Holotype male: China, Guizhou Province, Luodian County, Dongjia Town, Dongjia Village, Nameless Cave, 25°38.53'N, 106°54.67'E, 869 m a.s.l., 7 October 2019, Yun-Chun Li leg., in MCWNU (Ar-Ps-GZ-0055). Paratypes: 2 males, 4 females, collected with the holotype, in MCWNU (Ar-Ps-GZ-0008); 5 males, 2 females, Guizhou Province, Pingtang County, Tangbian Town, Baima Cave, 25°40'6.13"N, 106°45'53.89"E, 870 m a.s.l., 6 October 2019, Yun-Chun Li leg., in MCWNU (Ar-Ps-GZ-0010).

Troglobiont habitus. This new species is distinguished from other members of the genus Tyrannochthonius by the following combination of characters: carapace without eyes or eyespots, anterior margin with six setae; epistome absent; rallum composed of six blades; tergites I–IV with two setae; apex of coxa I with long and rounded anteromedial process, near the apex with a seta; chelal hand dorsal surface with chemosensory setae; fixed chelal finger with 24 or 25 teeth, movable chelal finger with 27–29 retrorse teeth. Pedipalpal femur (♂) 7.58–7.63×, (♀) 7.36–7.42× longer than broad, length (♂) 0.91–0.95 mm, (♀) 1.03–1.07 mm; chela (♂) 7.88–7.90×, (♀) 7.06–7.10 longer than deep, length (♂) 1.25–1.28 mm, (♀) 1.20–1.24 mm; ratio movable chelal finger/chelal hand (♂) 1.86–1.90×, (♀) 1.88–1.93×.

Latinized adjective, derived from the village of Dongjia, located near the type locality.

Adult male (Fig. 6A).

Pale yellow-orange, chelicera slightly darker, soft parts pale (Fig. 6A).

Carapace (Fig. 1A): 1.26–1.30× longer than broad, no eyes or eyespots; epistome absent; carapace surface smooth, lateral margins distinctly constricted posteriorly. With 18 setae arranged 6: 4: 4: 2: 2, anterolateral setae much shorter than others. Coxae: manducatory process pointed, with two distal setae, one long and the other slightly shorter. Pedipalpal coxa with three setae, coxa I 3, II 4, III 5, IV 5; intercoxal tubercle absent. Apex of coxa I with long and rounded anteromedial process, near the apex with a seta (Fig. 1E); coxae II with nine terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for ~ ½ their length (Fig. 1D). Chelicera (Fig. 1B): 1.82–1.85× longer than broad, hand with five setae and one lyrifissure dorsally, movable finger with one submedial seta. Cheliceral hand with moderate hispid granulation dorsally. Fixed finger with eight or nine teeth, distal one largest, decreasing in size proximally; movable finger with 12 or 13 teeth; galea absent. Serrula exterior with 22–25 blades. Rallum composed of six blades (Fig. 1C), distal blade weakly recumbent basally, with fine barbules and set apart from the other blades, the latter tightly grouped and with long pinnae. Pedipalp (Fig. 1H–J): all setae acuminate. Trochanter 1.56–1.61×, femur 7.58–7.63×, patella 2.73–2.76× longer than broad and with three lyrifissures (Fig. 1H). Femur 2.22–2.31× longer than patella. Chela 7.88–7.90×, hand 2.63–2.66× longer than deep; movable chelal finger 1.86–1.90× longer than hand. Chelal hand dorsal surface with a single row of five chemosensory setae between esb and ib/isb trichobothria; distal paraxial seta of hand not enlarged. Fingers straight in dorsal view (Fig. 1J). Fixed finger with 24 or 25 teeth, middle ones larger than those at both ends; movable finger with 27–29 retrorse teeth (Fig. 1I). Venom apparatus absent. Fixed chelal finger with eight trichobothria and movable finger with four, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb, and ist forming a straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly nearer to tip of fixed finger; dx situated distal to et; sb half-way between st and b; b and t situated subdistally, t situated at same level as est. Opisthosoma: tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy (I–XII): 2: 2: 2: 2: 3–4: 4: 4: 4: 4: 2: 0; sternal chaetotaxy (IV–XII): 12: 7: 7: 7: 7: 7: 7: 0: 2. Anterior genital operculum with ten setae, genital opening slit-like, with 14 or 15 setae on the right side and 18 on the left (Fig. 1K). Legs (Fig. 1F, G): leg I: trochanter 1.00–1.03×, femur 7.63–7.66× longer than deep and 1.91–1.97× longer than patella; patella 4.57–4.59×, tibia 4.33–4.37×, tarsus 13.40–13.44× longer than deep. Leg IV: trochanter 0.88–0.96×, femoropatella 3.83–3.87×, tibia 6.89–6.92× longer than deep, basitarsus 3.75–3.80×longer than deep, with a basal tactile seta (TS = 0.24–0.25), telotarsus 15.2–15.5×longer than deep and 2.53–2.55× longer than basitarsus, with a tactile seta near base (TS = 0.23–0.24). Arolia on legs I and IV shorter than claws.

