Holotype: China • ♂; Hongqi Reservoir, Mt Dawei, Pingbian County, Yunnan Province; 1550 m, 15-V-2016; Lu Qiu, Zhi-Wei Qiu leg; SWU-B-EC141501. Paratypes: China • 3♂♂; same data as holotype; SWU-B-EC141502-141504.

China • 1♂; Hongqi Reservoir, Mt Dawei, Pingbian County, Yunnan Province; 1550 m; 17-V-2016; Lu Qiu, Zhi-Wei Qiu leg.

This species is similar to M. satsumana (Asahina, 1979) in general appearance, but can be differentiated from the latter by the following characters: 1) median phallomere slender rod with base sharp, and apex expanded with three spines; while in the latter, base slightly expanded, and apex curved with some short spines; 2) subgenital plate not folded; while in the latter, folded inwards.

(mm). Male (n = 4), pronotum: length × width 1.6–2.1 × 2.6–2.9, tegmina length: 10.3–11.2, overall length: 12.5–13.1.

Male. Coloration: body yellowish-brown (Fig. 4A, B). Face pale yellowish-brown. Interocular space with a dark brown band. Ocelli spots white, interocellar space with a brown band. Antennae dark linen. Clypeus medium yellowish-brown (Fig. 4C). Maxillary palps dark yellowish-brown (Fig. 4E). Pronotal disc yellowish-brown with brown stripes and spots, and two lateral borders light brown (Fig. 4D). Tegmina medium brown, wings light linen (Fig. 4F, G). Abdomen yellowish-brown (Fig. 4B). Cerci dark yellowish-brown (Fig. 4K). Styli faint yellow (Fig. 4M). Head: vertex slightly exposed, interocular distance same length as antennal sockets space (Fig. 4C). Pronotum nearly trapezoidal, broader than long, the widest part after the midpoint, the front and posterior margins nearly straight, and the postero-lateral angle blunt and round; the disc with symmetrical irregular macules (Fig. 4D). The third and fourth palpi about same length, both obviously longer than the fifth, the fifth obviously expanded (Fig. 4E). Tegmina and wings: tegmina and wings fully developed, both extending beyond the end of abdomen (Fig. 4A, B). Tegmina with ScP simple, R multi-branched, M straight with five complete branches. Hind wings with ScP and RA expanded at apex; M straight and simple, without branches; CuA with four complete branches (Fig. 4F, G). Legs: anteroventral margin of front femur type B₂ (Fig. 4H). Pulvilli present on four proximal tarsomeres. Tarsal claws symmetrical and specialized, inner margin serrated, arolia present (Fig. 4I). Abdomen and genitalia: eighth abdominal tergum specialized with a tuft (Fig. 4J). Supra-anal plate transverse, posterior margin convex, the middle slightly concave. Paraprocts simple, similar, splitting into two pieces, apex with tufts (Fig. 4K). Subgenital plate asymmetrical, both lateral margins slightly concave. Styli similar, slender with spines; interstylar region obviously convex with small spines (Fig. 4M). Left phallomere complex, irregular bone-shaped, with two short spines (Fig. 4L). Median phallomere slender rod-shaped with base sharp, and apex expanded with three long spines; the accessory structure arched, at rightmost end brush-shaped (Fig. 4M). Hook phallomere on the right side, apex curved inwards with a short spine (Fig. 4N).

Figure 4. 

A–N Margattea deltodonta sp. nov., male A holotype, dorsal view B holotype, ventral view C head, ventral view D pronotum, dorsal view E maxillary palpi, ventral view F tegmen, dorsal view G hind wing, dorsal view H front femur, ventral view I tarsal claws J eighth abdominal terga K supra-anal plate and paraprocts, ventral view L left phallomere, dorsal view M subgenital plate and median phallomere, dorsal view N hook-like phallomere, dorsal view. Scale bars: 5 mm (A, B); 0.5 mm (C–H, J, K–N); 0.1 mm (I).

Female unknown.

The word “delt” and “odont” from Greek and means triangular, the species name “deltodontus” refers to the posterior margin of subgenital plate with small spines.

China (Yunnan).

