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Catalogue of the Diptera (Insecta) of Morocco— an annotated checklist, with distributions and a bibliography
expand article infoKawtar Kettani, Martin J. Ebejer§, David M. Ackland|, Gerhard Bächli, David Barraclough#, Miroslav Barták¤, Miguel Carles-Tolrá«, Milos Černý», Pierfilippo Cerretti˄, Peter Chandler˅, Mohamed Dakki¦, Christophe Daugeronˀ, Herman De Jongˁ, Josef Dils, Henry Disney, Boris Droz, Neal Evenhuis, Paul Gatt, Gustavo Graciolli, Igor Y. Grichanov, Jean-Paul Haenni, Martin Hauser, Oumnia Himmi¦, Iain MacGowan, Bruno Mathieu, Mohamed Mouna¦, Lorenzo Munari, Emilia P. Nartshuk, Oleg P. Negrobov, Pjotr Oosterbroekˁ, Thomas Pape‡‡, Adrian C. Pont|, Grigory V. Popov§§, Knut Rognes||, Marcela Skuhravᶶ, Vaclav Skuhravý¶¶, Martin Speight##, Guy Tomasovic¤¤, Bouchra Trari, Hans-Peter Tschorsnig««, Jean-Claude Vala»», Michael von Tschirnhaus˄˄, Rüdiger Wagner˅˅, Daniel Whitmore««, Andrzej J. Woźnica¦¦, Tadeusz Zatwarnickiˀˀ, Peter Zwickˁˁ
‡ Abdelmalek Essaadi University, Tetouan, Morocco
§ National Museum and Galleries of Wales, Cardiff, United Kingdom
| Oxford University Museum of Natural History, Oxford, United Kingdom
¶ Zoological Museum Winterthurerstr., Zürich, Switzerland
# University of Kwa Zulu-Natal, Durban, South Africa
¤ Czech University of Life Sciences Prague, Praha-Suchdol, Czech Republic
« Unaffiliated, Barcelona, Spain
» Unaffiliated, Halenkovice, Czech Republic
˄ Università di Roma “La Sapienza”, Rome, Italy
˅ Unaffiliated, Melksham, United Kingdom
¦ Université Mohammed V de Rabat, Rabat, Morocco
ˀ Muséum national d’Histoire naturelle, Paris, France
ˁ Naturalis Biodiversity Center, Leiden, Netherlands
₵ Unaffiliated, Hoevenen, Belgium
ℓ University of Cambridge, Cambridge, United Kingdom
₰ Unaffiliated, La Chaux-de-Fonds, Switzerland
₱ Bishop Museum, Honolulu, United States of America
₳ Unaffiliated, Wickford, United Kingdom
₴ Universidade Federal do Mato Grosso do Sul, Campo Grande, Brazil
₣ Institute of Plant Protection, St. Petersburg-Pushkin, Russia
₮ Muséum d’histoire naturelle, Neuchâtel, Switzerland
₦ California Department of Food & Agriculture, Sacramento, United States of America
₭ National Museums of Scotland, Edinburgh, United Kingdom
₲ Université de Strasbourg, Strasbourg, France
‽ Natural History Museum, Venezia, Italy
₩ Zoological Institute of Russian Academy of Sciences, St. Petersburg, Russia
₸ Voronezh State University, Voronezh, Russia
‡‡ Natural History Museum of Denmark, Copenhagen, Denmark
§§ I.I.Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, Kyiv, Ukraine
|| University of Stavanger, Stavanger, Norway
¶¶ Unaffiliated, Praha, Czech Republic
## Trinity College, Dublin, Ireland
¤¤ Université de Liège, Gembloux, Belgium
«« Staatliches Museum für Naturkunde Stuttgart, Stuttgart, Germany
»» Unaffiliated, Avignon, France
˄˄ Universität Bielefeld, Bielefeld, Germany
˅˅ Limnologische Fluss-Station des Max-Planck Instituts für Limnologie, Schlitz, Germany
¦¦ Wrocław University of Environmental & Life Sciences, Wrocław, Poland
ˀˀ Opole University, Opole, Poland
ˁˁ Unaffiliated, Schlitz, Germany
Open Access

Abstract

The faunistic knowledge of the Diptera of Morocco recorded from 1787 to 2021 is summarized and updated in this first catalogue of Moroccan Diptera species. A total of 3057 species, classified into 948 genera and 93 families (21 Nematocera and 72 Brachycera), are listed. Taxa (superfamily, family, genus and species) have been updated according to current interpretations, based on reviews in the literature, the expertise of authors and contributors, and recently conducted fieldwork. Data to compile this catalogue were primarily gathered from the literature. In total, 1225 references were consulted and some information was also obtained from online databases. Each family was reviewed and the checklist updated by the respective taxon expert(s), including the number of species that can be expected for that family in Morocco. For each valid species, synonyms known to have been used for published records from Morocco are listed under the currently accepted name. Where available, distribution within Morocco is also included. One new combination is proposed: Assuania melanoleuca (Séguy, 1941), comb. nov. (Chloropidae).

Keywords

Bibliography, classification, gnats, midges, new combination, new records, taxonomy, true flies

Foreword

Almost everyone is now aware that there is a biodiversity crisis, and even the popular press regularly reports on general declines in the diversity of species as well as more specific threats to the distribution and abundance of species. Despite this growing awareness of the importance of biodiversity to the health of our planet and thus to the welfare of humankind, relatively little attention has been paid to the glaringly obvious point that it is impossible to objectively assess changes such as declines in diversity in the absence of baseline data and a good taxonomic infrastructure. In other words, if you don’t know what you have to start with it is impossible to know what you are losing (or gaining). Recognition of invasions, extirpations, extinctions, range changes and changes in phenology all pivot on the documentation of what species occur in an area to begin with. Regional catalogues like this one provide both a baseline and a startling wake-up call; in this case with the announcements of 156 previously undocumented species, records of many hitherto unrecorded genera, and the first recognition of half a dozen families new to the country.

Documentation of the dipteran diversity of Morocco is of special significance both in terms of taxon and area. The taxon, Diptera or true flies, is one of the two or three largest orders of living things, with hundreds of thousands of undescribed species worldwide in addition to the approximately 160,000 currently named species. The Moroccan Diptera fauna is relatively well known compared to other parts of Africa, but nonetheless the current catalogue more than doubles the number of species known from the country as of the last inventories a couple of decades ago. This impressive advance in the documentation of Morocco’s significant dipteran fauna is of course in part due to the great dedication and Herculean effort on the part of the lead author, Prof. Kettani. But, it is also a great credit to the community of dipterists who rallied to the cause by meticulously cataloguing their chosen families.

Approximately 33 of the 60 contributors are retired entomologists or amateur dipterists who volunteered their time to contribute to this work. Sadly, a few have passed away since they made their contribution. It is both impressive and worrisome that more than half of those 60 top experts on the taxonomy of the region’s Diptera are retired or amateur, since there is a pressing and ongoing need for specialized expertise on all of the families of Diptera. Hopefully the framework, and the challenge, provided by this thorough description of our current state of knowledge of Moroccan flies will help and encourage new students of dipterology while at the same time providing a foundation for ongoing assessments of the dipteran diversity of North Africa.

I congratulate all involved for the production of such an important contribution to our understanding of the biodiversity of the region, and for the successful completion of the first full Diptera catalogue for any African country.

Stephen A. Marshall

Professor, University of Guelph, Canada

October 2020

Contributions

This catalogue was conceived by K. Kettani (KK) in 2011 and then further developed and planned together with Martin J. Ebejer (MJE). These plans included more fieldwork, predominantly by KK and MJE, with some contribution by students, carried out in 2012–2019, which resulted in about 1000 new species records for Morocco and 23 species (in 12 families) new to science. These records were published prior to the end of 2020 and the papers are included in the references and cited under the appropriate families. KK compiled the data and the preliminary list of species for each family. The taxonomic specialists (co-authors and contributors) revised any taxonomic issues, added data available to them and updated nomenclature. Contributors are thanked in the Acknowledgements, as well as mentioned in their sections of the checklist. The distributions of species in Morocco, with the appropriate citations, were added by KK. The introduction was written by KK and supplemented with additional information by MJE, who also checked the English language.

Introduction

Aims of the catalogue

Although the national study on biodiversity, Etude nationale sur la biodiversité (Dakki 1997; Mouna 1998), was the only important synthesis of entomological studies carried out in Morocco until now, important data on the Diptera are lacking. For example, no taxonomic rank such as subfamily and tribe, and no synonyms or revised generic combinations, names of museums or private collections that house species described from Morocco were included. No information was provided on the habitats and distribution of species in Morocco. Furthermore, the information resulting from the national study on biodiversity is now outdated. A large number of important recent publications, cited appropriately under each family, provide much additional data to the lists published in 1997 and 1998, almost doubling the number of known species. The main aims of this catalogue are to bring together all this information by providing an up-to-date checklist according to the current nomenclature and systematics, to provide synonyms used in older literature that dealt with the Moroccan fauna and to give regional distribution data. Finally, an essential comprehensive bibliography is provided.

This catalogue is the first for a North African country and it may prove useful also for neighbouring countries. It is hoped that it will provide a sound foundation for anyone choosing to study a group of Moroccan Diptera. It could indicate where the greatest need for investigation lies, both with regard to areas of the country that have been poorly studied and taxonomic groups (or families) that may benefit from more attention. It is undoubtedly not a complete inventory and many more discoveries are expected.

The faunistic knowledge of the Diptera of Morocco from 1787 to 2021 is summarized and updated, providing the first catalogue of the Moroccan species of Diptera. In total, 3057 species (975 Nematocera and 2082 Brachycera) in 93 families, classified into 949 genera (250 in Nematocera and 699 in Brachycera) are listed (Tables 1, 2).

Table 1.

Families of Nematocera with contributing specialists and numbers of taxa.

Family Specialist(s) Subfamilies Genera Species
Anisopodidae Haenni 1 1 3
Bibionidae Haenni 1 2 10
Blephariceridae Zwick 1 1 4
Cecidomyiidae Skuhravá & Skuhravý 3 31 57
Ceratopogonidae Mathieu 3 6 62
Chaoboridae Wagner 1 2 2
Chironomidae - 7 100 412
Culicidae Trari, Himmi & Dakki 2 7 43
Dixidae Wagner - 2 12
Keroplatidae Chandler 1 2 2
Limoniidae Oosterbroek 4 22 67
Mycetophilidae Chandler 5 25 64
Pediciidae Oosterbroek 1 2 6
Psychodidae Wagner 2 12 51
Ptychopteridae Wagner 1 1 5
Scatopsidae Haenni 3 11 13
Sciaridae Heller - 12 70
Simuliidae - 1 6 43
Thaumaleidae Wagner - 1 2
Tipulidae Oosterbroek & de Jong 2 3 39
Trichoceridae Krzemińska 1 1 8
Total: 250 Total: 975
Table 2.

