchewan: Larson Ranch, 16 km S Maple Creek: 8-IV-2010, sifting aspen choke-cherry leaf litter, D. Larson (DLC) 2 females; 18-VI-2010, D. Larson (DLC, LFC) 3 males, 2 females; 27-IV-2013, sifting willow, aspen, hawthorn litter near creek, D. Larson (DLC, LFC) 2 males, 6 females; 3-V-2013, aspen litter, D. Larson (DLC, LFC) 2 males, 2 females; 6-V-2013, sifting willow litter, D. Larson (DLC) 2 females; 10-V-2014, under fresh-cut aspen log rings, D. Larson (DLC) 2 males; 13-V-2014, under fresh-cut aspen log rings, D. Larson (DLC) 1 female; 14-V-2014, under fresh-cut aspen log rings, D. Larson (DLC) 3 females; 20-V-2008, D. Larson (DLC) 1 female; 5-6-VI-2013, maple litter, D. Larson (DLC) 3 females; 8-VI-2014, under fresh-cut aspen log rings, D. Larson (DLC) 1 male; 21-VI-2012, decaying bracket fungus on aspen, D. Larson (DLC) 3 females; 10-VIII-2012, aspen/maple litter, D. Larson (DLC) 2 males, 1 female; 8-IX-2012, compost, D. Larson (DLC) 3 females; 28-IX-2010, D. Larson (DLC) 1 male, 3 females; 20-X-2014, sifting willow leaf litter, D. Larson (DLC) 2 males, 7 females; Belanger Creek, Frenchman Valley, 18-X-2014, D. Larson (DLC) 5 males, 4 females; Cypress Hills Park, Center Block: Sucker Creek, 15-V-2013, sifting aspen litter, D. Larson (DLC) 1 male; 16-VI-2011, sifting wrack, D. Larson (DLC) 1 male; 18-VI-2012, sifting aspen litter, D. Larson (DLC) 3 females; Sucker Creek, 23-VI-2014, aspen woodland bracket/gilled fungi, D. Larson (DLC) 1 female; Saskatoon, 27-VII-1972, D. Larson (DLC) 1 female; 7-IX-2014, spruce-aspen, D. Larson (DLC) 1 female; Saskatoon, 7-X-1976, D. Larson (DLC) 1 female. |
References |
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The SK specimens were captured by sifting aspen litter, maple litter, aspen choke-cherry leaf litter, willow and aspen litter, hawthorn litter near creek, willow leaf litter, under fresh-cut aspen log rings, from decaying woodland bracket/gilled fungi, and from compost, in May through October. In Newfoundland, adults were frequently caught in pitfall traps in various forest types (birch, spruce-lichen, spruce-poplar, fir), in vegetation on coastal sand dunes, on shrubby limestone barrens and in disturbed fields amongst grass and weeds (
(for details and illustrations, see
Origin | Nearctic |
Distribution | Canada: NB, NF, ON, QC, SK. |
New provincial records | CANADA, Saskatchewan: Larson Ranch, Hwy 21, 16 km S Maple Creek: 27-IV-2013, sifting willow-aspen, hawthorn litter near creek, D. Larson (DLC) 1 male, 1 female; 20-V-2008, D. Larson (DLC) 1 female; 25-V-2013, D. Larson (DLC) 1 female; 12-VII-2012, wet grass and weed clippings, D. Larson (DLC) 2 females; 16-VIII-2012, new brome/alfalfa hay, D. Larson (DLC) 1 female; 8-IX-2012, compost, D. Larson (DLC) 1 female; Gull Lake, N town of Gull Lake, 17-V-2014, D. Larson (DLC) 1 female; Cypress Hills Park, Center Block: Highland Trail, 20-V-2013, moist spruce litter near stream, D. Larson (DLC) 2 females; 7-IX-2014, spruce-aspen, D. Larson (DLC) 1 male; 13-IX-2012, sifting spruce litter, D. Larson (DLC) 1 female; Loch Lomond, 21-IX-2011, spruce-aspen litter, D. Larson (DLC) 1 female; Lodgepole Trail, 24-IX-2014, decaying mushrooms, D. Larson (DLC) 1 female. |
References |
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In SK, specimens were captured from May through September from willow-aspen litter, hawthorn litter near creek, wet grass and weed clippings, moist spruce litter near stream, spruce litter, spruce-aspen litter, and in decaying mushrooms. In NB, adults were found in sphagnum moss and litter in calcareous eastern white cedar fens and in a black spruce forest (
Males of this species can be mixed up with those of M. breviuscula and positive identification may only be possible with female association as Mocyta are definitively identified by the shape of the spermatheca.
