Holotype : China • ♂; Tibet Autonomous Region, Medog County, Bari village; 29°20'13"N, 95°21'54"E; 1680 m a.s.l.; 29 July 2014; Yan Li leg; CAU. Paratypes: China • 2 ♂♂ 2 ♀♀; same data as holotype; CAU.

Antocha (A.) curvativa sp. nov. can be recognized by thorax with four more or less confluent stripes, wing having no stigma, basal section of M₃ which is about four times as long as m-m, posterior margin of tergite 9 having shallow median emargination and specific, stout outer gonostylus with tip distinctly flattened and nearly funnel-shaped. Aedeagal complex with interbase elongated, distally oval; paramere apically slender and curved ventrally; inner branch of paramere elongated, tip rounded.

Male. Body length 4.8–5.5 mm, wing length 4.3–4.8 mm, antenna length 0.9–1.1 mm.

Head (Fig. 2b). Dark brown, with brown setae. Antenna dark brown. Scape cylindrical; pedicel oval; flagellomeres oval, apically tapering and shortened. Setae on antenna brown. Rostrum light brown; palpus brown to dark brown; setae on rostrum and palpus brown.

Thorax (Fig. 2c). Pronotum dark brown. Prescutum and presutural scutum brown, with four more or less confluent dark brown stripes. Postsutural scutum brown; scutal lobes each with a darker brown spot. Scutellum brown, with side edges dark brown. Mediotergite dark brown. Pleuron brown (Fig. 2a). Legs with light brown coxae; trochanters light yellow with side edges brown; femora yellowish, becoming brown towards apex; tibiae and tarsal segments brown. Setae on legs brown. Wing light brown, without stigma; anal angle nearly right-angled (Fig. 2d). Veins light brown. Venation: Sc ending before fork of Rs, at about 2/3 of Rs; basal section of R₅ about as long as r-m; m-cu shortly before fork of M, distance approximately 1/4 its own length; basal section of M₃ about four times as long as m-m; cell m₁ about as long as cell dm. Halter with stem pale.

Figure 2. 

Antocha (Antocha) curvativa sp. nov. a habitus of male, lateral view b male head, lateral view c male thorax, dorsal view d male wing e aedeagal complex with gonocoxite and gonostyli, dorsal view f female ovipositor, lateral view. Scale bars: 1.0 mm (a); 0.3 mm (b, c); 0.5 mm (d); 0.2 mm (e, f).

Abdomen. Tergites 1–6 brown, tergites 7 and 8 dark brown. Sternites 1–6 light brown, sternites 7 and 8 dark brown.

Hypopygium (Figs 2e, 3). Brown. Posterior margin of tergite 9 with broad and shallow emargination (Fig. 3a). Gonocoxite nearly cylindrical, with long yellow setae (Figs 2e, 3). Outer gonostylus stout, apical half sclerotized, tip distinctly curved, flattened, nearly funnel-shaped. Inner gonostylus thick and fleshy. Interbase nearly V-shaped, distal part elongate and oval (Figs 2e, 3). Paramere with base rod-shaped, apical part slender, curved ventrally, and with tip sharp. Inner branch of paramere in the shape of elongated lobe with tip rounded (Figs 2e, 3). Aedeagus rod-shaped, curved ventrally (Figs 2e, 3).

Figure 3. 

Antocha (Antocha) curvativa sp. nov. a male hypopygium, dorsal view b male hypopygium, ventral view. Scale bar: 0.1 mm.

Female. Body length 5.0–5.5 mm, wing length 4.5–5.0 mm. Generally similar to male by body coloration.

Ovipositor (Fig. 2f). Tergite 10 dark brown, caudal part paler. Cercus brown with base darker, tip raised and tapering. Hypogynial valve light brown, reaching approximately 3/5 of cercus.

The specific name refers to the curved apical part of paramere.

China (Tibet).

The new species is similar to A. (A.) lacteibasis from China in having similar apical part of paramere and tip of inner branch of paramere being not bifid, but it can be easily distinguished by the basal section of vein M₃ being about four times as long as m-m (Fig. 2d) and the stout outer gonostylus with the tip sclerotized, funnel-shaped, inflated, and blunt (Figs 2e, 3). In A. (A.) lacteibasis, the basal section of vein M₃ is about twice as long as m-m, and the outer gonostylus narrow with acute tip (Alexander 1935; Markevičiūtė et al. 2019, 2021).