Holotype: China • ♂; Mt Daming, Guangxi Province; 2-VII-2015; Lu Qiu, Qi-Kun Bai leg; SWU-B-EC141201. Paratype: China • 1♂; same data as for holotype; SWU-B-EC141202.

This species is similar to M. flexa Wang et al., 2014 in general appearance and male genitalia, but it can be differentiated from the latter by the following characters: 1) interstylar region obviously convex with both sides curved inwards, three spines on each side, while in the latter, two sides curled up with 5–6 small thorns; 2) the left end of the accessory structure with a slender bone; the latter absent.

(mm). Male (n = 2), pronotum: length × width 1.6–2.1 × 2.6–2.9, tegmina length: 10.3–11.2, overall length: 12.5–13.1.

Male. Coloration: body yellowish-brown (Fig. 5A, B). Face yellowish-brown. Interocular space with a dark brown band. Ocelli spots white, interocelli space with a brown band. Antennae dark linen-colored. Clypeus dark yellowish-brown (Fig. 5C). Maxillary palps yellowish-brown (Fig. 5E). Pronotal disc yellowish-brown with dark brown stripes, and lateral borders light linen-colored (Fig. 5D). Tegmina pale yellow, wings medium brown (Fig. 5F, G). Abdomen pale yellowish-brown. Cerci pale yellowish-brown (Fig. 5K). Styli faint yellow (Fig. 5M). Head: vertex slightly exposed, distance between interocular same length as antennal sockets space (Fig. 5C). Pronotum nearly trapezoidal, broader than long, the widest part after the midpoint, the front and posterior margins nearly straight, and the postero-lateral angle blunt and round; the disc with symmetrical irregular macules (Fig. 5D). The third, fourth palpi of approximately same length, both obviously longer than the fifth, the fifth obviously expanded (Fig. 5E). Tegmina and wings: tegmina and wings fully developed, both extending beyond the end of abdomen (Fig. 5A, B). Tegmina with ScP simple, R multi-branched, M straight with six complete branches. Hind wings with ScP and RA expanded at apex; M straight and simple, without branches; CuA with five complete branches (Fig. 5F, G). Legs: anteroventral margin of front femur type B₃ (Fig. 5H). Pulvilli present on four proximal tarsomeres. Tarsal claws symmetrical and specialized, inner margin serrated, arolia present (Fig. 5I). Abdomen and genitalia: eighth abdominal tergum specialized with a tuft (Fig. 5J). Supra-anal plate transverse, posterior margin protruded. Paraprocts similar, splitting into two pieces, apex with tufts (Fig. 5K). Subgenital plate symmetrical, lateral borders flip inwards with spines and hairs. Styli similar, slender; interstylar region obviously convex, two sides convex and curved inwards, each side with three spines (Fig. 5M). Left phallomere complex, irregular bone-shaped, with a short spine (Fig. 5L). Median phallomere slender rod-shaped, obviously curved, apex with ordered long spines; the accessory structure arched, on at rightmost end with spines, left apex with a slender bone with apex sharp (Fig. 5M). Hook phallomere on the right side, apex curved inwards with a short spine (Fig. 5N).

Figure 5. 

A–N Margattea cuspidata sp. nov., male A holotype, dorsal view B holotype, ventral view C head, ventral view D pronotum, dorsal view E maxillary palpi, ventral view F tegmen, dorsal view G hind wing, dorsal view H front femur, ventral view I tarsal claws J eighth abdominal terga K supra-anal plate and paraprocts, ventral view L left phallomere, dorsal view M subgenital plate and median phallomere, dorsal view N hook-like phallomere, dorsal view. Scale bars: 5 mm (A, B); 0.5 mm (C–H, J, K–N); 0.1 mm (I).

Female unknown.

The latin name “cuspidatus” refers to interstylar region obviously convex, two sides convex and curved inwards.

China (Guangxi).

Holotype: China • ♂; Meizihu Reservoir, Pu’er City, Yunnan Province; 1400 m; 21-V-2016; Lu Qiu, Zhi-Wei Qiu leg; SWU-B-EC141301. Paratypes: China • 6♂♂; 1 ♀, same data as holotype SWU-B-EC141302-141308.