Figure 1. 

Tyrannochthonius dongjiensis sp. nov., holotype male (A–K) and paratype female (L) A carapace B left chelicera C rallum of left chelicera D coxal spines E process of left coxa I, ventral view F left leg I, lateral view G left leg IV, lateral view H palp (minus chela) I chela, retrolateral view J chela, dorsal view K male genital area L female genital area. Scale bars: 0.20 mm

Adult female (Fig. 6B).

Mostly the same as the holotype with the differences listed below.

Carapace : slightly longer than broad (1.13–1.15×). Chelicera: 2.30–2.33× longer than broad. Pedipalp: trochanter 1.80–1.86× longer than broad, femur 7.36–7.42× longer than broad, patella 2.63–2.70× longer than broad, femur 2.45–2.49× longer than patella. Chela 7.06–7.10× longer than deep, hand 2.41–2.46× longer than deep; movable finger 1.88–1.93× longer than hand. Opisthosoma: tergal chaetotaxy (I–XII): 2: 2: 2: 2: 4: 4: 4: 4: 4: 2: 2: 0; sternal chaetotaxy (IV–XII): 14: 12: 8: 7: 7: 9: 7: 0: 2. Anterior genital operculum with 9 + 14 setae on posterior margin (Fig. 1L).

Dimensions (mm, length/width or, in the case of the legs, chela, and chelal hand, length/depth).

Males (females in parentheses): body length 2.24–2.30 (2.49–2.56). Carapace 0.54–0.57/0.43–0.45 (0.53–0.55/0.47–0.48). Pedipalp: trochanter 0.25–0.28/0.16–0.18 (0.27–0.29/0.15–0.17), femur 0.91–0.95/0.12–0.14 (1.03–1.07/0.14–0.16), patella 0.41–0.44/0.15–0.17 (0.42–0.44/0.16–0.18), hand 0.42–0.45/0.16–0.17 (0.41–0.44/0.17–0.18), length of movable chelal finger 0.78–0.80 (0.77–0.79), chela 1.25–1.28/0.16–0.17 (1.20–1.24/0.17–0.18). Chelicera: 0.51–0.53/0.28–0.29 (0.53–0.55/0.23–0.24). Leg I: trochanter 0.15–0.17/0.15–0.16 (0.15–0.17/0.14–0.16), femur 0.61–0.64/0.08–0.09 (0.62–0.65/0.08–0.09), patella 0.32–0.35/0.07–0.08 (0.31–0.34/0.07–0.08), tibia 0.26–0.27/0.06–0.07 (0.27–0.29/0.06–0.07), tarsus 0.67–0.69/0.05–0.06 (0.66–0.68/0.05–0.06). Leg IV: trochanter 0.15–0.17/0.17–0.18 (0.17–0.19/0.14–0.16), femoropatella 0.88–0.92/0.23–0.25 (0.90–0.93/0.22–0.24), tibia 0.62–0.65/0.09–0.10 (0.63–0.65/0.09–0.10), basitarsus 0.30–0.32/0.08–0.09 (0.29–0.31/0.08–0.09), telotarsus 0.76–0.79/0.05–0.06 (0.79–0.82/0.06–0.07).

China (Guizhou).

Holotype male: China, Sichuan Province, Luzhou City, Gulin County, Shipping Town, Xiangding Village, Huaer Cave, 28°02.22'N, 106°01.43'E, 760 m a.s.l., 3 November 2019, Yun-Chun Li leg., in MCWNU (Ar-Ps-SC-0052). Paratypes: 4 males, 2 females, collected with the holotype, in MCWNU (Ar-Ps-SC-0001).

Troglobiont habitus. This new species is distinguished from other members of the genus Tyrannochthonius by the following combination of characters: carapace without eyes or eyespots, anterior margin with four setae; epistome very small; rallum composed of eight blades; tergites I–VI with four setae; chelal finger without intercalary teeth; coxae II with 12 terminally indented coxal spines on each side; chelal hand dorsal surface with chemosensory setae; apex of coxa I with long and rounded anteromedial process, near the apex without setae; movable finger retrolateral margins weakly curved between st and sb trichobothria; fixed chelal finger with 23 or 24 teeth, movable chelal finger with 14 or 15 macrodenticles and 7–9 vestigial teeth. Pedipalpal femur (♂) 8.92–8.95×, (♀) 8.54–8.59× longer than broad, length (♂) 1.16–1.19 mm, (♀) 1.11–1.17 mm; chela (♂) 7.00–7.07×, (♀) 8.67–8.69× longer than deep, length (♂) 1.61–1.64 mm, (♀) 1.56–1.58 mm; ratio movable chelal finger/chelal hand (♂) 1.56–1.59×, (♀) 1.52–1.55×.