Families of Brachycera with contributing specialists and numbers of taxa.

Family Specialist(s) Subfamilies Genera Species
Acroceridae Nartshuk 2 6 13
Agromyzidae Černý 2 12 62
Anthomyiidae Ackland 2 14 36
Anthomyzidae Ebejer - 2 2
Asilidae Tomasovic 9 42 131
Asteiidae Ebejer 1 2 5
Atelestidae Gatt - 1 1
Athericidae Mouna 1 1 2
Aulacigastridae Ebejer - 1 1
Bombyliidae Ebejer & Dils 11 48 248
Braulidae - 1 1 1
Calliphoridae Rognes 5 5 8
Camillidae Ebejer - 1 4
Canacidae Munari 2 3 15
Carnidae Ebejer - 1 3
Chamaemyiidae Ebejer 2 5 18
Chloropidae von Tschirnhaus & Ebejer 3 34 74
Chyromyidae Ebejer 2 3 22
Clusiidae Ebejer - 1 1
Coelopidae - 1 1 1
Conopidae - 5 10 34
Cryptochetidae Nartshuk - 1 1
Diastatidae Ebejer 2 2 2
Dolichopodidae Grichanov & Negrobov 12 36 112
Drosophilidae Bächli 2 9 26
Dryomyzidae - 1 1 1
Empididae Daugeron 3 8 40
Ephydridae Zatwarnicki 5 43 117
Fanniidae Pont - 1 10
Helcomyzidae - 1 1 1
Heleomyzidae Woźnica 3 5 19
Hippoboscidae Droz 3 10 17
Hybotidae Gatt 2 10 44
Lauxaniidae Ebejer 2 8 27
Lonchaeidae MacGowan 1 4 5
Lonchopteridae Barták 1 1 4
Micropezidae Ebejer - 1 1
Milichiidae - 3 6 8
Muscidae Pont 6 25 115
Mydidae Dikow 3 4 9
Mythicomyiidae Evenhuis 4 6 8
Nemestrinidae Barraclough 2 4 13
Nycteribiidae Graciolli 1 4 8
Odiniidae Ebejer 1 1 2
Oestridae Pape 2 5 10
Opomyzidae - - 2 5
Pallopteridae Ebejer - 1 1
Phoridae Disney - 2 3
Piophilidae Ebejer 1 3 3
Pipunculidae Ebejer 2 7 16
Platypezidae Ebejer 1 3 3
Platystomatidae Popov 1 2 4
Polleniidae Rognes - 1 12
Psilidae Ebejer - 1 1
Rhagionidae Ebejer 1 1 4
Rhiniidae Rognes 1 5 17
Rhinophoridae Pape 1 6 8
Sarcophagidae Whitmore & Pape 3 16 66
Scathophagidae - 1 2 3
Scenopinidae Carles-Tolrá 1 2 11
Sciomyzidae Vala 1 12 25
Sepsidae Haenni 1 4 12
Sphaeroceridae Gatt 3 29 67
Stratiomyidae Woodley 5 11 40
Streblidae Graciolli 1 2 2
Syrphidae Speight 2 50 166
Tabanidae - 3 11 69
Tachinidae Cerretti & Tschorsnig 4 88 147
Tephritidae Norrbom 3 29 69
Therevidae Hauser 2 9 27
Ulidiidae Ebejer 2 8 13
Vermileonidae Ebejer 1 2 6
Total: 698 Total: 2082

Brief history of Moroccan dipterology

The study of Diptera in Morocco is still far from being extensive despite the numerous investigations that have taken place so far. Until now, a synthesis of Moroccan Diptera had never been produced. The studies that have been devoted to this group of insects have been limited and sporadic. They were mainly concerned with either taxonomy and descriptions of new species, or faunal studies from specific areas of Morocco with little reference to the habitats of species. This was particularly so for early works such as Becker and Stein (1913), Séguy (1930a, 1934b, 1935a, 1941a, 1941d, 1949a, 1953a), Timon-David (1951), Vaillant (1956b) and, recently, Pârvu et al. (2006), Popescu-Mirceni (2011), and Ebejer et al. (2019). Very few recent works have provided relevant ecological data (Cassar et al. 2005, 2008). Becker and Stein (1913) were the first to publish a list of Moroccan Diptera, at that time comprising only 204 species. Séguy followed with a long series of valuable publications. In 1930 he produced his first list of Diptera, which included 471 species in 230 genera grouped into 37 families (Séguy 1930). Over the subsequent years, he contributed greatly to the knowledge of the Diptera of Morocco (Séguy 1926, 1934b, 1935a, 1941a, 1941d, 1949a, 1953a, 1957, 1961), where he also referenced an extensive bibliography. Later, Bailly-Choumara (1977) compiled a bibliography of Moroccan Diptera. It took until 1997–1998, with a national study on the biodiversity of Morocco co-ordinated by what was then the Ministry of the Environment, to update the list of the Moroccan insect fauna. This national study was dedicated to the compilation of the results of entomological research conducted in Morocco in order to highlight Moroccan biodiversity. It included a comprehensive inventory of published entomological research conducted in Morocco and was the last work to include a broad overview of Moroccan Diptera. It listed 623 species of aquatic Diptera in 210 genera and 25 families (Dakki 1997) and 928 species of terrestrial Diptera in 350 genera and 57 families (Mouna 1998).

Terrain and bioclimatic regions

Morocco, located in the westernmost corner of North Africa, constitutes a biogeographical crossroads between the Afrotropical and the western Palaearctic faunas, which may have allowed some exchange of genetic material. The country has a remarkable variety of bioclimates, ranging from humid in the Rif and the Middle and High Atlas to very arid in the Sahara in the south and to the sub-humid and semi-arid plains and foothills (Fig. 1). As a result of this diversity of landscapes and climates, there exists a great biological and ecological diversity in the country.

Figure 1. 

The major bioclimatic regions of the northern part of Morocco.

The geomorphological, orographic and bioclimatic diversity of the country is reflected in its seven distinct biogeographical areas: the Rif, the Atlantic Plain, Eastern Morocco, the Middle Atlas, the High Atlas, the Anti-Atlas and the Sahara. In the checklist, an abbreviated distribution (in bold) is given for each species according to these seven biogeographical areas (Fig. 2).

Figure 2. 

The seven biogeographical regions covering the studied area in the northern part of Morocco.

Rif: The Rif consists of a mountainous domain overlooking the Mediterranean Sea in the north of Morocco. It belongs to the Gibraltar Arc or Alborán Sea geological region, part of the Alpine orogenic belt, where the maximum altitude is 2,452 m in the Jbel Tidirrhine mountain range. The climate is mainly of the Mediterranean type and is characterised by high levels of precipitation. This domain is considered very rich and diverse in vegetation cover (Valdés et al. 2002), including nearly all of the Moroccan forest plant species (Médail and Quézel 1999) and integrating five bioclimatic stages.

Atlantic Plain (AP): Also known as the Moroccan Central Plateau, the Atlantic Plain is an ancient massif located in the northwest of Morocco, between the Atlantic coast and the Middle Atlas Mountains. It consists of a plateau surrounded by medium-altitude mountain massifs and covers an area of 8500 km². With an asymmetrical shape, the plateau culminates in the south-east at Jebel Mtourzgene (1627 m), located near the town of Oulmès. Its complex geomorphology presents tectonic troughs that have favoured the formation of lakes and rivers. The annual rainfall of this region, concentrated in winter and spring, totals an average of 500 mm, allowing the establishment of Holm Oak and Cork Oak forests in a semi-arid bioclimate (Piqué 1994). Towards the coast, the vegetation is mainly a steppe with Jujube shrubs and Dwarf Palms (Chamaerops humilis). The maritime fringe is underlined by cliffs and ancient consolidated dunes.

Eastern Morocco (EM): This is an arid region covering the Hauts Plateaux to the northeast of the country. It is the only region of Morocco subject to a true Mediterranean bioclimate, with some influence from the Sahara. The annual rainfall is less than 300 mm, encouraging the dominance of esparto steppe (Stipa tenacissima) (Piqué 1994). Its geomorphology is represented by a plateau at an altitude of 1000 m, which gradually decreases from 1500 to 500 m from the city of Midelt to the province of Guercif and is criss-crossed by the Moulouya wadi. In the northwestern part of Eastern Morocco is the Béni Snassen massif, beyond the Guercif depression.

Middle Atlas (MA): A mountainous massif located in the centre of Morocco, oriented from south-west to north-east and extending over 350 km (Piqué 1994). This mass of highlands is essentially composed of limestone layers, tabular and of medium altitude in the southwest (tabular Middle Atlas) and undulating in the northeast (folded Middle Atlas) to culminate at 3340 m at Jebel Bou Naceur, which overlooks the high plateaux of Eastern Morocco. The climate, generally subhumid to humid, is cold in winter, giving rise to cedar and Holm Oak forests covering the western slope, which is wetter than the eastern slope and where lakes are abundant and many rivers and streams have their source.

High Atlas (HA): The main mountainous massif of Morocco, oriented from west-south-west to east-north-east, is the High Atlas, stretching over more than 700 km from the Atlantic to the east of Morocco, with a north-south width of 50 to 100 km and reaching at Jebel Toubkal an altitude of 4165 m. The whole massif is high, incised by deep valleys that lead to passes through which the mountainous barrier is crossed: the Tizi-n’Test (2100 m) towards the Souss plain and the Tizi-n’Tichka (2200 m) towards the depression of Ouarzazate. The chain is formed by a massif of Palaeozoic rocks surrounded by sedimentary deposits. The relief of the High Atlas is divided into three different entities from west to east: the Western High Atlas, which is the oldest massif and is made up mainly of Jurassic or Cretaceous formations with deep valleys and culminates at Jebel Toubkal; the central High Atlas, which is an essentially limestone massif morphologically dominated by tabular zones culminating at an altitude of 2500 m, stretches between the towns of Azilal and Ouarzazate and is sheltered by the Jebel M’Goum (4071 m); and the Eastern High Atlas, which is formed by the vast high plateaux of the Upper Moulouya, stretches between the cities of Midelt and Imilchil and contains the Jebel Ayachi (3747 m) (Piqué 1994). The bioclimate of this area includes subhumid, humid and semi-arid zones, but with more than 800 mm of annual precipitation on the highest peaks and a significant snow cover from September to May, with temperatures dropping below -20 °C (AEFCS 1996) and bright sunshine even in winter. These climatic conditions have favoured large pine, Holm Oak, Thuja and thuriferous juniper forests (HCEFLCD 2017). The aridity of the southern slope of the High Atlas allows only steppe vegetation of sagebrush and esparto grass to grow.

