Holotype. ♂ (ZISP 1437), Georgia, Adjara, near Kobuleti, Ispani 2 bog, 41.86461°N, 41.79763°E, main bog, site 3 (600 m E of W Alnus belt), flat Sphagnum carpet, sample I-11, 10.V.2019, A. Przhiboro leg.

Paratypes. 1 ♂ (ZMMU TI-65), 1 ♀ (ZISP 1438), Georgia, Adjara, near Kobuleti, Ispani 1 bog, 41.85805°N, 41.78750°E, site 1 (150 m E of NW bog edge), dry Sphagnum hummocks, sample I-18, 11.V.2019, A. Przhiboro leg.; 1 ♂ (ISEA Ps. 001.0037), the same locality, habitat, and collector, sample I-37, 10.X.2019.

Carapace with moderately long and apically rounded epistome; two pairs of eyes present, lenses of posterior eyes with low convexity; movable cheliceral finger without large median tooth; pedipalpal femur and patella smooth (sometimes very rare and weak granules on outer side of femur); chelal hand with faint reticulate ornament; notch on median side of patella reaching almost middle of patellar club length; movable finger distinctly longer than hand⁺ but almost equal to femur in length; trichobothrium ist situated distal to middle of fixed chelal finger; distal half of fixed chelal finger with teeth almost equal in size and shape; anterolateral process of coxa of leg I pointed and slightly enlarged, its mediolateral process slightly prominent, denticulate; sternite II with 5 setae; pedipalpal femur 4.5–5.1× as long as broad (0.77–1.00/0.15–0.22), chelal hand⁺ 1.62 × as long as broad (0.62–0.71/0.38–40); movable chelal finger length 0.79–0.95.

♂ and ♀. Coloration: carapace distinctly darker than tergite I and slightly paler than pedipalp; pedipalpal segments uniformly coloured.

Carapace (Figs 4A, 6C, H, J): with posterior border partially unsclerotised and pale, covered with fine reticulate ornament (Fig. 6H), wider than long, 0.79 × (0.83–0.9 ×) as long as broad, with two pairs of corneate eyes, anterior eyes slightly larger than posterior ones; distance between anterior margin of anterior eye and anterior margin of carapace 0.05, diameter of anterior eye 0.10 (0.08–0.09), diameter of posterior eye 0.09 (0.07–0.09), distance between eyes 0.02; carapace with 27 (23–24) setae, its anterior margin with 6 (4–6) setae: 4 macrosetae and 2 marginal microsetae (one paratype without these microsetae); posterior margin with 9 (6–7) setae, chaetotaxy: m4m:6:6:9 (m4m:6:6:6, 4:6:6:7); epistome prominent, short and apically rounded, 0.025/0.040; glandular pores present; anterolateral corners with two small protuberances located below surface of carapace; carapace with 6 (5–6) microlyrifissures: one pair situated in ocular zone, close to anterior eyes, and two pairs located at posterior margin.

Tergites weakly sclerotised; all setae simple; tergite X with 2 (2–4) pseudotactile and 2 tactile setae; tergite XI with 4 tactile setae; chaetotaxy: 6:9:11:11:11:12:12:13:11:4T2T4:T2T2T2T:2 (6:8–9:10–11:11–12:11:12:11:10–12:11:4T1–2T4:T2T2T2T:2).

Sternites entirely smooth, weakly sclerotised; anterior operculum of males with 5 and posterior operculum of males with 17 (16–17) setae, of which 7 (6–7) setae located close to genital aperture (Fig. 4C); sternite II of female with 6 setae, sternite III of female: (3)18(3). All setae simple; those on sternites IV–XI uniseriate; chaetotaxy: 5:(3)17(3):(3)12(3):16:16:16:16:16:6T1T6:T2T3T2T:2 (5–6:(3)16–18(3):(3)12–13(3):15–17:14–15:15:14–16:13–15:4–6T1T4–6:T3T2T3T:2).

Internal genitalia with moderately long lateral and median genital sacs, median sac wrinkled, same length as lateral sacs; genital opening with 4+4 internal setae (Fig. 4D).

