Research Article |
Corresponding author: António Onofre Soares ( antonio.oc.soares@uac.pt ) Academic editor: Janakiraman Poorani
© 2021 António Onofre Soares, Hugo Renato Calado, José Carlos Franco, António Franquinho Aguiar, Miguel M. Andrade, Vera Zina, Olga M. C. C. Ameixa, Isabel Borges, Alexandra Magro.
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation:
Soares AO, Calado HR, Franco JC, Aguiar AF, Andrade MM, Zina V, Ameixa OMCC, Borges I, Magro A (2021) An annotated checklist of ladybeetle species (Coleoptera, Coccinellidae) of Portugal, including the Azores and Madeira Archipelagos. ZooKeys 1053: 107-144. https://doi.org/10.3897/zookeys.1053.64268
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A comprehensive annotated checklist of the ladybeetle species of Portugal, including the Azores and Madeira archipelagos, is presented. The Coccinellidae fauna comprises a total of 101 species: 83 from the Mainland, 39 from Madeira, and 32 from the Azores. The listed species are distributed among 2 sub-families and 13 tribes: within the subfamily Microweiseinae, Madeirodulini (1 species), Serangiini (2 species), and within the subfamily Coccinellinae, Azyini (1 species), Chilocorini (4 species), Coccidulini (7 species), Coccinellini (30 species), Epilachnini (4 species), Hyperaspidini (7 species), Noviini (2 species), Platynaspini (1 species), Scymnini (37 species), Stethorini (3 species), and Sticholotidini (2 species). The Portuguese fauna comprises 10 exotic species: 5 present in the Mainland, 7 in Madeira, and 6 in the Azores. Harmonia axyridis (Pallas, 1773) from Madeira, Propylea quatuordecimpunctata (Linnaeus, 1758) from the Azores, Delphastus catalinae (Horn, 1895) from the Azores and Madeira, Nephus (Geminosipho) reunioni (Fürsch, 1974) and Nephus (Nephus) voeltzkowi Weise, 1910 from Madeira and Microserangium sp. from the Mainland, are reported for the first time. Some species are considered doubtful records, as explained in the text. These results were obtained by compiling information on the available literature regarding ladybeetle species on the Portuguese mainland and insular territories, and original data.
Azores, Coccinellidae, Madeira, Palearctic Region, Portugal
The book by
Ladybeetles comprise about 375 genera (
Scientific evidence shows that human activities have huge negative impacts on biodiversity (e.g.,
Based on the literature and unpublished data, we present an updated, comprehensive checklist of ladybeetles of Portugal, including the Azores and Madeira Archipelagos.
In this work we follow the suprageneric classification of Coccinellidae proposed by
Current taxonomic affiliation follows
The species are listed in alphabetic order according to their valid tribe and genera. Species of the genera Nephus and Scymnus are listed under the respective subgenera because they are particularly important for their identification. Synonymy is mentioned but is restricted to the original name.
For each species, we specify the regions where the species were recorded (Mainland, Azores and/or Madeira), under “Distribution” and we provide brief notes about the status of the species in each region, possible taxonomic incongruences, and the current biogeographical (limits as in Löbl and Smetana 2007) distribution under “Comments”. New records are marked with a black spot (●), doubtful records with an asterisk (*) and exotic species with a dollar ($). The bibliographic references associated with each species recorded for Portugal are presented in Table
Checklist (by alphabetic order) and bibliographic references of Coccinellidae species recorded for Portugal, including the Azores and Madeira.
