This catalogue cites all the species of Chile in their valid and original combinations, provides details about the name-bearing types of all nominal species, and gives known distributions. It is based on the examination of virtually all of the approximately 450 publications listed in the References.

Valid names are arranged hierarchically and alphabetically according to the categories of subfamily, tribe, genus, subgenus, and species. Synonyms are given for valid names of genera, subgenera, and species, and are listed chronologically. Synonymic lists comprise taxa described from south of the United States, synonyms that have been used as valid names in the literature on Chilean Tachinidae, and (where known and listed last) misidentifications and incorrect spellings.

Each genus-group name is listed with the following information: genus name in italics and capital letters (and additionally in bold if valid, unless misidentified from Chile; e.g., Neoethilla Cerretti et al.), author, year (with letter if applicable), page, note in parentheses if applicable (e.g., junior homonym or proposed as subgenus), type species with author and date, form of type fixation, and country (or region, such as Europe, if country unknown) of the type locality of the type species in square brackets (the last not given for all generic names in O’Hara and Cerretti 2016). Each type species is cited in its original binomen (Recommendation 67B of the Code, ICZN 1999), and if that name is a synonym then it is followed by the valid name of the species in parentheses. We have invoked Article 70.3.2 of the Code (ICZN 1999) to fix the intended species as the type species for generic names that were based on misidentified type species. This maintains the concepts of these generic names as currently accepted and in prevailing usage. The genera so affected are listed below under “Summary of new taxonomic and nomenclatural changes”.

Type species were fixed by original designation, monotypy, subsequent designation, or in a few instances subsequent monotypy, except for type species newly fixed here for nominal genera based on misidentified type species. Fixation by original designation requires an explicit designation of a type species (Article 68.2 of the Code, ICZN 1999), so a new genus “proposed for” or “erected for” a single species has its type species fixed by monotypy. A new genus proposed before 1931 for a single species and accompanied by the expression “gen. n., sp. n.” or an equivalent also has its type species fixed by monotypy (Article 68.2.1). If, on the other hand, the new genus is proposed for more than one new species and the expression “gen. n., sp. n.” or an equivalent is applied to only one of the new species, then that species is fixed as type species by original designation (Article 68.2.1).

Species are listed by valid name followed by the available name(s) associated with it; i.e., the available name of the valid name plus synonyms. The valid name is represented by the valid specific epithet in bold and italics (in italics only if questionably recorded or misidentified from Chile; e.g., Archytas incertus (Macquart)) followed by the author, date (no letter suffix), and known distribution. Author and date are enclosed in parentheses if the species has moved from its original genus. The distribution is given first for the Neotropical Region and then for other regions as explained under “Geographic divisions” and “Distributional data”. Each available name is given in italics in its original combination and spelling followed by author, year (with letter suffix if applicable to match a publication listed in the References), page, and a note in parentheses if applicable (e.g., junior homonym or subsequent spelling). A questionable synonym is preceded by a question mark (e.g., “? Spathipalpus flavifrons Rondani”). Given next is name-bearing type information consisting of status (holotype, lectotype, neotype, or syntypes), sex (of single type, or number and sex of syntypes), type depository (in parentheses), and type locality. If a neotype or lectotype was designated then a citation is given to the designation. Additional information may be given in parentheses with the type depository to cite the number and sex of syntypes existing in a collection if that number is different from the information given in the original description, or if the original description did not provide details about the type series; also, a reference may be cited wherein information can be found about the name-bearing type.

A subsequent spelling of a generic or specific name can be an incorrect subsequent spelling (which is not an available name) or an unjustified emendation (which is an available name with its own author and date). Incorrect subsequent spellings are cited where known to us but others surely exist. An unjustified emendation is cited with an author and date following O’Hara and Cerretti (2016); a name only was given in O’Hara and Wood (2004) and O’Hara et al. (2009) except in rare instances.

Notes and/or references are often given after genus and species entries. Notes provide explanations of some sort; e.g., priority of names, composition of type series, justification for a new combination or new name. References have a standardised format consisting of a source followed by the information provided therein; e.g., first records from countries (as explained under “Distributional data”), redescriptions, keys, figures, type notes. These references attempt to trace the history of name usage and synonymy but do not cite every occurrence of a name in species lists (unless it is a first record from a country).

The following abbreviations are used:

Code International Code of Zoological Nomenclature, specifically the fourth edition published by the International Commission on Zoological Nomenclature in 1999; cited as ICZN 1999.

ICZN International Commission on Zoological Nomenclature.

JEOH James E. O’Hara.

DMW D. Monty Wood.

CRG Christian R. González.

We follow the same method developed by O’Hara et al. (2009) and followed by O’Hara and Cerretti (2016) for citing name-bearing type information for species described without a holotype designation in the original publication or without a subsequent lectotype or neotype designation. Details are provided about name-bearing types based on the content of the original descriptions and are not biased by existing type material in collections (that information being given in parentheses with the type depository). Our format for citing published data on name-bearing types other than a designated holotype, lectotype or neotype is as follows:

Type(s), male: One or more males. This citation is used for a species described from the male sex without indication of whether a single male (i.e., a holotype) or more than one male (i.e., syntypes) composed the type series.

Type(s), female: One or more females. See “Type(s), male”.

Type(s), unspecified sex: One or more specimens with no indication of sex.

Syntypes, [number] male[s] and [number] female[s] (e.g., “Syntypes, 3 males and 2 females”): Species described from an indicated number of males and females.

Syntypes, males and females: Species described from both sexes but the number of each sex was not given. A number in front of “males” with no number in front of “females” refers to the total number of males and females.

Syntypes, males: Species described from more than one male but without indication of the number of males.

Syntypes, females: Species described from more than one female but without indication of the number of females.

Syntypes, unspecified number and sex: Species described from more than one specimen but without indication of sex or number of specimens.

In following the foregoing format we have complied with Recommendation 73F of the Code (ICZN 1999), “Avoidance of assumption of holotype”, which states: “Where no holotype or syntype was fixed for a nominal species-group taxon established before 2000, and when it is possible that the nominal species-group taxon was based on more than one specimen, an author should proceed as though syntypes may exist and, where appropriate, should designate a lectotype rather than assume a holotype (see also Article 74.6)”. See O’Hara et al. (2009: 9–10) for a further discussion of this issue.

By following Recommendation 73F of the Code, assumed holotypes take on the status of syntypes. The recommendation favours “where appropriate” the designation of lectotypes. We have combined the spirit of Recommendation 73F and the provisions of Article 74.5 of the Code (ICZN 1999) to recognise certain published statements (as discussed in the next section) about assumed holotypes as lectotype fixations. This follows O’Hara et al. (2009) and O’Hara and Cerretti (2016) and is in our opinion the best way to reconcile assumed holotypes with the modern rules of nomenclature, while also giving credit of lectotype fixations to the authors who assumed holotypes.

There are two types of lectotypification in zoological nomenclature, explicit and implicit. In the former, a single syntype in a type series is designated as lectotype; in the latter, there is some form of statement that can be construed as the selection of a single name-bearing type. We follow O’Hara et al. (2009) in using the term “lectotype designation” for an explicit lectotypification and “lectotype fixation” for an implicit lectotypification. There is good reason to distinguish between the two because implicit lectotypifications are open to some interpretation, especially with respect to Article 74.5 of the Code (ICZN 1999: 82–83) that deals in part (see also Article 74.6) with lectotype designations before 2000:

“In a lectotype designation made before 2000, either the term ‘lectotype’, or an exact translation or equivalent expression (e.g. ‘the type’), must have been used or the author must have unambiguously selected a particular syntype to act as the unique name-bearing type of the taxon. When the original work reveals that the taxon had been based on more than one specimen, a subsequent use of the term ‘holotype’ does not constitute a valid lectotype designation unless the author, when wrongly using that term, explicitly indicated that he or she was selecting from the type series that particular specimen to serve as the name-bearing type”.

What constitutes a valid lectotypification (or lectotype fixation in our terminology) in the foregoing is largely dependent on how one interprets the passage about an author explicitly indicating “that he or she was selecting from the type series that particular specimen to serve as the name-bearing type”. At one end of the spectrum is the mere mention of a “holotype” or “type” by a subsequent author when the original type series clearly consisted of two or more syntypes. This statement does not constitute a lectotype fixation because the “holotype” is not distinguishable from other syntypes. At the other end of the spectrum is the mention of a “holotype” or “type” with accompanying details about its labelling, features, damage, etc. that clearly distinguishes that specimen from other syntypes; or perhaps there is only one type specimen in a collection and it is an “assumed holotype” (see section above) for a species described from an unspecified number of specimens. We considered these latter statements about a single type to qualify as lectotype fixations under Article 74.5 because they contain an explicit indication that an author accepted the cited “holotype” or “type” as the name-bearing type and restricted the term to a single recognisable specimen in a collection.

O’Hara and Cerretti (2016) recognised lectotype fixations in Townsend’s Manual of Myiology (Parts I–XII, 1934–1942), reversing the practice of O’Hara et al. (2009). We follow the former authors in recognising lectotype fixations in Manual of Myiology if there is a strong possibility of the lectotype being recognised in the stated collection. See O’Hara and Cerretti (2016: 11–12) for a more detailed discussion of this subject.

Type localities are cited first by country and then by location within the country from larger to smaller geographic area or place. Spellings of geographic areas and places follow The Times Comprehensive Atlas of the World (Times Books 2007), if found in that work. Modern names and spellings are given where these have been determined. Country and higher administrative subdivisions (i.e., regions and provinces of Chile, provinces of Argentina, states of Brazil, regions of Peru, etc.) are given only in their modern equivalents. For locality names that have changed since they were first published, the modern spelling is given first followed by the original spelling in square brackets and quotes; e.g., Puerto del Hambre [as “Port Famine”]. Elevations are cited in metres (m) or feet (ft) as given by the author. Coordinates given in an original publication are cited in parentheses after the type locality; e.g., Chile, Arica y Parinacota, Parinacota, Putre, 3530 m (18°12′S, 69°35′W). Coordinates are included for many type localities that we had difficulty locating and these are given in square brackets after the locality to distinguish them from coordinates provided by an author; e.g., Chile, Valparaíso, Marga Marga, Bosque Los Perales [as “Perales”, ca. 33°9′S, 71°18′W]. Criteria for citing type localities in Sweden are explained in O’Hara et al. (2009: 11). The few localities we could not find are given in quotes; e.g., Trinidad, “Legerville Mt.”. A variety of resources were used to locate type localities not found in The Times Comprehensive Atlas of the World including other atlases, maps, literature, and Internet searches (often for the locality and/or collector).

Type localities in Chile are preceded by region and province (e.g., Valparaíso, Marga Marga). Those in other countries are preceded by province (Argentina, Ecuador, etc.), state (Brazil, Mexico, etc.) or department (Peru, Uruguay, etc.), or by no higher administrative division (e.g., countries of the West Indies).

