Holotype: male, CHINA: Tibet, Baqen, 31°50.421'N, 094°18.016'E, 4575 m, 3 Jul. 2010, Ai-Min Shi and Yong-Sheng Pan leg. (MCWNU). Paratype: 21 males and 20 females, 4 paratypes (2 males, 2 females) in ZIN, same data as the holotype.

This new species can be distinguished based on the following: anterior margin of pronotum straight; protibia outer apical angles strongly elongated, inner surface with an obscure spur and strong setae at apical ½; parameres moderately narrowed apically, slightly sinuate near the apex ⅕ (in lateral view).

Named after the type locality, Baqen.

Body black; antennae, legs and palps brown; surface weakly shiny.

Male (Figs 1–8). Head broad, anterior margin of clypeus straight, fronto-clypeal suture slightly obscure; most of genae densely punctate, covered with recumbent hairs. Dorsal surface of head slight convex. Punctation of head fine, dense or moderately dense. Eyes transverse, with shallow emargination at anterior margin. Antennae when posteriorly extended, reaching pronotal base. Length (width) ratio of antennomeres from 2^nd to 11^th as follows: 14.0(11): 25.5(12.5): 14.0(12.5): 13.5(11.5): 15.0(12.0): 14.5(13.0): 17.5(15.0): 15.5(15.0): 15.5(15.5): 20.5(16.0) (n = 5).

Figures 1–10. 

Bioramix (Leipopleura) baqenensis sp. n.: 1 antenna of male 2 pronotum of male 3 protibiae and protarsomeres of male 4 mesotibiae and mesotarsomeres of male 5 metatibiae and metatarsomeres of male 6–7 apical part of aedeagus in dorsal and ventral views 8 apical part of aedeagus in lateral view 9–10 ovipositor in dorsal and ventral views.

Pronotum (Fig. 2) transverse, 1.35–1.51 (1.44 on average, n = 5) times as wide as long, widest before the middle, 1.72–1.82 (1.78 on average, n = 5) times as wide as head. Ratio of pronotal width at anterior margin to its maximum width and width at base (n = 5) 0.69: 1.00: 0.95 on average. Outer margin of pronotum shallowly sinuate in basal ½, more abruptly converging anteriorly in anterior ⅓, finely bordered along entire length. Anterior margin straight, bordered laterally; base weakly bisinuate, not bordered or bordered laterally. Anterior angles of pronotum widely obtuse; posterior ones nearly rectangular. Pronotal surface between outer margins convex, punctures larger and denser than those on head, finer at disc than laterally, lateral margins of pronotum weakly flattened. Intercoxal process not raised, sharply sloping behind procoxae. Prothoracic hypomeron with longitudinal wrinkles.

Elytra elongate-oval, 1.32–1.43 (1.36 on average, n=5) times as long as wide, maximum width before middle, 1.23–1.27 (1.25 on average, n=5) times as wide as pronotum. Outer margin of epipleural reaching sutural angle, visible dorsally only at base. Elytral surface between epipleura and sutural margin convex, with fine dense punctation and minute rugae, with traces of longitudinal prominences and also traces of smooth rounded prominences better developed in apical half. Epipleural surface with densely covered irregular wrinkles and small granules. Mesoventrite with rather sparse recumbent hairs; surface finely granulate and wrinkles. Abdominal ventrites with yellow hairs. Intercoxal process of abdomen rounded apically. First and second abdominal ventrites with shallow medial impressions, 4^th ventrite weakly concave at sides. Last abdominal ventrite with shallow depression and impunctate semicircular area at medial base.

Legs (Figs 3–5) robust, length (width) ratio of pro-, meso- and metafemora 69.0(25.0): 73.5(22.5): 88.0(24.5) (n = 5); that of corresponding tibiae 67.0(27.0): 63.5(17.5): 88.0(18.0) (n = 5). Protibiae 2.37–2.59 (2.49 on average, n = 5) times as long as wide, gradually widening in basal ⅔. Outer apical angles strongly elongate, underside concave. Inner surface with an obscure spur and strong setae in apical ½. Metatibia straight. Plantar surface of proximal pro- and mesotarsomeres 1-4 with setal brushes. Length (width) ratio of pro-, meso- and metatarsomeres from 1^st to 4^th as follows: 8.0 (7.0): 11.0 (15.5): 8.5 (14.5): 4.5 (12.5) (n = 5), 10.0 (8.0): 9.5(12.5): 7.0 (11.0): 5.0 (7.5) (n = 5) and 27.5 (6.5): 13.5 (6.0): 11.5 (5.5): 21.0 (5.0) (n = 5).

Aedeagus (Figs 6–8): length 2.9–3.0 mm, width 0.83 mm. Parameres 1.17 mm long and 0.67 mm wide, moderately narrowed apically, apical ⅕ slightly sinuate (in lateral view).