Species | References |
---|---|
Adalia bipunctata (Linnaeus, 1758) | 12, 27, 29, 30, 32, 39, 42, 47, 50, 52, 53, 55, 59, 60, 64, 67, 68, 74, 75, 77 |
Adalia decempunctata (Linnaeus, 1758) | 3, 4, 5, 9, 12, 14, 26, 30, 32, 39, 42, 47, 50, 52, 53, 55, 56, 57, 59, 60, 64, 65, 67, 68, 72, 73, 74, 75, 77, 82 |
Adalia testudinea (Wollaston, 1854) | 1, 2, 8, 14, 29, 67, 68 |
Anatis ocellata (Linnaeus, 1758) | 12, 67, 77 |
Anisosticta novemdecimpunctata (Linnaeus, 1758) | 13, 12, 39, 67, 77 |
Calvia decemguttata (Linnaeus, 1767) | 39, 67, 77 |
Calvia quatuordecimguttata (Linnaeus, 1758) | 12, 39, 77 |
Calvia quindecimguttata (Fabricius, 1777) | 39 |
Ceratomegilla notata (Laicharting, 1781) | 12, 39, 67 |
Ceratomegilla undecimnotata (Schneider, 1792) | 12, 39, 55, 67, 77 |
Chilocorus bipustulatus (Linnaeus, 1758) | 9, 12, 14, 26, 27, 29, 30, 39, 47, 50, 52, 53, 55, 59, 60, 64, 65, 66, 67, 68, 72, 75 |
Chnootriba elaterii (Rossi, 1794) | 31, 39, 67, 77 |
Clitostethus arcuatus (Rossi, 1794) | 1, 2, 8, 20, 29, 30, 39, 40, 47, 50, 52, 53, 55, 56, 57, 59, 64, 67, 68, 73, 75 |
Coccidula rufa (Herbst, 1783) | 18, 39, 77 |
Coccidula scutellata (Herbst, 1783) | 12, 39, 77 |
Coccinella genistae Wollaston, 1854 | 1, 2, 8, 14, 17, 29, 37, 42, 52, 67, 68 |
Coccinella septempunctata Linnaeus, 1758 | 9, 30, 32, 35, 39, 43, 47, 50, 55, 56, 57, 59, 60, 64, 66, 72, 73, 74, 75, 77 |
Coccinella undecimpunctata Linnaeus, 1758 | 3, 4, 5, 9, 30, 32, 35, 39, 45, 55, 59, 67, 77, 82 |
Coccinula quatuordecimpustulata (Linnaeus, 1758) | 1, 2, 39 |
Coccinula sinuatomarginata (Faldermann, 1837) | 29, 67 |
Coelopterus salinus Mulsant & Rey, 1852 | 67 |
Cryptolaemus montrouzieri Mulsant, 1853 | 39, 47, 50, 52, 53, 64 |
Delphastus catalinae (Horn, 1895) | Present study |
Eriopis connexa (Germar, 1824) | 44, 55, 59 |
Exochomus quadripustulatus (Linnaeus, 1758) | 19, 39, 47, 50, 52, 53, 64, 66, 67, 72, 75 |
Halyzia sedecimguttata (Linnaeus, 1758) | 12, 19, 39, 67, 77 |
Harmonia axyridis (Pallas, 1773) | Present study |
Harmonia quadripunctata (Pontoppidan, 1763) | 29, 30, 39, 40, 42, 47, 50, 64, 67, 68, 77 |
Henosepilachna angusticollis (Reiche, 1862) | 77 |
Henosepilachna argus (Geoffrey, 1785) | 12, 39, 67 |
Hippodamia variegata (Goeze, 1777) | 1, 2, 8, 11, 14, 21, 23, 25, 29, 32, 36, 39, 42, 50, 52, 53, 55, 56, 57, 64, 67, 68, 72, 75, 77 |
Hyperaspis concolor (Suffrian, 1843) | 67, 77 |
Hyperaspis duvergeri Fürsch, 1985 | 70 |
Hyperaspis hoffmannseggi (Gravenhorst, 1807) | 39 |
Hyperaspis illecebrosa Mulsant, 1846 | 16, 67, 77 |
Hyperaspis pantherina Fürsch, 1975 | 58, 61, 68, 69 |
Hyperaspis reppensis (Herbst, 1783) | 12, 47, 50, 52, 53, 60, 64, 66 |
Hyperaspis stigma (Olivier, 1808) | 67 |
Iberorhyzobius rondensis (Eizaguirre, 2004) | 62, 63, 74, 77, 78, 79 |
Madeirodula atlantica Szawaryn, Větrovec & Tomaszewska, 2020 | 85 |
Microserangium sp. | Present study |
Myrrha octodecimguttata (Linnaeus, 1758) | 6, 8, 14, 20, 29, 36, 38, 39, 42, 55, 59, 65, 67, 68,75, 77 |
Myzia oblongoguttata (Linnaeus, 1758) | 39, 66 |
Nephus (Bipunctatus) bisignatus (Boheman, 1850) | 38, 40, 49, 50, 52, 53, 55, 59, 64, 67, 72, 75, 77 |
Nephus (Bipunctatus) conjunctus (Wollaston, 1870) | 48, 49, 50, 52, 53, 64, 67, 68, 77 |
Nephus (Bipunctatus) peyerimhoffi (Sicard, 1923) | 49, 50, 64, 67, 77 |
Nephus (Geminosipho) reunioni (Fürsch, 1974) | 47, 48, 49, 50, 52, 53, 64, 67, 71, 77 |
Nephus (Nephus) binotatus (Brisout de Barneville, 1863) | 39, 47, 52, 53, 64 |
Nephus (Nephus) flavopictus (Wollaston, 1854) | 1, 2, 8, 14, 15, 17, 21, 22, 25, 30, 26, 27, 29, 32, 35, 40, 41, 42, 46, 55, 57, 59, 67, 68, 83 |