Latinized adjective, derived from the type locality, namely Huaer Cave.

Adult male (Fig. 6C).

Carapace, chelicera, pedipalps, and tergites I–VI reddish brown, remaining parts yellowish brown (Fig. 6C).

Carapace (Fig. 2A): 1.11–1.13× longer than broad, no eyes or eyespots; epistome very small, triangular; carapace surface smooth, lateral margins distinctly constricted posteriorly. With 18 setae arranged 4: 6: 4: 2: 2, anterolateral setae much shorter than others. Coxae: manducatory process pointed, with two distal setae, one long and the other slightly shorter. Pedipalpal coxa with three setae, coxa I 3, II 4, III 5, IV 5; intercoxal tubercle absent. Apex of coxa I with long and rounded anteromedial process, near the apex without setae (Fig. 2D); coxae II with 12 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for ~ ½ their length. Chelicera (Fig. 2B): 2.31–2.33× longer than broad, hand with five setae and one lyrifissure dorsally, movable finger with one submedial seta. Cheliceral hand with moderate hispid granulation dorsally. Fixed finger with 12 or 13 teeth, distal one largest, decreasing in size proximally; movable finger with 13 or 14 teeth; galea absent. Serrula exterior with 20–22 blades. Rallum composed of eight blades (Fig. 2C), distal blade weakly recumbent basally, with fine barbules and set apart from the other blades, the latter tightly grouped and with long pinnae. Pedipalp (Fig. 2E–G): all setae acuminate. Trochanter 1.25–1.30×, femur 8.92–8.95×, patella 2.75–2.78× longer than broad and with one lyrifissure. Femur 2.64–2.70× longer than patella. Chela 7.00–7.07×, hand 2.74–2.76× longer than deep; movable chelal finger 1.56–1.59× longer than hand. Chelal hand dorsal surface with a single row of seven chemosensory setae between esb and ib/isb trichobothria; distal paraxial seta of hand not enlarged. Fingers straight in dorsal view (Fig. 2G). Fixed finger with 23 or 24 teeth, middle ones larger than those at both ends; movable finger with 14 or 15 macrodenticles, base of finger with 7–9 very low, vestigial teeth (Fig. 2F). Venom apparatus absent. Movable finger retrolateral margins weakly curved between st and sb trichobothria. Fixed chelal finger with eight trichobothria and movable finger with four, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb, and ist forming a straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly nearer to tip of fixed finger; dx situated distal to et; sb near to st; b and t situated subdistally, t situated at same level as it. Opisthosoma: tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy (I–XII): 4: 4: 4: 4: 4: 4: 3: 3: 4: 4: 2: 0; sternal chaetotaxy (IV–XII): 10: 10: 9: 9: 9: 11: 8: 0: 2. Anterior genital operculum with ten setae, genital opening slit-like, with 14 or 15 marginal setae on each side (Fig. 2H). Legs: leg I: trochanter 1.58–1.59×, femur 8.25–8.30× longer than deep and 2.00–2.04× longer than patella; patella 4.71–4.75×, tibia 4.14–4.18×, tarsus 11.17–11.20× longer than deep. Leg IV: trochanter 1.15–1.18×, femoropatella 3.33–3.39×, tibia 6.70–6.72× longer than deep, basitarsus 3.75–3.79× longer than deep, with a basal tactile seta (TS = 0.21–0.22), telotarsus 12.67–12.70× longer than deep and 2.53–2.55× longer than basitarsus, with a tactile seta near base (TS = 0.19–0.20). Arolia on legs I and IV shorter than claws.

Figure 2. 

Tyrannochthonius huaerensis sp. nov., holotype male (A–H) and paratype female (I) A carapace B right chelicera C rallum of left chelicera D process of left coxa I, ventral view E palp (minus chela) F chela, retrolateral view G chela, dorsal view H male genital area I female genital area. Scale bars: 0.20 mm.

Adult female (Fig. 6D).

Mostly the same as the holotype with the differences listed below.

Carapace : slightly longer than broad (1.08–1.10×). Chelicera: 2.27–2.29× longer than broad. Pedipalp: trochanter 1.81–1.84× longer than broad, femur 8.54–8.59× longer than broad, patella 2.87–2.89× longer than broad, femur 2.58–2.60× longer than patella. Chela 8.67–8.69× longer than deep, hand 3.44–3.47× longer than deep; movable finger 1.52–1.55× longer than hand. Opisthosoma: tergal chaetotaxy (I–XII): 4: 4: 4: 4: 4: 4: 4: 4: 5: 4: 2: 0; sternal chaetotaxy (IV–XII): 10: 9: 9: 8: 10: 10: 8: 0: 2. Anterior genital operculum with 10 + 6 setae on posterior margin (Fig. 2I).

Dimensions (mm, length/width or, in the case of the legs, chela, and chelal hand, length/depth).