(for details and illustrations, see
Origin | Palaearctic, adventive in Canada |
Distribution | Canada: NB, NL, NS, ON, PE, QC, SK. USA: CA, LA, MA, MN, NE, NM, NY, PA, VT, TX |
New provincial records | CANADA, Saskatchewan: Saskatoon, 26-IX-1976, D. Larson (DLC) 1 male, 1 female. |
References |
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The SK specimens were captured from an unspecified habitat in September. In NL, one specimen was collected in October from an unspecified habitat (
(for details and illustrations, see
The SK male was captured in June by treading quaking moss, Typha, and Equisetum. The MB specimens were captured in June and August, from unspecified habitat (
(for diagnosis and illustrations, see
Origin | Nearctic |
Distribution | Canada: AB, SK. USA: AZ |
New provincial records | CANADA, Saskatchewan: Larson Ranch, Hwy 21, 16 km S Maple Creek, 9-V-2013, sifting willow/grass litter, D. Larson (DLC) 1 male; Cypress Lake Park, 16-VI-2011, sifting wrack, D. Larson (DLC) 1 female. |
Reference |
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In SK, one male was captured in May by sifting willow/grass litter, and one female was sifted from wrack on a lakeshore in June. In AB, adults were caught in window traps attached to aspen snags in a boreal aspen stand harvested 2 years previously, and in pitfall traps deployed in canola fields. Adults were collected in July (
(for diagnosis and illustrations, see
Origin | Nearctic |
Distribution | Canada: MB, NB, NT, ON, QC, SK, YT; USA: AK, IN |
New provincial records | CANADA, Saskatchewan: Cypress Hills Park, Center Block, Highland Trail, 10-VI-2013, treading quaking moss, Typha and Equisetum in June, D. Larson (DLC) 1 male. |
Reference |
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In SK, one male was captured in June by treading quaking moss, Typha and Equisetum.
(for diagnosis and illustrations, see
Origin | Nearctic |
Distribution | Canada: LB, NF, NS, ON, PE, QC, SK, YT. USA: CA, CO, CT, IA, KS, MO, NC, NJ, NM, NY, TX |
New provincial records | CANADA, Saskatchewan: Great Sand Hills, 50.9°N, 109.11°W, Bowie Ranch, 8-VII-2013, Larson (DLC) 1 female; Larson Ranch, 16 km S Maple Creek, 9-VII-2014, D. Larson (DLC) 1 female; 12 km NE Gull Lake, Scirpus wrack, saline pond, 25-V-2011, D. Larson (DLC) 1 male; Tompkins, Sidewood Rad, 15-IX-2014, D. Larson (DLC) 1 male. |
References |
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In SK, one specimen was found in Scirpus wrack on the shore of saline pond, and three others were found in unspecified habitats in May, July and September. In NB, one specimen was found under a cobblestone on moist sand on a lake margin (
(for diagnosis and illustrations, see
In SK, one female was captured in May from wind-drift on the lake. In NB, S. obscurata was found in flood debris on a river margin, on soil at the base of grass in a residential lawn, and captured in a Lindgren funnel trap in an old jack pine forest (
(for diagnosis and illustrations, see
Origin | Palaearctic, adventive in Canada |
Distribution | Canada: AB, BC, LB, NB, NF, NS, ON, QC, SK. USA: CA, MI, MN, NH, NY, OR |
New provincial records | CANADA, Saskatchewan: Cypress Hills Park, C Block, Sucker Creek, 23-VIII-2012, moose dung, D. Larson (DLC) 2 females; Larson Ranch, Hwy 21, 16 km S Maple Creek: 21-VI-2012, under bark of dead aspen, D. Larson (DLC) 1 female; 1-IX-2012, compost, D. Larson (DLC) 1 male. |
References |