Nephus (Nephus) quadrimaculatus (Herbst, 1783) | 39, 64 |
Nephus (Nephus) schatzmayri Canepari & Tedeschi, 1977 | 67, 70 |
Nephus (Nephus) ulbrichi Fürsch, 1977 | 49, 52, 53, 64 |
Nephus (Nephus) voeltzkowi Weise, 1910 | 84 |
Nephus (Sidis) depressiusculus (Wollaston, 1867) | 46, 68 |
Nephus (Sidis) hiekei (Fürsch, 1965) | 49, 50, 52, 53, 55, 56, 57, 64, 72, 77 |
Nephus (Sidis) pooti Fürsch, 1999 | 77 |
Novius cardinalis (Mulsant, 1850) | 30, 32, 35, 39, 42, 44, 45, 46, 47, 49, 50, 52, 53, 54, 55, 60, 64, 68 73, 67, 82 |
Novius cruentatus Mulsant, 1846 | 67 |
Oenopia conglobata (Linnaeus, 1758) | 12, 39, 47, 50, 52, 53, 59, 63, 65, 73, 74, 76 |
Oenopia doublieri (Mulsant, 1846) | 12, 39, 50, 52, 53, 64, 67, 72, 77, 81 |
Oenopia lyncea (Olivier, 1808) | 12, 39, 67 |
Parexochomus nigripennis (Erichson, 1843) | 67 |
Parexochomus nigromaculatus (Goeze, 1777) | 12, 39, 47, 50, 52, 53, 60, 64, 66, 72, 74, 75 |
Pharoscymnus decemplagiatus (Wollaston, 1857) | 2, 7, 14, 29, 32, 42, 46, 55, 67, 68, 82 |
Platynaspis luteorubra (Goeze, 1777) | 12, 39, 47, 50, 52, 53, 64, 72, 75 |
Propylea quatuordecimpunctata (Linnaeus, 1758) | 11, 39, 47, 50, 52, 53, 60, 64, 67, 72, 73, 75 |
Psyllobora vigintiduopunctata (Linnaeus, 1758) | 12, 39, 47, 50, 52, 53, 67, 77 |
Rhyzobius chrysomeloides (Herbst, 1792) | 10, 12, 36, 39, 47, 50, 52, 53, 55, 56, 57, 64, 65, 66, 67, 68, 72, 73, 74, 75, 82 |
Rhyzobius forestieri (Mulsant, 1853) | 80 |
Rhyzobius litura (Fabricius, 1787) | 1, 2, 3, 4, 5, 7, 8, 9, 11, 14, 25, 26, 27, 29, 30, 32, 35, 36, 39, 42, 47, 50, 52, 53 55, 56, 57, 64, 65, 67, 68, 72, 73, 75 |
Rhyzobius lophanthae (Blaisdell, 1892) | 36, 39, 42, 44, 46, 47, 50, 51, 52, 53, 55, 56, 59, 64, 65, 66, 67, 68, 72 |
Scymniscus fuerschi (Plaza, 1981) | 50, 52, 64 |
Scymniscus helgae (Fürsch, 1965) | 38, 39, 40, 55, 59, 72, 74, 75 |
Scymniscus semirufus (Weise, 1885) | 39, 47, 48, 64, 72, 77 |
Scymnus (Mimopullus) epistemoides Wollaston, 1867 | 10, 14, 29, 30, 40, 66, 67 |
Scymnus (Mimopullus) limnichoides Wollaston, 1854 | 1, 2, 8, 14,.30, 40, 51, 67, 68 |
Scymnus (Mimopullus) marinus (Mulsant, 1850) | 1, 2, 8, 12, 14, 30, 39, 40, 47, 50, 51, 52, 53, 64, 65, 68, 74, 75 |
Scymnus (Neopullus) ater Kugelann, 1794 | 13, 34, 39, 77 |
Scymnus (Neopullus) haemorrhoidalis Herbst, 1797 | 2, 8, 14, 40, 41, 55, 67, 68. |
Scymnus (Neopullus) limbatus Stephens, 1832 | 27, 29, 67, 68 |
Scymnus (Parapullus) abietis (Paykull, 1798) | 2, 11, 39, 56, 57, 68, 77 |
Scymnus (Pullus) auritus Thunberg, 1795 | 12, 39, 49, 50, 52, 53, 64, 67, 77 |
Scymnus (Pullus) subvillosus (Goeze, 1777) | 1, 2, 8, 9, 11, 12, 14, 26, 29, 32, 34, 35, 39, 43, 47, 50, 52, 53, 55, 59, 64, 65, 66, 67, 68, 72, 74, 75, 82 |
Scymnus (Pullus) suturalis Thunberg, 1795 | 24, 12, 27, 29, 30, 38, 39, 40, 46, 47, 50, 51, 52, 53, 55, 59, 73, 66, 67, 68, 82 |
Scymnus (Scymnus) apetzi Mulsant, 1846 | 1, 2, 4, 8, 14, 25, 27, 29, 32, 39, 40, 42, 46, 47, 50, 52, 53, 55, 64, 67, 68, 72, 74, 75 |
Scymnus (Scymnus) bivulnerus Baudi di Selve, 1894 | 39, 77 |
Scymnus (Scymnus) frontalis (Fabricius, 1787) | 12, 39, 60, 64, 67, 75, 77 |
Scymnus (Scymnus) interruptus (Goeze, 1777) | 5, 11, 32, 35, 39, 40, 43, 47, 50, 52, 53, 55, 57, 60, 64, 66, 67, 72, 74, 75, 82 |
Scymnus (Scymnus) laetificus Weise, 1879 | 77 |
Scymnus (Scymnus) nubilus Mulsant, 1850 | 2, 20, 30, 38, 39, 40, 43, 50, 52, 53, 55, 56, 57, 64, 65, 67, 68, 82 |
Scymnus (Scymnus) rubromaculatus (Goeze, 1777) | 27, 29, 30, 32, 40, 55, 67, 68 |