China • 2♀♀; Meizihu Reservoir, Pu’er City, Yunnan Province; 1400 m; 20-V-2016; Lu Qiu, Zhi-Wei Qiu leg.

This species is similar to M. mckittrickae Wang, Che & Wang, 2009 in general appearance, but it can be differentiated from the latter by the following characters: 1) interstylar region obviously convex, fishtail-shaped, while the latter slightly convex; 2) left phallomere complex, irregular bone-shaped, while in the latter, two sides of left phallomere sheet-like; and 3) median phallomere with one accessory structure, while the latter with two accessory structures.

(mm). Male (n = 4), pronotum: length × width 2.7–2.8 × 3.1–3.6, tegmina length: 10.4–12.6, overall length: 12.8–14.1. Female, pronotum: length × width 2.5–3.0 × 3.6–3.7, tegmina length: 9.0–9.1, overall length: 11.7–12.6.

Male. Coloration: body pale brown with yellowish-brown (Fig. 6A, B). Face pale yellowish-brown. Interocular space with a brown band. Ocelli spots white, interocellar space with a brown band. Antennae light linen-colored. Clypeus medium yellowish-brown (Fig. 6C). Maxillary palps dark yellowish-brown (Fig. 6E). Pronotal disc pale yellowish-brown with brown stripes and two lateral borders light yellowish-brown (Fig. 6D). Tegmina yellowish-brown and wings medium brown (Fig. 6F, G). Abdomen pale yellowish-brown. Cerci yellowish-brown (Fig. 6K). Styli faint yellow (Fig. 6M). Head: vertex slightly exposed, interocular distance interocular same length as antennal socket space (Fig. 6C). Pronotum nearly trapezoidal, broader than long, the widest part after the midpoint, the front and posterior margins nearly straight, and postero-lateral angle blunt and round; disc with symmetrical irregular macules (Fig. 6D). Third and fourth palpi of approximately same length, both obviously longer than the fifth, fifth palp obviously expanded (Fig. 6E). Tegmina and wings: tegmina and wings fully developed, both extending beyond the end of abdomen (Fig. 6A, B). Tegmina with ScP simple, R multi-branched, M straight with five complete branches. Hind wings with ScP and RA expanded at apex; M straight and simple, without branches; CuA with 7 complete branches (Fig. 6F, G). Legs: anteroventral margin of front femur type B₂ (Fig. 6H). Pulvilli present on four proximal tarsomeres. Tarsal claws symmetrical and specialized, inner margin serrated, arolia present (Fig. 6I). Abdomen and genitalia: eighth abdominal tergum specialized with a tuft (Fig. 6J). Supra-anal plate transverse, posterior margin protruded. Paraprocts similar, splitting into two pieces, apex with tufts (Fig. 6K). Subgenital plate symmetrical, the middle of front margin slightly concave; the base of two lateral margins concave, apex flips inwards with tufts. Styli similar, slender, distinctly separated; interstylar region obviously convex, fishtail-shaped, middle space slightly concave, two lateral angles convex with short spines (Fig. 6M). Left phallomere complex, irregular bone-shaped, with short spine (Fig. 6L). Median phallomere slender rod-shaped, apex splitting into two parts, each with some long spines (Fig. 6M). Hook phallomere on the right side, apex curved inwards with a short spine (Fig. 6N).

Figure 6. 

A–N Margattea caudata sp. nov., male A holotype, dorsal view B holotype, ventral view C head, ventral view D pronotum, dorsal view E maxillary palpi segments 3–5, ventral view F tegmen, dorsal view G hind wing, dorsal view H front femur, ventral view I tarsal claws J eighth abdominal terga K supra-anal plate and paraprocts, ventral view L left phallomere, dorsal view M subgenital plate and median phallomere, dorsal view N hook-like phallomere, dorsal view. Scale bars: 5 mm (A, B); 0.5 mm (C–H, J, K–N); 0.1 mm (I).

Female with tegmina and wings slightly reduced.

The latin name “caudatus” meaning “tail”, refers to the fishtail-shaped convexity on interstylar region.