Males (females in parentheses): body length 1.89–1.95 (1.87–1.99). Carapace 0.59–0.60/0.53–0.54 (0.53–0.55/0.49–0.51). Pedipalp: trochanter 0.20–0.22/0.16–0.18 (0.29–0.30/0.16–0.17), femur 1.16–1.19/0.13–0.15 (1.11–1.17/0.13–0.15), patella 0.44–0.45/0.16–0.17 (0.43–0.44/0.15–0.16), hand 0.63–0.65/0.23–0.25 (0.62–0.65/0.18–0.20), length of movable chelal finger 0.98–0.99 (0.94–0.97), chela 1.61–1.64/0.23–0.25 (1.56–1.58/0.18–0.20). Chelicera: 0.60–0.61/0.26–0.28 (0.59–0.60/0.26–0.27). Leg I: trochanter 0.19–0.20/0.12–0.14 (0.19–0.21/0.12–0.14), femur 0.66–0.69/0.08–0.09 (0.57–0.59/0.08–0.09), patella 0.33–0.35/0.07–0.08 (0.30–0.32/0.07–0.08), tibia 0.29–0.30/0.07–0.08 (0.29–0.30/0.06–0.07), tarsus 0.67–0.69/0.06–0.07 (0.65–0.67/0.05–0.06). Leg IV: trochanter 0.23–0.25/0.20–0.21 (0.17–0.19/0.14–0.16), femoropatella 0.90–0.92/0.27–0.29 (0.83–0.86/0.25–0.27), tibia 0.67–0.69/0.10–0.11 (0.62–0.64/0.10–0.11), basitarsus 0.30–0.32/0.08–0.09 (0.27–0.29/0.08–0.09), telotarsus 0.76–0.79/0.06–0.07 (0.70–0.74/0.05–0.06).

China (Sichuan).

Holotype male: China, Yunnan Province, Dali City, Nanjian County, Xiaowan Town, Huilongshan Village, Banpoyan Cave, 24°56.01'N, 100°18.87'E, 1990 m a.s.l., 23 August 2018, Yun-Chun Li leg., in MCWNU (Ar-Ps-YN-0079). Paratypes: 2 males, 7 females, collected with the holotype, in MCWNU (Ar-Ps-YN-0012).

Troglobiont habitus. This new species is distinguished from other members of the genus Tyrannochthonius by the following combination of characters: carapace without eyes or eyespots, anterior margin with four- setae; epistome present; tergites I–V with four setae; coxae II with eight terminally indented coxal spines on each side; apex of coxa I with long and rounded anteromedial process, near the apex with a seta; chelal hand dorsal surface with chemosensory setae; fixed chelal finger with 28 teeth and 16 or 17 intercalary teeth, movable chelal finger with 15 or 16 macrodenticles, 12 or 13 intercalary teeth and 5–7 vestigial teeth. Pedipalpal femur (♂) 4.87–4.90×, (♀) 5.33–5.37× longer than broad, length (♂) 0.73–0.76 mm, (♀) 0.80–0.83 mm; chela (♂) 5.61–5.66×, (♀) 6.37–6.40× longer than deep, length (♂) 1.01–1.09 mm, (♀) 1.21–1.25 mm; ratio movable chelal finger/chelal hand (♂) 1.75–1.80×, (♀) 1.80–1.83×.

Latinized adjective, derived from the village of Huilongshan, which is near the type locality.

Adult male (Fig. 7A).

Chelicera reddish brown, remaining parts yellowish brown (Fig. 7A).