Scymnus (Scymnus) rufipes (Fabricius, 1798) | 12, 39, 47, 54, 52, 53, 60, 64, 72, 67, 77 |
Scymnus (Scymnus) schmidti Fürsch, 1958 | 40, 55, 77 |
Scymnus (Scymnus) suffrianioides Sahlberg, 1913 | 33, 39, 64, 72 |
Sospita vigintiguttata (Linnaeus, 1758) | 12, 39, 77 |
Stethorus pusillus (Herbst, 1797) | 9, 19, 20, 26, 32, 35, 39, 40, 47, 50, 52, 53, 55, 59, 60, 64, 72, 73, 74, 75, 82 |
Stethorus tenerifensis Fürsch, 1987 | 42, 51, 67 |
Stethorus wollastoni Kapur, 1948 | 1, 2, 7, 8, 14, 15, 22, 28, 29, 32, 56, 57, 67, 68 |
Subcoccinella vigintiquatuorpunctata (Linnaeus, 1758) | 12, 39, 50, 52, 53, 67, 72 |
Tytthaspis sedecimpunctata (Linnaeus, 1761) | 12, 39, 52, 53, 67, 73, 75 |
Vibidia duodecimguttata (Poda von Neuhaus, 1761) | 12, 39, 67, 75 |
Madeirodula atlantica Szawaryn, Větrovec and Tomaszewska 2020 (following
Distribution. Madeira.
Comments. This is a recently described new tribe, genus and species, endemic to Madeira.
$ Delphastus catalinae (Horn, 1895)
= Cryptognatha catalinae
Distribution. Madeira● and Azores●.
Comments. A native species of Nearctic and Neotropical regions, currently established in the Palearctic region following introductions in biological control programs. Its presence in the Azores probably originated from deliberate releases for biological control of whiteflies. A large population was first recorded by Isabel Borges, from kales in a vegetable garden (S. Miguel Island 37°48'02"N, 25°36'42"W), August 2018, where both adults and larvae were abundant. In Madeira, Délia Cravo collected in October 2006 from Musa acuminata Colla (Funchal 32°39'26"N, 16°55'56"W) and José Jesus collected in September 2020 on Citrus reticulata Blanco, (Santana 32°48'27"N, 16°53'13"W). First records for Portugal.
$ Microserangium sp.
= Microserangium Miyatake, 1961
Distribution. Mainland●.
Comments. Oriental origin. First observations by Vera Zina, in 2012, Algarve. A few individuals were collected in August 2012, and May, July and November 2013 from the canopy of citrus (Carocha, Boliqueime 37°08'55.9"N, 8°08'11.6"W; Estibeira, Boliqueime 37°07'27"N, 8°07'16"W; Benafim 37°14'17"N, 8°06'36"W). First record for Portugal.
$ Cryptolaemus montrouzieri Mulsant, 1853
Distribution. Mainland.
Comments. Currently established in Palearctic, Afrotropical, Nearctic and Neotropical regions. This exotic species of Australian origin, used around the world for biological control since the 19th century, was introduced from France in the early 20th century for the biological control of the citrus mealybug, Planococcus citri (Risso) (
Chilocorus bipustulatus (Linnaeus, 1758)
= Coccinella bipustulata Linnaeus, 1758
Distribution. Mainland, Madeira and Azores.
Comments. Palearctic, Afrotropical, and Nearctic distribution.
Exochomus quadripustulatus (Linnaeus, 1758)
= Coccinella quadripustulata Linnaeus, 1758
Distribution. Mainland.
Comments. Palearctic and Nearctic distribution.
Parexochomus nigripennis (Erichson, 1843)
= Chilocorus nigripennis Erichson, 1843
Distribution. Mainland.
Comments. Palearctic and Afrotropical distribution.
Parexochomus nigromaculatus (Goeze, 1777)
= Coccinella nigromaculata Goeze, 1777
Distribution. Mainland
Comments. Palearctic distribution.
Coccidula rufa (Herbst, 1783)
= Dermestes rufus Herbst, 1783
Distribution. Mainland.
Comments. Palearctic distribution.
Coccidula scutellata (Herbst, 1783)
= Chrysomela scutellata Herbst, 1783
Distribution. Mainland.
Comments. Palearctic distribution.
Iberorhyzobius rondensis (Eizaguirre, 2004)
= Coccidula rondensis Eizaguirre, 2004
Distribution. Mainland.