Greater Antilles (part of the West Indies).

Bahamas; Cayman Islands (United Kingdom Overseas Territory); Cuba; Dominican Republic; Haiti; Jamaica; Puerto Rico; Turks & Caicos (United Kingdom Overseas Territory).

eastern Lesser Antilles (Leeward and Windward islands in the Lesser Antilles of the West Indies).

Anguilla (United Kingdom Overseas Territory); Antigua [Antigua & Barbuda] (including Redonda); Barbados; Dominica; Grenada; Guadeloupe (including Marie-Galante, La Désirade, Îles des Saintes) (France); Martinique (France); Montserrat (United Kingdom Overseas Territory); Saba (Netherlands); Saint-Barthélemy (France); Saint Kitts [Saint Kitts and Nevis]; Saint Lucia; Saint-Martin (comprising Saint Martin [France] and Sint Maarten [Netherlands]); Saint Vincent [Saint Vincent and The Grenadines]; Sint Eustatius (Netherlands); Virgin Islands (including the United States islands of Saint Thomas, Saint John, and Saint Croix, and the British Virgin Islands of Tortola, Virgin Gorda, Anegada, and Jost Van Dyke).

southern Lesser Antilles (islands north of the Venezuelan coast in the Lesser Antilles of the West Indies).

Aruba (Netherlands); Blanquilla (Venezuela); Bonaire (Netherlands); Curaçao (Netherlands); Los Roques Archipelago (Venezuela); Los Testigos (Venezuela); Margarita (including smaller neighbouring islands, principally La Tortuga, Coche, and Cubagua; all comprising Nueva Esparta state, Venezuela); Trinidad & Tobago.

Middle America (mainland Middle America).

Belize; Costa Rica; El Salvador; Guatemala; Honduras; Mexico; Nicaragua; Panama.

South America. [Cited as South America when more detailed distributional data is not available.]

Argentina; Bolivia; Brazil; Chile (excluding Juan Fernández Islands); Colombia; Ecuador (excluding Galápagos Islands); Falkland Islands (disputed United Kingdom Overseas Territory); French Guiana (France); Juan Fernández Islands (Chile); Galápagos Islands (Ecuador); Guyana; Paraguay; Peru; South Georgia (including the South Sandwich Islands; disputed United Kingdom Overseas Territory); Suriname; Uruguay; Venezuela.

The limits of the Nearctic Region follow O’Hara et al. (2020: 8, 18 [map 1]) and include the following subdivisions: Bermuda (United Kingdom Overseas Territory); Canada; Greenland (Denmark); United States [United States of America, as “USA” for type localities; Hawaii as part of Australasian and Oceanian regions].

See O’Hara et al. (2020: 11, 21 [map 4], 22 [map 5]) for the countries included in the broader subdivisions recognised here: Central Asia; China (Palaearctic part, sensu O’Hara et al. 2020); Europe; Japan (excluding Ryukyu Islands); Kazakhstan; Korean Peninsula; Middle East; Mongolia; North Africa; Russia; Transcaucasia.

This region is subdivided by country, as explained and mapped in O’Hara and Cerretti (2016: 15, 16 [fig. 1]) and O’Hara et al. (2020: 13, 23 [map 6]).

The Oriental Region is bounded on the north by the Palaearctic Region (O’Hara et al. 2020) and on the south by Weber’s Line (Evenhuis 1989: 31). See O’Hara et al. (2020: 15, 22 [map 5], 24 [map 7], 25 [map 8]) for a list of countries/subdivisions and maps.

The Australasian and Oceanian regions are bounded on the north by Weber’s Line (Evenhuis 1989: 31). See O’Hara et al. (2020: 16, 26 [map 9]) for a list of countries/subdivisions and maps.

A species recorded from all regions and subdivisions recognised here would be cited with the following distribution:

Neotropical: Greater Antilles (Bahamas, Cayman Islands, Cuba, Dominican Republic, Haiti, Jamaica, Puerto Rico, Turks & Caicos), eastern Lesser Antilles (Anguilla, Antigua, Barbados, Dominica, Grenada, Guadeloupe, Martinique, Montserrat, Saba, Saint-Barthélemy, Saint Kitts, Saint Lucia, Saint-Martin, Saint Vincent, Sint Eustatius, Virgin Islands), southern Lesser Antilles (Aruba, Blanquilla, Bonaire, Curaçao, Los Roques Archipelago, Los Testigos, Margarita, Trinidad & Tobago), Middle America (Belize, Costa Rica, El Salvador, Guatemala, Honduras, Mexico, Nicaragua, Panama), South America (Argentina, Bolivia, Brazil, Chile, Colombia, Ecuador, Falkland Islands, French Guiana, Juan Fernández Islands, Galápagos Islands, Guyana, Paraguay, Peru, South Georgia, Suriname, Uruguay, Venezuela). Nearctic: Bermuda, Canada, Greenland, United States. Palaearctic: Central Asia, China [Pal.], Europe, Japan, Kazakhstan, Korean Peninsula, Middle East, Mongolia, North Africa, Russia, Transcaucasia. Oriental: Andaman & Nicobar Islands, Bangladesh, Brunei, Bhutan, Cambodia, China [Orien.], Christmas & Cocos Islands, India, Indonesia [Orien.], Japan [Ryukyu Islands], Laos, Malaysia, Maldives etc., Myanmar, Nepal, Pakistan, Philippines, Singapore, Sri Lanka, Taiwan, Thailand, Vietnam. Australasian & Oceanian: Australia, Hawaii, Indonesia [Aust.], Melanesia, Micronesia, New Zealand, Papua New Guinea, Polynesia.

Distributions are cited at the country level within the Neotropical Region (as listed in Geographic Divisions section) for each species based on published records and our examination of specimens in CNC and UMCE. The principal sources for published records were Aldrich (1934) and the publications of Cortés (and co-authors, 1944–1992, mainly Chile) and Blanchard (1935–1966, mainly Argentina). The CNC has more than 3000 Chilean specimens but few have been reported in the literature. These mostly originated from the following sources: nearly 2000 collected by the esteemed Chilean entomologist Luis E. Peña [Guzmán] from 1959–1980 and 1994–1995 (privately sold to DMW over a period of years and recently donated to CNC); ca. 1000 collected by DMW and wife Grace between December 1993 and February 1994 (recently donated to CNC and including ca. 150 specimens from Argentina); and over 200 collected by JEOH in December 2015 (see Stireman et al. 2016).

A reference is cited, if known, for the first record of a species from countries different from the one(s) from which the species was described. Subsequent records from the same country are not generally given unless significant in some way. The first record is considered the most important because it is sometimes the source for later records even if it was based on a misidentification.

Two genera are newly recorded from Chile (one also newly recorded from Argentina).

Chaetoepalpus Vimmer & Soukup, 1940 (based on new records of Chaetoepalpus coquilleti Vimmer & Soukup, 1940). New records from Argentina and Chile.

Patelloa Townsend, 1916 (based on new synonymy of Macropatelloa Townsend, 1931 with Patelloa). New record from Chile.

The following species are newly recorded from Chile or other countries.

Lypha ornata Aldrich, 1934. New record from Chile.

Chaetoepalpus coquilleti Vimmer & Soukup, 1940. New records from Argentina and Chile.

Phytomyptera evanescens (Cortés, 1967). New record from Argentina.

Xanthobasis unicolor Aldrich, 1934. New record from Chile.

Species newly recognised as misidentified or misrecorded from Chile are listed here. The reasons for not recognising them from Chile are given under each name in the catalogue.

Archytas incertus (Macquart, 1851).–Not Chile [Argentina, Brazil, Paraguay, Uruguay].

Archytas seminiger (Wiedemann, 1830).–Not Chile [Brazil, Colombia].

Gonia crassicornis (Fabricius, 1794).–Not Chile [Brazil, Peru, Venezuela; also Middle America, West Indies and Nearctic].

Lespesia andina (Bigot, 1888), nomen dubium.–Not Chile [Cuba].

Lespesia archippivora (Riley, 1871).–Not Chile [widespread throughout the Nearctic Region and most of Middle and South America].

Neoethilla ignobilis (van der Wulp, 1890).–Not Chile [Mexico; United States].

Siphona (Siphona) geniculata (De Geer, 1776).–Not Chile [Nearctic (introduced), Palaearctic].

Winthemia quadripustulata (Fabricius, 1794).–Not Chile [Palaearctic; also Nearctic and Oriental].

Billaea Robineau-Desvoidy, 1830

Paratheresia rufiventris Townsend, 1929 and Sarcoprosena rufiventris Townsend, 1929 are secondary homonyms when placed together in Billaea. As the First Reviser (Article 24.2.2 of the Code, ICZN 1999), we fix Paratheresia rufiventris as the senior homonym.

Myiopharus Brauer & Bergenstamm, 1889

Mayophorinia angusta Townsend, 1927 and Metarrhinomyia angusta Townsend, 1927 are secondary homonyms when placed together in Myiopharus. As the First Reviser (Article 24.2.2 of the Code, ICZN 1999), we fix Mayophorinia angusta as the senior homonym.

Eight new names are proposed for preoccupied names that came to our attention during the preparation of this catalogue and belong to genera recorded from Chile. The preoccupied names do not concern Chilean species except for one but are renamed to avoid the confusion of having two Neotropical species with the same name in the same genus. The etymology of each new name is given in the catalogue. The country of the type locality of each preoccupied species name is given at the end of each entry.

Billaea rufescens O’Hara & Wood is proposed as a nomen novum for Sarcoprosena rufiventris Townsend, 1929, a name preoccupied in the genus Billaea Robineau-Desvoidy, 1830 by Paratheresia rufiventris Townsend, 1929 [Peru]. Nom. nov.

Billaea triquetrus O’Hara & Wood is proposed as a nomen novum for Sarcoprosena triangulifera Townsend, 1927, a name preoccupied in the genus Billaea Robineau-Desvoidy, 1830 by Dexia triangulifera Zetterstedt, 1844 [Peru]. Nom. nov.

Eucelatoria nudioculata O’Hara & Wood is proposed as a nomen novum for Eucelatorioidea nigripalpis Thompson, 1968, a name preoccupied in the genus Eucelatoria Townsend, 1909 by Chetolyga nigripalpis Bigot, 1889 [Trinidad]. Nom. nov.

Eucelatoria oblonga O’Hara & Wood is proposed as a nomen novum for Urodexodes elongatum Cortés & Campos, 1974, a name preoccupied in the genus Eucelatoria Townsend, 1909 by Exorista elongata van der Wulp, 1890 [Chile]. Nom. nov.