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The SK specimens were found in moose dung, under bark of dead aspen, and in compost in June, August and September. In NL, adults were collected in July using flight intercept traps in mixedwood forest and carrion traps on coastal shrubby barrens (
(for diagnosis and illustrations, see
Origin | Palaearctic, adventive in Canada |
Distribution | Canada: AB, BC, NB, ON, QC, SK. USA: IL, MA, MD, NJ, NY, UT, VA |
New provincial records | CANADA, Saskatchewan: Larson Ranch, Hwy 21, 16 km S Maple Creek: 1-IX-2012, compost, D. Larson (DLC, LFC) 1 female, 1 sex undetermined; 22-27-VI-2005, D. Larson (DLC) 1 sex undetermined; 17-IX-2012, compost, D. Larson (LFC) 1 male; Cypress Hills Lake: E dam, wind-drift, 9-V-2012, D. Larson (DLC) 1 sex undetermined; E end, sifting wrack, 31-VII-2012, D. Larson (DLC, LFC) 1 female, 5 sex undetermined; Crane Lake, NE Piapot., beach wrack, 28-VIII-2011, D. Larson (DLC, LFC) 1 male, 3 sex undetermined; Bigstick Lake, 16 km E Golden Prairie, 21-IX-2011, D. Larson (DLC) 1 sex undetermined; Saskatoon, 26-IX-1976, compost, D. Larson (DLC) 1 sex undetermined. |
References |
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The SK specimens were found in compost, wind drift, and beach wrack, from June through September. In North America, this species is associated with decaying plant material such as compost, mouldy corncobs, cornhusks, weeds, haystacks and rotting fungi (
This species is well established in northeastern and western North America (
(for diagnosis and illustrations, see
Origin | Nearctic |
Distribution | Canada: AB, BC, SK. USA: AZ, CO, ID, NM, UT |
New provincial records | CANADA, Saskatchewan: Larson Ranch, Hwy 21, 16 km S Maple Creek: 22-V-2008, D. Larson (DLC) 1 sex undetermined; 5-6-VI-2013, D. Larson (DLC) 1 sex undetermined; 15-30-VI-2006, D. Larson (DLC) 1 male; 18-VI-2001, D. Larson (LFC) 1 male; Cypress Hills, Center Block: Hidden Valley, 1-VI-1999, D. Larson (DLC) 1 male, 1 sex undetermined; 4-VI-2006, pine clearcut, D. Larson (DLC, LFC) 1 male, 1 female; Ski Lodge, 25-VI-2004, recently dead white spruce, D. Larson (DLC, LFC) 1 female, 2 sex undetermined; fire guard, 29-IX-2013, sifting moss and pine litter, D. Larson (DLC) 1 sex undetermined. |
References |
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The SK specimens were found in pine clearcut, on recently dead white spruce, and in moss and pine litter in May, June and September. Elsewhere, specimens were collected from under bark of logs, from leaf litter, flood debris and wet moss, from soil along a stream, from fungus (Fomitopsis pinicola, Fomes robineae), and from a squirrel midden (
Canada, Saskatchewan, Larson Ranch, Hwy 21, 16 km S Maple Creek, 12-IX-2013, mouldy aspen log, D. Larson (LFC). Paratypes. Canada, Saskatchewan, Larson Ranch, Hwy 21, 16 km S Maple Creek, 12-IX-2013, mouldy aspen log, D. Larson (DLC, LFC) 1 male, 2 females, 11 sex undetermined; Cypress Hills Pk., Center Block, Hidden Valley, 1-VI-1999, D. Larson (DLC) 1 female; Cypress Hills Pk., Center Block, Sucker Cr., 18-VII-2012, sifting aspen litter, D. Larson (DLC) 1 female.
A Latin feminine adjective pulchra, meaning beautiful, in reference to the body shape and beautiful colour of this species.
Body minute, narrowly oval, moderately convex, length 1.4-1.6 mm (Fig.
Agaricochara pulchra Klimaszewski & Larson, sp. n.: 73 habitus in dorsal view 74 median lobe of aedeagus in lateral view 75 male tergite VIII 76 male sternite VIII 77 female tergite VIII 78 female sternite VIII 79 spermatheca. Scale bar for habitus = 1 mm, and the remaining scale bars = 0.2 mm.
Known only from SK. This constitutes new genus record for Canadian fauna.