Carapace (Fig. 3A): 1.02–1.06× longer than broad, no eyes or eyespots; epistome small, triangular, with two setae flanking base; carapace surface smooth, lateral margins distinctly constricted posteriorly. With 18 setae arranged 4: 6: 4: 2: 2, anterolateral setae much shorter than others. Coxae: manducatory process pointed, with two distal setae, one long and the other slightly shorter. Pedipalpal coxa with three setae, coxa I 3, II 4, III 5, IV 5; intercoxal tubercle absent. Apex of coxa I with long and rounded anteromedial process, near the apex with a seta; coxae II with eight terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for ~ ½ their length. Chelicera (Fig. 3B): 2.25–2.29× longer than broad, hand with five setae and one lyrifissure dorsally, movable finger with one submedial seta. Cheliceral hand with moderate hispid granulation dorsally. Fixed finger with eight or nine teeth, distal one largest, decreasing in size proximally; movable finger with 7–9 teeth; galea absent. Serrula exterior with 19–21 blades. Rallum composed of eight blades (Fig. 3C), distal blade weakly recumbent basally, with fine barbules and set apart from the other blades, the latter tightly grouped and with long pinnae. Pedipalp (Fig. 3D–F): all setae acuminate. Trochanter 1.60–1.61×, femur 4.87–4.90×, patella 1.61–1.64× longer than broad and with one lyrifissure. Femur 2.52–2.55× longer than patella. Chela 5.61–5.66×, hand 2.00–2.10× longer than deep; movable chelal finger 1.75–1.80× longer than hand. Chelal hand dorsal surface with a single row of five chemosensory setae between esb and ib/isb trichobothria; distal paraxial seta of hand not enlarged. Fingers straight in dorsal view (Fig. 3F). Fixed finger with 28 teeth and 16 or 17 intercalary teeth, middle ones larger than those at both ends; movable finger with 15 or 16 macrodenticles and 12 or 13 intercalary teeth, base of finger with 5–7 very low, vestigial teeth (Fig. 3E). Venom apparatus absent. Fixed chelal finger with eight trichobothria and movable finger with four, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb, and ist forming a straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly nearer to tip of fixed finger; dx situated distal to et; sb near to st; b and t situated subdistally, t situated at same level as it. Opisthosoma: tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy (I–XII): 4: 4: 4: 4: 4: 6: 6: 6: 6: 4: 2: 0; sternal chaetotaxy (IV–XII): 8: 10: 6: 6: 6: 7: 7: 0: 2. Anterior genital operculum with ten setae, genital opening slit-like, with 11 or 12 marginal setae on each side (Fig. 3G). Legs: leg I: trochanter 1.68–1.70×, femur 6.50–6.58× longer than deep and 1.56–1.59× longer than patella; patella 4.17–4.20×, tibia 4.00–4.06×, tarsus 8.40–8.47× longer than deep. Leg IV: trochanter 1.00–1.07×, femoropatella 2.59–2.63×, tibia 4.40–4.47× longer than deep, basitarsus 2.71–2.74× longer than deep, with a basal tactile seta (TS = 0.15–0.17), telotarsus 9.60–9.66× longer than deep and 2.53–2.55× longer than basitarsus, with a tactile seta near base (TS = 0.15–0.16). Arolia on legs I and IV shorter than claws.

Figure 3. 

Tyrannochthonius huilongshanensis sp. nov., holotype male (A–G) and paratype female (H) A carapace B right chelicera C rallum of left chelicera D palp (minus chela) E chela, retrolateral view F chela, dorsal view G male genital area H female genital area. Scale bars: 0.20 mm.

Adult female (Fig. 7B).

Mostly the same as the holotype with the differences listed below.

Carapace : slightly longer than broad (0.90–0.99×). Chelicera: 2.13–2.17× longer than broad. Pedipalp: trochanter 1.86–1.88× longer than broad, femur 5.33–5.37× longer than broad, patella 1.94–1.98× longer than broad, femur 2.58–2.59× longer than patella. Chela 6.37–6.40× longer than deep, hand 2.11–2.15× longer than deep; movable finger 1.80–1.83× longer than hand. Opisthosoma: tergal chaetotaxy (I–XII): 4: 4: 4: 4: 5: 6: 6: 6: 5: 4: 2: 0; sternal chaetotaxy (IV–XII): 12: 10: 7: 8: 7: 7: 6: 0: 2. Anterior genital operculum with 10 + 17 setae on posterior margin (Fig. 3H).

Dimensions (mm, length/width or, in the case of the legs, chela, and chelal hand, length/depth).

Males (females in parentheses): body length 1.68–1.75 (1.89–1.95). Carapace 0.44–0.46/0.43–0.44 (0.45–0.49/0.50–0.51). Pedipalp: trochanter 0.24–0.26/0.15–0.17 (0.26–0.28/0.14–0.16), femur 0.73–0.76/0.15–0.17 (0.80–0.83/0.15–0.17), patella 0.29–0.31/0.18–0.19 (0.31–0.33/0.16–0.17), hand 0.36–0.40/0.18–0.20 (0.40–0.44/0.19–0.20), length of movable chelal finger 0.63–0.67 (0.72–0.76), chela 1.01–1.09/0.18–0.20 (1.21–1.25/0.19–0.20). Chelicera: 0.45–0.47/0.20–0.22 (0.51–0.54/0.24–0.26). Leg I: trochanter 0.17–0.19/0.10–0.11 (0.16–0.18/0.14–0.15), femur 0.39–0.42/0.06–0.07 (0.45–0.46/0.08–0.09), patella 0.25–0.27/0.06–0.07 (0.28–0.30/0.07–0.08), tibia 0.20–0.22/0.05–0.06 (0.22–0.25/0.06–0.07), tarsus 0.42–0.45/0.05–0.06 (0.49–0.53/0.05–0.06). Leg IV: trochanter 0.16–0.17/0.16–0.17 (0.21–0.22/0.15–0.17), femoropatella 0.57–0.59/0.22–0.24 (0.54–0.57/0.20–0.22), tibia 0.44–0.46/0.10–0.11 (0.43–0.46/0.11–0.12), basitarsus 0.19–0.21/0.07–0.08 (0.21–0.23/0.08–0.09), telotarsus 0.48–0.50/0.05–0.06 (0.50–0.54/0.05–0.06).