Comments. This is an endemic species of the Iberian Peninsula, associated with maritime pine forests, and is a specialist predator of the maritime pine bast scale, Matsucoccus feytaudi Ducasse (
Rhyzobius chrysomeloides (Herbst, 1792)
= Strongylus chrysomeloides Herbst, 1792
Distribution. Mainland, Madeira and Azores.
Comments. Palearctic distribution.
$ Rhyzobius forestieri (Mulsant, 1853)
= Platyomus forestieri Mulsant, 1853
Distribution. Azores.
Comments. Palaearctic, Nearctic and Australian distribution. This exotic species of Australian origin was introduced in Europe in the 1980´s for the biological control of scale insects (Coccoidea), and became established in different countries, including Italy, France, Greece and Albania (
Rhyzobius litura (Fabricius, 1787)
= Nitidula litura Fabricius, 1787
Distribution. Mainland, Madeira and Azores.
Comments. Palearctic distribution.
$ Rhyzobius lophanthae (Blaisdell, 1892)
= Scymnus lophanthae Blaisdell, 1892
Distribution. Mainland, Madeira and Azores.
Comments. Palaearctic, Afrotropical, Nearctic, Neotropical, and Australian distribution. This species, native to Queensland, Australia (
Adalia bipunctata (Linnaeus, 1758)
= Coccinella bipunctata Linnaeus, 1758
Distribution. Mainland, Madeira and Azores.
Comments. Worldwide distributed (Palaearctic, Afrotropical, Australian, Nearctic and Neotropical regions).
Adalia decempunctata (Linnaeus, 1758)
= Coccinella decempunctata Linnaeus, 1758
Distribution. Mainland, Madeira and Azores.
Comments. Palearctic distribution.
Adalia testudinea (Wollaston, 1854)
= Coccinella testudinea Wollaston, 1854
Distribution. Madeira.
Comments. Macaronesian endemic species.
Anatis ocellata (Linnaeus, 1758)
=Coccinella ocellata Linnaeus, 1758
Distribution. Mainland.
Comments. Palearctic distribution.
Anisosticta novemdecimpunctata (Linnaeus, 1758)
= Coccinella novemdecimpunctata Linnaeus, 1758
Distribution. Mainland.
Comments. Palearctic distribution.
Calvia decemguttata (Linnaeus, 1767)
= Coccinella decemguttata Linnaeus, 1767
Distribution. Mainland.
Comments. Palearctic distribution.
Calvia quatuordecimguttata (Linnaeus, 1758)
= Coccinella quatuordecimguttata Linnaeus, 1758
Distribution. Mainland.
Comments. Palearctic, Nearctic and Oriental distribution.
Calvia quindecimguttata (Fabricius, 1777)
= Coccinella quindecimguttata Fabricius, 1777
Distribution. Mainland.
Comments. Palearctic distribution.
Ceratomegilla notata (Laicharting, 1781)
= Coccinella notata Laicharting, 1781
Distribution. Mainland.
Comments. Palearctic distribution.
Ceratomegilla undecimnotata (Schneider, 1792)
= Coccinella undecimnotata Schneider, 1792
Distribution. Mainland and Azores.
Comments. Palearctic distribution.
Coccinella genistae Wollaston, 1854
Distribution. Madeira.
Comments. Macaronesian endemic species.
Coccinella septempunctata Linnaeus, 1758
Distribution. Mainland, Madeira and Azores*.
Comments. Palearctic, Afrotropical, Nearctic and Oriental distribution. Relatively important species in cereal crops in the Azores, especially in the first half of the 20th century, having eventually disappeared when these crops became scarce (
Coccinella undecimpunctata Linnaeus, 1758
Distribution. Mainland and Azores.
Comments. Palaeartic, Australian and Nearctic distribution. In the Azores, it is a threatened species due to anthropogenic pressures on the coastal areas (
Coccinula quatuordecimpustulata (Linnaeus, 1758)
= Coccinella quatuordecimpustulata Linnaeus, 1758
Distribution. Mainland and Madeira*.
Comments. Palearctic and Afrotropical distribution. Although previously recorded in the Madeira archipelago, there are doubts regarding its present occurrence. It might have been introduced but did not establish (Franquinho Aguiar, personal communication).
Coccinula sinuatomarginata (Faldermann, 1837)
= Coccinella sinuatomarginata Faldermann, 1837
Distribution. Mainland and Madeira*.
Comments. Palearctic distribution. Although previously recorded for the Madeira archipelago, there are doubts as to its present occurrence. It might have been introduced but did not establish (Franquinho Aguiar, personal communication).
$ Eriopis connexa (Germar, 1824)
= Coccinella connexa Germar, 1824
Distribution. Azores*.
Comments. Of Neotropical origin where it is very common. Although previously reported for the Azores, it did not become established (fide A. O. Soares, after intensive surveys).
Halyzia sedecimguttata (Linnaeus, 1758)
= Coccinella sedecimguttata Linnaeus, 1758
Distribution. Mainland.