Adults were collected from mouldy aspen logs in September and by sifting aspen litter in July.
as a subgenus of Gyrophaena Mannerheim, but
(for diagnosis and illustrations, see
Origin | Nearctic |
Distribution | Canada: NB, SK. USA: DC, IL, IN, KA, MI, NY, WA, WI |
New provincial records | CANADA, Saskatchewan: Larson Ranch, Hwy 21, 16 km S Maple Creek, 29-VIII-2014, D. Larson (DLC) 1 male. |
References |
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The SK specimen was collected in August from unspecified habitat. In NB, adults were captured in gilled mushrooms in mixed and hardwood forests from July through September by sifting mushrooms and aspirating specimens (
(for diagnosis and illustrations, see
Origin | Nearctic |
Distribution | Canada: MB, ON, SK. USA: IL, KS, ME, MN, MO, NY, WI |
New provincial records | CANADA, Saskatchewan: Cypress Hills Park, Block Fire, Sucker Creek, 23-VI-2014, aspen woodland bracket/gilled fungi, D. Larson (LFC) 1 male; Maple Creek, Hwy 21, 16 km S, 18-VII-2003, D. Larson (DLC) 1 male, 1 female. |
References |
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Two SK specimens were found in aspen woodland on bracket/gilled fungi, in June and July. In NB, specimens were collected by sifting in June from sun-exposed gilled mushrooms on stump in 8.5-year-old regenerating mixed forest and red oak (
(for diagnosis and illustrations, see
Origin | Nearctic |
Distribution | Canada: AB, BC, NB, NF, NS, ON, QC, SK |
New provincial records | CANADA, Saskatchewan: Cypress Hills Park, Center Block, 1-VI-2004, Hooper & Larson (DLC) 1 male, 1 sex undetermined; Larson Ranch, Hwy 21, 16 km S Maple Creek: 27-IV-2013, sifting willow, aspen, hawthorn near creek, D. Larson (DLC, LFC) 1 male, 6 sex undetermined; 14-V-2014, under bark/in polypore fungus on aspen, D. Larson (DLC) 3 sex undetermined; 5-6-VI-2013, maple litter, D. Larson (DLC) 2 sex undetermined; 6-VI-2013, D. Larson (DLC) 1 sex undetermined; 8-VI-2007, under bark/in polypore fungus on aspen, D. Larson (DLC) 1 female, 1 sex undetermined; 21-VI-2012, under bark of dead aspen, D. Larson (DLC) 1 sex undetermined. |
References |
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The SK specimens were collected from willow, aspen, and hawthorn litter near creek, under bark of dead aspen, in polypore fungus on aspen, in May and June. Elsewhere, two specimens were captured in May on Polyporus betulinus, one by general sweeping in deciduous forest, and one in June in red spruce/hemlock mature forest (
(for illustrations, see
The SK male was found in September in decaying mushrooms.
(for illustrations, see
Origin | Nearctic |
Distribution | Canada: ON, SK, YT |
New provincial records | CANADA, Saskatchewan: Cypress Hills Park, Center Block: Lodgepole Trail, 24-IX-D. Larson (DLC) 1 female; Belanger Creek, 14-X-2014, mossy hummocks bordering marsh and spruce forest, D. Larson (DLC, LFC) 4 females; Belanger Creek, Frenchman Valley, 18-X-2014, mossy hummocks near creek, D. Larson (LFC) 1 male. |
References |
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In SK, specimens were collected from mossy hummocks at the border between a marsh and spruce forest, and mossy hummocks near creek, in September and October.
Body length 1.4–1.5 mm, compact, subparallel, piceous to nearly black, with legs/tarsi, three basal antennal articles, maxillary palps, posterior edge of elytra, and tip of abdomen yellowish brown (Fig.
Origin | Palaearctic, adventive in Canada |
Distribution | Canada: SK |
New North American, Canadian and provincial records | CANADA: Saskatchewan, Larson Ranch, Hwy 21, 16 km S Maple Creek: 14-V-2013 (DLC) 1 female; 22-27-VI-2005 (DLC) 1 female; 16-VIII-2012, new brome/alfalfa hay, D. Larson (DLC, LFC) 2 males, 5 females; 1-IX-2012, compost, D. Larson (DLC, LFC) 5 males, 9 females. |
References |
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The SK specimens were found in compost and new brome/alfalfa hay. Collecting period: June, August and September
Oligota inflata is a Palaearctic species known from Europe, N. Africa, Congo, Egypt, and Brazil. It is reported here for the first time from North America.