Lespesia thompsoni O’Hara & Wood is proposed as a nomen novum for Sturmiopsoidea obscura Thompson, 1966, a name preoccupied in the genus Lespesia Robineau-Desvoidy, 1863 by Eurigaster obscurus Bigot, 1857 [Cuba]. Nom. nov.

Myiopharus charapensis O’Hara & Wood is proposed as a nomen novum for Metarrhinomyia angusta Townsend, 1927, a name preoccupied in the genus Myiopharus Brauer & Bergenstamm, 1889 by Mayophorinia angusta Townsend, 1927 [Peru]. Nom. nov.

Myiopharus incognitus O’Hara & Wood is proposed as a nomen novum for Stenochaeta claripalpis Thompson, 1968, a name preoccupied in the genus Myiopharus Brauer & Bergenstamm, 1889 by Neoxynopsoidea claripalpis Thompson, 1968 [Trinidad]. Nom. nov.

Myiopharus rufopalpus O’Hara & Wood is proposed as a nomen novum for Paralispe palpalis Townsend, 1929, a name preoccupied in the genus Myiopharus Brauer & Bergenstamm, 1889 by Myioxynops palpalis Townsend, 1927 [Peru]. Nom. nov.

Article 70.3.2 of the Code (ICZN 1999) allows the type species of a nominal genus to be fixed as the species intended by the original author if the type species designated by that author was misidentified. We have invoked Article 70.3.2 for the three instances of misidentified type species in this catalogue that had not been dealt with previously (e.g., O’Hara and Wood 2004) to preserve the current concepts of the genera involved. Type species are fixed for the following nominal genera (see catalogue for further details).

Parafabricia Brauer & Bergenstamm, 1894: 612 [also 1895: 76]. Type species newly fixed as Parafabricia perplexa Townsend, 1931. Synonym of Archytas Jaennicke, 1867.

Tachinodes Brauer & Bergenstamm, 1889: 133 [also 1889: 65]. Type species newly fixed as Jurinia metallica Robineau-Desvoidy, 1830. Synonym of Archytas Jaennicke, 1867.

Willistonia Brauer & Bergenstamm, 1889: 97 [also 1890: 29]. Type species newly fixed as Willistonia aldrichi Townsend, 1931. Synonym of Belvosia Robineau-Desvoidy, 1830.

Lectotypes are designated for four nominal species (see Lectotype Designations section).

Echinomyia pygmaea Macquart, 1851. This is a valid name in the genus Peleteria Robineau-Desvoidy, 1830, as Peleteria pygmaea (Macquart).

Gonia chilensis Macquart, 1844. This is a junior synonym in the genus Gonia Meigen, 1803. The valid name of the species is Gonia pallens Wiedemann, 1830.

Masicera auriceps Macquart, 1844. This is a valid name in the genus Lespesia Robineau-Desvoidy, 1863, as Lespesia auriceps (Macquart).

Prosopochoeta nitidiventris Macquart, 1851. This is a valid name in the genus Prosopochaeta Macquart, 1851.

New and revived combinations proposed in this work are listed below. These are based

on the study of type material, authoritatively identified specimens, and/or descriptions

and figures in the literature, mostly by DMW.

Blepharipeza andina Bigot, 1888 is moved from an unplaced species in “Sturmiini” or Tachinidae to Lespesia Robineau-Desvoidy, 1863 as a nomen dubium. Distribution: Cuba (not Chile as published). Comb. nov.

Camposodes evanescens Cortés, 1967 is moved from its original placement in Camposodes Cortés, 1967 to Phytomyptera Rondani, 1845. Distribution: Argentina, Chile. Comb. nov.

Ectophasiopsis ypiranga Dios & Nihei, 2017 is moved from its original placement in Ectophasiopsis Townsend, 1915 to Trichopoda Berthold, 1827 (and assigned to subgenus Galactomyia Townsend, 1908). Distribution: Argentina, Brazil. Comb. nov.

Embiomyia australis Aldrich, 1934 (type species of Embiomyia Aldrich, 1934) is moved from its original placement in Embiomyia to Steleoneura Stein, 1924 (with Embiomyia in synonymy). Distribution: Argentina, Chile. Comb. nov.

Eurigaster modestus Bigot, 1857 is moved from its position in unplaced species of Exoristinae (as “Goniinae”) by Guimarães (1971: 215) to Lespesia Robineau-Desvoidy, 1863. Distribution: Cuba. Comb. nov.

Eurigaster obscurus Bigot, 1857 is moved from its position in unplaced species of Exoristinae (as “Goniinae”) by Guimarães (1971: 215) to Lespesia Robineau-Desvoidy, 1863. Distribution: Cuba. Comb. nov.

Macropatelloa tanumeana Townsend, 1931 (type species of Macropatelloa Townsend, 1931) is moved from its original placement in Macropatelloa to Patelloa Townsend, 1916 (with Macropatelloa in synonymy). Distribution: Argentina, Chile. Comb. nov.

Masicera insignis van der Wulp, 1882 is moved from its placement in Sturmia Robineau-Desvoidy, 1830 by previous authors (e.g., Cortés and Hichins 1969: 59; Guimarães 1971: 192; Henry 1987: 200) to Drino Robineau-Desvoidy, 1863. Distribution: Argentina, Chile. Comb. nov.

Parasetigena hichinsi Cortés, 1967 is moved from its original placement in Parasetigena Brauer & Bergenstamm, 1891 to Chetogena Rondani, 1856. Distribution: Chile. Comb. nov.

Parasetigena porteri Brèthes, 1920 is moved from its placement in Stomatotachina Townsend, 1931 by previous authors (e.g., Guimarães 1971: 160; Mulieri et al. 2013: 169) to Chetogena Rondani, 1856. Distribution: Chile. Comb. nov.

Phorocera calyptrata Aldrich, 1934 is moved from uncertain placements by Guimarães (1971: 152, unplaced in Blondeliini), Henry (1987: 206, in Phorocera Robineau-Desvoidy, 1830 but genus unplaced in Tachinidae) and González (1992b: 183, in Phorocera but genus unplaced in Exoristinae [as “Goniinae”]) to Admontia Brauer & Bergenstamm, 1889. Distribution: Argentina, Chile. Comb. nov.

Poliops auratus Campos, 1953 is moved from its original placement in Poliops Aldrich, 1934 to Admontia Brauer & Bergenstamm, 1889. Distribution: Chile. Comb. nov.

Poliops striatus Aldrich, 1934 is moved from its original placement in Poliops Aldrich, 1934 to Admontia Brauer & Bergenstamm, 1889. Distribution: Argentina, Chile. Comb. nov.

Ruiziella frontosa Cortés, 1951 is moved from its original placement in Ruiziella Cortés, 1951 to Chaetoepalpus Vimmer & Soukup, 1940, where it is placed in synonymy with C. coquilleti Vimmer & Soukup, 1940, syn. nov. Distribution of C. coquilleti: Argentina, Chile, Peru. Comb. nov.

Ruiziella luctuosa Cortés, 1951 is moved from its original placement in Ruiziella Cortés, 1951 to Chaetoepalpus Vimmer & Soukup, 1940. Distribution: Argentina, Chile. Comb. nov.

Sarcoprosena luteola Cortés & Campos, 1974 is moved from its original placement in Sarcoprosena Townsend, 1927 to Billaea Robineau-Desvoidy, 1830. Distribution: Chile. Comb. nov.

Sarcoprosena rufiventris Townsend, 1929 is moved from its original placement in Sarcoprosena Townsend, 1927 to Billaea Robineau-Desvoidy, 1830. Comb. nov. The name S. rufiventris is a junior secondary homonym of Paratheresia rufiventris Townsend, 1929 when placed in Billaea and is renamed herein as Billaea rufescens O’Hara & Wood, nom. nov. Distribution: Peru.

Sarcoprosena triangulifera Townsend, 1927 (type species of Sarcoprosena Townsend, 1927) is moved from its original placement in Sarcoprosena to Billaea Robineau-Desvoidy, 1830 (with Sarcoprosena in synonymy). Comb. nov. The name S. triangulifera is a junior secondary homonym of Dexia triangulifera Zetterstedt, 1844 when placed in Billaea and is renamed herein as Billaea triquetrus O’Hara & Wood, nom. nov. Distribution: Peru.

Saundersia aurea Giglio-Tos, 1893 is moved from its placement in Epalpus Rondani, 1850 by Guimarães (1971: 64) to “Unplaced species of Tachinini”. Distribution: Mexico. Comb. nov.

Schistostephana aurifrons Townsend, 1919 (type species of Schistostephana Townsend, 1919) is moved from its original placement in Schistostephana to Billaea Robineau-Desvoidy, 1830 (with Schistostephana in synonymy). Distribution: Peru. Comb. nov.

Siphoactia charapensis Townsend, 1927 (type species of Siphoactia Townsend, 1927) is moved from its original placement in Siphoactia to Clausicella Rondani, 1856 (with Siphoactia in synonymy). Distribution: Peru. Comb. nov.

Siphoactia peregrina Cortés & Campos, 1971 is moved from its original placement in Siphoactia Townsend, 1927 to Clausicella Rondani, 1856. Distribution: Chile. Comb. nov.

Stomatotachina splendida Townsend, 1931 (type species of Stomatotachina Townsend, 1931) is moved from its original placement in Stomatotachina to Chetogena Rondani, 1856 (with Stomatotachina in synonymy). Stomatotachina splendida continues to be treated as a junior subjective synonym of Parasetigena porteri Brèthes, 1920 (see above). Comb. nov.

Sturmia festiva Cortés, 1944 is moved from its original placement in Sturmia Robineau-Desvoidy, 1830 to Drino Robineau-Desvoidy, 1863. Distribution: Argentina, Chile. Comb. nov.

Sturmiopsoidea obscura Thompson, 1966 (type species of Sturmiopsoidea Thompson, 1966) is moved from its original placement in Sturmiopsoidea to Lespesia Robineau-Desvoidy, 1863 (with Sturmiopsoidea in synonymy). Comb. nov. The name S. obscura is a junior secondary homonym of Eurigaster obscurus Bigot, 1857 when placed in Lespesia and is renamed herein as Lespesia thompsoni O’Hara & Wood, nom. nov. Distribution: Trinidad.

Trichopoda arcuata Bigot, 1876 is returned to Trichopoda Berthold, 1827 (and assigned to subgenus Galactomyia Townsend, 1908) from its placement in Ectophasiopsis Townsend, 1915 by previous authors (e.g., Aldrich 1934: 12; Guimarães 1971: 12; Dios and Nihei 2017: 5). Distribution: Argentina, Chile. Comb. revived.

Trichopoda gradata Wiedemann, 1830 is returned to Trichopoda Berthold, 1827 (and assigned to subgenus Galactomyia Townsend, 1908) from its placement in Ectophasiopsis Townsend, 1915 by Dios and Nihei (2017: 10). Distribution: Argentina, Brazil, Uruguay. Comb. revived.