Pleural membrane granulated. Chelicera (Figs 5D–F, 6F): hand with 7 acuminate setae; galea knob-like, with poorly developed hyaline convexity, sub-galeal seta situated distal to middle (T = 0.7); hand covered with faint reticulate pattern with pointed corners (Fig. 5D); small tooth (tis) under interior seta (is) present, triangular (Fig. 6F); fixed finger with 18 (17–19) teeth reaching finger base; movable finger with 7 (8–10) teeth reaching distal to middle of segment, large median tooth absent; serrula interior with 22 (22–25) blades, serrula exterior with 26 (25–28) blades; rallum with 8 blades, two distal ones denticulate, 6 posterior blades simple, smooth and acuminate, 2 or 3 proximal blades smallest.

Coxae (Fig. 4B): pedipalpal coxa excluding manducatory process with 7 or 8 setae, manducatory process with 4 acuminate setae, seta at base of manducatory process longest; coxa I with moderately long triangular, sclerotised, apically pointed anterolateral process (0.06/0.04), and with denticulate mediolateral process; coxal chaetotaxy of legs: 6:6–8:7:10–11 (5–6:6–8:7:10–11). Each coxa of legs with one lyrifissure, pedipalpal coxa with one maxillary lyrifissure.

Pedipalp (Figs 5A–C, 6B, D, E, G, I): femur and patella smooth (in one paratype, outer side of femur finely granulate), hand of chela covered with fine reticulate ornament (Fig. 6D), pedicel of chela with distinct ornament; trochanter with small dorsal tubercle, 2.0 × (1.60–1.85 ×) as long as broad; femur with short pedicel, its margins of femur without tubercles, some setae mostly located in basal half of segment, without enlarged alveoli, one glandular pore located dorsodistally, 4.5 × (5–5.1 ×) as long as broad; patella with short and stout pedicel (0.20–0.22), patella distinctly shorter than femur, 1.43 × (1.54–1.66 ×) as long as broad, with 3 lyrifissures basally and 2 lyrifissures distally, notch on median side reaching very close to middle of club length. Chela⁺ 3.8 × (3.68–3.87 ×) and chela- 3.7 × (3.52–3.75 ×) as long as broad; movable finger distinctly longer than hand⁺, 1.31 × (1.27–1.33 ×), but almost equal to femur in length, 0.95 × (0.95–1.02 ×); hand⁺ 1.62 × (1.63–1.77 ×) and hand- 1.52 × (1.57–1.62 ×) as long as broad; retrolateral surface of hand with 3 glandular pores located around trichobothria eb and esb (two pores close to each other, one distant); fixed finger with 3 lyrifissures: fa located close to base in retrolateral view, fb located slightly proximal to ib, fd located distal to it; movable finger with 3 lyrifissures in retrolateral view: ma₂ located closer to trichobothrium b than to sb, ma₁ nearly at same level with sb (slightly proximal to sb), ma₃ closer to st than to sb, ma₁ slightly closer to ma₃ than to ma₂; one or two sensilla pc located between trichobothria sb and st, close to dental canals (one sensillum closer to sb, other closer to st; latter not present in all specimens); fixed finger with 48 (48–51) contiguous triangular teeth reaching level of trichobothrium ib, two outer distal teeth smallest, other teeth almost equal in size, all teeth with dental canal; movable finger with 45 (44–47) contiguous teeth not reaching level of trichobothrium b, teeth slightly reduced in size from tip of base of, distal 10–12 teeth triangular, other blunt and apically rounded, teeth with dental canal (except basal 6 or 7 small teeth); nodus ramosus of venom duct in fixed chelal finger short, situated very close to finger tip.

Trichobothriotaxy: fixed finger with 8 trichobothria, movable finger with 4 trichobothria; fixed finger with trichobothria esb and eb located close to each other and slightly proximal to ib, with isb on retrolateral surface, with ib closer to isb than to esb (in lateral and dorsal views), trichobothrium ist distinctly closer to est than to isb, distance ist–est similar to distance est–it, but less than twice as long as distance est–et in lateral view, trichobothrium ist situated distal to middle of finger (T = 0.57–0.60), ist located distinctly proximal to t, et located posterior to it, at approximately the same distance between est and it (in lateral and dorsal views), ist situated distal to middle of finger; movable finger with trichobothrium st situated slightly closer to t than to sb, trichobothrium sb slightly closer to b than to st (at almost equal distances), distance b–sb almost equal to distance st–t.