(for illustrations, see
Origin | Nearctic |
Distribution | Canada: AB, BC, MB, NB, NF, NS, ON, QC, SK. USA: MI, MO, NH, NY, OR |
New provincial records | CANADA, Saskatchewan: Larson Ranch, 16 km S Maple Creek, 1-15-VI-2005, D. Larson (DLC) 1 sex undetermined; Cypress Hills Park, Center Block, 13-VI-2003, D. Larson (DLC, LFC) 2 sex undetermined |
References |
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The SK specimens were collected in June from unspecified habitat.
(for illustrations, see
Origin | Nearctic |
Distribution | Canada: AB, BC, SK, YT. USA: CA |
New provincial records | CANADA, Saskatchewan: Cypress Hills Park, Center Block, Sucker Creek, 20-V-2013, lodgepole pine litter, D. Larson (DLC) 1 male |
References |
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In SK, one specimen was collected in May from lodgepole pine litter. Elsewhere, adults were captured in clear-cut Sitka spruce forest on Vancouver Island and in moss and gravel at the edge of small pools at other localities in the interior of British Columbia (
(for illustrations, see
Origin | Nearctic |
Distribution | Canada: NB, NS, NT, ON, QC, SK. USA: NY |
New provincial records | CANADA, Saskatchewan, Larson Ranch, Hwy 21, 16 km S Maple Creek: 25-VI-2008, carrion trap, D. Larson (DLC) 1 female; 4-VIII-1998, D. Larson (DLC) 1 female; 27-VIII-2012, pigeon coop, D. Larson (DLC) 1 male |
References |
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In SK, one specimen was collected from pigeon coop, one from carrion trap, and one from unspecified habitat. In NB, Hylota ochracea was a common inhabitant of barred owl nests (
(for illustrations, see
Origin | Nearctic |
Distribution | Canada: LB, NB, NF, NS, ON, QC, SK, YT |
New provincial records | CANADA, Saskatchewan, Larson Ranch, Hwy 21, 16 km S Maple Creek: Apr., 27-IV-2013, sifting willow, aspen, hawthorn litter near creek, D. Larson (DLC) 1 male, 1 female; 21-VI-2012, under bark of dead aspen, D. Larson (DLC) 1 female; 20-X-2014, sifting willow leaf litter, D. Larson (DLC) 1 female. |
References |
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In SK, specimens were captured in willow, aspen, and hawthorn litter near creek, and under bark. In New Brunswick, adults were captured in moist leaf litter on the margin of a vernal pond in a mixed forest, among leaves and sedges on pond margin, in moist grass litter and sphagnum in Carex marsh, among sedges along margin of small spring-fed brook in a mature hardwood forest and among leaf litter and grass on hummocks in a wet alder (Alnus sp.) swamp (
Canada, Saskatchewan, Larson Ranch, Hwy 21, 16 km S Maple Creek, 22-IV-2012, D. Larson (LFC). Paratype. Canada, Saskatchewan, Larson Ranch, Hwy 21, 16 km S Maple Creek, 1-IV-2012, D. Larson (
The name of this species is derived from Latin feminine adjective domestica-, meaning domestic, in reference to the capture of the type specimens in the vicinity of the farmstead.
Body length 3.4-3.6 mm, narrowly subparallel, broadest at posterior elytra, abdomen subparallel (Fig.
The two males were captured in April in an unspecified habitat near a farmstead.
This species is very similar externaly to O. irrasa Mäklin, from which it may be distinguished by the shape of tubus of median lobe of aedeagus with slightly sinuate ventral margin and triangular apical part in lateral view (Fig.