China (Yunnan).

Holotype: China • ♂; Wangtianshu Scenery Spot, Mengla County, Xishuangbanna Prefecture, Yunnan Province; 720 m; 23-V-2016; Lu Qiu, Zhi-Wei Qiu leg; SWU-B-EC141401. Paratypes: China • ♂; Gougu Tropical Rainforest, Xishuangbanna Tropical Botanical Garden (CAS), Menglun Town, Jinghong City, Yunnan Province; 570 m; 26-V-2016; Lu Qiu, Zhi-Wei Qiu leg; SWU-B-EC141402.

China • 1♂; Lvshilin (Green Stone Forest), Xishuangbanna Tropical Botanical Garden (CAS), Menglun Town, Jinghong City, Yunnan Province; 25-V-2016; Lu Qiu, Zhi-Wei Qiu leg.

This species is similar to M. flexa Wang et al., 2014 in male genitalia, but it can be differentiated from the latter by the following characters: 1) interstylar region obviously irregularly convex, the left part obviously larger than the right, while in the latter, interstylar region obviously regularly convex; 2) the left part of the accessory structure of median phallomere with a brush, absent in the latter.

(mm). Male (n = 3), pronotum: length × width 2.4–2.6 × 3.2–3.5, tegmina length: 9.3–9.9, overall length: 11.2–11.9.

Male. Coloration: body yellowish-brown with pale brown (Fig. 7A, B). Face yellowish-brown. Interocular space with a dark brown band. Ocellar spots white and small, interocellar space with a brown band. Antennae light linen- colored. Clypeus medium yellowish-brown (Fig. 7C). Maxillary palps light yellowish-brown to yellowish-brown (Fig. 7E). Pronotal disc yellowish-brown with brown stripes and two lateral borders light yellow (Fig. 7D). Tegmina light fawn, wings and legs pale brown (Fig. 7F, G). Abdomen light linen with pale yellowish-brown. Cerci pale brown (Fig. 7K). Styli light yellowish-brown (Fig. 7M). Head: vertex slightly exposed, distance between interocular same length antennal sockets space (Fig. 7C). Pronotum nearly trapezoidal, broader than long, the widest part after the midpoint, the front and posterior margins nearly straight, and postero-lateral angle blunt and round; disc with symmetrical irregular macules (Fig. 7D). Third and fourth palpi of approximately the same length, both obviously longer than fifth, fifth palp obviously expanded (Fig. 7E). Tegmina and wings: tegmina and wings developed, both extending the end of abdomen (Fig. 7A-B). Tegmina with ScP simple, R multi-branched, M straight with seven complete branches. Hind wings with ScP and RA expanded at apex; M straight and simple, without branches; CuA with five complete branches (Fig. 7F, G). Legs: anteroventral margin of front femur type B₃ (Fig. 7H). Pulvilli present on four proximal tarsomeres. Tarsal claws symmetrical and specialized, inner margin serrated, arolia present (Fig. 7I). Abdomen and genitalia: eighth abdominal tergum specialized with a tuft (Fig. 7J). Supra-anal plate transverse, posterior margin convex. Paraprocts simple, similar, splitting into two pieces, base with tufts (Fig. 7K). Subgenital plate asymmetrical. Styli similar, slender, distinctly separated; interstylar region obviously irregularly convex, middle part concave, two lateral angles spherical with some short thorns, left angle obviously larger than right (Fig. 7M). Left phallomere complex, irregular bone-shaped, with a long spine (Fig. 7L). Median phallomere slender rod-shaped, base splitting into some long spines; the accessory structure arched, at leftmost end with a brush (Fig. 7M). Hook phallomere on right side, base curved inwards with a short spine (Fig. 7N).

Figure 7. 

A–N Margattea disparilis sp. nov., male A holotype, dorsal view B holotype, ventral view C head, ventral view D pronotum, dorsal view E maxillary palpi segments 3–5, ventral view F tegmen, dorsal view G hind wing, dorsal view H front femur, ventral view I tarsal claws J eighth abdominal terga K supra-anal plate and paraprocts, ventral view L left phallomere, dorsal view M subgenital plate and median phallomere, dorsal view N hook-like phallomere, dorsal view. Scale bars: 5 mm (A, B); 0.5 mm (C–H, J, K–N); 0.1 mm (I).