Legs (Figs 4E, F, 6A): all claws of legs with small dorsal accessory tooth (Figs 4E, 6A), arolia simple and shorter than claws. Leg I femur 4.00 × (4.44–5.50 ×) and patella 3.30 × (3.30–3.40 ×) as long as deep, femur 1.21 × (1.48–1.66 ×) as long as patella, tibia 4.12 × (4.28–4.37 ×), metatarsus 3.00 × (3.12–3.14 ×) and tarsus 4.28 × (4.0–4.28 ×) as long as deep, tarsus 1.25 × (1.20–1.27 ×) as long as metatarsus; leg IV femur 2.00 × (1.76–2.00 ×), patella 2.22 × (2.05–2.63 ×), femur+patella 4.22 × (3.94–4.50 ×), tibia 6.0 × (5.80–6.36 ×), metatarsus 3.75 × (3.75–4.12 ×) and tarsus 5.71 × (5.42–5.75 ×) as long as deep, tarsus 1.33 × (1.26–1.39 ×) as long as metatarsus; tibia IV with long tactile seta situated slightly proximal to middle (T = 0.40), metatarsus IV with long tactile seta situated basally (T = 0.125), tarsus with tactile seta situated proximal to middle (T = 0.38); sub-terminal setae branched, basal ramus short and smooth (Fig. 4F).

Sexual dimorphism not pronounced. Measurements: body length 2.95 (2.87–2.90); carapace 0.65/0.82 (0.64–0.65/0.72–0.77); chelicera 0.45/0.24 (0.40–0.50/0.22–0.26), movable finger of chelicera 0.28 (0.26–0.30); pedipalp: trochanter 0.40/0.20 (0.37–0.40/0.20–0.25), femur 0.90/0.20 (0.77–1.00/0.15–0.20), patella 0.63/0.24 (0.50–0.60/0.20–0.23), chela⁺ 1.52 (1.40–1.55), chela- 1.48 (1.34–1.50), hand⁺ 0.65/0.40 (0.62–0.71/0.38–0.40), hand- 0.61 (0.60–0.65), movable finger 0.86 (0.79–0.95); leg I: trochanter 0.18/0.15 (0.18–0.22/0.15), femur 0.40/0.10 (0.40–0.55/0.09–0.10), patella 0.33/0.10 (0.27–0.33/0.08–0.10), tibia 0.33/0.08 (0.30–0.35/0.07–0.08), metatarsus 0.24/0.08 (0.22–0.25/0.07–0.08), tarsus 0.30/0.07 (0.28–0.30/0.07); leg IV: trochanter 0.40/0.15 (0.40/0.18), femur 0.36/0.18 (0.30–0.40/0.17–0.20), patella 0.40/0.18 (0.37–0.50/0.18–0.19), femur+patella 0.76 (0.67–0.90), tibia 0.60/0.10 (0.58–0.70/0.10–0.11), metatarsus 0.30/0.08 (0.30–0.33/0.08), tarsus 0.40/0.07 (0.38–0.46/0.07–0.08).

The new species is most similar to N. (N.) vilcekii and N. (N.) speleophilum to the following characters: tarsal claw IV with dorsal accessory tooth, palpal femur without granulate, trichobothrium ist located distinctly proximal to t. The new species differs from N. (N.) vilcekii in the following characters: shorter pedipalpal femur (0.77–1.00 vs. 1.35–1.72 in N. (N.) vilcekii), shorter hand⁺ (0.62–0.71 vs. 1.07–1.45) and movable finger (0.79–0.95 vs. 1.32–1.57), higher ratio of length of movable finger to hand⁺ (1.27–1.33 × vs. 1.06–1.23 ×), lower ratio of length to width of chelal hand⁺ (1.62–1.77 × vs. 1.81–2.15 ×), longer epistome (moderately long vs. short), smaller number of setae on sternite II (5 vs. 10–15), longer anterolateral process on coxa I (0.06/0.04 vs. 0.04/0.05), smaller number of setae on pedipalpal and leg coxae (see Table 1), smaller number of setae on manducatory process of pedipalp (4 vs. 5), and the presence/absence of large median tooth on movable finger of chelicera (absent vs. present) (Krumpál 1983; Nassirkhani and Snegovaya 2021). The new species differs from N. (N.) speleophilum in the following characters: shorter chela⁺ (1.40–1.55vs. 1.72–1.80 in N. (N.) speleophilum), hand⁺ (0.62–0.71 vs. 0.73–0.80), and movable finger (0.79–0.95 vs. 1.12–1.20), lower ratio movable finger/hand⁺ (1.27–1.33 × vs. 1.45–1.53 ×), sternite II with 5 setae vs. 11 or 12, longer (as long as lateral sacs) median sac of male genitalia vs. short (lateral sac longer than median) (Krumpál 1986; Nassirkhani et al. 2020).