New and revived generic and specific synonymies are proposed for the names below. As with the new and revived combinations listed above, they result from the study of type material, authoritatively identified specimens, and/or descriptions and figures in the literature, mostly by DMW.

Camposodes Cortés, 1967, formerly treated as a genus (e.g., Guimarães 1971: 166; Cortés 1984: 378), is synonymised with Phytomyptera Rondani, 1845. Syn. nov.

Ectophasiopsis Townsend, 1915, formerly treated as a genus (e.g., Aldrich 1934: 11; Guimarães 1971: 12; Dios and Nihei 2017: 4), is synonymised with Trichopoda Berthold, 1827, subgenus Galactomyia Townsend, 1908. Syn. nov.

Embiomyia Aldrich, 1934, formerly treated as a genus (e.g., Cortés and Hichins 1969: 32; Guimarães 1971: 85), is synonymised with Steleoneura Stein, 1924. Syn. nov.

Fabricia andicola Bigot, 1888, treated as a junior synonym of Peleteria filipalpis (Rondani, 1863) by Guimarães (1971: 44), is returned to synonymy with Peleteria robusta (Wiedemann, 1830) as proposed earlier by Guimarães (1962: 484). Syn. revived.

Macropatelloa Townsend, 1931, formerly treated as a genus (e.g., Cortés 1986: 144, 158), González (1992b: 179), is synonymised with Patelloa Townsend, 1916. Syn. nov.

Peleteria inca Curran, 1925, treated as a junior synonym of Peleteria filipalpis (Rondani, 1863) by Guimarães (1971: 44), is returned to synonymy with Peleteria robusta (Wiedemann, 1830) as proposed earlier by Guimarães (1962: 484). Syn. revived.

Poliops Aldrich, 1934, formerly treated as a genus (e.g., Guimarães 1971: 169; Cortés 1979: 81), is synonymised with Admontia Brauer & Bergenstamm, 1889. Syn. nov.

Ruiziella Cortés, 1951, formerly treated as a genus (e.g., Guimarães 1971: 45; Cortés and Campos 1974: 116), is synonymised with Chaetoepalpus Vimmer & Soukup, 1940. Syn. nov.

Ruiziella frontosa Cortés, 1951, formerly treated as a valid species of Ruiziella (e.g., Guimarães 1971: 45), is synonymised with Chaetoepalpus coquilleti Vimmer & Soukup, 1940 in the genus Chaetoepalpus Vimmer & Soukup, 1940. Comb. nov., syn. nov.

Sarcoprosena Townsend, 1927, formerly treated as a genus (e.g., Guimarães 1971: 38; Cortés 1984: 381), is synonymised with Billaea Robineau-Desvoidy, 1830. Syn. nov.

Schistostephana Townsend, 1919, formerly treated as a genus (e.g., Guimarães 1971: 38), is synonymised with Billaea Robineau-Desvoidy, 1830. Syn. nov.

Siphoactia Townsend, 1927, formerly treated as a genus (e.g., Guimarães 1971: 170; Cortés 1984: 381), is synonymised with Clausicella Rondani, 1856. Syn. nov.

Stomatotachina Townsend, 1931, formerly treated as a genus (e.g., Guimarães 1971: 160; Mulieri et al. 2013: 169; Nihei 2015: 1), is synonymised with Chetogena Rondani, 1856. Syn. nov.

Sturmiopsoidea Thompson, 1966, formerly treated as a genus (e.g., Guimarães 1971: 192), is synonymised with Lespesia Robineau-Desvoidy, 1863. Syn. nov.

GONZALEZODORIA Cortés, 1967b: 18. Type species: Gonzalezodoria gonioides Cortés, 1967, by original designation [Chile].

Notes: Gonzalezodoria gonioides is a small dark tachinid with a globous abdomen that Cortés (1967b: 18) noted would run to Dexiinae in the keys of Mesnil (1939), or to the Prosenidae in the family key of Townsend (1936b). In Aldrich’s (1934) key to Tachinidae of Patagonia and South Chile it runs to Myiophasia Brauer & Bergenstamm, 1891 (now a synonym of Gnadochaeta Macquart, 1851) but “con el cual no tiene ningun parecido” [“with which it has no resemblance”] according to Cortés (1967b: 20). Guimarães (1971: 26) did not agree and placed Gonzalezodoria in the Myiophasiini along with a dozen species of Myiophasia and five monotypic genera. We have examined five females of G. gonioides in CNC with the following data: Chile, Los Lagos, Volcán Osorno, La Picada, 600 m, 1980, L. Peña (CNC1546958, CNC1546966–CNC1546969). They clearly belong to the “Oestrophasiini” sensu Guimarães (1977a), differing from the genera treated therein in having setae on the parafacial that are continuous with the orbitals on the fronto-orbital plate (as shown in figs 1–2 in Cortés 1967b: 21) and lacking banding on the wing. Guimarães (1977a: 216) remarked that the male terminalia of Oestrophasiini “are very similar to those of the Old World Dufouriini … but it will be many years before the interrelationships of this difficult group … becomes clear”. The recent molecular phylogeny of Tachinidae of Stireman et al. (2019: 9 [fig. 4]) places Oestrophasia in the Dufouriini and Gonzalezodoria belongs there as well.

The male of G. gonioides has not been described but a single male in CNC with the following data might be the male of this species: Santiago Metro. Region, Mirador de Los Tres Valles, 1820 m, 7.xii.2015, J.E. O’Hara [CNC487604]. This male is a good match morphologically with the aforementioned females but the colouration is different. The females have a yellow head and thorax and a black abdomen; the male has a yellow head except for black underlying the continuous orbital and parafacial setae, black thorax except for yellow scutellum, and yellow abdomen except for black syntergite 1+2, median vitta and bands posteriorly on tergites 3–5.

gonioides Cortés, 1967.—Neotropical: South America (Chile). (Fig. 4b)

Gonzalezodoria gonioides Cortés, 1967b: 19. Holotype female (EEAM). Type locality: Chile, Coquimbo, Limarí, 15 km southwest of Pachingo, near Parque Nacional Bosque Fray Jorge, 110–250 m.

XANTHOBASIS Aldrich, 1934: 110. Type species: Xanthobasis angustifrons Aldrich, 1934, by original designation [Argentina].

PROXANTHOBASIS Blanchard, 1966b: 219. Type species: Proxanthobasis rufipes Blanchard, 1966, by original designation [Argentina].

Note: Cortés (1973a: 101) synonymised Proxanthobasis with Xanthobasis and Cortés (1986: 147) later commented that two additional monotypic genera described from Argentina, Neoxanthobasis Blanchard, 1966 and Paraxanthobasis Blanchard, 1966, may also be synonyms. We did not examine specimens of these last two nominal genera and cannot comment on the merits of this proposition but we do include all three nominal genera in the Eutrixini based on their presumed close relationship.

References: Aldrich (1934: 110), key to three Patagonian species; Townsend (1936c: 47), diagnosis of Ebeniini and key to genera (including Xanthobasis); Townsend (1939c: 163), redescription of Xanthobasis; Cortés (1973a: 101), synonymy of Proxanthobasis with Xanthobasis; Cortés (1986: 143, 147), in key to tachinid genera of Aysén and Magallanes regions, comments on likely additional generic synonymy.

angustifrons Aldrich, 1934.—Neotropical: South America (Argentina, Chile).

Xanthobasis angustifrons Aldrich, 1934: 111. Holotype male (USNM). Type locality: Argentina, Río Negro, Lago Gutiérrez.

Note: Xanthobasis angustifrons was recorded from both Argentina and Chile in the original description.

References: Blanchard (1966b: 225), taxonomic notes; Cortés (1973a: 101), taxonomic notes.

rufescens (Blanchard, 1966).—Neotropical: South America (Argentina, Chile). (Fig. 4c)

Proxanthobasis rufescens Blanchard, 1966b: 222. Holotype male (not located). Type locality: Argentina, Río Negro, San Carlos de Bariloche [as “Bariloche”].

Reference: Cortés (1973a: 101), moved to Xanthobasis, first record from Chile.

unicolor Aldrich, 1934.—Neotropical: South America (Argentina, Chile). New record from Chile.

Xanthobasis unicolor Aldrich, 1934: 112. Holotype male (NHMUK). Type locality: Argentina, Río Negro, Lago Gutiérrez.

Note: Xanthobasis unicolor is recorded from Chile for the first time based on material from Rofuco [in Los Ríos Region, Valdivia Province] in MZSP identified by R. Cortés (ex. unpublished notes of Cortés in UMCE examined by CRG).

References: Blanchard (1966b: 218), redescription and wing figure, assigned to Paraxanthobasis Blanchard, 1966; Guimarães (1971: 110), as Paraxanthobasis unicolor; Cortés (1986: 147), as Xanthobasis unicolor.

ACTINOPLAGIA Blanchard, 1940: 234. Type species: Actinoplagia koehleri Blanchard, 1940, by original designation [Argentina].

Reference: Cortés (1967b: 12), key to separate Actinoplagia Blanchard and Chaetodemoticus Brauer & Bergenstamm.

koehleri Blanchard, 1940.—Neotropical: South America (Argentina, Chile, Uruguay).

Actinoplagia koehleri Blanchard, 1940: 234. Holotype male (MACN). Type locality: Argentina, Buenos Aires, Arrecifes.

References: Parker et al. (1951 [pages unknown], also 1953: 53, 66), first record from Uruguay; Parker (1953: 62), figures of first instar larva and puparium; Blanchard (1963: 170), redescription, head figures; Cortés (1967b: 12), first record from Chile; Mulieri et al. (2013: 165), notes on holotype in MACN.

APHELOGASTER Aldrich, 1934: 22 (junior homonym of Aphelogaster Kolbe, 1897). Type species: Aphelogaster coracella Aldrich, 1934, by original designation [Argentina].

ALDRICHIOPA Guimarães, 1971: 165 (nomen novum for Aphelogaster Aldrich, 1934).

References: Townsend (1936c: 129), diagnosis of adults and immatures of Actiini and key to genera (including Aphelogaster); Townsend (1940a: 192), redescription of Aphelogaster.

coracella (Aldrich, 1934).—Neotropical: South America (Argentina, Chile).

Aphelogaster coracella Aldrich, 1934: 23. Holotype male (NHMUK). Type locality: Argentina, Río Negro, Lago Gutiérrez.

Reference: Townsend (1940a: 192), first record from Chile.

ALEXOGLOBLINIA Cortés, 1945b: 256. Type species: Metopomuscopteryx shannoni Aldrich, 1934, by original designation [Argentina].

shannoni (Aldrich, 1934).—Neotropical: South America (Argentina, Chile).

Metopomuscopteryx shannoni Aldrich, 1934: 46. Holotype male (USNM). Type locality: Argentina, Río Negro, Lago Nahuel Huapí.