Anti-Atlas (AA): This is the pre-Saharan area, which extends north of the Wadi Draa and rises to an average altitude of 2000 m. It is a semi-arid region with Jebel Bani and Jebel Ouarkziz as its main elevations, and is home to numerous oases. The western part of the Anti-Atlas is characterised by the Saharan plateau, with a temperate winter except in the high mountains. On the other hand, in the eastern part of the Anti-Atlas, the greatest arid area is marked by the northward extension of the Saharan landscape and a bioclimate of cool and temperate winters. These bioclimatic stages are characterised by plant communities such as wooded steppes with Acacia raddiana, Artemisia herba alba and Stipa tenuissima, interspersed with thuriferous juniper (Juniperus thurifera) at the top of the slopes (Piqué 1994). The vegetation is also represented by two formations that partially interpenetrate each other: groves of the Argan tree (Argania spinosa) and the Euphorbia steppe (Euphorbia echinus).

Sahara (SA): This area in the south of the country consists of a wide arid zone with a typical Saharan bioclimate of high daytime temperatures and very low precipitation. From a geological point of view it is part of the West African Craton and its margins. It is mainly ancient terrain with Precambrian basement rock and Palaeozoic cover (Piqué 1994). This area is represented by a peneplain of very modest altitudes from 200 to 500 m, established on ancient crystalline terrain. Precipitation from July to September is very low, of the order of 100 mm.

The dipteran species richness of each biogeographical region of Morocco is summarised in Table 3, which highlights the great disparity between the different regions and reflects the lack of knowledge about this insect order in some areas of Morocco. The Rif region in the north of the country has the greatest recorded diversity, but this undoubtedly reflects a greater sampling effort in this area, with large parts of the country still very poorly explored for Diptera. The catalogue does not include any data from the southern part of the Moroccan Sahara.

Table 3.

Species richness of Diptera families each biogeographical region in Morocco. Rif; AP – Atlantic Plain; EM – Eastern Morocco; MA – Middle Atlas; HA – High Atlas; AA – Anti-Atlas; SA – Sahara. * No precise localities known.

Rif AP EM MA HA AA SA
Acroceridae 7 1 0 1 3 0 0
Agromyzidae 7 15 0 10 22 10 0
Anisopodidae *
Anthomyiidae 4 12 0 13 2 4 1
Anthomyzidae 1 0 0 0 1 0 0
Asilidae 34 27 4 35 25 24 5
Asteiidae 0 3 0 0 1 1 0
Atelestidae 1 0 0 0 0 0 0
Athericidae 0 0 0 1 2 0 0
Aulacigastridae 1 0 0 0 0 0 0
Bibionidae 4 2 0 3 1 2 2
Blephariceridae 1 0 0 0 3 0 0
Bombyliidae 35 80 12 59 48 56 19
Braulidae *
Calliphoridae 3 8 2 4 4 0 0
Camillidae 1 2 0 0 1 0 0
Canacidae 6 10 0 0 0 6 0
Carnidae 1 0 0 0 0 0 0
Cecidomyiidae 8 27 21 25 20 7 4
Ceratopogonidae 15 43 14 35 33 24 36
Chamaemyiidae 5 5 0 4 8 6 0
Chaoboridae *
Chironomidae 230 30 7 105 120 14 1
Chloropidae 35 22 0 7 6 25 1
Chyromyidae 6 6 0 0 1 12 1
Clusiidae 1 0 0 0 0 0 0
Coelopidae 1 0 0 0 0 0 0
Conopidae 7 13 0 16 7 9 0
Cryptochetidae 0 0 0 1 0 0 0
Culicidae 32 41 24 30 29 22 11
Diastatidae 0 1 0 0 1 0 0
Dixidae 2 2 0 2 5 0 0
Dolichopodidae 72 28 3 6 18 26 0
Drosophilidae 11 14 0 8 12 5 1
Dryomyzidae 1 0 0 0 0 0 0
Empididae 4 3 0 13 20 1 0
Ephydridae 47 44 1 60 9 20 1
Fanniidae 4 2 0 2 5 0 0
Helcomyzidae 1 0 0 0 0 0 0
Heleomyzidae 11 3 0 2 2 3 0
Hippoboscidae 0 2 3 4 4 4 1
Hybotidae 20 7 3 8 7 8 0
Keroplatidae 1 0 0 0 0 0 0
Lauxaniidae 23 3 2 1 8 2 0
Limoniidae 43 9 4 5 34 5 0
Lonchaeidae 3 3 0 1 1 0 0
Lonchopteridae 0 0 0 1 1 0 0
Micropezidae 1 0 0 0 0 0 0
Milichiidae 2 1 0 0 4 3 1
Muscidae 25 47 13 38 68 20 6
Mycetophilidae 60 3 7 3 0 0 0
Mydidae 0 3 2 1 0 3 1
Mythicomyiidae 2 0 0 0 1 2 1
Nemestrinidae 1 1 1 5 3 0 1
Nycteribiidae 6 5 0 6 0 2 1
Odiniidae 1 0 0 0 0 0 0
Oestridae 0 2 1 1 1 0 1
Opomyzidae 1 1 0 1 2 0 0
Pallopteridae 0 0 0 1 0 0 0
Pediciidae 3 1 0 0 0 0 0
Phoridae 0 2 0 0 0 0 0
Piophilidae 2 1 0 1 1 0 0
Pipunculidae 13 1 0 0 0 0 0
Platypezidae 2 1 0 0 0 0 0
Platystomatidae 0 2 1 1 0 0 0
Polleniidae 1 4 2 5 5 1 1
Psilidae 1 0 0 1 0 0 0
Psychodidae 25 12 12 12 34 9 2
Ptychopteridae *
Rhagionidae 2 0 0 0 2 0 0
Rhiniidae 2 5 3 5 0 5 4
Rhinophoridae 6 2 0 1 0 0 0
Sarcophagidae 11 19 1 13 8 9 2
Scathophagidae 3 1 0 1 0 0 0
Scatopsidae 10 2 0 0 1 3 0
Scenopinidae 3 4 1 1 0 0 1
Sciaridae 29 12 2 8 45 9 3
Sciomyzidae 8 4 2 11 18 3 0
Sepsidae 3 6 0 4 7 3 0
Simuliidae 32 8 0 19 32 6 0
Sphaeroceridae 42 3 2 10 1 3 3
Stratiomyidae 20 13 5 13 6 1 0
Streblidae 1 1 0 0 0 1 0
Syrphidae 70 49 11 82 64 25 4
Tabanidae 26 27 11 35 21 7 1
Tachinidae 18 27 4 46 43 29 3
Tephritidae 34 13 4 16 13 23 3
Thaumaleidae 0 0 0 0 2 0 0
Therevidae 9 7 2 9 4 5 2
Tipulidae 28 7 0 16 18 2 0
Trichoceridae 7 1 3 1 0 0 0
Ulidiidae 6 2 0 1 4 4 1
Vermileonidae 1 2 1 1 4 0 0
Total species 1126 770 191 831 876 474 126

Sources of data

Data for the present study were gathered from the literature, supplemented by information taken from a number of websites. The cut-off date for inclusion of species published in the literature is 30 June 2021. Taxonomic, nomenclatural, and distributional data for the present catalogue were obtained from:

  • studies by various foreign and Moroccan researchers published between 1787 and 2021, including theses, books, catalogues, checklists and notes from the 1997–1998 synthesis studies on Morocco’s biodiversity.
  • data from websites and organisations such as Global Species, the Global Biodiversity Information Facility, the BioSystematic Database of World Diptera, Systema Dipterorum, Catalogue of Life, Systema Naturae 2000 and others, or from relevant homepages specific to particular families.
  • the collection of Diptera in the National Museum of Morocco (Muséum National d’Histoire Naturelle à l’Institut Scientifique, Université Mohammed V, Rabat).
  • collaboration with specialists who communicated reliable identifications from their personal collections or provided data for species deposited in foreign museums.
  • published records of specimens in the personal collection of the lead author (PCKK), deposited at Abdelmalek Essaadi University, Tetouan, identified with the assistance of specialists.

Classification

The arrangement of families in this catalogue (Table 4) is mainly based on Pape et al. (2011) and Wiegmann et al. (2011). All families are listed alphabetically within each superfamily. Nomenclature follows the most recent version of Systema Dipterorum (Evenhuis and Pape 2021) and the Polleniidae are listed as a separate family (Gisondi et al. 2020).

Table 4.

Diptera classification used in the catalogue.

Suborder NEMATOCERA
Tipuloidea Limoniidae
Pediciidae
Tipulidae
Trichoceridoidea Trichoceridae
Psychodoidea Psychodidae
Scatopsoidea Scatopsidae
Ptychopteridae
Culicoidea Chaoboridae
Culicidae
Dixidae
Chironomoidea Ceratopogonidae
Chironomidae
Simuliidae
Thaumaleidae
Blephariceridae
Bibionoidea Anisopodidae
Bibionidae
Sciaroidea Cecidomyiidae
Keroplatidae
Mycetophilidae
Sciaridae
Suborder BRACHYCERA
Stratiomyoidea Stratiomyidae
Tabanoidea Athericidae
Rhagionidae
Tabanidae
Vermileonidae
Nemestrinoidea Acroceridae
Nemestrinidae
Asiloidea Asilidae
Bombyliidae
Mydidae
Mythicomyiidae
Scenopinidae
Therevidae
Empidoidea Atelestidae
Empididae
Dolichopodidae
Hybotidae
Platypezoidea Phoridae
Platypezidae
Lonchopteridae
Syrphoidea Pipunculidae
Syrphidae
Conopoidea Conopidae
Nerioidea Micropezidae
Tanypezoidea Psilidae
Tephritoidea Lonchaeidae
Pallopteridae
Piophilidae
Platystomatidae
Tephritidae
Ulidiidae
Lauxanioidea Chamaemyiidae
Lauxaniidae
Sciomyzoidea Coelopidae
Dryomyzidae
Helcomyzidae
Sciomyzidae
Sepsidae
Opomyzoidea Agromyzidae
Anthomyzidae
Asteiidae
Aulacigastridae
Clusiidae
Odiniidae
Opomyzidae
Carnoidea Canacidae
Carnidae
Chloropidae
Milichiidae
Sphaeroceroidea Chyromyidae
Heleomyzidae
Sphaeroceridae
Ephydroidea Braulidae
Camillidae
Cryptochetidae
Diastatidae
Drosophilidae
Ephydridae
Hippoboscoidea Hippoboscidae
Nycteribiidae
Streblidae
Muscoidea Anthomyiidae
Fanniidae
Muscoidea Muscidae
Scathophagidae
Oestroidea Calliphoridae
Oestridae
Polleniidae
Rhiniidae
Rhinophoridae
Sarcophagidae
Tachinidae

Arrangement of the catalogue

The catalogue includes all names of taxonomically valid species of the Order Diptera so far known to occur in Morocco. For each family, synonyms and alternative combinations that have been published as records of Moroccan provenance are listed under the currently accepted name.