Known only from two adjacent Sphagnum bogs, Ispani 1 and Ispani 2, situated near the town of Kobuleti in the Republic of Adjara (Georgia).

In Ispani 1 bog, N. (N.) adjaricum sp. nov. was collected in its northwestern part (site 1), with large relatively dry hummocks (20–40 cm high, 1–2 m wide) and moist flat spaces between them (Fig. 2A). The new species was collected only from hummocks. The hummocks consisted mainly of loose Sphagnum (with predominant S. papillosum, S. austinii, and S. palustre; in particular, the latter species was predominant in sample I-37, where one paratype was collected; Fig. 2B), with common Juncus effusus L., Molinia caerulea arundinacea (Schrank) K. Richt., Rhododendron spp., and Pteridium tauricum V.I. Krecz. Sphagnum was dry to slightly wet, with pH 2.9–3.0 and mineralization of 0.04–0.08 ppt.

In Ispani 2 bog, N. (N.) adjaricum sp. nov. was collected from a flat site in its (site 3; Fig. 1A) covered by a flat carpet of Sphagnum, with S. papillosum and S. palustre predominant, and with the thinned stands of M. caerulea arundinacea (Fig. 1B). Sphagnum was wet to moist, with a pH of 4.0 and mineralization of 0.07 ppt.

The new species seems to be rare, considering that only 1–3 specimens were collected at each site, in 10 quantitative samples (sample size of 0.05 m²). As distinct from other pseudoscorpion species, all specimens of N. (N.) adjaricum sp. nov. were collected only from quantitative samples and no specimens were collected by other techniques (sifting, pitfall trapping, and/or rearing from substrata in the lab).

The Latin adjective adjaricum indicates that the new species is named after the Republic of Adjara (= Ajaria), Georgia, where it was discovered.

Holotype ♂ (SNM 28SR), Russia, North Ossetia–Alania, Prigorodnyi Distr., near Karmadon, 1.VI.1976, Mikuláš J. Lisický leg.

Carapace with short and apically rounded epistome; two pair of eyes present, lenses of posterior eyes with low convexity; movable cheliceral finger with large median tooth; pedipalpal segments smooth, covered with fine reticulate ornament; notch on median side of pedipalpal patella not extending from distal third of patellar club length; movable finger longer than hand⁺, but almost equal to femur in length; trichobothrium ist situated distal to middle of fixed chelal finger; distal half of fixed chelal finger with teeth almost equal in size and shape; anterolateral process of coxa of leg I pointed and relatively short, with mediolateral process slightly prominent, denticulate; sternite II with 15 setae; pedipalpal femur 4.41 × as long as broad (1.41/0.32), chelal hand⁺ 1.89 × as long as broad (1.19/0.63); movable chelal finger length 1.40.

♂. Coloration: carapace reddish brown, opisthosoma and legs paler.

Carapace (Figs 7A, 9A–C): without transverse furrows, covered with reticulate ornament (Fig. 9A); 0.71 × as long as broad, with two pairs of corneate eyes (Figs 7A, 9B), anterior eyes slightly larger than posterior ones (0.11 vs. 0.09); distance between anterior margin of anterior eye and anterior margin of carapace 0.09, diameter of anterior eye 0.07, diameter of posterior eye 0.08, distance between eyes 0.02; carapace with 24 setae, anterior margin with 6 macrosetae, posterior margin with 7 setae; chaetotaxy: 6:7:4:7; all anterior setae almost equal in length (Fig. 7A); epistome short and apically rounded, 0.03/0.05 (Figs 7A, 9C); glandular pores present (Fig. 7A); anterolateral corners with two small protuberances; carapace with 6 microlyrifissures: one pair situated in ocular zone, close to anterior eyes, and two pairs located at posterior margin (Fig. 7A).

Tergites undivided, posterior ones damaged; all setae acuminate; chaetotaxy of tergites I–IX: 5:6:11:12:11:12:12:13:12.

Sternites undivided, posterior ones damaged; all setae acuminate; anterior operculum with 15 setae and one lyrifissure, posterior operculum with 38 setae, of which 19 located close to genital aperture and two lyrifissures (Fig. 8A); chaetotaxy of sternites II–IX: 15:(5)38(6):(4)16(4):19:17:15:15:15.