Holotype : China • ♂; Sichuan province, Mount Omei; 2134 m a.s.l.; 17 July 1931; Franck leg; USNM. Other material examined: China • 8 ♂♂ 6 ♀♀; Tibet Autonomous Region, Chayu County, Xiachayu Farm Hydropower Station; 28°30'19"N, 97°01'25"E; 1520 m a.s.l.; 8 July 2016; Shaolin Han leg; CAU.

Antocha (A.) setigera can be recognized by thorax with four brown stripes, wing lacking a stigma, basal section of M₃ as long as m-m, and slightly curved outer gonostylus with blackened, blunt tip. Aedeagal complex with interbase distally small and parameres apically fused, arch-shaped.

Male. Body length 4.5–5.0 mm, wing length 5.0–6.0 mm, antenna length 1.5–1.8 mm.

Head (Fig. 4b). Dark brown with brown setae. Antenna brown with light brown scape. Scape cylindrical; pedicel and flagellomeres elongate oval, apically tapering; terminal segment short, about half as short as other segments. Rostrum yellow; palpus light brown; setae on rostrum and palpus brown.

Figure 4. 

Antocha (Antocha) setigera a habitus of male, lateral view b male head, lateral view c male thorax, dorsal view d male wing e aedeagus complex with gonocoxite and gonostyli, dorsal view f female ovipositor, lateral view. Scale bars: 1.0 mm (a); 0.2 mm (b, c); 0.5 mm (d); 0.2 mm (e, f).

Thorax (Fig. 4c). Pronotum brown. Prescutum and presutural scutum brown four brown stripes. The central stripes fused in the anterior third, the rest is separated by pale narrow vitta. Postsutural scutum dark brown, middle area yellow, scutal lobes each with brown spot. Scutellum pale yellow, with side edges dark brown. Mediotergite brown to dark brown. Pleuron brown (Fig. 4a). Legs with fore coxa brown; mid coxa brownish yellow; hind coxa yellow; trochanters yellow; femora and tibiae brownish yellow; tarsi brown with terminal segments darker brown. Wing light brown, without stigma; anal angle nearly right-angled (Fig. 4d). Veins brown. Venation: Sc ending nearly at fork of Rs; basal section of R₅ about twice as long as r-m; m-cu shortly before fork of M, distance approximately 1/3 its own length; basal section of M₃ as long as m-m; cell m₁ longer than cell dm. Halter pale with stem light yellow.

Abdomen. Tergites dark brown. Sternites 1–6 brown with side edges yellow, sternites 7 and 8 dark brown.

Hypopygium (Figs 4e, 5). Yellow. Posterior margin of tergite 9 convex with middle slightly emarginate (Fig. 5a). Gonocoxite nearly cylindrical with long brown setae (Figs 4e, 5). Outer gonostylus slightly curved, with blackened distal half and tip sclerotized and blunt. Inner gonostylus slightly curved. Interbase distally flattened, small and horn-like with tip sharp (Figs 4e, 5). Parameres apically fused, arch-shaped. Inner branch of paramere elongated, with tip narrowly obtuse (Figs 4e, 5). Aedeagus curved ventrally, tip bifid (Figs 4e, 5).

Figure 5. 

Antocha (Antocha) setigera a male hypopygium, dorsal view b male hypopygium, ventral view. Scale bar: 0.1 mm.

Female. Body length 4.5–5.5 mm, wing length 5.0–6.0 mm. Generally similar to male by body coloration.

Ovipositor (Fig. 4f). Tergite 10 yellowish brown with base darker. Cercus brown, tip raised and tapering, apex acute. Hypogynial valve yellowish, reaching sub-tip of cercus.

China (Sichuan, Tibet).

In China, this species was previously only known in Sichuan province and is now recorded in Tibet for the first time. For descriptions and illustrations of this species, also see Alexander (1933b) and Markevičiūtė et al. (2019, 2021).