The transliteration of Moroccan place names in the literature, maps, and sometimes on data labels presented a small problem. Although it would have been preferable to use standard names, this would have meant changing some place names from their original spellings in the literature. Most transliteration used the French language for spelling and Roman letters, but not all. English was sometimes used, and occasionally also Spanish. Readers are therefore advised to take this into account when searching for place names given in the catalogue and in the cited references. Some common examples are “Jebel”, “Jbel”, “Djebel” or “Dj.”, used for “mountain”, or “Kbir” and “Kebir”, for “large” or “great”. The article “the” in Arabic is El or L; it is sometimes separated by a hyphen from its associated noun or joined to it, sometimes capitalised while the noun is not, both are capitalised, or only the noun is. When the article “el” is followed by a noun beginning with a certain consonant, like n, r, s and t, the l is changed to match the noun’s initial, for example “Er-Rifiyine”. In transliteration, Arabic speakers would make this change, whereas non-Arabic speakers may not. In cases where the article is joined to the noun, the resulting versions may appear as being different place names; for example, El Rachidia, Er Rachidia and Errachidia are alternative transliterations of the same place name. In some entries, the French terms “affluent” and “aval affluent” are left as in the original citation, because “tributary” may not be an accurate translation in all instances. For some of the sites, the French term refers to a downstream overflow of water into a side tributary or waterfall, whereas for others it refers to an actual tributary feeding into a main river or stream.

Older authors were imprecise with localities, and many references did not give the coordinates of localities from which species were collected. We could not reliably provide accurate coordinates for each species. Where known, species depositories are included. Individual family checklists may include, at the end, taxonomic notes and comments on species doubtfully present in the country. New records are indicated with an asterisk (*) and the relevant data listed at the end of the family section.

Limitations

One of the major difficulties encountered while compiling the catalogue was the limited ability to check old identifications. In some cases, identifications were accepted as given in the original sources, even when doubtful, as it was usually not possible to re-examine specimens. The inaccessibility of some references was also an obstacle to updating the checklists for some families.

Collections cited

The following museums and collections house some of the specimens mentioned in this catalogue. The National Museum of Natural History of Morocco is located in the Institut Scientifique, Université Mohammed V, Rabat. However, the Diptera collection housed in that museum is not yet included in any world database such as the GBIF Registry of Scientific Collections, https://www.gbif.org/grscicoll, or the Insect and Spider collections of the World Website, http://hbs.bishopmuseum.org/codens/.

HNHM Hungarian Natural History Museum, Budapest, Hungary;

INHS Illinois Natural History Survey, Urbana, USA;

IRSNB Institut Royal des Sciences Naturelles de Belgique, Brussels, Belgium;

MfN Museum für Naturkunde, Berlin, Germany;

MHNN Muséum d’Histoire Naturelle de Neuchâtel, Neuchâtel, Switzerland;

MISR Museum National d’Histoire Naturelle à l’Institut Scientifique, Université Mohammed V, Rabat, Morocco;

MNCN Museo Nacional de Ciencias Naturales, Madrid, Spain;

MNHN Muséum National d’Histoire Naturelle, Paris, France;

NHMD Natural History Museum of Denmark, Copenhagen, Denmark;

NHMUK Natural History Museum, London, United Kingdom;

NMWC National Museum of Wales, Cardiff, United Kingdom;

OUMNH Oxford University Museum of Natural History, Oxford, United Kingdom;

RMNH Rijksmuseum voor Natuurlijke Historie, Leiden, The Netherlands;

SMNS Staatliches Museum für Naturkunde Stuttgart, Stuttgart, Germany;

ZMUM Zoological Museum of Moscow University, Moscow, Russia;

ZSM Zoologische Staatssammlung, Munich, Germany;

ZSM Zoologische Staatssammlung, Munich, Germany.

The personal collection of Dr Michael von Tschirnhaus has been donated to the ZSM, but is currently on loan to Dr von Tschirnhaus.

Other abbreviations

HT Holotype;

NPH National Park of Al Hoceima;

NPHAO National Park of Haut Atlas Oriental;

NPI National Park of Ifrane;

NPTNational Park of Talassemtane;

PCJD Private Collection of Jos Dils, Belgium;

PCKK Private Collection of Kawtar Kettani, Morocco;

PNPB Project of Natural Park of Bouhachem.

Acknowledgements

The authors are deeply grateful to the reviewers for their constructive comments, which helped to improve the manuscript. Our sincere thanks are expressed to the subject editor, Torsten Dikow, for his significant contribution and support as editor.

We are most grateful to the following specialists who generously contributed in various ways to this catalogue by sharing information and knowledge of their group or by reviewing a family section: Patrick Ashe (Dublin, Ireland)—Chironomidae; Torsten Dikow (Washington, D.C., USA)—Mydidae; the late Amnon Freidberg (Tel Aviv, Israel)—Tephritidae; Kai Heller (Heikendorf, Germany)—Sciaridae; Gail Kampmeier (Champaign, USA)—Therevidae; Valery Korneyev (Kiev, Ukraine)—Tephritidae; Ewa Krzemińska (Kraków, Poland)—Trichoceridae; Allen Norrbom (Washington, D.C., USA)—Tephritidae; Adrian R. Plant (Mahasarakham, Thailand)—Empididae, Hemerodrominae; Bradley J. Sinclair (Ottawa, Canada)—Empididae, Clinocerinae; Jens-Hermann Stuke (Leer, Germany)—Conopidae; Norman Woodley (Hereford, AZ, USA)—Stratiomyidae. Mohamed Mouna and Mohamed Dakki are acknowledged and thanked for their encouragement and inspiration that led the first author to commence this project, as well as for generously donating to her a large bibliography on the subject. Mohamed Mouna was instrumental in ensuring that she had full access to the collections and library and full cooperation from the staff of these departments. He is immensely thanked by the first author for donating to her his library, consisting of many books and publications.

Mohammed Fekhaoui, Abdellatif Bayed, Mohamed-Aziz El Agbani, Mohamed Arahou from the administration of Museum National d’Histoire Naturelle à l’Institut Scientifique, Université Mohammed V, Rabat are warmly thanked for their valuable assistance in providing access to the library and Diptera collections of the museum. Thanks also to Amina Zouida, the librarian at the said institute, for her tireless and precious help in searching for obscure publications, and to Abderrahmane Mataame (responsible for the zoological collections of the museum) for his kind and patient assistance during the first author’s frequent visits to the museum. Yassine Fekrani is acknowledged for his kind help producing the maps.

Suborder NEMATOCERA

Tipuloidea

LIMONIIDAE

K. Kettani, P. Oosterbroek

Number of species: 67. Expected: 85

Faunistic knowledge of the family in Morocco: moderate

Chioneinae

Baeoura Alexander, 1924

Baeoura ebenina Starý, 1981

Driauach and Belqat 2015, Rif, Oued Tazarine (Mezine village); Dayat near Aïn Afersiw; Driauach and Belqat 2016; Oosterbroek 2020 (CCW: Rif)

Baeoura staryi Driauach & Belqat, 2015

Driauach and Belqat 2015, Rif, Jnane Niche; Driauach and Belqat 2016; Oosterbroek 2020 (CCW: Rif)

Cheilotrichia Rossi, 1848

Cheilotrichia (Empeda) cinerascens (Meigen, 1804)

Driauach et al. 2013, Rif, Ikadjiouen; Driauach and Belqat 2016, Rif, Oued Ouara, Oued Amsa, Oued Maggou, Aïn Quanquben (Jebel Bou Bessoui), EM, Grotte du Chameau (Zegzel, Béni Snassen); Oosterbroek 2020 (CCW: Rif; EM)

Cheilotrichia (Empeda) fuscohalterata (Strobl, 1906)

Driauach and Belqat 2016, Rif, Dayat Fifi, tributary of Oued El Fondak, Barrage Ajras, Dayat Mghara, Oued Zarka, Oued Tizekhte; Oosterbroek 2020 (CCW: Rif)

Cheilotrichia (Empeda) minima (Strobl, 1898)

Driauach and Belqat 2016, Rif, Oued Zendoula, Oued Jnane Niche; Oosterbroek 2020 (CCW: Rif)

Ellipteroides Becker, 1907

Ellipteroides (Ellipteroides) lateralis (Macquart, 1835)

= Gonomyia lateralis Macquart, in Pierre 1922a: 22, Dakki 1997: 62

= Gonomyia cincta Egger, in Séguy 1941a: 26

Pierre 1922a, AP, Dradek (near Rabat); Lackschewitz 1940a, HA, Tachdirt (2200–2900 m); Séguy 1941a, HA, Tachdirt (Toubkal, 2500 m); Savchenko et al. 1992; Dakki 1997; Starý and Freidberg 2007; Driauach et al. 2013; Driauach and Belqat 2016, Rif, Dayat Afrate; Oosterbroek 2020 (CCW: Rif)

Ellipteroides (Protogonomyia) alboscutellatus (von Roser, 1840)

Lachschewitz 1940a, HA; Savchenko et al. 1992; Starý and Oosterbroek 2008; Starý 2009a; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Ellipteroides (Protogonomyia) hutsoni (Starý, 1971)

Starý 1971, HA, Jebel Ayachi; Savchenko et al. 1992, HA, Jebel Ayachi; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Erioconopa Starý, 1976

Erioconopa diuturna (Walker, 1848)

Driauach et al. 2013; Driauach and Belqat 2016, Rif, Dayat Jebel Zemzem, Oued Mezine, Aïn Bab Tariouant; Oosterbroek 2020 (CCW: Rif)

Erioconopa symplectoides (Kuntze, 1914)

= Dactylolabis symplectoides Egger, in Pierre 1922a: 23

Pierre 1922a, HA, Marrakech; Savchenko et al. 1992; Starý and Oosterbroek 2008, HA; Gavryushin pers. comm. 2012, HA; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW: HA)