Comments. Palearctic distribution.
$ Harmonia axyridis (Pallas, 1773)
= Coccinella axyridis Pallas, 1773
Distribution. Madeira●.
Comments. Worldwide distribution. Several specimens (adults, larvae and pupae) collected by Miguel M. Andrade, in September 2019, from Enterolobium sp. (Funchal 32°38'39"N, 16°55'31"W), Graça Freitas and Franquinho Aguiar, collected in September and October 2020, from Annona cherimola Mill. (Funchal 32°39'47"N, 16°53'41"W). Harmonia axyridis is the most invasive insect of the world (
Harmonia quadripunctata (Pontoppidan, 1763)
= Coccinella quadripunctata Pontoppidan, 1763
Distribution. Mainland, Madeira.
Comments. Palearctic and Nearctic distribution.
Hippodamia variegata (Goeze, 1777)
= Coccinella variegata Goeze, 1777
Distribution. Mainland, Madeira and Azores.
Comments. Palearctic, Afrotropical, Nearctic and Oriental distribution.
Myrrha octodecimguttata (Linnaeus, 1758)
= Coccinella octodecimguttata Linnaeus, 1758
Distribution. Mainland, Madeira and Azores.
Comments. Palearctic distribution.
Myzia oblongoguttata (Linnaeus, 1758)
= Coccinella oblongoguttata Linnaeus, 1758
Distribution. Mainland.
Comments. Palearctic distribution.
Oenopia conglobata (Linnaeus, 1758)
= Coccinella conglobata Linnaeus, 1758
Distribution. Mainland.
Comments. Palearctic distribution.
Oenopia doublieri (Mulsant, 1846)
= Harmonia doublieri Mulsant, 1846
Distribution. Mainland and Azores.
Comments. Palearctic distribution.
Oenopia lyncea (Olivier, 1808)
= Coccinella lyncea Olivier, 1808
Distribution. Mainland.
Comments. Palearctic distribution.
Propylea quatuordecimpunctata (Linnaeus, 1758)
= Coccinella quatuordecimpunctata Linnaeus, 1758
Distribution. Mainland and Azores●.
Comments. Palearctic and Nearctic distribution. First record for the Azores. Several adults were collected in July 2019 by António O. Soares and Isabel Borges, in a vegetable garden, of the parish of Castelo Branco (GPS coordinates: 38°31'23.2"N, 28°41'21.0"W), Faial Island.
Psyllobora vigintiduopunctata (Linnaeus, 1758)
= Coccinella vigintiduopunctata Linnaeus, 1758
Distribution. Mainland.
Comments. Palearctic distribution.
Sospita vigintiguttata (Linnaeus, 1758)
= Coccinella vigintiguttata Linnaeus, 1758
Distribution. Mainland.
Comments. Palearctic distribution.
Tytthaspis sedecimpunctata (Linnaeus, 1761)
= Coccinella sedecimpunctata Linnaeus, 1761
Distribution. Mainland.
Comments. Palearctic distribution.
Vibidia duodecimguttata (Poda von Neuhaus, 1761)
= Coccinella duodecimguttata Poda von Neuhaus, 1761
Distribution. Mainland.
Comments. Palearctic and Oriental distribution.
Chnootriba elaterii (Rossi, 1794)
= Coccinella elaterii Rossi, 1794
Distribution. Mainland.
Comments. Palearctic and Afrotropical distribution. Reported by
Henosepilachna angusticollis (Reiche, 1862)
= Epilachna angusticollis Reiche, 1862
Distribution. Mainland.
Comments. Distributed in the Mediterranean region.
Henosepilachna argus (Geoffrey, 1785)
= Coccinella argus Geoffrey, 1785
Distribution. Mainland.
Comments. Palearctic and Afrotropical distribution.
Subcoccinella vigintiquatuorpunctata (Linnaeus, 1758)
= Coccinella vigintiquatuorpunctata Linnaeus, 1758
Distribution. Mainland.
Comments. Palearctic and Nearctic distribution.
Hyperaspis concolor (Suffrian, 1843)
= Coccinella concolor Suffrian, 1843
Distribution. Mainland.
Comments. Palearctic distribution.
Hyperaspis duvergeri Fürsch, 1985
Distribution. Mainland.
Comments. Palearctic distribution.
Hyperaspis hoffmannseggi (Gravenhorst, 1807)
= Coccinella hoffmannseggi Gravenhorst, 1807
Distribution. Mainland.
Comments. Palearctic distribution.
Hyperaspis illecebrosa Mulsant, 1846
Distribution. Mainland.
Comments. Palearctic distribution.
$ Hyperaspis pantherina Fürsch, 1975, following
Distribution. Madeira.
Comments. Established in Palearctic/Madeira, originally from Afrotropical region. The first record for Europe was in 2002, in Madeira archipelago (
Hyperaspis reppensis (Herbst, 1783)
= Coccinella reppensis Herbst, 1783
Distribution. Mainland*.