China • 9 ♂♂ 1 ♀; Tibet Autonomous Region, Bayi District, Pailong; 30°01'25"N, 95°00'32"E; 2003 m a.s.l.; 20 June 2018; Liang Wang leg.; CAU.

Antocha (A.) spiralis can be recognized by thorax having no stripes, wing without stigma, basal section of M₃ about one and half times as long as m-m, posterior margin of tergite 9 having shallow, median emargination and slightly curved, blackened in distal 2/3 of outer gonostylus. Aedeagal complex with interbase nearly U-shaped; paramere apically slender and twisted into a spiral; inner branch of paramere with bifid tip.

Male. Body length 4.5–5.5 mm, wing length 5.0–6.0 mm, antenna length 1.0–1.2 mm.

Head (Fig. 6b). Black with brown setae. Antenna brown, with dark brown pedicel. Scape cylindrical; pedicel and flagellomeres oval; terminal two segments slender. Rostrum yellow; palpus light brown; setae on rostrum and palpus brown.

Figure 6. 

Antocha (Antocha) spiralis a habitus of male, lateral view b male head, lateral view c male thorax, dorsal view d male wing e aedeagus complex with gonocoxite and gonostyli, dorsal view f female ovipositor, lateral view. Scale bars: 1.0 mm (a); 0.3 mm (b); 0.4 mm (c); 0.6 mm (d); 0.2 mm (e, f).

Thorax (Fig. 6c). Pronotum brown. Prescutum and presutural scutum dark brown, without stripe. Postsutural scutum brownish yellow in middle; scutal lobes each with brown spot. Scutellum brown with middle of base brownish yellow. Mediotergite dark brown, with middle brownish yellow. Pleuron brown (Fig. 6a). Legs with fore and mid coxae brown; hind coxa yellow; trochanters yellow with side edges brown; femora brownish yellow to brown; remaining segments dark brown. Wing light brown, without stigma; anal angle nearly right-angled (Fig. 6d). Veins brown. Venation: Sc ending before fork of Rs, at about 5/6 of Rs; basal section of R₅ about 1½ times as long as r-m; m-cu shortly before fork of M, distance approximately 1/4 its own length; basal section of M₃ about 1½ as long as m-m; cell m₁ longer than cell dm. Halter pale.

Abdomen. Tergites 1–6 brown, tergites 7 and 8 dark brown. Sternites 1–6 brownish yellow to light brown; sternites 7 and 8 dark brown.

Hypopygium (Figs 6e, 7). Posterior margin of tergite 9 with broad and shallow emargination (Fig. 7a). Gonocoxite nearly cylindrical with brown setae (Figs 6e, 7). Outer gonostylus slightly curved; base yellowish brown; distal 2/3 blackened, narrowing towards obtuse apex. Interbase nearly U-shaped, distal part flattened and oval (Figs 6e, 7). Paramere with rod-shaped base; apical part slender and twisted into spiral; tip sharp. Inner branch of paramere with tip bifid; outer tooth longer than inner one (Figs 6e, 7). Aedeagus with two projections near tip (Figs 6e, 7).

Figure 7. 

Antocha (Antocha) spiralis a male hypopygium, dorsal view b male hypopygium, ventral view. Scale bar: 0.1 mm.

Female. Body length 4.5–5.5 mm, wing length 5.0–6.0 mm. Generally similar to male by body coloration.

Ovipositor (Fig. 6f). Tergite 10 brownish yellow, with brown base. Cercus yellowish brown, with base darker, slender, and curved; tip raised and tapering. Hypogynial valve yellow, reaching approximately middle of cercus.

China (Sichuan, Tibet), India.

In China, this species was previously only known in Sichuan and is now recorded in Tibet for the first time. For descriptions and illustrations of this species, also see Alexander (1932) and Markevičiūtė et al. (2019, 2021). Both A. (A.) spiralis and A. (A.) bella from China have the twisted structure of the hypopygium. In A. (A.) spiralis, the apical part of the paramere is slender and twisted into a spiral, and the tip of the inner branch of the paramere is bifid (Figs 6e, 7), while in A. (A.) bella, the tip of the inner branch of the paramere is twisted (Markevičiūtė et al. 2019).