Erioptera Meigen, 1803

Erioptera (Erioptera) fuscipennis Meigen, 1818

Pierre 1922a, AP, Casablanca (Oued Guerera); Pierre 1922b, HA, Savchenko et al. 1992; Dakki 1997; Driauach et al. 2013, Rif, Sidi Brahim Ben Arrif (Bab Hachef Aissa); Driauach and Belqat 2016, Rif, Aïn El Ma Bared, Oued Amsemlil, Oued Maggou (Bridge), Oued Tkarae, Dayat near Aïn Afersiw, Aïn Afersiw, Oued Abou Bnar, Oued Maggou (Zaouiet El Habtiyne), Dayat Amsemlil, Dayat Aïn Jdi­oui; Dayat Afrate, Oued Mezine; Oosterbroek 2020 (CCW: Rif, HA)

Erioptera (Erioptera) lutea lutea Meigen, 1804

Driauach and Belqat 2016, Rif, Aïn Boughaba; Oosterbroek 2020 (CCW: Rif)

Erioptera (Erioptera) transmarina Bergroth, 1889

= Mesocyphona transmarina Bergroth, in Pierre 1922a: 22, Pierre 1922b: 148, Dakki 1997: 62

Pierre 1922a, HA, Marrakech; Pierre 1922b, HA, Tannaout (1000 m), Marrakech; Dakki 1997; Savchenko et al. 1992; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Gonomyia Meigen, 1818

Gonomyia (Gonomyia) abscondita Lackschewitz, 1935

Driauach and Belqat 2016, Rif, maison forestière; Oosterbroek 2020 (CCW: Rif)

Gonomyia (Gonomyia) sicula Lackschewitz, 1940

Driauach and Belqat 2016, Rif, Oued Kbir, Dayat Jebel Zemzem, Aïn Sidi Brahim Ben Arrif, Dayat Tazia, Aïn El Maounzil; Oosterbroek 2020 (CCW: Rif)

Gonomyia (Gonomyia) subtenella Savchenko, 1972

Starý and Oosterbroek 2008, HA, Massif Toubkal, Aghbalou, 43 km S Marrakech (1000 m); Starý 2009a; Driauach et al. 2013; Driauach and Belqat 2016, Rif, Oued Jnane Niche, Oued Sidi Yahia Aârab, EM, Oued Béni Ouachekradi; Oosterbroek 2020 (CCW: Rif, EM)

Gonomyia (Gonomyia) tenella (Meigen, 1818)

Pierre 1924a, HA, Asni (1200 m); Séguy 1930a, HA, Asni; Savchenko et al. 1992; Dakki 1997; Starý 2009a; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Hoplolabis Osten-Sacken, 1869

Hoplolabis (Parilisia) obtusiapex (Savchenko, 1982)

Starý 2006, HA, Oasis Meski, AA; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Hoplolabis (Parilisia) punctigera (Lackschewitz, 1940)

Starý 2006, HA, Oasis Meski, AA; Koçak and Kemal 2010; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Hoplolabis (Parilisia) sororcula (Lackschewitz, 1940)

Driauach et al. 2013; Driauach and Belqat 2016, Rif, Barrage Ajras, Oued Ouringa, Oued El Kanar, Oued Maggou; Oosterbroek 2020 (CCW: Rif)

Idiocera Dale, 1842

Idiocera (Euptilostena) jucunda (Loew, 1873)

= Gonomyia jucunda Loew, in Pierre 1924a: 201, Séguy 1930a: 22

Pierre 1924a, HA, Asni (1200 m); Séguy 1930a, HA, Asni; Savchenko et al. 1992; Dakki 1997; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Idiocera (Idiocera) ampullifera (Starý, 1979)

Driauach and Belqat 2016, AA, Oued Zag; Oosterbroek 2020 (CCW: AA)

Idiocera (Idiocera) pulchripennis (Loew, 1856)

= Gonomyia sexpunctata Dale, in Pierre 1922b: 148

= Gonomyia pulchripennis (Loew), in Dakki 1997: 62

Pierre 1922b, AP, Atlantic coast; Ramdani 1981, AP, Merja Sidi Boughaba; Savchenko et al. 1992; Dakki 1997; Koçak and Kemal 2010; Driauach et al. 2013; Driauach and Belqat 2016, Rif, Oued Jnane Niche, Oued Aârkoub, Aïn Jdioui; Oosterbroek 2020 (CCW)

Idiocera (Idiocera) sziladyi (Lackschewitz, 1940)

Driauach and Belqat 2016, Rif, Oued Zarka; Oosterbroek 2020 (CCW: Rif)

Ilisia Rondani, 1856

Ilisia maculata (Meigen, 1804)

Driauach and Belqat 2016, Rif, Oued Maggou, EM, Oued Tafoughalt; Oosterbroek 2020 (CCW: Rif; EM)

Molophilus Curtis, 1833

Molophilus (Molophilus) ibericus Starý, 2011

Starý 2011, HA; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Molophilus (Molophilus) obscurus (Meigen, 1818)

Starý and Oosterbroek 2008, HA, Massif Toubkal, Oukaimeden, 2500–2800 m; Driauach et al. 2013; Driauach and Belqat 2016, Rif, tributary of Oued Ouara, tributary of Oued Maggou, Dayat El Ânassar, Dayat Jebel Zemzem, Aïn El Ma Bared, Dayat Rmali, Oued Amsemlil, Aïn Mâaze, Dayat Afrate, Dayat Lemtahane; Oosterbroek 2020 (CCW: Rif)

Molophilus (Molophilus) propinquus propinquus (Egger, 1863)

Savchenko et al. 1992; Starý and Oosterbroek 2008, HA, Oukaimeden (2500–2800 m); Starý 2011; Driauach et al. 2013; Driauach and Belqat 2016, Rif, Oued Ouara, Oued Farda, tributary of Oued Taida, Oued Maggou; Oosterbroek 2020 (CCW: Rif)

Molophilus (Molophilus) testaceus Lackschewitz, 1940

Driauach and Belqat 2016, Rif, Dayat Amsemlil, Dayat Lemtahane, Marj El kheyl, Oued Tkarae, Dayat near Aïn Afersiw, Dayat Mezine, Dayat Tazia, Dayat Rmali, Dayat Amsemlil, Dayat near Aïn Afersiw; Oosterbroek 2020 (CCW: Rif)

Symplecta Meigen, 1830

Symplecta (Symplecta) grata Loew, 1873

Ebejer et al. 2020, Rif, Aïn Jdioui (Tahaddart, 8 m)

Symplecta (Symplecta) hybrida (Meigen, 1804)

Lackshewitz 1940a, HA, Goundafa (1200 m); Séguy 1941a, HA, Taroudant; Savchenko et al. 1992; Oosterbroek et al. 2007; Koçak and Kemal 2010; Driauach et al. 2013; Driauach and Belqat 2016, Rif, Barrage Ajras, Oued El Kanar, Oued Aârkoub, Oued Jnane Niche, MA, Barrage Allal El Fassi; Oosterbroek 2020 (CCW: Rif; MA)

Symplecta (Trimicra) pilipes (Fabricius, 1787)

= Trimicra pilipes Fabricius, in Pierre 1922a: 23

= Trimicra andalusiaca Strobl, in Pierre 1922b: 149

= Trimicra hirsutipes Macquart, in Séguy 1930a: 22

Pierre 1922a, AP, Dradek (near Rabat), HA, Marrakech; Pierre 1922b, MA, Volubilis, HA, Oued Tensift; Séguy 1930a; Séguy 1941d, AA, Taroudant; Savchenko et al. 1992; Dakki 1997; Pârvu and Zaharia 2007; Driauach et al. 2013; Driauach and Belqat 2016, Rif, tributary of Oued Hachef, Oued Aârk­ob, Oued El Kanar, Oued Mezine, AP, Aïn Chouk (Larache); Oosterbroek 2020 (CCW: Rif)

Tasiocera (Dasymolophilus) murina (Meigen, 1818)

Driauach and Belqat 2016, Rif, Oued Farda, Oued Amsemlil; Oosterbroek 2020 (CCW: Rif)

Dactylolabinae

Dactylolabis Osten-Sacken, 1860

Dactylolabis (Dactylolabis) symplectoidea Egger, 1863

Pierre 1922a, AP, Around Casablanca (coastal meseta); Savchenko et al. 1992; Dakki 1997; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Limnophilinae

Austrolimnophila Alexander, 1920

Austrolimnophila (Austrolimnophila) latistyla Starý, 1977

Driauach and Belqat 2016, Rif, Oued Maggou; Oosterbroek 2020 (CCW: Rif)

Dicranophragma Osten-Sacken, 1860

Dicranophragma (Brachylimnophila) adjunctum (Walker, 1848)

= Neolimnomyia adjuncta (Walker, 1848), in Pârvu et al. 2006: 273

Pârvu et al. 2006, AP, Merja Zerga; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Dicranophragma (Brachylimnophila) nemorale (Meigen, 1818)

Lackschewitz 1940b, HA, Tachdirt (2200–2700 m); Savchenko et al. 1992; Driauach et al. 2013; Driauach and Belqat 2016, Rif, Aïn Sidi Brahim Ben Arrif, tributary Oued Ouara, Oued Maggou; Oosterbroek 2020 (CCW: Rif)

Eloeophila Rondani, 1856

Eloeophila maroccana Starý, 2009

Starý 2009b, HA, Okaïmeden (2500–2800 m); Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Euphylidorea Alexander, 1972

Euphylidorea (Euphylidorea) crocotula (Séguy, 1941)

= Phylidorea crocotula (Séguy), in Séguy 1941a: 28

Séguy 1941a, HA, Tachdirt (Toubkal, 2500 m); Savchenko et al. 1992, HA, Tachdirt (Toubkal); Starý and Oosterbroek 2008, HA, Oukaimeden (2500 m–2800 m), Imlil (1400 m); Driauach et al. 2013; Driauach and Belqat 2016, Rif, Aïn Bab Tariouant; Oosterbroek 2020 (CCW: Rif; HA)

Euphylidorea (Euphylidorea) dispar (Meigen, 1818)

Driauach and Belqat 2016, Rif, Oued at 15 km from Fifi; Oosterbroek 2020 (CCW: Rif)

Euphylidorea (Euphylidorea) lineola (Meigen, 1804)

Lackschewitz 1940b, HA; Savchenko et al. 1992; Starý and Freidberg 2007; Driauch et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Hexatoma Latreille, 1809

Hexatoma (Hexatoma) bicolor (Meigen, 1818)

Driauach and Belqat 2016, Rif, Oued at 15 km from Fifi, tributary of Oued Ouara, Oued Madissouka, Oued Maggou, Aïn Quanquben (Jebel Bou Bessoui), Oued Tkarae, Oued Tamerte; Oosterbroek 2020 (CCW: Rif)

Hexatoma (Hexatoma) gaedii (Meigen, 1830)