China (Sichuan).

Holotype male: China, Yunnan Province, Zhaotong City, Zhenxiong County, Wude Town, Xinzhai Village, Daguoquan Cave, 27°35.90'N, 104°46.25'E, 1301 m a.s.l., 8 April 2017, Yun-Chun Li leg., in MCWNU (Ar-Ps-YN-0080). Paratypes: 1 male, 6 females, 6 tritonymphs, collected with the holotype, in MCWNU (Ar-Ps-YN-0007).

Troglobiont habitus. This new species is distinguished from other members of the genus Tyrannochthonius by the following combination of characters: carapace without eyes or eyespots, anterior margin with five or six setae; epistome present; tergites V–X with four setae; coxae II with 12 terminally indented coxal spines on each side; apex of coxa I with long and rounded anteromedial process, near the apex without setae; chelal hand dorsal surface with chemosensory setae; fixed chelal finger with 26 teeth, movable chelal finger with 34 or 35 teeth. Pedipalpal femur (♂) 6.94–6.97×, (♀) 6.71–6.77× longer than broad, length (♂) 1.18–1.21 mm, (♀) 1.14–1.18 mm; chela (♂) 7.90–7.91×, (♀) 6.44–6.42× longer than deep, length (♂) 1.66–1.68 mm, (♀) 1.61–1.64 mm; ratio movable chelal finger/chelal hand (♂) 1.61–1.64×, (♀) 1.76–1.80×.

Latinized adjective, derived from the village of Xinzhai, located near the type locality.

Adult male (Fig. 7C).

Carapace and chelicera reddish brown, remaining parts yellowish brown (Fig. 7C).

Carapace (Fig. 4A): 0.98–1.01× longer than broad, no eyes or eyespots; epistome very pointed and small, triangular; carapace surface smooth, lateral margins weakly constricted posteriorly. With 17 or 18 setae arranged 5–6: 4: 4: 2: 2, anterolateral setae much shorter than others. Coxae: manducatory process pointed, with two distal setae, one long and the other slightly shorter. Pedipalpal coxa with three setae, coxa I 3, II 4, III 5, IV 5; intercoxal tubercle absent. Apex of coxa I with long and rounded anteromedial process, near the apex without setae; coxae II with 12 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for ~ ½ their length. Chelicera (Fig. 4B): 2.59–2.61× longer than broad, hand with five setae and one lyrifissure dorsally, movable finger with one submedial seta. Cheliceral hand with moderate hispid granulation dorsally. Fixed finger with 16 teeth, distal one largest, decreasing in size proximally; movable finger with 14 or 15 teeth; galea absent. Serrula exterior with 23 or 24 blades. Rallum composed of eight blades (Fig. 4C), distal blade weakly recumbent basally, with fine barbules and set apart from the other blades, the latter tightly grouped and with long pinnae. Pedipalp (Fig. 4E–G): all setae acuminate. Trochanter 1.56–1.59×, femur 6.94–6.97×, patella 1.83–1.86× longer than broad and with four lyrifissures. Femur 2.68–2.70× longer than patella. Chela 7.90–7.91×, hand 2.90–2.93× longer than deep; movable chelal finger 1.61–1.64× longer than hand. Chelal hand dorsal surface with a single row of five chemosensory setae between esb and ib/isb trichobothria; distal paraxial seta of hand not enlarged. Fingers straight in dorsal view (Fig. 4G). Fixed finger with 26 teeth, middle ones larger than those at both ends; movable finger with 34 or 35 teeth (Fig. 4F). Venom apparatus absent. Fixed chelal finger with eight trichobothria and movable finger with four, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb, and ist forming a straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly nearer to tip of fixed finger; dx situated distal to et; sb near to st; b and t situated subdistally, t situated at same level as est. Opisthosoma: tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy (I–XII): 4: 4: 4: 4: 4: 4: 4: 4: 4: 4: 2: 0; sternal chaetotaxy (IV–XII): 12: 10: 10: 9: 9: 9: 7: 0: 2. Anterior genital operculum with nine setae, genital opening slit-like, with 15 or 16 marginal setae on each side (Fig. 4H). Legs: leg I: trochanter 1.43–1.44×, femur 6.60–6.62× longer than deep and 1.78–1.79× longer than patella; patella 4.11–4.14×, tibia 3.88–3.92×, tarsus 9.57–9.60× longer than deep. Leg IV: trochanter 1.05–1.07×, femoropatella 3.83–3.85×, tibia 5.91–5.93× longer than deep, basitarsus 3.75–3.76× longer than deep, with a basal tactile seta (TS = 0.20–0.21), telotarsus 12.17–12.20× longer than deep and 2.43–2.45× longer than basitarsus, with a tactile seta near base (TS = 0.18–0.19). Arolia on legs I and IV shorter than claws.