Female unknown.

The latin name “disparilis” refers to the interstylar region obviously irregular convex.

China (Yunnan).

Holotype: China • ♂; Meizihu Reservoir, Pu’er City, Yunnan Province; 20-V-2016; Lu Qiu, Zhi-Wei Qiu leg; SWU-B-EC141801. Paratypes: China • 2♂; same date as for holotype SWU-B-EC141802-141803.

China • 1♀; Meizihu Reservoir, Pu’er City, Yunnan Province; 20-V-2016; Lu Qiu, Zhi-Wei Qiu leg. • 1♀; Meizihu Reservoir, Pu’er City, Yunnan Province; 21-V-2016; Lu Qiu, Zhi-Wei Qiu leg.

This species is similar to M. nimbata (Shelford, 1907) in male genitalia, but it can be differentiated from the latter by the following characters: 1) median phallomere base with a curved spine, while in the latter, with two curved spines; 2) a long piece of bone extends from the right side of the accessory structure, while absent in the latter; and 3) left phallomere with four long spines; the latter with two long spines.

(mm). Male (n = 3), pronotum: length × width 2.5–2.6 × 3.2–3.9, tegmina length: 11.7–12.3, overall length: 14.0–14.1. Female, pronotum: length × width 2.6–2.7 × 3.2–3.4, tegmina length: 11.3–11.5, overall length: 13.4–13.6.

Male. Coloration: body pale yellowish-brown with brown (Fig. 8A, B). Face yellowish-brown. Interocular space with a dark brown band. Ocelli spots white and big, interocelli space with a brown band. Antennae pale yellowish-brown. Clypeus medium yellowish-brown. Maxillary palps light yellowish-brown (Fig. 8E). Pronotal disc brownish-gray with brown stripes and two lateral borders light grey (Fig. 8D). Tegmina pale yellowish-brown, wings grey brown (Fig. 8F, G). Abdomen yellowish-brown. Cerci pale yellowish-brown to pale brown (Fig. 8K). Styli faint yellow (Fig. 8M). Head: vertex slightly exposed, interocular distance same length as antennal socket space (Fig. 8C). Pronotum nearly trapezoidal, broader than long, widest part after midpoint, front and posterior margins nearly straight, and postero-lateral angle blunt and round; disc with symmetrical irregular macules (Fig. 8D). Third and fourth palpi of approximately same length, both obviously longer than fifth palp, fifth palp obviously expanded (Fig. 8E). Tegmina and wings: tegmina and wings fully developed, both extending beyond the end of abdomen (Fig. 8A, B). Tegmina with ScP simple, R multi-branched, M straight with seven complete branches. Hind wings with ScP and RA expanded at base; M straight and simple, without branches; CuA with five complete branches (Fig. 8F, G). Legs: anteroventral margin of front femur type B₂ (Fig. 8H). Pulvilli present on four proximal tarsomeres. Tarsal claws symmetrical and specialized, inner margin serrated, arolia present (Fig. 8I). Abdomen and genitalia: eighth abdominal tergum specialized with a tuft (Fig. 8J). Supra-anal plate transverse, posterior margin convex. Paraprocts simple, similar, splitting into two pieces (Fig. 8K). Subgenital plate symmetrical. Styli similar, slender, distinctly separated (Fig. 8M). Left phallomere complex, irregular bone-shaped, with four spines (Fig. 8L). Median phallomere slender rod-shaped, apex with a curved spine; the accessory structure arched, at rightmost end blunt (Fig. 8M). Hook phallomere on the right side, base curved inwards with a short spine (Fig. 8N).

Figure 8. 

A–N Margattea transversa sp. nov., male A holotype, dorsal view B holotype, ventral view C head, ventral view D pronotum, dorsal view E maxillary palpi segments 3–5, ventral view F tegmen, dorsal view G hind wing, dorsal view H front femur, ventral view I tarsal claws J eighth abdominal terga K supra-anal plate and paraprocts, ventral view L left phallomere, dorsal view M subgenital plate and median phallomere, dorsal view N hook-like phallomere, dorsal view. Scale bars: 5 mm (A, B); 0.5 mm (C–H, J, K–N); 0.1 mm (I).