Female (Figs 9–10). Body wider. Antennae and epipleural carina shorter than in male. Pronotum 1.60–1.72 (1.65 on average, n = 5) times as wide as long, 1.76–1.81 (1.78 on average, n = 5) times as wide as head. Ratio of pronotal width at anterior margin to its maximum width and width at base 0.71: 1.00: 0.95. Elytra 1.27–1.35 (1.32 on average, n = 5) times as long as wide, 1.29–1.40 (1.35 on average, n = 5) times as wide as pronotum. Plantar surface of pro- and mesotarsomeres without setal brush. Ovipositor in dorsal view with golden setae at apical ⅓, tuft of setae (with 3–4 long setae) present near apical margin.

Male body length 9.3–10.4 mm, width 4.7–5.0 mm; female body length 9.1–9.8 mm, width 4.7–5.2 mm.

China: Tibet (Tanggula Shan, Baqen).

Bioramix (Leipopleura) baqenensis Li & Egorov, sp. n. is similar to Bioramix (Leipopleura) nagquana (Meng & Ren, 2005) based on the following characters: length of antennae (when posteriorly extended, reaching pronotal base), anterior margin of clypeus straight, pronotum widest near middle, and metatibia straight.

Holotype: male, CHINA: Tibet, Nyainrong, 32°06.763'N, 92°17.171'E, 4728 m, 6 Aug. 2010, Yun-Chun Li and Yong-Sheng Pan leg. (MCWNU). Paratype: 4 males and 3 females, 3 paratypes (2 males, 1 females) in ZIN, same data as the holotype.

This new species can be distinguished based on the following: shorter antennae (when posteriorly extended, not reaching pronotal base); anterior margin of pronotum emarginated, and metatibia weakly incurved.

Named after the type locality, Nyainrong.

Body black; antennae, legs and palps brown; surface weakly shiny.

Male (Figs 11–18). Head broad, anterior margin of clypeus weakly, but noticeably arcuate in the middle; fronto-clypeal suture slightly obscure; most of genae densely punctate, covered with recumbent hairs. Dorsal surface of head slight convex. Punctation of head coarse large, dense or moderately dense. Antennae very shorter, when posteriorly extended, not reaching posterior ½ of pronotum. Length (width) ratio of antennomeres from 2^nd to 11^th as follows: 11.7(10.3): 20.7(10.0): 12.7(9.3): 12.3(10.0): 13.7(10.3): 13.3(10.3): 14.0(13.3): 12.0(13.0): 13.0(14.0): 17.0(14.7) (n = 3).

Figures 11–20. 

Bioramix (Leipopleura) nyainrongensis sp. n.: 11 antenna of male 12 pronotum of male 13 protibiae and protarsomeres of male 14 mesotibiae and mesotarsomeres of male 15 metatibiae and metatarsomeres of male 16–17 apical part of aedeagus in dorsal and ventral views 18 apical part of aedeagus in lateral view 19–20 ovipositor in dorsal and ventral views.

Pronotum (Fig. 12) transverse, 1.49–1.63 (1.55 on average, n = 3) times as wide as long, widest before in the middle, 1.75–1.87 (1.83 on average, n = 3) times as wide as head. Ratio of pronotal width at anterior margin to its maximum width and width at base (n = 3) 0.66: 1.00: 0.96 on average. Outer margins of pronotum acutely convex, bordered along entire length. Anterior margin emarginate, bordered laterally; base weakly bisinuate, not bordered or bordered laterally. Anterior angles weakly obtuse; posterior ones nearly rectangular. Pronotal surface between outer margins convex, punctures smaller and denser than those on head, finer at disc than laterally, lateral margins of pronotum weakly flattened. Intercoxal process not raised, sharply sloping behind procoxae. Prothoracic hypomeron with longitudinal wrinkles.

Elytra elongate-oval, 1.27–1.33 (1.30 on average, n = 3) times as long as wide, maximum width before middle, 1.10–1.20 (1.14 on average, n = 3) times as wide as pronotum. Outer margin of epipleural reaching sutural angle, visible dorsally only at base. Elytral surface between epipleura and sutural margin convex, with traces of longitudinal carina, elytra apex sharply declined. Epipleural surface covered with dense irregular wrinkles and sparse shallow punctures. Lateral carina of elytra (outer margin of pseudepipleura) visible in dorsal view only anteriorly, explanate on humeri, merging with epipleura, reaching sutural angle. Mesoventrite with rather sparse recumbent hairs; surface finely granulate and wrinkles. Abdominal ventrites with yellow hairs. intercoxal process of abdomen rounded apically. First and second abdominal ventrites with shallow medial impressions.