(for illustrations, see
Origin | Nearctic |
Distribution | Canada: AB, SK, YT. USA: AK, OR |
New provincial records | CANADA, Saskatchewan, Larson Ranch, Hwy 21, 16 km S Maple Creek: Apr., 22-IV-2010, dam, D. Larson (DLC) 1 female; 28-IV-2011, on snowbank (DLC) 1 female; 15-VII-2014, decaying polypore mushrooms (DLC) 1 male; 7-X-2010, (LFC) 1 male; Cypress Hills Park, Center Block fire guard, 8-VIII-2013: gilled mushroom, D. Larson (DLC, LFC) 1 male, 5 females; 18-VIII-2014, old polypore fungus on dead lodgepole pine stump (DLC) 2 males, 1 female; Highland Trail, 2-X-2014, gilled mushroom (LFC) 1; 7-X-2014, spruce-aspen (DLC) 1 female; 10-X-2013, decaying mushrooms (DLC) 2 females. |
References |
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In SK, specimens were captured on decaying and old polypore mushrooms in lodgepole pine and spruce-aspen habitats in March, July, August and September. One specimen was captured on snowbank in March. Elsewhere, adults were captured from May through August with most of the specimens taken in August (
(for illustrations, see
Origin | Nearctic |
Distribution | Canada: BC, MB, SK. USA: CO |
New provincial records | CANADA, Saskatchewan, Larson Ranch, Hwy 21, 16 km S Maple Creek: 17-VI-2005, flood debris, D. Larson (DLC) 1 male; 15-30-VIII-2005, D. Larson (DLC) 1 female. |
References |
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In SK, specimens were captured in June and August, one male was found in flood debris along the margin of a seasonal creek. Elsewhere, adults were captured in July and August in Arctic habitats or in the Rocky Mountains (853-2896 m) (Klimaszewki et al. 2006).
(for illustrations, see
Origin | Nearctic |
Distribution | Canada: LB, ON, SK. USA: MA, NC, PA |
New provincial records | CANADA, Saskatchewan, Larson Ranch, Hwy 21, 16 km S Maple Creek: 30-VIII-2014, D. Larson (DLC) 1 female. |
References | As Tetralecopora: |
In SK, one female was captured in August from unspecified habitat. In NF, adults were collected from rocks/gravel at a stream margin in early August (
(for diagnosis and illustrations, see
Origin | Palaearctic, adventive in North America; possibly introduced separately in eastern Canada and western WA |
Distribution | Canada: AB, BC, NB, NF, NS, ON, QC, SK. USA: WA; Palaearctic: Europe |
New provincial records | CANADA, Saskatchewan, Cypress Hills Park, Center Block: Lodgepole Trail, 18-IX-2012, pine/spruce litter near stream, D. Larson (DLC) 1 male; fire guard, 29-X-2013, under fresh-cut pine slabs, D. Larson (DLC) 1 male; Sucker Creek, 1-VI-2012, under bark of recently killed aspen, D. Larson (DLC) 1 female. |
References |
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In SK, specimens were captured in pine/spruce litter near stream, under fresh-cut pine slabs, and under bark of recently killed aspen. In AB, adults were collected from dead or dying white spruce in aggregated retention patches surrounded by different levels of dispersed retention, using emergence traps and window traps (
(for diagnosis and illustrations, see
Origin | Nearctic |
Distribution | Canada: AB, BC, QC, ON, SK |
New provincial records | CANADA, Saskatchewan, Cypress Hills Park, Center Block: fire guard, 29-X-2013, under fresh-cut pine slabs, D. Larson (DLC) 1 male, 1 female. |
References |
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In SK, adults were captured under fresh-cut pine slabs. In AB, adults were collected from dead or dying white spruce in aggregated retention patches surrounded by different levels of dispersed retention, using window traps (
(for diagnosis and illustrations, see
Origin | Palaearctic, adventive in North America |
Distribution | Canada: AB, BC, NB, NS, QC, ON, SK. USA: CA, MA. Palaearctic: Europe, the Mediterranean, Caucasus, Siberia, Japan |
New provincial records | CANADA, Saskatchewan, Larson Ranch, Hwy 21, 16 km S Maple Creek, 30-V-2014, D. Larson (DLC) 1 male; Cypress Hills Park, Center Block, Sucker Creek, 1-4-VI-2012, under bark of recently killed aspen, D. Larson (DLC) 1 male. |