Reference: Stireman et al. (2016: 34), first record from Chile (Araucanía Region, Parque Nacional Conguillío, Laguna Conguillío), single female in Stireman collection.

ALPINOPLAGIA Townsend, 1931d: 475. Type species: Alpinoplagia boliviana Townsend, 1931, by original designation [Bolivia].

References: Townsend (1936b: 232), diagnosis of adults and immatures of Voriini and key to genera (including Alpinoplagia); Townsend (1939a: 373), redescription; Cortés and Campos (1971: 23, 1974: 114) and Cortés (1984: 380), in keys to tachinid genera of Tarapacá and Antofagasta regions; Cortés and González (1989: 116), in key to genera of Chilean Voriini.

boliviana Townsend, 1931.—Neotropical: South America (Bolivia, Chile).

Alpinoplagia boliviana Townsend, 1931d: 476. Holotype female (NHMW). Type locality: Bolivia, La Paz, Cerro Sillutincara [as “Cuesta de Cillutincara”, ca. 16°17′S, 67°53′W], 11,000 ft.

Reference: Cortés and Campos (1971: 46), first record from Chile, redescription, head figure.

References: Cortés and Valencia (1972: 66), key to the two subgenera of Ateloglutus and the three species of new subgenus Proteloglutus; Cortés (1984: 378), in key to tachinid genera of Tarapacá and Antofagasta regions; Cortés (1986: 143), in key to tachinid genera of Aysén and Magallanes regions; González (1989), review, key to Chilean species; Cortés and González (1989: 116), in key to genera of Chilean Voriini.

CHAETODEMOTICUS Brauer & Bergenstamm, 1891: 385 [also 1891: 81]. Type species: Demoticus chilensis Schiner, 1868, by monotypy [Chile].

References: Townsend (1936b: 218), diagnosis of adults and immatures of Germariini and key to genera (including Chaetodemoticus); Townsend (1939a: 319), redescription; Cortés (1967b: 12), key to separate Chaetodemoticus and Actinoplagia Blanchard; Cortés and Campos (1971: 25, 1974: 115) and Cortés (1984: 380), in keys to tachinid genera of Tarapacá and Antofagasta regions.

chilensis (Schiner, 1868).—Neotropical: South America (Chile).

Demoticus chilensis Schiner, 1868: 324. Holotype male [not female as published, Aldrich 1927b: 5] (NHMW). Type locality: Chile.

References: Aldrich (1927b: 5), redescription of holotype; Cortés (1945c: 24), redescription; Cortés and Campos (1971: 56, 61), notes, head figure.

CHILOCLISTA Townsend, 1931c: 334. Type species: Chiloclista bicolor Townsend, 1931, by original designation [Chile].

References: Townsend (1936c: 13), diagnosis of adults and immatures of Macquartiini and key to genera (including Chiloclista); Townsend (1939c: 30), redescription.

bicolor Townsend, 1931.—Neotropical: South America (Chile).

Chiloclista bicolor Townsend, 1931c: 334. Holotype male (USNM). Type locality: Chile, O’Higgins, Cardenal Caro, Tanumé [ca. 34°13′S, 71°55′W].

Reference: Stireman et al. (2016: 37), habitus images.

CORACOMYIA Aldrich, 1934: 21. Type species: Coracomyia crassicornis Aldrich, 1934, by original designation [Argentina].

References: Townsend (1936c: 129), diagnosis of adults and immatures of Actiini and key to genera (including Coracomyia); Townsend (1940a: 206), redescription; Cortés (1986: 143), in key to tachinid genera of Aysén and Magallanes regions; Cortés and González (1989: 116), in key to genera of Chilean Voriini.

crassicornis Aldrich, 1934.—Neotropical: South America (Argentina, Chile).

Coracomyia crassicornis Aldrich, 1934: 22. Holotype male (NHMUK). Type locality: Argentina, Río Negro, Lago Nahuel Huapí.

Reference: Cortés (1976: 8), partial redescription including first description of female, first record from Chile.

woodi Cortés, 1976.—Neotropical: South America (Chile).

Coracomyia woodi Cortés, 1976: 8. Holotype male (MEUC). Type locality: Chile, Los Lagos, Osorno, Parque Nacional Puyehue, Paso Cardenal Antonio Samoré [as “Paso Puyehue”], 1200 m [ca. 40°42′S, 71°57′W].

CYRTOPHLOEBA Rondani, 1856: 207. Type species: Tachina ruricola Meigen, 1824, by original designation [Europe].

EUCYRTOPHLOEBA Townsend, 1916e: 316. Type species: Eucyrtophloeba rhois Townsend, 1916, by original designation [Mexico].

OPSOPHAGUS Aldrich, 1926a: 15. Type species: Opsophagus ornatus Aldrich, 1926, by original designation [Peru].

CYRTHOPHLEBA. Incorrect subsequent spelling of Cyrtophloeba Rondani, 1856 (Rondani 1857: 13) (see O’Hara et al. 2011: 68).

CYRTOPHLEBA. Incorrect original spelling of Cyrtophloeba Rondani, 1856 (Rondani 1856: 68) (see O’Hara et al. 2011: 69).

References: Coquillett (1910: 530), type species of Cyrtophloeba (as “Cyrtophleba”); Aldrich (1934: 3, 32), in key to Patagonian genera, taxonomic notes (as Opsophagus); Townsend (1936b: 232), diagnosis of adults and immatures of Voriini and key to genera (including Cyrtophloeba [as “Cyrtophleba”], Eucyrtophloeba and Opsophagus); Townsend (1939a: 378, 379, 393), redescriptions of Cyrtophloeba (as “Cyrtophleba”), Eucyrtophloeba and Opsophagus; Caltagirone (1966: 63), key to Neotropical species (as Opsophagus); Cortés (1986: 143), in key to tachinid genera of Aysén and Magallanes regions (as Opsophagus); Cortés and González (1989: 116), in key to genera of Chilean Voriini (as Opsophagus); Wood and Zumbado (2010: 1402), synonymy of Opsophagus with Cyrtophloeba (as “Cyrtophleba”).

cortesi (Caltagirone, 1966).—Neotropical: South America (Argentina, Chile).

Opsophagus cortesi Caltagirone, 1966: 64. Holotype male (INLA). Type locality: Chile, Maule, Talca, Gualleco.

References: Cortés (1967b: 12), taxonomic notes; Gramajo (1998: 95), first record from Argentina.

nigripalpis (Aldrich, 1926).—Neotropical: South America (Argentina, Chile, Ecuador).

Opsophagus nigripalpis Aldrich, 1926a: 16. Holotype male (USNM). Type locality: Chile, Valparaíso, Marga Marga, Bosque Los Perales [as “Perales”, ca. 33°9′S, 71°18′W].

References: Aldrich (1934: 33), redescription, taxonomic notes, first record from Argentina; Cortés (1979: 80), taxonomic notes; Cortés (1980: 106), first record from Ecuador.

DISCHOTRICHIA Cortés, 1944f: 54. Type species: Dischotrichia caelibata Cortés, 1944, by original designation [Chile].

Reference: Cortés (1975: 36), in key to related genera.

caelibata Cortés, 1944.—Neotropical: South America (Chile).

Dischotrichia caelibata Cortés, 1944f: 56. Holotype male (USNM). Type locality: Chile, Valparaíso, Marga Marga.

Reference: Campos (1953: 25), first description of female.

GANOPLEURON Aldrich, 1934: 118. Type species: Ganopleuron divergens Aldrich, 1934, by original designation [Chile].

divergens Aldrich, 1934.—Neotropical: South America (Argentina, Chile).

Ganopleuron divergens Aldrich, 1934: 119. Holotype female (NHMUK). Type locality: Chile, Los Lagos, Chiloé, Castro.

References: Townsend (1936c: 13), diagnosis of adults and immatures of Macquartiini and key to genera (including Ganopleuron); Townsend (1939c: 39), redescription; Cortés (1963: 243), notes on holotype in NHMUK; Cortés (1973a: 98), first description of male; Gramajo (1998: 94), first record from Argentina.

LAFUENTEMYIA Marnef, 1965: 243. Type species: Lafuentemyia yanezi Marnef, 1965, by original designation [Chile].

Reference: Cortés (1975: 36), in key to genera with a modified hind femur in male.

yanezi Marnef, 1965.—Neotropical: South America (Chile).

Lafuentemyia yanezi Marnef, 1965: 246. Holotype male (UVVC). Type locality: Chile, Valparaíso, Valparaíso, Reserva Nacional Lago Peñuelas.

Reference: Cortés (1975: 36), taxonomic notes.

MYIOCHAETA Cortés, 1967b: 24. Type species: Myiochaeta marnefi Cortés, 1967, by original designation [Chile].

Reference: Cortés and González (1989: 116), in key to genera of Chilean Voriini.

marnefi Cortés, 1967.—Neotropical: South America (Chile).

Myiochaeta marnefi Cortés, 1967b: 25. Holotype male (EEAM). Type locality: Chile, Metropolitana de Santiago, Santiago, Maipú, Rinconada, Quebrada de La Plata, 510 m.

NEOCHAETOPLAGIA Blanchard, 1963: 173. Type species: Neochaetoplagia pastranai Blanchard, 1963, by original designation [Argentina].

Reference: Cortés and González (1989: 116), in key to genera of Chilean Voriini.

pastranai Blanchard, 1963.—Neotropical: South America (Argentina, Chile).

Neochaetoplagia pastranai Blanchard, 1963: 173. Holotype male (presumed lost, Mulieri et al. 2013: 168). Type locality: Argentina, Buenos Aires, Buenos Aires [as “Capital Federal”].

References: Cortés and González (1989: 118), first record from Chile; Mulieri et al. (2013: 168), notes on type series.

NOTHOVORIA Cortés & González, 1989: 120. Type species: Nothovoria praestans Cortés & González, 1989, by original designation [Chile].

Reference: Cortés and González (1989: 116), in key to genera of Chilean Voriini.

praestans Cortés & González, 1989.—Neotropical: South America (Chile).

Nothovoria praestans Cortés & González, 1989: 120. Holotype female (UMCE). Type locality: Chile, Tarapacá, Iquique, 40 km from Iquique, Pampa del Tamarugal, Estación Refresco, 1200 m.

PHAEODEMA Aldrich, 1934: 145. Type species: Phaeodema mystacina Aldrich, 1934, by original designation [Chile].

References: Townsend (1936c: 47), diagnosis of Ebeniini and key to genera (including Phaeodema); Townsend (1939c: 160), redescription; Cortés (1986: 143), in key to tachinid genera of Aysén and Magallanes regions.

mystacina Aldrich, 1934.—Neotropical: South America (Argentina, Chile).

Phaeodema mystacina Aldrich, 1934: 145. Holotype male (NHMUK). Type locality: Chile, Los Lagos, Llanquihue, Puerto Montt.

