Camptoscaphiella fulva Caporiacco, 1934, by monotypy.

Males of this genus can easily be separated from all other oonopid genera by the heart-shaped sternum with conical projection on the anterolateral corners, spination of the first two legs (tibia I and II with four pairs of long spines, and metatarsus I and II with two pairs of long spines); the extremely large, club-shaped palpal patella; and a cymbium that is not fused with the bulb. Females of this genus are similar to those of Ischnothyreus Simon, 1893, but can be separated by lacking the distinct, darkly sclerotized, strongly winding duct and uniquely shaped atrium (revised from Baehr and Harvey 2013).

China (Yunnan), New Caledonia, South Asia (Bhutan, India, Pakistan, Sri Lanka), Southeast Asia (Thailand).

Holotype ♂ China, Yunnan, Menglun, XTBG, Paramichelia baillonii plantation; 21°53.823'N, 101°17.072'E; 613 m; pitfall traps; 1–15 May 2007; Guo Zheng leg. (SYNU-481). Paratypes [same data as holotype except where indicated] 1♂ (SYNU-482); 1♂ (SYNU-494); 1♂1♀, pitfall traps, 16–31 June 2007 (SYNU-483–484); 1♂, pitfall traps, 1–9 Dec. 2006 (SYNU-485); 1♂, pitfall traps, 1–9 Dec. 2006 (SYNU-486); 1♀ (SYNU-487); 1♂1♀, XTBG, Secondary tropical montane evergreen broad-leaved forest, 21°54.813'N, 101°12.634'E, 876 m, pitfall traps, 16–24 Sep. 2006, Guo Zheng leg. (SYNU-488–489); 1♂1♀, pitfall traps, 1–15 July 2007 (SYNU-490–491); 1♂1♀, pitfall traps, 16–31 June 2007 (SYNU-492–493).

This new species is similar to C. schwendingeri Baehr, 2010 (female unknown), but can be distinguished by the long, strongly-curved setae between the cheliceral paturons (Figs 1F, G, 2A–C) and a uniformly coloured carapace (vs setae absent and the carapace with a longitudinal brown stripe (Baehr and Ubick 2010: figs 327–331)). Females of this new species can be distinguished from congeners by the star-shaped structure of the endogyne and the strongly-curved copulatory duct (Fig. 3H–J).

Figure 1. 

Camptoscaphiella changxu sp. nov., male holotype (SYNU-481) A, C, E habitus in dorsal, ventral, and lateral views B, D, F, G prosoma in dorsal, ventral, lateral, and anterior views H, I left palp in prolateral and retrolateral views. Scale bars: 0.4 mm (A, C, E); 0.2 mm (B, D, F–I).

Male (holotype): Body: uniformly colored, yellow; habitus as in Fig. 1A, C, E; length 1.29. Carapace (Fig. 1B, F): 0.65 long, 0.59 wide; pars cephalica strongly elevated in lateral view, surface of elevated portion and sides of pars cephalica finely reticulate. Eyes (Fig. 1B, G): ALE 0.057; PME 0.049; PLE 0.049; ALE circular, PME oval, PLE oval; posterior eye row procurved from both above and front; ALE separated by less than one radius. Clypeus (Fig. 1F, G): margin unmodified, straight in front view, sloping forward in lateral view. Mouthparts (Figs 1F, G, 2A–C): with several long, strongly-curved setae between paturons. Sternum (Fig. 1D): as long as wide, pale orange, surface finely reticulate. Abdomen (Fig. 1A, C, E): 0.67 long, 0.41 wide; oval, scuta pale orange; dorsal scutum covering about 2⁄3 of abdomen length, between 1/2 and 3⁄4 of abdomen width, not fused to epigastric scutum; postgastric scutum small, just near epigastric furrow. Legs: pale orange. Palp (Figs 1H, I, 2D–K): reddish-brown; patella extremely long club-shaped, about 4.3 times of the femur length, and 2.1 times of the bulb length; cymbium narrow in dorsal view; distal part of bulb with a broad ventral process (vp) and a narrow dorsal process (dp), the dorsal one with basally wing-shaped appendices (wa).

Figure 2. 

Camptoscaphiella changxu sp. nov., male holotype (SYNU-481), SEM A–C prosoma in anterior, lateral, and ventral views D–F left palp, prolateral, retrolateral, and dorsal views G, J, K distal part of bulb, dorsal, prolateral, and retrolateral views H, I left palpal bulb, prolateral and retrolateral views. Abbreviations: dp = dorsal process; po = pore; vp = ventral process; wa = wing-shaped appendices. Scale bars: 0.2 mm (A–E); 0.1 mm (F, H, I); 0.05 mm (G, J, K).

Female (SYNU-484): Body: habitus as in Fig. 3A–C; length 1.37. Carapace: 0.62 long, 0.57 wide. Eyes: ALE 0.065; PME 0.049; PLE 0.057. Mouthparts: chelicerae unmodified. Abdomen: 0.79 long, 0.52 wide. Epigastric area (Fig. 3H, I): surface without external features. Endogyne (Fig. 3J): with an anterior star-shaped structure (ss); copulatory duct (cd) very long, strongly curved; apodemes thread-shaped (a).