Internal genitalia (Fig. 9D) with moderately long lateral and median genital sacks, median sack wrinkled; genital opening with 8+7 internal setae.

Pleural membrane granulated (Fig. 9E). Chelicera (Figs 7B, C, 9F): 1.51 × as long as broad; hand with 7 acuminate setae and two lyrifissures (Fig. 7B); galea knob-like, with poorly developed hyaline convexity; sub-galeal seta situated distal to middle (T = 0.11) of movable finger; small tooth (tis) under interior seta is present, blunt (Fig. 9F); hand covered with faint reticulate pattern with pointed corners; fixed finger with 13 teeth reaching finger base, movable finger with 6 teeth reaching middle of finger, large median tooth present; serrula interior with 20 blades, serrula exterior with 27 blades; rallum (Fig. 7C) with 8 blades, two distal ones denticulate, 6 posterior blades simple, smooth and acuminate, 3 proximal blades smallest.

Coxae (Figs 7D, 9G): pedipalpal coxa excluding manducatory process with 10 setae, manducatory process with 5 acuminate setae; coxa I with short triangular, sclerotised, apically pointed anterolateral process (0.04/0.05) (Fig. 9G), mediolateral process denticulate; coxal chaetotaxy of legs: 10–11:10–12:8–9:13, all setae acuminate; lyrifissures: two on each pedipalpal coxa, one on each coxa I–III of legs, not visible on pedal coxae IV; pedipalpal coxa with one maxillary lyrifissure.

Pedipalp (Figs 8B, C, 9H–K): pedipalpal segments smooth (Fig. 9H); all setae acuminate; trochanter with small dorsal tubercle and one lyrifissure (Fig. 8B), 2.03 × as long as broad; femur with short pedicel, margins of femur without tubercles, 5 glandular pores present (Fig. 8B), 4.41 × as long as broad; patella with short pedicel, 3 glandular pores present (Fig. 8B), 2.67 × as long as broad, with 3 lyrifissures basally and 2 distally (Fig. 8B); chela⁺ 3.76 × and chela- 3.57 × as long as broad; movable finger distinctly longer than hand⁺, 1.18 ×, but almost equal to femur in length, 0.99 ×; hand⁺ 1.89 × and hand- 1.68 × as long as broad, pedicel with faint reticulate pattern with pointed corners (Fig. 9I); retrolateral surface of hand with one glandular pore located near trichobothrium eb (Fig. 8C); fixed finger with one lyrifissure: fb located slightly proximal to trichobothrium ib (Fig. 9J), fa and fd not visible; lyrifissures on movable finger not visible; two coupled sensilla pc situated between st and sb (Figs 8C, 9K); fixed finger with 65 contiguous triangular teeth reaching level of trichobothrium ib, teeth almost equal in size; movable finger with 54 contiguous teeth not reaching level of trichobothrium b, teeth blunt and apically rounded; nodus ramosus of venom duct in fixed chelal finger short, situated very close to finger tip (Fig. 8C).

Trichobothriotaxy (Fig. 8C): fixed finger with 8 trichobothria, movable finger with 4 ones; fixed finger with close-set trichobothria esb and eb located slightly proximal to ib, with isb on retrolateral surface, ib closer to isb than to esb, ist distinctly closer to est than to isb, distance ist–est approximately the same as distance est–it, but less than twice as long as distance est–et in lateral view, trichobothrium ist situated distal to middle of finger, ist located distinctly proximal to t, et located proximal to it, at approximately the same distance between est and it (in lateral and dorsal views), ist situated distal to middle of finger; movable finger with trichobothrium st situated slightly closer to t than to sb, trichobothrium sb slightly closer to b than to st, distance b–sb almost equal to distance st–t.

Legs (Fig. 7E): all claws of legs with small dorsal accessory tooth, arolia simple and shorter than claws. Leg I trochanter 1.42 ×, femur 5.79 ×, patella 4.07 ×, tibia 5.82 ×, metatarsus 3.80 × and tarsus 5.40 × as long as deep. Leg IV trochanter 2.11 ×, femoropatella 3.75 ×, tibia 6.33 ×, metatarsus 4.23 × and tarsus 6.82 × as long as deep; tibia IV with long tactile seta situated slightly proximal to middle (T = 0.21 in length, 0.52 from base), metatarsus IV with long tactile seta situated basally (visible only base of trichobothrium, 0.08 from base), tarsus IV with tactile seta situated approximately in middle of segment (T = 0.35, 0.37 from base); sub-terminal setae branched.