Comments. Palearctic distribution. According to
Hyperaspis stigma (Olivier, 1808)
= Coccinella stigma Olivier, 1808
Distribution. Mainland.
Comments. Palearctic distribution.
$ Novius cardinalis (Mulsant, 1850), following
= Vedalia cardinalis Mulsant, 1850
Distribution. Mainland, Madeira and Azores.
Comments. Established in Palaeartic, Afrotropical, Nearctic and Neotropical, Oriental. Native to Australian region. This exotic species was introduced in California and South Africa in the 1890´s for the control of the cottony cushion scale, Icerya purchasi Maskell (
Novius cruentatus Mulsant, 1846
Distribution. Mainland.
Comments. Palearctic distribution.
Platynaspis luteorubra (Goeze, 1777)
= Coccinella luteorubra Goeze, 1777
Distribution. Mainland.
Comments. Palearctic distribution.
Clitostethus arcuatus (Rossi, 1794)
=Coccinella arcuata Rossi, 1794
Distribution. Mainland, Madeira and Azores.
Comments. Palearctic, Afrotropical, Nearctic distribution.
Nephus (Bipunctatus) bisignatus (Boheman, 1850)
= Scymnus bisignatus Boheman, 1850
Distribution. Mainland and Azores.
Comments. Palearctic distribution.
Nephus (Bipunctatus) conjunctus (Wollaston, 1870)
= Scymnus conjunctus Wollaston, 1870
Distribution. Mainland and Madeira.
Comments. Palearctic and Afrotropical distribution. This species was first reported as N. (bipunctatus) includens (Kirsch 1871) based on adults emerged from larvae collected in 1988 from citrus fruits infested with mealybugs, in the South of Portugal (Algarve) (
Nephus (Bipunctatus) peyerimhoffi (Sicard, 1923)
= Scymnus peyerimhoffi Sicard, 1923
Distribution. Mainland.
Comments. Palearctic and Afrotropical distribution.
$ Nephus (Geminosipho) reunioni (Fürsch, 1974a)
= Scymnus reunioni Fürsch, 1974a
Distribution. Mainland, Azores and Madeira●.
Comments. Palearctic and Afrotropical distribution. The first record in Europe was in France in 1983 (
Nephus (Nephus) binotatus (Brisout de Barneville, 1863)
= Scymnus binotatus Brisout de Barneville, 1863
Distribution. Mainland.
Comments. Palearctic distribution.
Nephus (Nephus) flavopictus (Wollaston, 1854)
= Scymnus flavopictus Wollaston, 1854
Distribution. Madeira and Azores.
Comments. Macaronesian endemic species.
Nephus (Nephus) quadrimaculatus (Herbst, 1783)
= Sphaeridium quadrimaculatum Herbst, 1783
Distribution. Mainland.
Comments. Palearctic distribution.
Nephus (Nephus) schatzmayri Canepari & Tedeschi, 1977
Distribution. Mainland.
Comments. Palearctic distribution.
Nephus (Nephus) ulbrichi Fürsch, 1977
Distribution. Mainland.
Comments. Palearctic distribution.
$ Nephus (Nephus) voeltzkowi Weise, 1910, following
Distribution. Azores and Madeira●.
Comments. Afrotropical origin. Very recently, two parthenogenetic populations of this species were found in the Azores and Mascarene archipelagos, becoming the first reported case of asexuality in the Coccinellidae (
Nephus (Sidis) depressiusculus (Wollaston, 1867)
= Scymnus depressiusculus Wollaston, 1867
Distribution. Madeira.
Comments. Palearctic and Afrotropical distribution.
Nephus (Sidis) hiekei (Fürsch, 1965)
= Scymnus hiekei Fürsch, 1965
Distribution. Mainland.
Comments. Palearctic distribution.
Nephus (Sidis) pooti Fürsch, 1999
Distribution. Mainland.
Comments. Palearctic distribution.
Scymniscus fuerschi (Plaza, 1981)
= Nephus fuerschi Plaza, 1981
Distribution. Mainland.
Comments. Palearctic distribution.
Scymniscus helgae (Fürsch, 1965)
= Scymnus helgae Fürsch, 1965
Distribution. Mainland and Azores.
Comments. Palearctic distribution.
Scymniscus semirufus (Weise, 1885)
=Scymnus semirufus Weise, 1885
Distribution. Mainland.
Comments. Palearctic distribution.
Scymnus (Mimopullus) epistemoides Wollaston, 1867
Distribution. Madeira.
Comments. Palearctic distribution.
Scymnus (Mimopullus) limnichoides Wollaston, 1854
Distribution. Madeira.
Comments. Palearctic distribution.
Scymnus (Mimopullus) marinus (Mulsant, 1850)
= Rhyzobius marinus Mulsant, 1850
Distribution. Mainland and Madeira.
Comments. Palearctic distribution.