Legs (Figs 13–15), length (width) ratio of pro-, meso- and metafemora 53.3(19.7): 59.7(18.7): 70.7(19.3) (n = 3); that of corresponding tibiae 53.3(20.7): 54.3(14.0): 72.7(14.3) (n = 3). Protibiae 2.55–2.60 (2.58 on average, n = 3) times as long as wide, gradually widening towards apex, Outer apical angles weakly elongate, underside concave. pro-, meso- and metatibiae with densely golden hairs at apex of inner surface, protibiae with obviously a spur. metatibia weakly incurved. Plantar surface of proximal pro- and mesotarsomeres 1-4 with setal brushes. Length (width) ratio of pro-, meso- and metatarsomeres from 1^st to 4^th as follows: 8.0(6.7): 7.7 (13.3): 6.3 (13.7): 4.7 (10.0) (n = 3), 8.0 (7.0): 8.0 (12.3): 6.3 (10.0): 4.7 (6.3) (n = 3) and 21.7 (6.7): 10.3 (6.0): 8.3 (5.7): 16.0 (6.0) (n = 3).

Aedeagus (Figs 16–18): length 2.4–2.5 mm, width 0.67 mm. Parameres 0.95 mm long and 0.57 mm wide. Parameres noticeably narrowed toward apex in dorsal view, but weakly widened in distal ¼, distinctly sinuate (in lateral view).

Female (Figs 19–20). Body longer and wider. Pronotum 1.57–1.59 (1.58 on average, n = 3) times as wide as long, 1.75–1.77 (1.76 on average, n = 3) times as wide as head. Ratio of pronotal width at anterior margin to its maximum width and width at base (n = 3) 0.67: 1.00: 0.98. Elytra 1.30–1.32 (1.31 on average, n = 3) times as long as wide, 1.19–1.21 (1.20 on average, n = 3) times as wide as pronotum. Plantar surface of pro- and mesotarsomeres without setal brush. Ovipositor in dorsal view with golden setae at apical ¼, and nearly apex shorter setae formation looped pile, within a strong long setae highlight, inner apical surface of with densely setae.

Male body length 7.9–8.1 mm, width 3.9–4.0 mm; female body length 8.1–8.2 mm, width 4.0–4.1 mm.

China: Tibet (Tanggula Shan, Nyainrong).

Bioramix (Leipopleura) nyainrongensis Li & Egorov, sp. n. is similar to Bioramix (Leipopleura) baqenensis Li & Egorov, sp. n. and Bioramix (Leipopleura) nagquana (Meng & Ren, 2005) based on the following characters: pronotum (widest near middle), posterior margin of pronotum base weakly bisinuate, elytra elongate-oval (widest near middle), and outer margin of epipleural visible dorsally only at base.

Holotype: male, CHINA: Tibet, Banbar, 30°55.964'N, 094°42.482'E, 3730 m, 25 Jul. 2010, Ai-Min Shi and Yong-Sheng Pan leg. (MCWNU). Paratype: 59 males and 51 females, 4 paratypes (2 males, 2 females) in ZIN, same data as the holotype.

This new species can be distinguished based on the following: shorter antennae (when posteriorly extended, not reaching pronotal base), anterior margin of pronotum straight, parameres distinctly sinuate near the apex ⅙ (in lateral view).

Named after the type locality, Banbar.

Body dark brown; surface weakly shiny.

Male (Figs 21–28). Head broad, anterior margin of clypeus straight, fronto-clypeal suture slightly obscure; most of genae densely punctate, covered with recumbent hairs. Dorsal surface of head slight convex. Punctation of head fine, dense or moderately dense. Eyes transverse, with shallow emargination at anterior margin. Antennae short, when posteriorly extended, not apices reaching base of pronotum. Length (width) ratio of antennomeres from 2^nd to 11^th as follows: 15.8(11.6): 27.2(12.2): 18.4(12.0): 17.4(12.0): 17.6(12.0): 16.8(12.2): 18.8(15.6): 15.8(14.8): 15.8(14.6): 22.4(14.0) (n = 5).

Figures 21–30. 

Bioramix (Leipopleura) banbarensis sp. n.: 21 antenna of male 22 pronotum of male 23 protibiae and protarsomeres of male 24 mesotibiae and mesotarsomeres of male 25 metatibiae and metatarsomeres of male 26–27 apical part of aedeagus in dorsal and ventral views 28 apical part of aedeagus in lateral view 29–30 ovipositor in dorsal and ventral views.

Figures 31–36. 

31–32Bioramix (Leipopleura) baqenensis sp. n.: 31 male, length 9.3–10.4 mm 32 female, length 9.1–9.8 mm 33–34B. (L.) nyainrongensis sp. n.: 33 male, length 7.9–8.1 mm 34 female, length 8.1–8.2 mm 35–36B. (L.) banbarensis sp. n.: 35 male, length 9.9–10.6 mm 36 female, length 10.0–10.5 mm.