Reference: Gramajo (1998: 94), first record from Argentina.

PIRIONA Aldrich, 1928b: 24. Type species: Piriona fasciculata Aldrich, 1928, by original designation [Chile].

References: Aldrich (1934: 3, 44), in key to Patagonian genera, taxonomic notes; Townsend (1936c: 13), diagnosis of adults and immatures of Macquartiini and key to genera (including Piriona); Townsend (1939c: 58), redescription; Cortés (1975: 36), in key to genera with a modified hind femur in male.

fasciculata Aldrich, 1928.—Neotropical: South America (Argentina, Chile).

Piriona fasciculata Aldrich, 1928b: 24. Holotype male (USNM). Type locality: Chile, Valparaíso, Marga Marga.

Note: Aldrich (1928b: 25) described Piriona fasciculata from five males and one female collected from three localities in Argentina and Chile. Aldrich cited a male “Type” in USNM with “Cat. No. 41385” but did not give the type locality. The holotype in USNM was examined by DMW and is from Marga Marga Province in Chile (see also Aldrich 1934: 45).

References: Aldrich (1934: 45), redescription; Cortés (1963: 243), notes on specimens in NHMUK; Cortés (1975: 36), taxonomic notes.

PROSOPOCHAETA Macquart, 1851: 183 [also 1851: 210] (as “Prosopochoeta”, see note). Type species: Prosopochaeta nitidiventris Macquart, 1851, by original designation [Chile].

PUNACLISTA Townsend, 1915e: 406. Type species: Punaclista setosa Townsend, 1915, by original designation [Peru].

PROSOPOCHOETA. Incorrect original spelling of Prosopochaeta Macquart, 1851 (Macquart 1851: 183, see note).

Notes: The name Prosopochaeta Macquart, 1851 was originally published as Prosopochoeta but subsequent authors (e.g., Aldrich 1934; Cortés and Hichins 1969; Guimarães 1971; González 1992b) used the spelling Prosopochaeta. This changed spelling would normally be considered an incorrect subsequent spelling but because it is in prevailing usage and is attributed to Macquart (1851), it is deemed to be the correct original spelling (Article 33.3.1 of the Code, ICZN 1999).

Macquart (1851: 184 [also 1851: 211]) noted about his new genus Prosopochaeta (as Prosopochoeta), “Le type de ce genre est du Chili” [“The type of this genus is from Chile”]. This statement is accepted as a type species designation for Prosopochaeta of the single included species, Prosopochaeta nitidiventris Macquart, from Chile.

References: Aldrich (1934: 5, 115), in key to Patagonian genera, synonymy of Punaclista with Prosopochaeta, redescription; Townsend (1936b: 121), diagnosis of Trichoprosopini and key to genera (including Prosopochaeta); Townsend (1936c: 13), diagnosis of adults and immatures of Macquartiini and key to genera (including Punaclista); Townsend (1938: 299), redescription of Prosopochaeta; Townsend (1939c: 62), redescription of Punaclista; Parker (1953: 66), figures of puparium of Prosopochaeta sp.; Cortés and Campos (1971: 45), taxonomic notes including reinstating synonymy of Punaclista with Prosopochaeta; Cortés and Campos (1971: 26, 1974: 116) and Cortés (1984: 381), in keys to tachinid genera of Tarapacá and Antofagasta regions; Cortés (1986: 142), in key to tachinid genera of Aysén and Magallanes regions.

anomala Aldrich, 1934.—Neotropical: South America (Argentina, Chile).

Prosopochaeta anomala Aldrich, 1934: 118. Holotype male (USNM). Type locality: Argentina, Río Negro, eastern end of Lago Nahuel Huapí, Jones Estancia.

Note: Prosopochaeta anomala was recorded from both Argentina and Chile in the original description.

caliginosa Cortés & Campos, 1971.—Neotropical: South America (Argentina, Chile).

Prosopochaeta caliginosa Cortés & Campos, 1971: 43. Holotype male (EEAM). Type locality: Chile, Antofagasta, Antofagasta, north of Quebrada de Paposo, 200–1000 m (25°03′S, 70°25′W) (coordinates and elevation given on p. 12).

Reference: Cortés (1979: 81), first record from Argentina.

nitidiventris Macquart, 1851.—Neotropical: South America (Argentina, Chile).

Prosopochoeta nitidiventris Macquart, 1851: 184 [also 1851: 211]. Lectotype male (MNHN), by designation herein (see Lectotype Designations section). Type locality: Chile (“Coquimbo, etc.” according to Blanchard 1854: 424).

Notes: Punaclista setosa Townsend, 1915 from Peru was treated as a synonym of Prosopochoeta nitidiventris Macquart by Aldrich (1934: 116) but is currently recognised as a valid species of Prosopochaeta (e.g., Cortés and Campos 1971: 46; Guimarães 1971: 100).

References: Aldrich (1934: 116), redescription, head figure, first record from Argentina; Verbeke (1962: 103), description of male terminalia.

TRICHODISCHIA Bigot, 1885a: 237. Nomen nudum.

TRICHODISCHIA Bigot, 1885b: xlv [also 1885b: xlv, Bull. Soc. Ent. France]. Type species: Trichodischia soror Bigot, 1885, by subsequent designation of Townsend (1916b: 9) [Argentina].

TRICHORAEA Cortés, 1975: 37. Type species: Trichodischia caerulea Bigot, 1885, by original designation [Argentina].

TRICODISCHIA. Incorrect subsequent spelling of Trichodischia Bigot, 1885 (Henry 1987: 194).

Note: The two Bigot species Trichodischia caerulea and T. soror were treated as generically different by Cortés (1975: 37) and the former was assigned to new genus Trichoraea Cortés. Subsequent authors have continued to treat the original combination Trichodischia caerulea as valid (e.g., Guimarães 1977c: 76; Henry 1987: 195; Molina-Ochoa et al. 2003: 269) and this classification is followed here, but without further study.

References: Townsend (1936c: 13), diagnosis of adults and immatures of Macquartiini and key to genera (including Trichodischia); Townsend (1939c: 68), redescription of Trichodischia; Parker (1953: 66), figures of first instar larva and puparium of Trichodischia sp., from Argentina; Cortés (1969: 98), key to separate the two species; Cortés (1975: 36), Trichodischia and Trichoraea in key to genera with a modified hind femur in male.

caerulea Bigot, 1885.—Neotropical: South America (Argentina, Chile, Uruguay).

Trichodischia caerulea Bigot, 1885b: xlv [also 1885b: xlv, Bull. Soc. Ent. France]. Syntypes, 3 specimens as “♀?” (2 males and 1 female in NHMUK, examined by DMW). Type locality: Argentina, Buenos Aires, Buenos Aires.

caerulai. Incorrect subsequent spelling of caerulea Bigot, 1885 (Blanchard 1963: 184).

caerulia. Incorrect subsequent spelling of caerulea Bigot, 1885 (Blanchard 1963: 185).

coerulea. Incorrect subsequent spelling of caerulea Bigot, 1885 (Brauer 1899: 498).

References: Cortés (1969: 98), separation from Trichodischia soror, first record from Chile; Cortés (1975: 36, 37), first record from Uruguay (distribution given for Trichoraea in key, a monotypic genus based on T. caerulea), taxonomic notes.

soror Bigot, 1885.—Neotropical: South America (Argentina, Brazil, Chile, Uruguay).

Trichodischia soror Bigot, 1885b: xlvi [also 1885b: xlvi, Bull. Soc. Ent. France]. Holotype male (NHMUK). Type locality: Argentina, Buenos Aires, Buenos Aires.

Trichodischia caerulea of Cortés (1944f: 51, 1946: 175) and Blanchard (1963: 184), not Bigot, 1885. Misidentification (Cortés 1969: 97, 1975: 37).

References: Cortés (1944f: 54), first records from Chile and Uruguay, misidentified as Trichodischia caerulea; Cortés (1969: 98), separation from T. caerulea; Cortés (1975: 37), taxonomic notes; Cortés (1980: 105), first record from Brazil.

VELARDEMYIA Valencia, 1972a: 364. Type species: Velardemyia ica Valencia, 1972, by original designation [Peru].

Reference: Cortés and González (1989: 116), in key to genera of Chilean Voriini.

ica Valencia, 1972.—Neotropical: South America (Chile, Peru).

Velardemyia ica Valencia, 1972a: 364. Holotype male (SENASA, Lozada et al. 2005: 460). Type locality: Peru, Ica, Arrabales.

Reference: Cortés and González (1989: 122), first record from Chile.

VORIA Robineau-Desvoidy, 1830: 195. Type species: Voria latifrons Robineau-Desvoidy, 1830 (= Tachina ruralis Fallén, 1810), monotypy [France].

PLAGIA Meigen, 1838: 201. Type species: Tachina verticalis Meigen, 1824 (= Tachina ruralis Fallén, 1810), by subsequent designation of Rondani (1856: 69) [Europe].

XENOPLAGIA Townsend, 1914a: 13. Type species: Xenoplagia setosa Townsend, 1914, by original designation [Peru].

ITAVORIA Townsend, 1931d: 474. Type species: Itavoria aurescens Townsend, 1931, by original designation [Brazil].

References: Coquillett (1910: 591, 619), type species of Plagia and Voria (with Plagia [and others] in synonymy with Voria); Townsend (1936b: 232), diagnosis of adults and immatures of Voriini and key to genera (including Itavoria, Voria and Xenoplagia); Townsend (1936c: 280), Plagia as synonym of Voria; Townsend (1939a: 385, 402, 403), redescriptions of Itavoria, Voria (with Plagia in synonymy) and Xenoplagia; Mesnil (1974: 1261), redescription (with Plagia in synonymy), taxonomic notes; Cortés and Campos (1974: 113) and Cortés (1984: 379), Voria in keys to tachinid genera of Tarapacá and Antofagasta regions; Cortés and González (1989: 116), Voria in key to genera of Chilean Voriini; Fleming et al. (2017: 7), synonymy of Itavoria and Xenoplagia with Voria [also the synonymy with Voria of the Afrotropical and Oriental genus Hystricovoria Townsend, 1928 and its synonyms Afrovoria Curran, 1938 and Anavoria Mesnil, 1953, but the present authors follow O’Hara and Cerretti (2016: 56) in recognising Hystricovoria as a valid genus].

ruralis (Fallén, 1810).—Neotropical: southern Lesser Antilles (Trinidad & Tobago), Middle America (Mexico, Nicaragua), South America (Argentina, Brazil, Chile, Colombia, Peru, Uruguay, Venezuela). Nearctic: Canada, United States. Palaearctic: Central Asia, China [Pal.], Europe, Japan, Korean Peninsula, Middle East, Mongolia, Russia, Transcaucasia. Afrotropical: Kenya to South Africa, Yemen. Oriental: China (Yunnan), India, Nepal, Pakistan, Taiwan. Australasian & Oceanian: Australia, Papua New Guinea.