Scymnus (Neopullus) ater Kugelann, 1794
Distribution. Mainland.
Comments. Palearctic distribution.
Scymnus (Neopullus) haemorrhoidalis Herbst, 1797
Distribution. Madeira and Azores.
Comments. Palearctic distribution.
Scymnus (Neopullus) limbatus Stephens, 1832
Distribution. Madeira.
Comments. Palearctic distribution.
Scymnus (Parapullus) abietis (Paykull, 1798)
= Coccinella abietis Paykull, 1798
Distribution. Mainland and Madeira.
Comments. Palearctic distribution.
Scymnus (Pullus) auritus Thunberg, 1795
Distribution. Mainland.
Comments. Palearctic distribution.
Scymnus (Pullus) subvillosus (Goeze, 1777)
= Coccinella subvillosa Goeze, 1777
Distribution. Mainland, Madeira and Azores.
Comments. Palearctic and Afrotropical distribution.
Scymnus (Pullus) suturalis Thunberg, 1795
Distribution. Mainland, Madeira and Azores.
Comments. Palearctic and Nearctic distribution.
Scymnus (Scymnus) apetzi Mulsant, 1846
Distribution. Mainland.
Comments. Palearctic distribution.
Scymnus (Scymnus) bivulnerus Baudi di Selve, 1894
Distribution. Mainland.
Comments. Palearctic distribution.
Scymnus (Scymnus) frontalis (Fabricius, 1787)
= Coccinella frontalis Fabricius, 1787
Distribution. Mainland.
Comments. Palearctic distribution.
Scymnus (Scymnus) interruptus (Goeze, 1777)
= Coccinella interrupta Goeze, 1777
Distribution. Mainland, Madeira and Azores.
Comments. Palearctic distribution.
Scymnus (Scymnus) laetificus Weise, 1879
Distribution. Mainland.
Comments. Palearctic distribution. However, it only occurs in the Western Mediterranean.
Scymnus (Scymnus) nubilus Mulsant, 1850
Distribution. Mainland, Madeira and Azores.
Comments. Palearctic distribution. On Portugal’s mainland, this species was wrongly identified as Scymnus levaillanti Mulsant.
Scymnus (Scymnus) rubromaculatus (Goeze, 1777)
= Coccinella rubromaculata Goeze, 1777
Distribution. Mainland, Madeira and Azores.
Comments. Palearctic and Afrotropical distribution.
Scymnus (Scymnus) rufipes (Fabricius, 1798)
= Coccinella rufipes Fabricius, 1798
Distribution. Mainland.
Comments. Palearctic distribution.
Scymnus (Scymnus) schmidti Fürsch, 1958
Distribution. Mainland and Azores.
Comments. Palearctic distribution.
Scymnus (Scymnus) suffrianioides Sahlberg, 1913
Distribution. Mainland.
Comments. Palearctic distribution.
Stethorus pusillus (Herbst, 1797)
= Scymnus pusillus Herbst, 1797
Distribution. Mainland and Azores.
Comments. Palearctic distribution.
Stethorus tenerifensis Fürsch, 1987
Distribution. Madeira.
Comments. Macaronesian endemic species.
Stethorus wollastoni Kapur, 1948
Distribution. Madeira.
Comments. Macaronesian endemic species.
Coelopterus salinus Mulsant & Rey, 1852
Distribution. Mainland.
Comments. Palearctic and Afrotropical distribution.
Pharoscymnus decemplagiatus (Wollaston, 1857)
= Scymnus decemplagiatus Wollaston, 1857
Distribution. Madeira and Azores.
Comments. Palearctic distribution.
In Memoriam of Armando Américo Cardoso Raimundo (2.V.1942–9.Xi.2019): A farewell to colleague and friend.
Thanks are due to Claudio Canepari for helping in the identification of Microserangium sp. and Jaroslav Větrovec of Harmonia axyridis Pallas from Madeira.
This study was financed by FEDER in 85% and by Azorean Public funds by 15% through Operational Program Azores 2020, under the following projects AZORESBIOPORTAL –PORBIOTA (ACORES-01-0145-FEDER-000072), and under the project ECO2 – TUTA (ACORES-01-0145-FEDER-000081). The Forest Research Centre is funded by Fundação para a Ciência e a Tecnologia I.P. (FCT), Portugal (UIDB/00239/2020). Thanks are also due to FCT/MCES for financial support to CESAM (UIDB/50017/2020+UIDP/50017/2020) through national funds, and the co-funding by the FEDER, within the PT2020 Partnership Agreement and Compete 2020. OMCCA is funded by national funds (OE), through FCT, in the scope of the framework contract foreseen in the numbers 4, 5 and 6 of the article 23, of the Decree-Law 57/2016, of August 29, changed by Law 57/2017, of July 19. AM was supported by the “Laboratoires d’Excellence” LabEx TULIP (ANR-10-LABX-41). The Open Access of this manuscript was supported by the project FCT-UIDB/00329/2020-2024.