Female same as male.

The latin name “transversus” refers to the interocular space having a dark brown transverse band.

China (Yunnan).

Holotype: China • ♂; Meizihu Reservoir, Pu’er City, Yunnan Province; 1400 m; 21-V-2016; Lu Qiu, Zhi-Wei Qiu leg; SWU-B-EC141701. Paratype: China • 6♂♂; same data as holotype; SWU-B-EC141702-141707.

China • 2♀; Meizihu Reservoir, Pu’er City, Yunnan Province; 1400 m; 20-V-2016; Lu Qiu, Zhi-Wei Qiu leg.

This species closely resembles Margattea transversa sp. nov., but they can be distinguished by the following characteristics: 1) Left phallomere of the former with three long spines, while the latter with four long spines; 2) In the former, median phallomere apex with a slightly curved spine, while the median phallomere apex of latter with a distinct curved spine. In addition, this species is also similar to M. nimbata (Shelford, 1907) in general appearance, but it can be differentiated from the latter by the following characters: 1) median phallomere base with a curved spine, while in the latter, with two curved spines; 2) A long piece of bone extends from the right side of the accessory structure, while absent in the latter; and 3) left phallomere with three long spines; the latter with two long spines.

(mm). Male (n = 5), pronotum: length × width 2.7–2.8 × 3.1–3.6, tegmina length: 10.4–12.6, overall length: 12.8–14.1.

Male. Coloration: body pale yellowish-brown with yellowish-brown (Fig. 9A, B). Face dark yellowish-brown. Interocular space with a brown band. Ocelli spots white. Antennae yellowish-brown. Clypeus pale brown (Fig. 9C). Maxillary palps light linen-colored (Fig. 9E). Pronotal disc light linen-colored with brown stripes and two lateral borders yellowish-white (Fig. 9D). Tegmina pale yellowish-brown, wings medium brown (Fig. 9F, G). Abdomen cream-colored to pale brown. Cerci yellowish brown (Fig. 9K). Styli faint yellow (Fig. 9M). Head: vertex slightly exposed, distance between interocular shorter than antennal socket space (Fig. 9C). Pronotum nearly trapezoidal, broader than long, the widest part after the midpoint, the front and posterior margins nearly straight, and the postero-lateral angle blunt and round; disc with symmetrical irregular maculae (Fig. 9D). Third and fourth palpi of approximately same length, both obviously longer than fifth palp, fifth palp obviously expanded (Fig. 9E). Tegmina and wings: tegmina and wings fully developed, both extending beyond the end of abdomen (Fig. 9A, B). Tegmina with ScP simple, R multi-branched, M straight with seven complete branches. Hind wings with ScP and RA expanded at apex; M straight and simple without branches; CuA with five complete branches (Fig. 9F, G). Legs: anteroventral margin of front femur type B₂ (Fig. 9H). Pulvilli present on four proximal tarsomeres. Tarsal claws symmetrical and specialized, inner margin serrated, arolia present (Fig. 9I). Abdomen and genitalia: eighth abdominal tergum specialized with a tuft (Fig. 9J). Supra-anal plate transverse, posterior margin convex. Paraprocts simple, splitting into two pieces, apex with tufts (Fig. 9K). Subgenital plate symmetrical. Styli similar, slender, distinctly separated; interstylar region slightly convex (Fig. 9M). Left phallomere complex, irregular bone-shaped, with three long spines (Fig. 9L). Median phallomere slender rod-shaped, apex with a slightly curved spine; accessory structure arched, a long piece of bone extends from right side of accessory structure (Fig. 9M). Hook phallomere on right side, apex curved inwards with a short spine (Fig. 9N).

Figure 9. 