Measurements: body length 3.65; carapace 0.92/1.30; chelicera 0.59/0.39, movable finger of chelicera 0.42; pedipalp: trochanter 0.69/0.34, femur 1.41/0.32, patella 1.04/0.39, chela⁺ 2.37, chela- 2.25, hand⁺ 1.19/0.63, hand- 1.06, movable finger 1.40; leg I: trochanter 0.34/0.24, femur 0.81/0.14, patella 0.57/0.14, tibia 0.64/0.11, metatarsus 0.38/0.10, tarsus 0.54/0.10; leg IV: trochanter 0.57/0.27, femur+patella 1.35/0.36, tibia 1.14/0.18, metatarsus 0.55/0.13, tarsus 0.75/0.11.

The states of several characters reported here slightly differ from those in the original description of N. (N.) vilcekii (Krumpál 1983), viz. the number of setae on the posterior carapace margin (7 vs. 8); number of setae on the tergites III (11 vs. 10), V (11 vs. 12) and VIII (13 vs. 12); number of setae on the sternites III [(5)38(6) vs. (6)37(6)], VI (17 vs. 16), VIII and IX (each with 15 vs. each with 14); number of internal setae in the genital opening (8 + 7 vs. 7 + 7); number of teeth on the fixed and movable cheliceral fingers (13 and 6 vs. 11 and 4); number of setae on the coxae II of legs (10–12 vs. 11–12) and pedal coxae III (8–9 vs. 9–11) and IV (13 vs. 12–13); body length (3.65 vs. 3.04); carapace (0.92/1.30 vs. 0.69/0.95); pedipalpal trochanter (0.69/0.34 vs. 0.60/0.32). The numbers of setae on two last tergites and sternites are missing in the present redescription because these segments were damaged on the slide mount, but the respective data are given in the original description (tergites X with 12, XI with 10 and sternites X with 16, XI with 12 setae; see Krumpál 1983). Several new characters were added in the present redescription, namely, the measurements and ratio of length to width of chelicera, measurements and ratio of length to depth of trochanters of leg I and leg IV, presence of glandular pores, number of lyrifissures on carapace, number of blades on serrula exterior and interior, presence of tooth tis on chelicera, number of lyrifissures on palps and coxae.

The holotype differs from the specimens described by Nassirkhani and Snegovaya (2021) in the following characters: the number of setae on the anterior carapace margin (6 vs. 4), number of setae on the sternites II (15 vs. 10–14) and III (38 vs. 39–47); number of setae around male genital aperture on the sternite III (19 vs. 21–27), number of internal setae in the genital opening (8 + 7 vs. 5–6+5–6), number of teeth on fixed and movable cheliceral fingers (13 and 6 vs. 15–18 and 10–12), chela⁺ (3.76 × vs. 4.00–4.33 ×) and chela- of the male (3.57 × vs. 3.84–4.12 ×); chelal hand⁺ (1.89 × vs. 1.98–2.15 ×) and hand- of the male (1.68 × vs. 1.73–1.89 ×), number of the pc sensilla (two, between st and sb, vs. one, proximal to sb), number of teeth on the movable chelal finger of male (54 vs. 56–62), and number of setae on the coxa IV (13 vs. 15–22). The states of most characters mentioned here are within the variability range of this species (Krumpál 1983; Nassirkhani and Snegovaya 2021).

The species is known only from two localities in the Republic of North Ossetia–Alania (Russia) in the North Caucasus: the type locality near Karmadon (ca. 1500 m a.s.l.), and another locality in the valley of the Terek River in the environs of Mozdok (115 m a.s.l.) that was reported by Nassirkhani and Snegovaya (2021). The localities are situated at different altitudes, hence they should strongly differ from each other in the landscape and climatic conditions.

The habitat of N. (N.) vilcekii near Karmadon is unclear (Krumpál 1983). It is questionable whether the species is confined to mire or wet habitats. The material from the environs of Mozdok that was used by Nassirkhani and Snegovaya (2021) for redescription was collected in a floodplain, in litter consisting of leaves of broad-leaved trees mixed with wood pieces. Hence, it is possible that N. (N.) vilcekii is a hygrophilous species.