References |
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In SK, one male was captured under bark of recently killed aspen. In AB, adults were reared from white spruce logs in early and intermediate decay stages in white spruce dominated stands (
(for diagnosis and illustrations, see
Origin | Nearctic |
Distribution | Canada: AB, BC, NB, NF, NS, NT, QC, ON, SK, YT. USA: AZ, MA, WA, WI |
New provincial records | CANADA, Saskatchewan, Larson Ranch, Hwy 21, 16 km S Maple Creek, 12-IX-2013, mouldy aspen log, D. Larson (DLC) 1 female; Cypress Hills Park, Center Block, fire guard: 10-IX-2013, newly cut lodgepole pine log, D. Larson (DLC, LFC) 3 males, 3 females; 8-VIII-2013, Ips tunnels in lodgepole pine (DLC) 3 males, 1 female; 26-VIII-2014, under bark of lodgepole pine (DLC) 1 male. |
References |
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In SK, adults were captured from mouldy aspen log, newly cut lodgepole pine log, and in Ips tunnels in lodgepole pine. In eastern Canada, P. tacomae was collected in Lindgren funnel traps from Pinus strobus, Pinus resinosa, Pinus banksiana, Picea glauca, and A. saccharum stands (
(for diagnosis and illustrations, see
Origin | Nearctic |
Distribution | Canada: BC, NB, NS, NT, QC, ON, SK, YT. USA: CA |
New provincial records | CANADA, Saskatchewan, Cypress Hills Park, Lodgepole Trail, 18-IX-2012, under bark of lodgepole pine, D. Larson (DLC) 1 male. |
References |
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In SK, one specimen was captured under bark of lodgepole pine. In QC, specimens were captured in Abies balsamea stands: old-growth stands, undetermined age stands, in Picea glauca stand, and Populus tremuloides with Picea glauca stand (
(for diagnosis and illustrations, see
Origin | Nearctic |
Distribution | Canada: AB, BC, NB, NF, NS, NT, QC, ON, SK, YT. USA: AK, CA, MN |
New provincial records | CANADA, Saskatchewan, Cypress Hills Park, Center Block, fire guard: 8-VIII-2013, gilled mushrooms, D. Larson (DLC) 2 males, 1 female, 1 sex undetermined; 10-IX-2013, decaying mushrooms, D. Larson (DLC) 1 female; 10-VIII-2004, lodgepole pine, D. Larson (DLC) 1 sex undetermined; 18-VIII-2014, old polypore fungus on dead lodgepole pine stump, D. Larson (DLC) 2 males, 2 females, 3 sex undetermined; Cypress Hills Park, Lodgepole Trail, 21-VIII-2013, dry and decaying mushrooms, D. Larson (DLC) 2 males. |
References |
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In SK, adults were captured from gilled mushrooms, dry and decaying mushrooms, old polypore fungus on dead lodgepole pine stump and on lodgepole pine. Elsewhere, adults of S. californica were collected from July through September by means of passive pitfall traps, Luminoc pit-light traps, Malaise traps and by sifting forest litter, wet moss on forest floor, marten dung on moss, and mushrooms (
1 | Median lobe of aedeagus with narrowly triangular apical part forming dorsally distinct angular projection in apical half of tubus in lateral view (see Fig. 5N, in |
Brachyusa americana (Fenyes) |
– | Median lobe of aedeagus with narrowly triangular apical part without angular dorsal projection in apical half of tubus in lateral view (Figs |
2 |
2 | Body broad (Fig. |
Brachyusa saskatchewanae Klimaszewski & Larson, sp. n. |
– | Body moderately narrow (Fig. |
Brachyusa helenae Casey |
Brachyusa helenae Klimaszewski & Larson, sp. n.: 92 habitus in dorsal view 93 median lobe of aedeagus in lateral view 94 male tergite VIII 95 male sternite VIII 96 female tergite VIII 97 female sternite VIII 98 spermatheca. Scale bar for habitus = 1 mm, and the remaining scale bars = 0.2 mm.
Brachyusa saskatchewanae Klimaszewski & Larson, sp. n.: 99 habitus in dorsal view 100 median lobe of aedeagus in lateral view 101 male tergite VIII 102 male sternite VIII 103 female tergite VIII 104 female sternite VIII 105 spermatheca. Scale bar for habitus = 1 mm, and the remaining scale bars = 0.2 mm.
