Short Communication |
Corresponding author: Christophe Gouraud ( collection.baillon@gmail.com ) Academic editor: George Sangster
© 2016 Christophe Gouraud, Sylke Frahnert, Anita Gamauf, Steven van der Mije.
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation:
Gouraud C, Frahnert S, Gamauf A, van der Mije S (2016) Review of the type series of Pterocles exustus Temminck, 1825 (Aves, Pterocliformes, Pteroclidae) and designation of a lectotype. ZooKeys 580: 145-152. https://doi.org/10.3897/zookeys.580.7892
|
The type locality of Pterocles exustus Temminck, 1825, is ‘West coast of Africa, Egypt and Nubia’. This is problematic because it includes the type locality of Pterocles exustus floweri (Nicoll, 1921), which is Fayum, Egypt. In the interest of clarification and to preserve stability of nomenclature, a non-Egyptian specimen from the type series of P. exustus is designated as lectotype of the taxon, and the type locality is restricted to Senegal.
Chestnut-bellied Sandgrouse, lectotypification, type locality
The Chestnut-bellied Sandgrouse (Pterocles exustus) is a sedentary and nomadic species that naturally inhabits bare semi-deserts from Senegambia and Mauritania to Egypt, Sudan, Ethiopia and Somalia, as well as in Kenya, Tanzania, Arabian Peninsula and further east to Iran, Pakistan and India (
P. e. exustus Temminck in Temminck and Laugier de Chartrouse, 1825. Senegal, Gambia and Mauritania to Sudan,
P. e. floweri Nicoll, 1921. Endemic to the Nile Valley (Egypt). This subspecies was believed to be extinct until its recent rediscovery (
P. e. ellioti Bogdanov, 1881. SE Sudan E to Eritrea, N Ethiopia, Somalia. Includes somalicus Hartert, 1900,
P. e. olivascens (
P. e. erlangeri (
P. e. hindustan R. Meinertzhagen, 1923. SE Iran, Pakistan and India. The name hindustan was introduced by Meinertzhagen to replace Pterocles exustus orientalis Hartert, 1900, pre-occupied by Tetrao orientalis Linnaeus, 1758. The type designated by Meinertzhagen has no type status as this author simply introduced a nomen novum (see
In his description, Temminck (in Temminck and Laugier de Chartrouse 1825: pl. 354 and 360 + text) wrote:
The Sandgrouse described herein was sent in great number from West coast of Africa… Since then, Berlin and Frankfurt’s collections received specimens from travellers who explore Egypt for zoological discoveries purposes. Specimens received from this country by the Prussian naturalists [Friedrich Wilhelm Hemprich (1796-1825) and Christian Gottfried Ehrenberg (1795-1876)] and Mn. Rüppell [Wilhelm Peter Eduard Simon Rüppell (1794-1884)] do not differ from those received from Senegal [translated from French].
The author ended with:
Inhabits the West coast of Africa, Egypt and Nubia. Museums of Leiden, Paris, Berlin, Wien and Frankfurt [translated from French].
All specimens used by Temminck to describe his Pterocles exustus constitute a type series and therefore all these specimens are syntypes (Art. 73.2 of the Code, see
Naturalis Biodiversity Center, Leiden (formerly
Naturhistorisches Museum, Wien (hereafter
Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Berlin (hereafter
Specimens of Pterocles exustus were sent by Hemprich and Ehrenberg to Berlin in their shipments numbered 7, 8 and 10 (
There is another questionable specimen (
Musée George Sand et de la Vallée Noire, La Châtre (hereafter MLC): MLC.2011.0.1184 (female) from Upper Egypt (collected and given by Rüppell), designated as “Probable syntype” by
The Forschunginstitut und Naturmuseum Senckenberg (formerly Senckenberg Museum Frankfurt am Main, hereafter
Distributions of ellioti, olivascens, erlangeri and hindustan are well established and do not present any nomenclatural or taxonomical issues in respect to the nominate subspecies. However, the situation between floweri and exustus is problematic and necessitates a review. When describing his floweri, from Upper Egypt and Fayum,
From the above discussion, the lectotype should be designated amongst specimens
Pterocles exustus Temminck in Temminck and Laugier de Chartrouse 1825: pl. 354 and 360 + text.
Lectotype (hereby designated):
Paralectotype:
Paralectotype:
Paralectotype:
Paralectotype:
Paralectotype:
Probable paralectotype: MLC.2011.0.1184, adult female (mount), Upper Egypt, Egypt. Collected by/for Rüppell.
We are grateful to Robert Prŷs-Jones who drew our attention to the issue discussed here and valuably commented an earlier version of the manuscript. Gerald Mayr provided useful information about material housed in