Pronotum (Fig. 22) transverse, 1.41–1.51 (1.47 on average, n = 5) times as wide as long, widest in the middle, 1.69–1.79 (1.75 on average, n = 5) times as wide as head. Ratio of pronotal width at anterior margin to its maximum width and width at base (n = 5) 0.64: 1.00: 0.88 on average. Outer margins of pronotum acutely convex, bordered along entire length. Anterior margin straight, bordered laterally; base weakly bisinuate, not bordered. Anterior angles of pronotum widely obtuse, posterior ones weakly obtuse. Pronotal surface between outer margins convex, with trace of slight depression. Punctuation on disc similar to that on frons, mainly dense, coarser and denser at sides. Prosternum with rather sparse hairs directed backwards and small granules. Intercoxal process not raised, sharply sloping behind procoxae. Prothoracic hypomeron with obvious longitudinal wrinkles and coarse granules.

Elytra elongate-oval, 1.28–1.45 (1.35 on average, n = 5) times as long as wide, widest nearly in the middle, 1.21–1.34 (1.28 on average, n = 5) times as wide as pronotum. Outer margin of epipleural not reaching sutural angle, visible dorsally only at base. Elytral surface between epipleura and sutural margin convex, with sparse and shallow punctures, irregular fine wrinkles and traces of longitudinal carina. Elytra margin setae and abdomen setae subequal length, elytra apex sharply declined. Mesosternum with rather sparse recumbent hairs, surface finely granulate and wrinkles. Abdominal ventrites with yellow hairs. First and second abdominal ventrites with shallow medial impressions. Last abdominal ventrite with depression and impunctate semicircular area at medial base, apical margin widely rounded.

Legs (Figs 23–25) robust, length (width) ratio of pro-, meso- and metafemora 70.4(25.0): 76.2(23.0): 93.2(24.6) (n = 5); that of corresponding tibiae 68.4(25.6): 68.6(19.4): 98.4(19.2) (n = 5). Protibiae 2.45–2.95 (2.78 on average, n = 5) times as long as wide, gradually widening towards apex, apical margin extended but not forming sharp triangle, underside concave. Pro- and mesotibia subequal length. Protibia with an obscure spur, metatibia nearly straight. Plantar surface of proximal pro- and mesotarsomeres 1-4 with setal brushes. Length (Width) ratio of pro-, meso- and metatarsomeres from 1^st to 4^th as follows: 8.6(8.2): 10.4 (17.4): 8.2 (16.0): 5.4 (9.8) (n = 5), 9.2 (8.2): 9.6 (15.4): 8.4 (12.8): 6.0 (7.4) (n = 5) and 28.2 (7.8): 13.4 (6.6): 10.6 (6.0): 21.6 (5.8) (n = 5).

Aedeagus (Figs 26–28): length 3.1–3.2 mm, width 0.93 mm. Parameres 1.15 mm long and 0.70 mm wide, moderately narrowed apically, apical ⅙ distinctly sinuate (in lateral view).

Female (Figs 29–30). Body longer and wider. Pronotum 1.50–1.57 (1.54 on average, n = 5) times as wide as long, 1.79–1.88 (1.82 on average, n = 5) times as wide as head. Ratio of pronotal width at anterior margin to its maximum width and width at base (n = 5) 0.70: 1.00: 0.95. Elytra 1.32–1.39 (1.35 on average, n = 5) times as long as wide, 1.24–1.34 (1.28 on average, n = 5) times as wide as pronotum. Plantar surface of pro- and mesotarsomeres without setal brush. Ovipositor in dorsal view with golden setae at apical ⅖, tuft of setae (with 7–8 long setae) present near apical margin.

Male body length 9.9–10.6 mm, width 4.9–5.4 mm; female body length 10.0–10.5 mm, width 5.0–5.4 mm.

China: Tibet (Nyainqentanglha Shan (Nyenchen Tanglha Mountains), Banbar).

Bioramix (Leipopleura) banbarensis Li & Egorov, sp. n. is similar to Bioramix (Leipopleura) crypticoides (Reitter, 1887) based on the following characters: pronotum widest near middle, anterior and posterior angles of pronotum obtuse, base weakly bisinuate, protibia apical margin extended not sharp apices, and metatibia nearly straight.

The world fauna of the tribe Platyscelidini comprises 8 genera, 28 subgenera (Egorov 2004, 2009) and found in Palaearctic area. moreover, its southern border coincides with the part of Palaearctic southern border in the Himalayas. The majority of species live in the steppe and mountain habitats, lesser number in semideserts. The mountain regions of Asia, such as the Tien Shan, the Pamirs, the Hindu Kush, the Karakorum Range, the unlun Shan, the Himalayas, as well as Northern and Central China mountains are the main centers of species diversity. The classification of the tribe, developed in detail by Kaszab (1940) and Egorov (2004, 2009).