(for diagnosis, see
Tetralina
filitarsus
Casey, 1911: 225. Holotype (male): USA, Montana, Kalispell, June, Wickham, Type
Origin | Nearctic |
Distribution | Canada: LB, NB, NF, NT, ON, SK, YT. USA: AK, MT |
New provincial records | CANADA, Saskatchewan, Cypress Hills Park: Center Block, Lodgepole Trail, 18-IX-2012, pine/spruce litter near stream, D. Larson (LFC) 1 female; Loch Lomond, 29-VIII-2011, D. Larson (DLC) 1 female. |
References |
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In SK, one specimen was captured in pine/spruce litter near stream, and another in an unspecified habitat in August and September. In LB, adults were collected in July and August on sand and gravel on the banks of the Churchill River (
The two SK females agree in colour, body shape, morphology of tergite and sternite VIII, and spermatheca with the type of B. helenae and the recently examined specimens from NF and NB. We have studied the types of B. alutacea (Casey), B. filitarsis (Casey) and B. helenae (Casey). The genital illustrations of B. americana (Fenyes), recorded from BC, are provided by
Canada, Saskatchewan, Bear Creek at Crane Lake, near Piapot, 18-VIII-2011, D. Larson (LFC). Paratypes. Canada, Saskatchewan, Grasslands National Park, Frenchman River at Ecotour Rd., 26-VII-2004, sandy-clay river bank, D. Larson (DLC) 1 male; Bigstick Lake, 16 km E Golden Prairie, 21-IX-2011, D. Larson (DLC, LFC) 4 females.
The name of this species, saskatchewanae-, is a feminine adjective derived from the name of the province of Saskatchewan, where the type series was found.
Body narrowly oval, length 2.3-2.5 mm, slightly flattened; uniformly black with light brown tarsi (Fig.
Brachyusa saskatchewanae may be distinguished from other Brachyusa species by its uniformly black and narrow body, sinuate lateral margins of pronotum, and the genitalic features described above (Figs
Known only from SK.
All SK specimens were captured near water with some on sandy-clay river bank. They were mainly collected by splashing water onto the bank, which caused the beetles to run up the bank.
(for diagnosis and illustrations, see
In SK, one female was captured in June from sandy-clay river bank. In NB, two specimens were captured in June, one from under debris on muddy soil near a small pool in a silver maple forest, and the other from under debris on clay and sand mix at river margin (
(for diagnosis and illustrations, see
Origin | Nearctic |
Distribution | Canada: NB, SK |
New provincial records | CANADA, Saskatchewan, Grassland National Park, W Block, oxbow N jct Ecotour Tr-Frenchman River, 13-VI-2009, D. Larson (DLC) 2 males, 3 females; Grassland National Park, W Block, Ecotour stop 3, shallow oxbow pond, 11-VI-2009, D. Larson (DLC) 1 male, 2 females; Bigstick Lake, N Maple Creek, 4-VIII-2012, organic mud/sedges, rushes, etc. near water, D. Larson (DLC, LFC) 2 males, 4 females; Bigstick Lake, 16 km E Golden Prairie, 21-IX-2011, D. Larson (DLC, LFC) 4 males, 4 females; Larson Ranch, Hwy 21, 16 km S Maple Creek: 10-VI-1998, D. Larson (DLC) 1 male; dam, 28-VIII-2011, D. Larson (DLC) 1 male; 3-IX-2011, D. Larson (DLC) 1 male; Harris Res., 10 km S Maple Creek, wind-drift, 12-V-2012, D. Larson (DLC) 1 male; Cypress Hills Park, C Block, fire break, 10-VI-2011, under bark of lodgepole pine, D. Larson (DLC) 1 male; Cypress Lake, east dam, 12-VI-1998, D. Larson (DLC) 1 male; Cypress Lake Park, sifting wrack, 16-VI-2011, D. Larson (DLC) 1 male; Cypress Lake E end, sifting wrack, 31-VII-2012, D. Larson (DLC) 2 males, 1 female; Cypress Lake E dam, wind-drift, 9-V-2012, DE Larson (DLC) 1 male, 2 females. |
References |
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In SK, specimens were captured from May through September from shallow oxbow pond, organic mud/sedges, rushes, etc. near water, under bark of lodgepole pine, wind-drift, and by sifting wrack. In NB, adults were captured in May from moist leaves near margin of vernal pond in silver maple (Acer saccharinum L.) swamp, and in June from flood debris at the margin of the Saint John River (
We thank Pamela Cheers, English Editor (LFC), who edited the first draft of the manuscript, and Diane Paquet (LFC) for formatting it. We appreciate the help of Amélie Gilbert and Philippe Fortin (LFC) who dissected the specimens used in this study. Benoit Godin, Whitehorse, Yukon, provided a few additional specimens for this study. Volker Assing, Hannover, Germany, kindly provided some genital images of some European species for comparison. We are greatfull to Shockley Floyd (