Tachina ruralis Fallén, 1810: 265. Lectotype male (NHRS), by designation of Crosskey (1973: 163). Type locality: Sweden, Skåne, Äsperöd [as “Esperöd”].

Plagia americana van der Wulp, 1890c: 102. Syntypes, males and females (NHMUK). Type localities: Mexico, Veracruz (Orizaba), Guerrero (Venta del Zopilote [ca. 17°46′N, 99°32′W], 2800 ft; Xocomanatlán [as “Xucumanatlan”, ca. 17°34′N, 99°37′W], 7000 ft; Omiltemi [as “Omilteme”, ca. 17°33′N, 99°41′W], 8000 ft), and Tabasco (Teapa).

Plagia mexicana Giglio-Tos, 1893: 5. Type(s), female (MZUT). Type locality: Mexico.

Voria brasiliana Townsend, 1929: 380. Syntypes, “many males and females” (USNM). Type locality: Brazil, São Paulo, Itaquaquecetuba.

Voria ayerzai Blanchard, 1937: 47 (as “Voria ayerzai, (Brethes)”). Nomen nudum.

Voria ayerzai Blanchard, 1943c: 157 (as “Plagia ayerzai, Brèthes in lit.”). Syntypes, 3 males and females (MLPA). Type locality: Argentina, Buenos Aires [province or city].

Notes: The mention of a “Ht” for Tachina ruralis from Sweden in NHRS by Townsend (1939a: 402) is not accepted as a lectotype fixation because the specimen in question is not distinguishable from the other specimens in the type series.

Voria ruralis as here interpreted is almost certainly a species complex (see also Fleming et al. 2017). This complex in the New World may not include the true Voria ruralis described from the Palaearctic Region by Fallén (1810). Some of the names listed here in synonymy may represent distinct species.

References: Aldrich (1926a: 14), synonymy of Plagia americana with Tachina ruralis; Cortés (1944d: 142), Voria brasiliana and Voria ayerzai as possible synonyms of Tachina ruralis, first record from Chile; Parker (1953: 64), figures of first instar larva and puparium; Thompson (1961: 28), synonymy including Plagia mexicana as a questionable synonym of Tachina ruralis, redescription, first record from Trinidad; Blanchard (1963: 169), taxonomic notes, record from Argentina; Guimarães (1971: 93), synonymy; Nihei (2016: 911), first record from Colombia.

SCHLINGERMYIA Cortés, 1967b: 20. Type species: Schlingermyia venusta Cortés, 1967, by original designation [Chile].

Note: This genus was not placed beyond subfamily Dexiinae by the original author, Cortés (1967: 20). Guimarães (1971: 100) listed Schlingermyia under subfamily Dexiinae, tribe Macquartiini, following Townsend’s (1936c: 13) concept of the tribe. Most of the Neotropical genera then assigned to the Macquartiini are currently placed in the dexiine tribes Dexiini and Voriini.

venusta Cortés, 1967.—Neotropical: South America (Chile).

Schlingermyia venusta Cortés, 1967b: 22. Holotype male (EEAM). Type locality: Chile, Valparaíso, Marga Marga, Bosque Los Perales [as “Los Perales”, ca. 33°9′S, 71°18′W].

GRAVENHORSTIA Robineau-Desvoidy, 1863a: 924 (junior homonym of Gravenhorstia Boie, 1836). Type species: Gravenhorstia longicornis Robineau-Desvoidy, 1863 (= Tachina grandicornis Zetterstedt, 1849), by original designation [France].

ADMONTIA Brauer & Bergenstamm, 1889: 104 [also 1890: 36]. Type species: Admontia podomyia Brauer & Bergenstamm, 1889, by monotypy [Austria, Germany, Italy, Poland, Germany and Czech Republic].

TRICHOPAREIA Brauer & Bergenstamm, 1889: 103 [also 1890: 35]. Type species: Tachina seria Meigen, 1824, by monotypy [Germany].

AUSTROSTAUROCHAETA Townsend, 1931d: 476. Type species: Degeeria antarctica Thomson, 1869, by original designation [probably Chile].

POLIOPS Aldrich, 1934: 94. Type species: Poliops striatus Aldrich, 1934, by original designation [Argentina]. Syn. nov.

Notes: The relative priority of Admontia Brauer & Bergenstamm, 1889 and Trichopareia Brauer & Bergenstamm, 1889, when the two are treated as synonyms, was established by Strobl (1910: 137), as the First Reviser (Article 24.2.2 of the Code, ICZN 1999).

The new synonymy of Poliops with Admontia is explained below under Admontia striata.

References: Coquillett (1910: 503), type species of Admontia (as synonym of Hyperecteina Schiner, 1861); Aldrich (1934: 4, 94, 95), Admontia and Poliops in key to Patagonian genera, synonymy including Trichopareia and Austrostaurochaeta with Admontia, taxonomic notes, key to six Patagonian species of Admontia; Townsend (1936c: 129, 154), diagnosis of adults and immatures of Actiini and key to genera (including Austrostaurochaeta and Poliops), diagnosis of adults and immatures of Trichopareiini and key to genera (including Admontia and Trichopareia); Townsend (1940a: 193, 254, 301, 304), redescriptions of Admontia, Austrostaurochaeta, Poliops and Trichopareia; Cortés (1973a: 100), separation of Admontia and Austrostaurochaeta from Notomanes Aldrich; Wood (1985: 14, 17), in key to the Blondeliini of North and Central America and the West Indies, synonymy including Austrostaurochaeta with Admontia, diagnosis, taxonomic notes; Cortés (1986: 144), in key to tachinid genera of Aysén and Magallanes regions.

antarctica (Thomson, 1869).—Neotropical: South America (Argentina, Chile).

Degeeria antarctica Thomson, 1869: 527. Lectotype male (NHRS), by fixation of Townsend (1931b: 183) (examination of “Male Ht” from Patagonia in NHRS is regarded as a lectotype fixation). Type locality: “Patagonia” (most likely Chile, Magallanes y de la Antártica Chilena, Magallanes, Puerto del Hambre [frequently as “Port Famine”], based on localities where insects were collected during the voyage of the Swedish frigate Eugenie, including the lectotype of Degeeria antarctica; see Persson 1971: 168).

References: Aldrich (1934: 97), redescription, first record from Argentina; Cortés (1973a: 97), taxonomic notes.

aurata (Campos, 1953).—Neotropical: South America (Chile). Comb. nov.

Poliops auratus Campos, 1953: 27. Holotype male (MNNC). Type locality: Chile, Biobío, Concepción, Tomé.

Note: Poliops auratus is assumed to have the same generic features as the type species of Poliops, P. striatus, and for this reason is transferred to Admontia. The transfer of P. striatus to Admontia is discussed below under A. striata.

calyptrata (Aldrich, 1934).—Neotropical: South America (Argentina, Chile). Comb. nov. (Fig. 4d)

Phorocera calyptrata Aldrich, 1934: 73. Holotype male (USNM). Type locality: Argentina, Río Negro, Lago Correntoso.

Note: Phorocera calyptrata Aldrich has the Admontia features of a setose facial ridge, haired parafacial and tiny fore claws in the female but the eye has scattered long hairs rather than the usual bare condition. We interpret the species as an aberrant member of the Admontia lineage and move it here to Admontia from its prior indefinite placements in Tachinidae (see References below).

References: Aldrich (1934: 69), in key to Patagonian species of Phorocera Robineau-Desvoidy, 1830 (s. lato); Cortés (1945d: 158), in key to Chilean species of Phorocera Robineau-Desvoidy, 1830 (s. lato) and Parasetigena Brauer & Bergenstamm, 1891, known from Argentina and not Chile; Guimarães (1971: 152), as unplaced species of Blondeliini; Henry (1987: 206), first record from Chile (in Phorocera but genus unplaced in Tachinidae); González (1992b: 183), record from Chile, as Phorocera calyptrata (sensu previous authors).

communis Aldrich, 1934.—Neotropical: South America (Argentina, Chile).

Admontia communis Aldrich, 1934: 99. Holotype, unspecified sex (NHMUK). Type locality: Argentina, Río Negro, eastern end of Lago Nahuel Huapí.

Admontia communis albescens Aldrich, 1934: 100. Syntypes, 5 males (NHMUK). Type localities: Argentina, Río Negro, eastern end of Lago Nahuel Huapí and San Carlos de Bariloche [as “Bariloche”].

Notes: Admontia communis was recorded from both Argentina and Chile in the original description. Cortés and Hichins (1969: 16) gave the depository for the holotype of A. communis as USNM, in error.

debilis Aldrich, 1934.—Neotropical: South America (Argentina, Chile).

Admontia debilis Aldrich, 1934: 102. Holotype male (NHMUK). Type locality: Chile, Los Lagos, Llanquihue, Casa Pangue.

finisterrae Cortés, 1986.—Neotropical: South America (Chile).

Admontia finisterrae Cortés, 1986: 155. Holotype male (MEUC). Type locality: Chile, Magallanes y de la Antártica Chilena, Antártica Chilena, Islas Hermite, Isla Deceit, Caleta Toledo.

flavibasis Aldrich, 1934.—Neotropical: South America (Argentina, Chile).

Admontia flavibasis Aldrich, 1934: 103. Holotype female (USNM). Type locality: Argentina, Río Negro, Lago Gutiérrez.

Reference: González (1992b: 178), first record from Chile.

pictiventris Aldrich, 1934.—Neotropical: South America (Argentina, Chile).

Admontia pictiventris Aldrich, 1934: 100. Holotype male (NHMUK). Type locality: Chile, Los Lagos, Llanquihue, Peulla.

Reference: Gramajo (1998: 96), first record from Argentina.

striata (Aldrich, 1934).—Neotropical: South America (Argentina, Chile). Comb. nov.

Poliops striatus Aldrich, 1934: 94. Holotype male (NHMUK). Type locality: Argentina, Río Negro, eastern end of Lago Nahuel Huapí.

Notes: Poliops striatus was recorded from both Argentina and Chile in the original description.

Aldrich (1934: 94) described Poliops as a monotypic genus based on the new species P. striatus, noting: “Very similar to Admontia, but costal spine about as long as distance between auxiliary and first vein on costa, and third vein at base with a single setula of about the same length”. A male homotype of P. striatus in CNC that DMW compared to the holotype in NHMUK possesses the usual features of the Blondeliini and diagnostic characters of Admontia (Wood 1985: 18), including bare eye, setose facial ridge, and a few hairs on the upper parafacial below lowest frontal seta. A single setula at the base of wing vein R₄₊₅ is not unique in Admontia to the two species originally described in Poliops and was even given in the original description as a characteristic of A. finisterrae Cortés, 1986.