Lackschewitz 1940b, HA, Tachdirt (2200 m–2700 m); Savchenko et al. 1992; Starý and Freidberg 2007; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW: HA)

Pseudolimnophila Alexander, 1919

Pseudolimnophila (Pseudolimnophila) sepium (Verrall, 1886)

Driauach et al. 2013, Rif, Guelta Tazia; Driauach and Belqat 2016, Rif, Dayat Tazia, Aïn Sidi Brahim Ben Ar­rif, Aïn Afersiw, Oued Maggou, Dayat Tazia, Oued Taida, Aïn Sidi Brahim Ben Arrif; Oosterbroek 2020 (CCW: Rif)

Limoniinae

Dicranomyia Stephens, 1829

Dicranomyia (Dicranomyia) affinis (Schummel, 1829)

Driauach et al. 2013; Driauach and Belqat 2016, Rif, Oued Amsemlil, Dayat Lemtahane, Oued Tkarae, Marj El Kheyl, Aïn Sidi Brahim Ben Arrif, Dayat Tazia, Aïn El Maounzil, Aïn El Malâab, Dayat near Aïn Afersiw, Aïn Bab Tariouant; Oosterbroek 2020 (CCW: Rif)

Dicranomyia (Dicranomyia) chorea (Meigen, 1818)

Pierre 1922b, HA, Haute Réghaya, Tannaout, Asni (1000 m–1250 m); Savchenko et al. 1992; Dakki 1997; Starý and Freidberg 2007; Driauach et al. 2013; Driauach and Belqat 2016, Rif, Oued Farda, Oued Kelâa, Aïn Ras el Ma, Oued Jnane Niche, Cascade Chrafate, maison forestière, Oued El Kanar, tributary of Oued Zarka, HA, Oued Sidi Fares (National Park of Toubkal); Oosterbroek 2020 (CCW: Rif; HA)

Dicranomyia (Dicranomyia) didyma (Meigen, 1804)

Pierre 1922b, HA, Haute Réghaya (2000 m); Dakki 1997; Savchenko et al. 1992; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW: HA)

Dicranomyia (Dicranomyia) goritiensis (Mik, 1864)

Lackschewitz 1940b; Vaillant 1956b; Savchenko et al. 1992; Driauach et al. 2013; Driauach and Belqat 2016, Rif, Oued Maggou, Cascade Chrafate, Oued El Koub, Cascade Zarka, Âounsar Aheramen, maison forestière; Oosterbroek 2020 (CCW: Rif)

Dicranomyia (Dicranomyia) longicollis (Macquart, 1846)

= Telecephala longicollis (Macquart), in Pierre 1922a: 21, Pierre 1922b: 148, Séguy 1930a: 22

Séguy 1930a, AP, Dradek (near Rabat); Dakki 1997; Pierre 1922a, AP, Dradek, HA, Marrakech; Pierre 1922b, AP, Rabat; Savchenko et al. 1992; Driauach et al. 2013; Driauach and Belqat 2016, Rif, Oued Aârate, Barrage Moulay Bouchta, Oued Kbir, MA, Barrage Allal El Fassi; Oosterbroek 2020 (CCW)

Dicranomyia (Dicranomyia) mitis (Meigen, 1830)

= Dicranomyia hygropetrica Vaillant, in Vaillant 1956b: 42

Vaillant 1956b, HA, Asif Tessaout (M’Goum), Izourar, Tahanaout, Tamesrit, Imi-N’Ifri, Aguelmous, Sidi Chamarouch, Lac Tamhda (Anremer), Oukaimeden; Savchenko et al. 1992; Pârvu et al. 2006, AA, near Agadir, Souss plain to High Atlas occidental; Pârvu and Zaharia 2007; Gavryushin pers. comm. 2012, HA; Driauach et al. 2013, Rif, Tazia, Tisgris; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Dicranomyia (Dicranomyia) modesta (Meigen, 1818)

Driauach and Belqat 2016, Rif, Oued Farda, Oued El Kanar, Oued Jnane Niche, Oued Sidi Yahia Aârab; Oosterbroek 2020 (CCW: Rif)

Dicranomyia (Dicranomyia) novemmaculata (Strobl, 1906)

Driauach and Belqat 2016, Rif, tributary of Oued el Fondak, Oued Aârate; Oosterbroek 2020 (CCW: Rif)

Dicranomyia (Dicranomyia) ventralis (Schummel, 1829)

Driauach and Belqat 2016, Rif, Lake Badriouen; Oosterbroek 2020 (CCW: Rif)

Dicranomyia (Glochina) sericata (Meigen, 1830)

Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Dicranomyia (Melanolimonia) hamata Becker, 1908

Driauach and Belqat 2016, Rif, tributary of Oued Kbir, Oued Aârate, Aïn Sidi Brahim Ben Arrif, Dayat Tazia; Oosterbroek 2020 (CCW: Rif)

Dicranomyia (Melanolimonia) morio (Fabricius, 1787)

= Limonia pauliani Séguy, in Séguy 1941a: 26

Séguy 1941a, HA, Tachdirt (Toubkal, 2500 m); Savchenko et al. 1992, HA, Tachdirt (Toubkal); Driauach et al. 2013; Driauach and Belqat 2016, Rif, Seguia Lemtahane, Dayat near Aïn Afersiw, Dayat Jebel Zemzem; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Dicranomyia majuscula Pierre, 19241

Pierre 1924a, HA, Haut Imminen (2400 m); Séguy 1930a, HA, Haut Imminen; Dakki 1997; Savchenko et al. 1992, HA, Haut Imminen; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Dicranoptycha Osten-Sacken, 1860

Dicranoptycha fuscescens (Schummel, 1829)

Eiroa 2000, Rif; Driauach et al. 2013; Driauach and Belqat 2016, Rif, Oued Mlilah, Oued Zendoula, AP, Oued Loukous; Oosterbroek 2020 (CCW)

Geranomyia Haliday, 1833

Geranomyia caloptera (Mik, 1867)

Driauach et al. 2013, HA, Setti Fatma; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Geranomyia obscura Strobl, 1900

Vaillant 1956b, HA, Lac Tamhda (Anremer), Oukaimeden, Izourar, Sidi Chamarouch, Tamesrit; Savchenko et al. 1992; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW: HA)

Helius Lepeletier & Serville, 1828

Helius (Helius) hispanicus Lackschewitz, 1928

Starý and Oosterbroek 2008, HA, Massif Toubkal, Imlil (17 km S Asni, 1700–1900 m); Driauach et al. 2013; Driauach and Belqat 2016, Rif, Oued Amsemlil; Oosterbroek 2020 (CCW: Rif)

Helius (Helius) pallirostris Edwards, 1921

Driauach and Belqat 2016, AP, Aïn Chouk (Larache); Oosterbroek 2020 (CCW: Rif)

Limonia Meigen, 1803

Limonia flavipes (Fabricius, 1787)

Pierre 1922b, HA, Haute Réghaya, Asni (1250 m); Savchenko et al. 1992; Dakki 1997; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Limonia hercegovinae (Strobl, 1898)

Starý and Oosterbroek 2008, MA, Ifrane (1700 m), HA; Gavryushin pers. comm. 2012, HA; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW: HA)

Limonia macrostigma (Schummel, 1829)

Starý and Oosterbroek 2008, HA, 5 km from Oukaimeden (2350 m); Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Limonia nubeculosa Meigen, 1804

Pierre 1922b, HA, Haute Réghaya, Asni (1250 m); Savchenko et al. 1992; Dakki 1997; Gavryushin pers. comm. 2012, HA; Driauach and Belqat 2016, Rif, tributary of Oued El Fondak, Aïn Ras el Ma, Aïn Boughaba, Âounsar Aheramen, Aïn Takhninjoute, maison forestière, Oued Madissouka, Aïn Quan­quben (Jebel Bou Bessoui), EM, Oued Azila; Oosterbroek 2020 (CCW: Rif; HA)

Limonia phragmitidis (Schrank, 1781)

Starý and Oosterbroek 2008, MA, Ifrane (1700 m); Driauach and Belqat 2016, Rif, Aïn Quanquben (Jebel Bou Bessoui); Oosterbroek 2020 (CCW)

PEDICIIDAE

K. Kettani, P. Oosterbroek

Number of species: 6. Expected: 10

Faunistic knowledge of the family in Morocco: moderate

Pediciinae

Dicranota Zetterstedt, 1838

Dicranota (Dicranota) bimaculata (Schummel, 1829)

Pierre 1922b, HA, Haute Réghaya (1800 m); Savchenko et al. 1992 (? Morocco); Dakki 1997; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Dicranota (Dicranota) irregularis Pierre, 1921

Pierre 1922b, HA, Haute Réghaya (1800 m); Savchenko et al. 1992, HA, Cirque d’Arround (Haute Réghaya); Dakki 1997; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Dicranota (Ludicia) claripennis (Verrall, 1888)

Driauach and Belqat 2016, Rif, Oued Amsemlil, maison forestière; Oosterbroek 2020 (CCW)

Dicranota (Paradicranota) candelisequa Starý, 1981

Pârvu et al. 2006, AP, Merja Zerga; Driauach et al. 2013; Driauach and Belqat 2016; Oosterbroek 2020 (CCW)

Dicranota (Paradicranota) landrocki Czižek, 1931

Driauach et al. 2013, Rif, Fifi (1252 m); Driauach and Belqat 2016, Rif, Oued Ouara, Oued Taida, Aïn Sidi Brahim Ben Arrif, Âounsar Aheramen, Oued Tizekhte, Oued Mezine, maison forestière, Aïn Bab Tariouant, HA, Imlil (Assif Haouz); Oosterbroek 2020 (CCW)

Tricyphona Zetterstedt, 1838

Tricyphona (Tricyphona) immaculata (Meigen, 1804)

Driauach and Belqat 2016, Rif, maison forestière, Dayat Lemtahane; Oosterbroek 2020 (CCW)

TIPULIDAE

K. Kettani, P. Oosterbroek, H. de Jong

Number of species: 39. Expected: 42

Faunistic knowledge of the family in Morocco: moderate

Dolichopezinae

Dolichopeza Curtis, 1825

Dolichopeza (Dolichopeza) hispanica Mannheims, 1951

Theowald and Oosterbroek 1980, HA, Aghbalou, Oukaimeden, Imlil, Tadmant; Oosterbroek and Theowald 1992; Oosterbroek and Lantsov 2011, HA, Oukaimeden (2300 m), Aghbalou (Massif Toubkal), 43 km S Marrakech (1000 m), Imlil (17 km S Asni, 1700–1900 m), Tadmant (17 km E Asni); Adghir et al. 2018, Rif, Kitane, Aîn Ras el Ma (Chefchaouen); Oosterbroek 2020 (CCW)