A–N Margattea paratransversa sp. nov., male A holotype, dorsal view B holotype, ventral view C head, ventral view D pronotum, dorsal view E maxillary palpi, ventral view F tegmen, dorsal view G hind wing, dorsal view H front femur, ventral view I tarsal claws J eighth abdominal terga K supra-anal plate and paraprocts, ventral view L left phallomere, dorsal view M subgenital plate and median phallomere, dorsal view N hook-like phallomere, dorsal view. Scale bars: 5 mm (A, B); 0.5 mm (C–H, J, K–N); 0.1 mm (I).

Female similar as male.

The species name “paratransversa” reflects its similarity to M. transversa sp. nov.

China (Yunnan).

Holotype: China • ♂; Wangtianshu Scenery Spot, Mengla County, Xishuangbanna Prefecture, Yunnan Province; 23-V-2016; Lu Qiu, Zhi-Wei Qiu leg; SWU-B-EC141601. Paratype: China • 2 ♂♂, same data as for holotype; SWU-B-EC141602-141603.

This species is similar to M. brevialata (Caudell, 1927) in male genitalia, but it can be differentiated from the latter by the following characters: 1) median phallomere slender rod, apex forked without spines; while in the latter, one side of splitting apex with 2 long spines; 2) left phallomere without a spine; the latter with a spine; and 3) supra-anal plate symmetrical, the front and the posterior margin straight; while in the latter, posterior margin convex, the middle part concave.

(mm). Male (n = 3), pronotum: length × width 2.1–2.2 × 3.0–3.2, tegmina length: 9.9–10.9, overall length: 11.6–12.0.

Male. Coloration: body pale yellow with yellowish-brown (Fig. 10A, B). Face pale yellowish-brown. Interocular space with a brown band. Ocelli spots white. Antennae pale yellowish-brown. Clypeus medium yellowish-brown (Fig. 10C). Maxillary palps light linen-colored (Fig. 10E). Pronotal disc pale yellowish-brown with yellowish-brown stripes, and two lateral light linen-colored borders (Fig. 10D). Tegmina light yellowish-brown, wings brownish grey (Fig. 10F, G). Abdomen cream-colored. Cerci yellowish-brown to pale brown (Fig. 10K). Styli faint yellow (Fig. 10L). Head: vertex slightly exposed, distance between interocular shorter than antennal sockets space (Fig. 10C). Pronotum nearly trapezoidal, broader than long, the widest part after midpoint, front and posterior margins nearly straight, and postero-lateral angle blunt and round; disc with symmetrical irregular stripes (Fig. 10D). Third and fourth palpi of approximately same length, both obviously longer than fifth palp, fifth palp obviously expanded (Fig. 10E). Tegmina and wings: tegmina and wings fully developed, both extending beyond the end of abdomen (Fig. 10A, B). Tegmina with Scp simple, R multi-branched, M straight with five complete branches. Hind wings with ScP and RA expanded at apex; M straight and simple, without branches; CuA with six complete branches (Fig. 10F, G). Legs: anteroventral margin of front femur type B₂ (Fig. 10H). Pulvilli present on four proximal tarsomeres. Tarsal claws symmetrical and specialized, inner margin serrated, arolia present (Fig. 10I). Abdomen and genitalia: eighth abdominal tergum specialized with a tuft (Fig. 10J). Supra-anal plate transverse. Paraprocts simple, similar, splitting into two pieces (Fig. 10K). Subgenital plate symmetrical. Styli similar, slender, distinctly separated. Left phallomere complex, irregular bone-shaped. Median phallomere slender rod-shaped with apex forked; the accessory structure arched, at leftmost end with a brush. Hook phallomere on right side, apex curved inwards with a short spine (Fig. 10L).

Figure 10. 

A–L Margattea bicruris sp. nov., male A holotype, dorsal view B holotype, ventral view C head, ventral view D pronotum, dorsal view E maxillary palpi segments 3–5, ventral view F tegmen, dorsal view G hind wing, dorsal view H front femur, ventral view I tarsal claws J eighth abdominal terga K supra-anal plate and paraprocts, ventral view L subgenital plate and phallomeres, dorsal view. Scale bars: 5 mm (A, B); 0.5 mm (C–H, J, K–L); 0.1 mm (I).

Female unknown.

The Latin name “bicruris” refers to the median phallomere having the base forked.

China (Yunnan).