EUCELATORIA Townsend, 1909: 249. Type species: Tachina armigera Coquillett, 1889, by monotypy [United States].

SPATHIMYIA Townsend, 1912b: 318. Type species: Spathimyia ferox Townsend, 1912, by original designation [Peru].

XIPHOMYIA Townsend, 1917: 125. Type species: Xiphomyia gladiatrix Townsend, 1917, by original designation [Panama].

URODEXODES Townsend, 1919b: 572. Type species: Urodexodes charapensis Townsend, 1919, by original designation [Peru].

MACHAIROMASICERA Townsend, 1919b: 577. Type species: Machairomasicera carinata Townsend, 1919, by original designation [Ecuador].

LIXINIA Curran, 1926: 108. Type species: Lixinia jamaicensis Curran, 1926, by original designation [Jamaica].

TINALYDELLA Townsend, 1927a: 265. Type species: Tinalydella tinensis Townsend, 1927, by original designation [Peru].

OROPHOROCERA Townsend, 1927a: 267. Type species: Orophorocera ocellaris Townsend, 1927, by original designation [Peru].

HYPOMYOTHYRIA Townsend, 1927a: 276. Type species: Hypomyothyria hypodermica Townsend, 1927, by original designation [Brazil].

EUCELATORIOPSIS Townsend, 1927a: 276. Type species: Eucelatoriopsis teffeensis Townsend, 1927, by original designation [Brazil].

HELIOLYDELLA Townsend, 1927a: 277. Type species: Heliolydella aurata Townsend, 1927, by original designation [Brazil].

TACHINOPHYTOPSIS Townsend, 1927a: 277. Type species: Tachinophytopsis carinata Townsend, 1927 (junior secondary homonym of Machairomasicera carinata Townsend, 1919; = Eucelatoria paracarinata Nihei & Dios, 2016), by original designation [Brazil].

HEMILYDELLA Townsend, 1927a: 278. Type species: Hemilydella fasciata Townsend, 1927, by original designation [Peru].

LYDELLOHOUGHIA Townsend, 1927a: 280. Type species: Lydellohoughia nana Townsend, 1927, by original designation [Brazil].

EUPTILODEGEERIA Townsend, 1931d: 465. Type species: Hypostena obumbrata van der Wulp, 1890, by original designation [Mexico].

COROZALIA Curran, 1934: 465. Type species: Corozalia longula Curran, 1934, by original designation [Panama].

CELATORIOPSIS Blanchard, 1963: 228. Type species: Celatoriopsis eucelatorioides Blanchard, 1963, by original designation [Argentina].

EUCELATORIOIDEA Thompson, 1968: 176. Type species: Eucelatorioidea nigripalpis Thompson, 1968 (junior secondary homonym of Chetolyga nigripalpis Bigot, 1889; = Eucelatoria nudioculata O’Hara & Wood, nom. nov., see below), by original designation [Trinidad & Tobago].

DEXODIMYIA Thompson, 1968: 181. Type species: Dexodimyia discalis Thompson, 1968, by original designation [Trinidad & Tobago].

PSEUDOCELATORIA Thompson, 1968: 190. Type species: Pseudocelatoria robusta Thompson, 1968, by original designation [Trinidad & Tobago].

HELIODEXODES Thompson, 1968: 197. Type species: Heliodexodes argenteus Thompson, 1968, by original designation [Trinidad & Tobago].

DEXODIOPSIS Thompson, 1968: 202. Type species: Dexodiopsis aurea Thompson, 1968, by original designation [Trinidad & Tobago].

Notes: The relative priority of Eucelatorioidea Thompson, 1968, Dexodimyia Thompson, 1968, Pseudocelatoria Thompson, 1968, Heliodexodes Thompson, 1968 and Dexodiopsis Thompson, 1968, when the five are treated as synonyms, has not been established and is not of concern while all are junior synonyms of Eucelatoria Townsend, 1909 (as proposed by Wood 1985: 40).

References: Townsend (1936c: 86, 237), diagnosis of adults and immatures of Compsilurini and key to genera (including Eucelatoria, Eucelatoriopsis, Euptilodegeeria, Heliolydella, Hemilydella, Hypomyothyria, Machairomasicera, Orophorocera, Spathimyia, Tachinophytopsis, Tinalydella and Urodexodes), diagnosis of adults and immatures of Trypherini and key to genera (including Corozalia, Lixinia, Lydellohoughia and Xiphomyia); Townsend (1940a: 48, 50, 53, 54, 55, 56, 63, 76, 94, 97, 98, 100), redescriptions of Eucelatoria, Eucelatoriopsis, Euptilodegeeria, Heliolydella, Hemilydella, Hypomyothyria, Machairomasicera, Orophorocera, Spathimyia, Tachinophytopsis, Tinalydella and Urodexodes; Townsend (1941: 258, 279, 281, 327), redescriptions of Corozalia, Lixinia, Lydellohoughia and Xiphomyia; Thompson (1968: 174, 176), revision of Eucelatoria species of Trinidad, as nine genera (five new) collectively termed the “Trinidad compsilurines”; Cortés and Campos (1971: 26, 1974: 115, 116), Hemilydella and Eucelatoria in keys to tachinid genera of Tarapacá and Antofagasta regions; Sabrosky (1981: 3), in key to genera of Blondeliini in which females possess an abdominal keel and sharp, curved piercer (key also including the following generic names later synonymised with Eucelatoria: Eucelatoriopsis, Heliodexodes, Heliolydella, Hemilydella, Lydellohoughia, Machairomasicera, Spathimyia, Tinalydella, Urodexodes and Xiphomyia), synonymy of Celatoriopsis with Eucelatoria; Cortés (1984: 381, 382), Eucelatoria, Hemilydella and Urodexodes in key to tachinid genera of Tarapacá and Antofagasta regions; Wood (1985: 13, 40), in key to the Blondeliini of North and Central America and the West Indies, new synonymy of all generic names listed above with Eucelatoria (with the exception of the previously synonymised Celatoriopsis), diagnosis, taxonomic notes.

australis Townsend, 1911.—Neotropical: eastern Lesser Antilles (Saint Vincent), southern Lesser Antilles (Trinidad & Tobago), South America (Brazil, Chile, Peru).

Eucelatoria australis Townsend, 1911: 140, based on female reproductive system [1912b: 315, adult description]. Lectotype female (USNM), by fixation of Townsend (1912b: 316) (description of female “Type” [dissection TD 4025] from Piura in USNM is regarded as a lectotype fixation). Type locality: Peru, Piura, Piura.

Note: Aldrich (1927a: 19) synonymised Compsilura oppugnator Walton, 1914 from Puerto Rico with Eucelatoria australis Townsend and this synonymy was followed by Guimarães (1971: 133). Sabrosky (1981) recognised Eucelatoria oppugnator as valid and Wood (1985: 44) did also but with the note: “[? = australis (c/f Aldrich 1927a: 19)]”.

References: Aldrich (1927a: 19), first record from St. Vincent; Sauer (1946: 21), first record from Brazil; Thompson (1968: 200), redescription, first record from Trinidad; Cortés and Campos (1971: 81), first record from Chile; Cortés (1984: 386), taxonomic notes; Vergara de Sánchez (1987: 10), redescription, figures of male and female terminalia.

digitata Sabrosky, 1981.—Neotropical: South America (Chile, Peru).

Eucelatoria digitata Sabrosky, 1981: 11. Holotype male (USNM). Type locality: Peru, Lima, San Diego.

Note: Eucelatoria digitata was recorded from both Chile and Peru in the original description.

References: Cortés (1984: 386), taxonomic notes; Vergara de Sánchez (1987: 10, 11), redescription, figures of male and female terminalia.

fasciata (Townsend, 1927).—Neotropical: southern Lesser Antilles (Trinidad & Tobago), South America (Chile, Peru).

Hemilydella fasciata Townsend, 1927a: 315. Holotype male (USNM). Type locality: Peru, Piura, Río Macará, La Tina, on border with Ecuador, 1370 ft.

nudioculata O’Hara & Wood, nom. nov.—Not Chile [Trinidad].

Eucelatorioidea nigripalpis Thompson, 1968: 177 (junior secondary homonym of Chetolyga nigripalpis Bigot, 1889). Holotype female (CNC). Type locality: Trinidad.

Eucelatoria nudioculata O’Hara & Wood, nom. nov. for Eucelatorioidea nigripalpis Thompson, 1968.

Note: Eucelatorioidea nigripalpis Thompson, 1968 from Trinidad, the type species of Eucelatorioidea Thompson, 1968, became a junior secondary homonym of Chetolyga nigripalpis Bigot, 1889 from Mexico when transferred to Eucelatoria Townsend, 1909 by Wood (1985: 44). Both names were treated as valid in that work and the junior homonym was not renamed “pending a revision of the genus” (Wood 1985: 44). This situation has continued to the present and both names are listed as valid in the most recent version of the checklist of world Tachinidae (O’Hara et al. 2020: 225). In the interests of nomenclatural stability, we hereby propose the new name Eucelatoria nudioculata to replace the preoccupied name Eucelatorioidea nigripalpis Thompson. The same type material applies to the new name. The specific epithet nudioculata refers to the bare eye that was noted by Thompson (1968: 176) as a characteristic of his new genus Eucelatorioidea, for which E. nigripalpis was designated type species.

oblonga O’Hara & Wood, nom. nov.—Neotropical: South America (Chile).

Urodexodes elongatum Cortés & Campos, 1974: 124 (junior secondary homonym of Exorista elongata van der Wulp, 1890). Holotype male (MEUC). Type locality: Chile, Arica y Parinacota, Parinacota, Belén, 3500 m.

Eucelatoria oblonga O’Hara & Wood, nom. nov. for Urodexodes elongatum Cortés & Campos, 1974.

Note: Urodexodes elongatum Cortés & Campos, 1974, from Chile, is a junior secondary homonym of Exorista elongata van der Wulp, 1890, the valid name of a Costa Rican species of Eucelatoria (Wood 1985: 43). The two species names are listed as valid in the most recent version of the checklist of world Tachinidae (O’Hara et al. 2020: 224). In the interests of nomenclatural stability, we hereby propose the new name Eucelatoria oblonga to replace the preoccupied name Urodexodes elongatum Cortés & Campos. The same type material applies to the new name. The specific epithet oblonga, Latin for longer than broad, refers to the elongated appearance of the species.

parkeri (Sabrosky, 1952).—Neotropical: South America (Argentina, Brazil, Chile, Uruguay).

Eucelatoriopsis parkeri Sabrosky, 1952: 325. Holotype male (USNM). Type locality: Uruguay, Montevideo, Montevideo.

References: Cortés (1967b: 12), first record from Chile; Guimarães (1977c: 35), first record from Brazil; Boldt et al. (1991: 843), first record from Argentina.