Tipulinae

Nephrotoma Meigen, 1803

Nephrotoma alluaudi (Pierre, 1922)

= Pachyrhina lunulicornis Schummel, in Pierre 1922a: 24

= Pachyrhina alluaudi Pierre, in Pierre 1922b: 150, Dakki 1997: 62

= Pales alluaudi (Pierre), in Mannheims 1951: 47

Pierre 1922b, HA, Tannaout (1000 m); Mannheims 1951, Rif, Beni Seddat, AP, Lagune Guedira; HA, Taddert north of Marrakech, Goundafa (1200 m); AA, Llano Amarillo, Tlata Reisana; Oosterbroek 1979b, Rif, HA, Imlil (1400 m), Tizi-n’Tichka (2200 m); Theowald and Oosterbroek 1980, AP, Rabat, Guedira lagoon, MA, Immouzer, Ifrane, Timahdit, Aghbalou, Tizi-n’Zou, HA, Marrakech, Goundafa, Taddert, Dayat, Tizi-n’Test, Tizi-n’Tichka, Asni, Imlil, Oukaimeden, Setti Fatma, Tinmel, Acif Tifni, AA, Taroudant, Mikdana, Sidi Said bou Merdoul, Tlata Reisana, Llano Amarillo; Eiroa 1990, MA, Azrou, Ajabo; Oosterbroek and Theowald 1992, HA, Tannaout; Dakki 1997; Mouna 1998; Adghir et al. 2018, Rif, Ras el Ma (Chefchaouen), Oued Laou, Dardara, Douar Mouarâa, Oued Zandoula, Laghmari-Rmal, Douar Louamera, Douar Laheyayda, Oued Jnane Niche, Cabo Negro, Oued Beni Said, Oued El Kanar, Oued Amsemlil; Oosterbroek 2020 (CCW) – MISR (HA), MHNV, MNCNM, MAKB

Nephrotoma appendiculata pertenua Oosterbroek, 1978

Oosterbroek 1978, Rif, 9 km SW Chefchaouen; Theowald and Oosterbroek 1980, Rif, Dardara, MA, Fès, Immouzer, Khemisset; Oosterbroek and Theowald 1992; de Jong 1993, 1998, Rif; Adghir et al. 2018, Rif, Oued Ametrasse, Dayat Aïn Jdioui, Barrage Moulay Bouchta, Oued Sahil, Dayat Jebel Zemzem, Aïn Sidi Brahim Ben Arrif, Oued Nakhla, Oued Tizekhte, Lot Hemmadi, Douar Ayacha, Douar Louamera, Dayat Mezine, Aïn El Malâab, Aïn Takhninjoute, Dayat Tazia, Oued Taida, maison forestière Tazia, Tourbière Amesmlil, Oued El Hamma, Oued Kbir, Jebel Lakraâ; Oosterbroek 2020 (CCW)

Nephrotoma astigma Pierre, 1925

Theowald and Oosterbroek 1980, Rif, Dardara, MA, Taza; Adghir et al. 2018, Rif, Oued Tabandout, Etang Maggou, Aïn El Malâab, Douar Remla; Oosterbroek 2020 (CCW)

Nephrotoma fontana Oosterbroek, 1978

de Jong 1998, Rif; Adghir et al. 2018, Rif, Ketama, maison forestière Tazia, Dayat Tazia; Oosterbroek 2020 (CCW)

Nephrotoma guestfalica vaillanti de Jong, Adghir & Bosch, 2021

de Jong et al. 2021, Rif, Ras el Ma (Chefchaouen, 500 m), Fomento, Oued Laou (6 km north-west of Chefchaouen, 200 m), Jebel Tissouka (3 km south of Chefchaouen, 500 m; 3 km south of Chefchaouen, 700 m; 4 km south-east of Chefchaouen, 700–800 m; 5 km southeast of Chefchaouen, 700–900 m; 5 km south of Chefchaouen, 900–1000 m), Bab Taza (25 km south-east of Chefchaouen, 750–800 m), Dardara (10 km south of Chefchaouen, 300 m), Oued Martil, Nakhla, Âounsar Aheramen, Boumerouil, Oued Maggou, Oued Maggou (Aïn Ras el Ma), Douar Kitane, Oued Sidi Yahia Aârab, Oued Tamerte, Oued Zandoula, Oued El Hamma, Lot Hemmadi, Oued Tamerte, Oued Sidi Mohamed Saâda, Oued El Koub, Douar Iholebatine, Dayat Tazia, Belouazen, Oued Lemtahane, Oued Siflaou, Oued Amsemlil, AP, Oued Loukous, Aïn el-Aouda, MA, Ifrane (road to Mischliffen, 1680 m), N.S. and W of Ifrane (1400–1800 m), Khemisset, Azrou, Oum-er-Rbia, HA, M’semrir (bord de l’Oued), Rich, Haute Rhégaya (identified by Mannheims in 1951 as surcoufi)

Nephrotoma luteata (Meigen, 1818)

Oosterbroek 1979a, Rif, Targuist, Chefchaouen, HA, Kasba Taguendaft, near Oukaimeden; Theowald and Oosterbroek 1980, Rif, Targuist, Chefchaouen, HA, Kasba Taguendaft, near Oukaimeden; Oosterbroek and Theowald 1992; Pârvu et al. 2006, AP, Merja Zerga; Adghir et al. 2018, Rif, Oued Laou, Dardera, Oued Nakhla; Oosterbroek 2020 (CCW)

Nephrotoma subanalis (Mannheims, 1951)

= ? Pachyrhina analis Schummel, in Pierre 1922a: 24, Pierre 1922b: 150

= Pales subanalis Mannheims, in Mannheims 1951: 56

Mannheims 1951, HA, Tachdirt (2200–2900 m); Vaillant 1956b, HA, Oukaimeden (2250 m); Oosterbroek 1979c, HA, Tachdirt (2200–2900 m); Theowald and Oosterbroek 1980, HA, Tachdirt, Oukaimeden, M’Goum, Tadmant, Imlil, Tizi-N’Tichka; Oosterbroek and Theowald 1992, HA, Tachdirt; Oosterbroek 2020 (CCW)

Nephrotoma submaculosa Edwards, 1928

Oosterbroek 1982, Rif, Ketama, Dardara, MA, Azrou; Oosterbroek and Theowald 1992; de Jong 1998, Rif, Atlas; Oosterbroek 2011; Adghir et al. 2018, Rif, Jebel Dahedouh, Ketama, Aïn Sidi Brahim Ben Arrif, Oued Taida, tributary of Oued Ouara, Sidi Chouiref, Dayat Tazia; Oosterbroek 2020 (CCW)

Nephrotoma sullingtonensis Edwards, 1938

Oosterbroek 1978; Theowald and Oosterbroek 1980, Rif, Bab Berred; Oosterbroek 1982 Rif, Bab Berred; de Jong 1993, 1998, Rif; Oosterbroek and Theowald 1992; Adghir et al. 2018, Rif, 4 km SE Ketama, Barrage Moulay Bouchta, Oued Aârate, Aïn Takhninjoute, Oued Jbara, Aïn El Malâab, Aïn El Ma Bared, Oued Lemtahane, maison forestière Tazia, Oued Taida, Dayat Tazia; Oosterbroek 2020 (CCW)

Tipula Linnaeus, 1758

Tipula (Acutipula) anormalipennis Pierre, 1924

Pierre 1924b, HA, Haut Imminen; Séguy 1930a, HA, Haut Imminen; Séguy 1941a, HA, Tachdirt (2500 m), Haut Imminen; Mannheims 1952, HA, Tachdirt (2500 m), Imminen (2400–2500 m), Arround (1950 m); Vaillant 1956b, HA, Lake of Tamhda (Anremer, 2900 m); Theowald and Oosterbroek 1980, HA, Anremer, Haut-Immenen, Tachdirt, Arround, Oukaimeden; Vermoolen 1983, HA, Haut Imminen, Oukaimeden (2500–2800 m), Arround (1950 m), Tachdirt (2500 m); Oosterbroek and Theowald 1992; de Jong 1994a, HA; de Jong 1998, HA; Dakki 1997; Oosterbroek 2020 (CCW)

Tipula (Acutipula) repentina Mannheims, 1952

= Tipula maxima Poda, in Séguy 1941a: 26

Séguy 1941a, HA, Tachdirt (2500 m); Mannheims 1952, HA, Tachdirt (2200–2700 m); Vaillant 1956b, HA, Lac Tamhda (Anremer, 2900), M’Goum (2500 m); Theowald and Oosterbroek 1980, HA, Anremer, M’Goum, Tachdirt, Tizi-N’Tichka, Asni, Oukaimeden, Setti Fatma, Imlil, Tadmant; Vermoolen 1983, MA, Ifrane, HA, Androment, M’Goum, Tadmant, Tizi-N’Test, Setti Fatma, Oukaimeden, Tizi-N’Tichka; Oosterbroek and Theowald 1992, HA, Tachdirt; de Jong 1994a, MA, HA; de Jong 1998, HA; Dakki 1997; Oosterbroek 2020 (CCW)

Tipula (Acutipula) rifensis Theowald & Oosterbroek, 1980

Theowald and Oosterbroek 1980, Rif, Targuist; Vermoolen 1983, Rif, Targuist, Tidiguin (90 km E. of Ouezzane, 2350 m); Oosterbroek and Theowald 1992, Rif; de Jong 1994a, 1998, Rif; Adghir et al. 2018; Oosterbroek 2020 (CCW)

Tipula (Emodotipula) leo Dufour, 1991

= Tipula (Emodotipula) obscuriventris Strobl2, in Oosterbroek and Theowald 1992: 99

Oosterbroek and Theowald 1992 (?); Dufour 2003, Rif; Adghir et al. 2018, Rif, Jebel Tissouka; Oosterbroek 2020 (CCW)

Tipula (Lunatipula) bivittata Pierre, 1922

Pierre 1922a, AP, Maâmora between Kénitra and Oued Beth, Dradek (near Rabat); Pierre 1922b, AP, Rabat; Mannheims 1968; Theowald and Oosterbroek 1980, AP, forest of Maâmora, Dradek; Oosterbroek and Theowald 1992, AP, forest of Maâmora, Dradek; Dakki 1997; Oosterbroek 2020 (CCW) – MISR (AP, Kénitra, Dradek)

Tipula (Lunatipula) cinereicolor Pierre, 1924

Pierre 1924b, HA, Haut Imminen; Séguy 1930a, HA, Tachdirt (3100–3200 m); Theowald 1973, HA, Haut Imminen (2400 m); Theowald and Oosterbroek 1980, MA, Ifrane, HA, Oukaimeden, Tachdirt, Haut-Imminen; Eiroa 1990, MA, Ajabo; Oosterbroek and Theowald 1992, HA, Imminen; Dakki 1997; Oosterbroek 2020 (CCW)