Figure 4. 

Tyrannochthonius xinzhaiensis sp. nov., holotype male (A–H) and paratype female (I) A carapace B right chelicera C rallum of left chelicera D process of left coxa I, ventral view E palp (minus chela) F chela, retrolateral view G chela, dorsal view H male genital area I female genital area. Scale bars: 0.20 mm.

Adult female (Fig. 7D).

Mostly the same as the holotype with the differences listed below.

Carapace : slightly longer than broad (1.00–1.02×). With 18 setae, including six on the anterior margin and two on the posterior margin. Chelicera: 2.26–2.27× longer than broad. Pedipalp: trochanter 1.82–1.86× longer than broad, femur 6.71–6.77× longer than broad, patella 1.76–1.79× longer than broad, femur 3.08–3.12× longer than patella. Chela 6.44–6.42× longer than deep, hand 2.20–2.22× longer than deep; movable finger 1.76–1.80× longer than hand. Opisthosoma: tergal chaetotaxy (I–XII): 4: 4: 3: 3: 4: 4: 4: 4: 4: 4: 2: 0; sternal chaetotaxy (IV–XII): 14: 10: 9: 9: 9: 9: 7: 0: 2. Anterior genital operculum with 9 + 6 setae on posterior margin (Fig. 4I).

Dimensions (mm, length/width or, in the case of the legs, chela, and chelal hand, length/depth).

Males (females in parentheses): body length 2.76–2.85 (2.69–2.88). Carapace 0.59–0.61/0.60–0.61 (0.58–0.60/0.58–0.59). Pedipalp: trochanter 0.25–0.26/0.16–0.17 (0.31–0.34/0.17–0.19), femur 1.18–1.21/0.17–0.19 (1.14–1.18/0.17–0.19), patella 0.44–0.46/0.24–0.26 (0.37–0.40/0.21–0.23), hand 0.61–0.63/0.21–0.22 (0.55–0.58/0.25–0.26), length of movable chelal finger 0.98–1.00 (0.97–1.01), chela 1.66–1.68/0.21–0.22 (1.61–1.64/0.25–0.26). Chelicera: 0.70–0.73/0.27–0.29 (0.61–0.64/0.27–0.29). Leg I: trochanter 0.20–0.22/0.14–0.15 (0.16–0.18/0.14–0.15), femur 0.66–0.68/0.10–0.11 (0.62–0.65/0.08–0.09), patella 0.37–0.39/0.09–0.10 (0.31–0.34/0.06–0.07), tibia 0.31–0.32/0.08–0.09 (0.31–0.33/0.06–0.07), tarsus 0.67–0.69/0.07–0.08 (0.64–0.67/0.06–0.07). Leg IV: trochanter 0.21–0.23/0.20–0.21 (0.19–0.21/0.17–0.19), femoropatella 0.92–0.95/0.24–0.26 (0.83–0.86/0.22–0.24), tibia 0.65–0.67/0.11–0.12 (0.57–0.60/0.09–0.10), basitarsus 0.30–0.31/0.08–0.09 (0.27–0.29/0.07–0.08), telotarsus 0.73–0.75/0.06–0.07 (0.68–0.70/0.05–0.06).

China (Yunnan).

Holotype male: China, Yunnan Province, Lushui City, Fugong County, Shiyueliang Town, Lishadi Village, Yamu River, Nameless Cave, 27°07.69'N, 98°51.61'E, 1500 m a.s.l., 18 August 2018, Yun-Chun Li leg., in MCWNU (Ar-Ps-YN-0078). Paratypes: 1 male, collected with the holotype, in MCWNU (Ar-Ps-YN-0014).

(male, female unknown). Troglobiont habitus. This new species is distinguished from other members of the genus Tyrannochthonius by the following combination of characters: carapace without eyes or eyespots, anterior margin with four setae; epistome present; tergites II–VI with four setae; coxae II with ten terminally indented coxal spines on each side; apex of coxa I with long and rounded anteromedial process, near the apex with a seta; chelal hand dorsal surface with chemosensory setae. Fixed chelal finger with 25 teeth and 20 intercalary teeth, movable chelal finger with 22–24 teeth and three or four intercalary teeth. Pedipalpal femur 6.06–6.07× longer than broad, length 0.97–0.99 mm; chela 7.63–7.66× longer than deep, length 1.45–1.46 mm; ratio movable chelal finger/chelal hand 1.91–1.92×.

Latinized adjective, derived from the river of Yamuhe, which is near the type locality.

Adult male (Fig. 8).

Chelicera reddish brown, carapace and opisthosoma brown, remaining parts yellowish brown (Fig. 8).