Figure 3. 

Camptoscaphiella changxu sp. nov., female paratype (SYNU-484) A–C habitus in dorsal, ventral, and lateral views D–G prosoma in dorsal, ventral, anterior, and lateral views H epigastric region, ventral view I, J endogyne (cleared in lactic acid), ventral and dorsal views. Abbreviations: a= apodemes; cd = copulatory duct; ss = star-shaped structure. Scale bars: 0.4 mm (A–C); 0.2 mm (D–G); 0.1 mm (H–J).

The specific name is derived from Chinese pinyin, “changxu”, which means “macrochaeta”, referring to the long, curved setae between male cheliceral paturons; noun in apposition.

The male chelicerae are unmodified in species considered to belong to the genus (Baehr and Ubick 2010). The long, strongly-curved setae between male cheliceral paturon of this species are unique in this genus, even in the entire order.

Known only from the type locality.

Holotype ♂ China, Yunnan, Baoshan City, Longling County, Datianba Village, Xianren Cave; 24°358.09'N, 99°037.93'E; 3 Oct. 2020; Yejie Lin Leg. (SYNU-479). Paratype 1♀: same data as holotype (SYNU-480).

This new species is similar to C. sinensis, but can be distinguished by the flat carapace (Fig. 4F), the presence of dorsal and ventral abdominal scuta (Fig. 4A, C, E), the unmodified cymbium (Fig. 5E), and the short processes of tip of the bulb (Fig. 5G, H). Camptoscaphiella sinensis has the highest point of carapace at posterior 2/3, the abdomen lacking scuta, the tip of the cymbium with a pair of enlarged tubular setae, and the tip of bulb with elongated processes (Deeleman-Reinhold 1995: figs 1–5).

Figure 4. 

Camptoscaphiella linyejiei sp. nov., male holotype (SYNU-479) A, C, E habitus in dorsal, ventral, and lateral views B, D, F, G prosoma in dorsal, ventral, lateral, and anterior views H, I left palp in prolateral and retrolateral views. Scale bars: 0.4 mm (A, C, E); 0.2 mm (B, D, F–I).

Male (holotype): Body: uniformly yellow; habitus as in Fig. 4A, C, E; length 1.53. Carapace (Fig. 4B, F): 0.71 long, 0.56 wide; pars cephalica slightly elevated in lateral view, surface of elevated portion and sides of pars cephalica finely reticulated. Eyes (Fig. 4B, G): reduced, with only remnants. Clypeus (Fig. 4F, G): margin unmodified, straight in anterior view, sloping forward in lateral view. Mouthparts (Fig. 4D, G): chelicerae slightly divergent, anterior-median part of the endites strongly sclerotized. Sternum (Fig. 4D): pale orange, surface finely reticulated. Abdomen (Fig. 4A, C, E): 0.77 long, 0.48 wide; dorsal scutum covering 2⁄3 of abdomen length, 1⁄2 of abdomen width. Legs: pale orange. Palp (Figs 4H, I, 5A–H): reddish-brown; patella extremely long club-shaped, about 4.3 times of the femur length, and 2.2 times of the bulb length; cymbium narrow in dorsal view; distal part of bulb with a rectangular prolateral process (pp), a round retrolateral process (rp) and a small ventral process (vsp).

Figure 5. 

Camptoscaphiella linyejiei sp. nov., male holotype (SYNU-479), SEM A–C left palp, prolateral, retrolateral, and dorsal views D, G, H distal part of bulb, dorsal, prolateral, and retrolateral views E, F left palpal bulb, prolateral and retrolateral views. Abbreviations: pp = prolateral process; rp = retrolateral process; vsp = ventral small process. Scale bars: 0.2 mm (A, B); 0.1 mm (C, E, F); 0.03 mm (D, G, H).

Female (SYNU-480): Body: habitus as in Fig. 6A–C; length 1.75. Carapace: 0.76 long, 0.61 wide. Abdomen: 1.04 long, 0.63 wide. Epigastric area (Fig. 6H, I): surface without external features. Endogyne (Fig. 6J): with two transverse sclerites (trs) and a median stick-shaped sclerite (ssc); apodemes (a) thread-shaped.

Figure 6. 

Camptoscaphiella linyejiei sp. nov., female paratype (SYNU-480) A–C habitus in dorsal, ventral, and lateral views D–G prosoma in dorsal, ventral, lateral, and anterior views H abdomen, ventral view I, J endogyne (cleared in lactic acid), ventral and dorsal views. Abbreviations: a = apodemes; ssc = stick-shaped sclerite; trs = transverse sclerites. Scale bars: 0.4 mm (A–C); 0.2 mm (D–J).

The specific name is named after Mr Yejie Lin, the collector of the type specimens; noun (name) in genitive case.

Known only from the type locality.