Holotype : China • ♂; Tibet Autonomous Region, Medog County, 80k; 29°28'47"N, 96°05'19"E; 2104 m a.s.l.; 30 July 2014; Tingting Zhang leg; CAU. Paratypes: China • 1 ♂ 2 ♀♀; Tibet Autonomous Region, Medog County, 80k; 29°28'47"N, 96°05'19"E; 2104 m a.s.l.; 1 Aug. 2014; Tingting Zhang leg; CAU.

Antocha (A.) tibetana sp. nov. can be recognized by thorax with three dark brown stripes, wing having indistinct stigma, basal section of M₃ about twice as long as m-m, posterior margin of tergite with shallow emargination and outer gonostylus apically claw-shaped. Aedeagal complex with interbase distally horn-like; paramere apically flattened and triangular; inner branch of paramere with tip bifid into two teeth.

Male. Body length 4.5–5.0 mm, wing length 5.1–5.5 mm, antenna length 1.0–1.2 mm.

Head (Fig. 8b). Dark brown, with brown setae. Antenna brown, with brown setae. Scape nearly cylindrical; pedicel oval; flagellomeres oval, apically shortened. Rostrum and palpus brown, with brown setae.

Figure 8. 

Antocha (Antocha) tibetana sp. nov. a habitus of male, lateral view b male head, lateral view c male thorax, dorsal view d male wing e aedeagus complex with gonocoxite and gonostyli, dorsal view f female ovipositor, lateral view. Scale bars: 1.0 mm (a); 0.3 mm (b, c); 0.5 mm (d); 0.2 mm (e, f).

Thorax (Fig. 8c). Pronotum brown. Prescutum and presutural scutum brownish yellow, with three dark brown stripes. Postsutural scutum brownish yellow; scutal lobes each with a brown spot. Scutellum brown, with middle brownish yellow. Mediotergite brown with side edges light brown. Pleuron brownish yellow (Fig. 8a). Legs with coxae and trochanters yellow; rest of segments brownish yellow. Wing light brownish yellow, with very indistinct stigma; anal angle nearly right-angled (Fig. 8d). Veins brownish yellow. Venation: Sc ending before fork of Rs, at about 5/6 of Rs; basal section of R₅ about 1½ as long as r-m; m-cu shortly before fork of M, distance approximately 1/3 its own length; basal section of M₃ about twice as long as m-m; cell m₁ longer than cell dm. Halter pale with stem light yellow.

Abdomen. Tergites 1–6 brown, tergites 7 and 8 dark brown. Sternites 1–6 brownish yellow; sternites 7 and 8 dark brown.

Hypopygium (Figs 8e, 9). Yellow. Posterior margin of tergite 9 with shallow emargination (Fig. 9a). Gonocoxite nearly cylindrical with brown setae (Figs 8e, 9). Outer gonostylus apically black with tip curved, claw-shaped. Inner gonostylus nearly straight with tip rounded. Interbase nearly V-shaped, distal part flattened and horn-like, with tip blunt (Figs 8e, 9). Paramere with base rod-shaped; apical part flattened, triangular in shape. Inner branch of paramere with tip bifid, two teeth almost equal in length (Figs 8e, 9). Aedeagus rod-shaped, curved ventrally (Figs 8e, 9).

Figure 9. 

Antocha (Antocha) tibetana sp. nov. a male hypopygium, dorsal view b male hypopygium, ventral view. Scale bar: 0.1 mm.

Female. Body length 5.0–5.3 mm, wing length 5.5–5.7 mm. Generally similar to male by body coloration.

Ovipositor (Fig. 8f). Tergite 10 yellowish. Cercus pale yellow, with base darker; tip raised and tapering. Hypogynial valve yellowish, reaching approximately middle of cercus.

The species is named after the type locality, Tibet.

China (Tibet).

The new species is somewhat similar to A. (A.) spiralis from China and India with the similar wing venation and bifid tip of inner branch of paramere, but it can be easily distinguished by the three stripes on the thorax (Fig. 8c) and the triangular apex of the paramere (Figs 8e, 9). In A. (A.) spiralis, the thorax has no obvious longitudinal stripes (Fig. 6c), while the apex of the paramere is slender and twisted into a spiral (Figs 6e, 7).