Carapace (Fig. 5A): 1.06–1.08× longer than broad, no eyes or eyespots; epistome very pointed and small, triangular; carapace surface smooth, lateral margins weakly constricted posteriorly. With 18 setae, including four on anterior margin and two on posterior margin, anterolateral setae much shorter than others. Coxae: manducatory process pointed, with two distal setae, one long and the other slightly shorter. Pedipalpal coxa with three setae, coxa I 3, II 4, III 5, IV 5; intercoxal tubercle absent. Apex of coxa I with long and rounded anteromedial process, near the apex with a seta; coxae II with ten terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for ~ ½ their length (Fig. 5D). Chelicera (Fig. 5B): 2.31–2.33× longer than broad, hand with five setae and two lyrifissures dorsally, movable finger with one submedial seta. Cheliceral hand with moderate hispid granulation dorsally. Fixed finger with eight or nine teeth, distal one largest, decreasing in size proximally; movable finger with 12 or 13 teeth; galea absent. Serrula exterior with 20 or 21 blades. Rallum composed of eight blades (Fig. 5C), distal blade weakly recumbent basally, with fine barbules and set apart from the other blades, the latter tightly grouped and with long pinnae. Pedipalp (Fig. 5F–H): all setae acuminate. Trochanter 1.01–1.04×, femur 6.06–6.07×, patella 2.38–2.40× longer than broad. Femur 2.55–2.56× longer than patella. Chela 7.63–7.66×, hand 2.47–2.50× longer than deep; movable chelal finger 1.91–1.92× longer than hand. Chelal hand dorsal surface with a single row of five chemosensory setae between esb and ib/isb trichobothria; distal paraxial seta of hand not enlarged. Fingers straight in dorsal view (Fig. 5H). Fixed finger with 25 teeth and 20 intercalary teeth, middle ones larger than those at both ends; movable finger with 22–24 teeth and three or four intercalary teeth (Fig. 5G). Venom apparatus absent. Fixed chelal finger with eight trichobothria and movable finger with four, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb, and ist forming a straight oblique row at base of fixed chelal finger; it slightly distal to est, situated subdistally; et slightly nearer to tip of fixed finger; dx situated distal to et; sb near to st; b and t situated subdistally, t situated at same level as est. Opisthosoma: tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy (I–XII): 3: 4: 4: 4: 4: 4: 6: 5: 5: 5: 2: 0; sternal chaetotaxy (IV–XII): 12: 10: 7: 7: 7: 7: 6: 0: 2. Anterior genital operculum with ten, genital opening slit-like, with 15 or 16 marginal setae on each side (Fig. 5I). Legs: leg I: trochanter 1.38–1.40×, femur 6.63–6.65× longer than deep and 1.77–1.79× longer than patella; patella 4.29–4.30×, tibia 5.20–5.22×, tarsus 11.80–11.81× longer than deep. Leg IV: trochanter 1.06–1.07×, femoropatella 3.00–3.02×, tibia 5.70–5.71× longer than deep, basitarsus 3.00–3.01× longer than deep, with a basal tactile seta (TS = 0.20–0.21), telotarsus 12.80–12.81× longer than deep and 2.67–2.69× longer than basitarsus, with a tactile seta near base (TS = 0.19–0.20). Arolia on legs I and IV shorter than claws.

Figure 5. 

Tyrannochthonius yamuhensis sp. nov., holotype male (A–I) A carapace B right chelicera C rallum of left chelicera D coxal spines E process of right coxa I, ventral view F palp (minus chela) G chela, retrolateral view H chela, dorsal view I male genital area. Scale bars: 0.20 mm.

Dimensions (mm, length/width or, in the case of the legs, chela, and chelal hand, length/depth).

Males: body length 2.25–2.30. Carapace 0.56–0.57/0.53–0.54. Pedipalp: trochanter 0.14–0.15/0.14–0.15, femur 0.97–0.99/0.16–0.18, patella 0.38–0.39/0.16–0.17, hand 0.47–0.49/0.19–0.20, length of movable chelal finger 0.90–0.92, chela 1.45–1.46/0.19–0.20. Leg I: trochanter 0.18–0.19/0.13–0.15, femur 0.53–0.55/0.08–0.09, patella 0.30–0.31/0.07–0.08, tibia 0.26–0.28/0.05–0.06, tarsus 0.59–0.60/0.05–0.06. Leg IV: trochanter 0.18–0.20/0.17–0.18, femoropatella 0.75–0.77/0.25–0.26, tibia 0.57–0.59/0.10–0.11, metatarsus 0.24–0.25/0.08–0.09, tarsus 0.64–0.66/0.05–0.06.

Figure 6. 

A, B Tyrannochthonius dongjiensis sp. nov., dorsal views A holotype male B paratype female C, D T. huaerensis sp. nov., dorsal views C holotype male D paratype female. Scale bar: 1.00 mm (A–D).

Figure 7. 

A, B Tyrannochthonius huilongshanensis sp. nov., dorsal views A holotype male B paratype female C, D T. xinzhaiensis sp. nov., dorsal views C holotype male D paratype female. Scale bar: 1.00 mm (A–D).

China (Yunnan).