Tipula (Lunatipula) cornicula Pierre, 1922

Pierre 1922b, HA, Arround (2000 m); Séguy 1930a, HA, Tachdirt (3100–3200 m); Theowald 1973, HA, Cirque d’Arround (2000 m), Tachdirt (2200–2700 m), Goundafa (1200 m); Theowald and Oosterbroek 1980, HA, Oukaimeden, Tachdirt, Goundafa; Oosterbroek and Theowald 1992, HA, Tachdirt; Dakki 1997; Oosterbroek 2020 (CCW) – MISR (HA)

Tipula (Lunatipula) fabiola Mannheims, 1968

Theowald and Oosterbroek 1980, Rif, Bab Berred, Ras El Ma (Chefchaouen), MA, Jebel Abad, Ifrane; Oosterbroek and Theowald 1992; Adghir et al. 2018; Oosterbroek 2020 (CCW)

Tipula (Lunatipula) hermes Theischinger, 1977

Theischinger 1977, Rif, north of Ouezzane; Theowald 1980; Theowald and Oosterbroek 1980, Rif, north of Ouezzane; Oosterbroek and Theowald 1992, Rif, Ouezzane; Adghir et al. 2018, Rif, 4 km SE Ketama, Oued Ametrasse, Aïn El Ma Bared; Oosterbroek 2020 (CCW)

Tipula (Lunatipula) iberica iberica Mannheims, 1963

= Tipula lunata Linnaeus, in Pierre 1922b: 149, Séguy 1930a: 23

Pierre 1922b, HA, Haute Réghaya, Tannaout (1000 m); Séguy 1930a, HA, Haut Imminen; Mannheims 1963, MA; Theowald and Oosterbroek 1980, Rif, Bab Berred, MA, Ifrane, Taounate; Eiroa 1990, MA, Ifrane; Oosterbroek and Theowald 1992; Oosterbroek 2009; Adghir et al. 2018, Rif, Ketama; Oosterbroek 2020 (CCW) – MISR (HA)

Tipula (Lunatipula) iberica spinula Theischinger, 1980

Theischinger 1980, HA, Oukaimeden (1300–2800 m); Theowald and Oosterbroek 1980, HA, Oukaimeden; Oosterbroek and Theowald 1992, HA, Oukaimeden; Oosterbroek 2020 (CCW)

Tipula (Lunatipula) peliostigma peliostigma Schummel, 1833

Eiroa 1990, MA, Azrou; Mouna 1997; Oosterbroek 2020 (CCW)

Tipula (Lunatipula) pjotri de Jong & Adghir, 2018

Adghir et al. 2018, Rif, Jebel El Kelâa (Talassemtane, 1340 m)

Tipula (Lunatipula) pseudocinerascens Strobl, 1906

Adghir et al. 2018, Rif, Stehat, Oued Taida, Perdicaris Park, Dayat Tazia

Tipula (Lunatipula) rocina Theischinger, 1979

Theowald and Oosterbroek 1980, Rif, Tétouan; Oosterbroek and Theowald 1992; Oosterbroek 2009; Adghir et al. 2018; Oosterbroek 2020 (CCW)

Tipula (Lunatipula) selenaria Mannheims, 1967

Mannheims 1967, MA, Jebel Tazzeka (1500–1989 m), HA, Goundafa (1200 m); Theowald and Oosterbroek 1980, MA, Tazzeka, HA, Foum Keneg, Goundafa, Oukaimeden; Eiroa 1990, MA, Ajabo; Oosterbroek and Theowald 1992, HA, Goundafa; de Jong 1995, MA, HA, Oukaimeden; Oosterbroek 2020 (CCW)

Tipula (Lunatipula) stimulosa Mannheims, 1973

Adghir et al. 2018, Rif, Jebel El Kelâa (Talassemtane, 1340 m)

Tipula (Lunatipula) subfalcata Mannheims, 1967

de Jong 1995, 1998, Rif; Adghir et al. 2018, Rif, Jebel Tissouka, Oued Tamerte, Oued El Koub; Oosterbroek 2020 (CCW)

Tipula (Lunatipula) subpustulata Mannheims, 1963

= Tipula pustulata Pierre, in Pierre 1922b: 150

Pierre 1922b, AP, Mogador; Vaillant 1956b, HA, M’Goun (2500 m); Mannheims 1963, AP, Aïn el Aouda, MA, Jebel Tazzeka (1600–1989 m), HA, Goundafa (1200 m), Tachdirt (2200–2900 m), AA, Lac Goulmima; Theowald 1972; Theowald and Oosterbroek 1980, AP, Aïn el Aouda, MA, Iebel Tazzeka, HA, M’Goum, Goundafa, Tachdirt, Oukaimeden, Imlil; Eiroa 1990, MA, Ajabo; Oosterbroek and Theowald 1992, HA, Goundafa; Dakki 1997; Adghir et al. 2018, Rif, 4 km SE Ketama, Aïn El Ma Bared; Oosterbroek 2020 (CCW) – MISR (AP, Mogador)

Tipula (Lunatipula) tazzekai Theowald, 1973

Theowald 1973, MA, Jebel Tazzeka (1600–1989 m); Theowald and Oosterbroek 1980, MA, Jebel Tazzeka; Eiroa 1990, MA, Jebel Hebri; Oosterbroek and Theowald 1992, MA, Jebel Tazzeka; Oosterbroek 2020 (CCW)

Tipula (Savtshenkia) atlas Pierre, 1924

Pierre 1924b, HA, Tachdirt (3100–3250 m); Séguy 1930a, HA, Tachdirt (3100–3200 m); Vaillant 1956b, HA, Cascade Siroua (3000 m), M’Goun (2500 m), Toubkal (3350 m), Lake of Tamhda (Anremer, 2900 m); Mannheims 1964, HA; Theowald 1973, HA; Theowald 1980, HA; Theowald and Oosterbroek 1980, HA, Toubkal, Sources de Tessaouts, M’Goum, Siroua, Anremer, Tachdirt, Oukaimeden, Tizi-N’Tichka, Tadmant; Eiroa 1990, MA, Oum-Er-Rbia; Oosterbroek and Theowald 1992, HA, Tachdirt; de Jong 1994b; de Jong 1998, Atlas; Dakki 1997; Oosterbroek 2020 (CCW)

Tipula (Savtshenkia) breviantennata Lackschewitz, 1933

de Jong 1998, Rif; Adghir et al. 2018, Rif, Ras el Ma (Chefchaouen), Oued Maggou, Douar Aouzighen; Oosterbroek 2020 (CCW)

Tipula (Savtshenkia) confusa van der Wulp, 1883

Adghir et al. 2018, Rif, maison forestière (Talassemtane)

Tipula (Savtshenkia) rufina rufina Meigen, 1818

Theowald and Oosterbroek 1980, HA, Oukaimeden; Theowald and Oosterbroek 1983, Rif, Atlas; Oosterbroek and Theowald 1992; Adghir et al. 2018, Rif, Ras el Ma (Chefchaouen); Oosterbroek 2020 (CCW)

Tipula (Tipula) mediterranea Lackschewitz, 1930

Vaillant 1956b, HA, Oukaimeden (2250 m), M’Goun (2500 m); Mannheims 1952; Theowald and Oosterbroek 1980, MA, Ifrane, HA, Oukaimeden, M’Goum, Asni, M’Semrir, Ifni, Bab-Rou-Idie, Tizi-N’Tichka, Setti Fatma; Theowald 1984, Rif; Eiroa 1990, MA, Azrou, Oum-Er-Rbia; Oosterbroek and Theowald 1992; Pârvu et al. 2006 AP, Merja Zerga; Adghir et al. 2018, Rif, Ras el Ma (Chefchaouen), Jebel Tissouka, Bab Taza, Dardara, 4 km SE Ketama, Aïn Afersiw, Douar Kitane, Dayat Jebel Zemzem, Wilaya (Tétouan), Aïn El Ma Bared, Oued Tamerte, Douar Louamera, Tourbière Amesmlil, Dayat Mezine, Hejar Nehal, tributary of Oued Ouara, Oued Tkarae, Oued Jnane Niche, Dayat Afrate, Oued Ametrasse; Oosterbroek 2020 (CCW)

Tipula (Tipula) oleracea Linnaeus, 1758

Mannheims 1952, Rif, Tlata Ketama; Theowald and Oosterbroek 1980, Rif, Tlata Ketama; Theowald 1984, Rif; Oosterbroek and Theowald 1992; Dakki 1997; Pârvu and Zaharia 2007; Oosterbroek 2011; Adghir et al. 2018, Rif, Oued Laou, Dardara, Jebel Tissouka, Ksar Rimal, 15 km from Fifi, Oued Aârate, Oued El Hamma, Douar Louamera, Oued Zaouya, Près de Beni Said, Wilaya (Tétouan), Tourbière Amesmlil, Oued Smir, Aïn Jdida; Oosterbroek 2020 (CCW) – MISR

Tipula (Vestiplex) vaillanti vaillanti Theowald, 1977

Adghir et al. 2018, Rif, Jebel El Kelâa (Talassemtane, 1340 m), Douar Kitane

Tipula (Yamatotipula) afriberia afriberia Theowald & Oosterbroek, 1980

Theowald and Oosterbroek 1980, HA, Oukaimeden; Oosterbroek and Theowald 1992, HA, Oukaimeden; Oosterbroek 1994a; Adghir et al. 2018, Rif, Jebel Tissouka, Dardara, Douar Mokedassen, Oued Zarka; Oosterbroek 2020 (CCW)

Tipula (Yamatotipula) barbarensis Theowald & Oosterbroek, 1980

= Tipula lateralis Meigen, in Pierre 1922a: 23, Pierre 1922b: 150, Séguy 1930a: 23, Séguy 1941a: 26, Mannheims 1952: 98 (in part), Vaillant 1956b: 238

Pierre 1922a, AP, Dradek (Rabat), MA, Azrou (riverside of Oued Tigrigra); Pierre 1922b, AP, Mogador, MA, Beni Méllal, HA, Asni; Séguy 1930a; Séguy 1941a, HA, Imi n’Ouaka (1500 m); Mannheims 1952; Vaillant 1956b, HA, Oukaimeden; Theowald and Oosterbroek 1980, Rif, Dardara, MA, Ifrane, Aghbalou, HA, Asni, Oukaimeden, Setti Fatma, Imlil, Tizi-N’Tichka, Tadmant, Tachdirt; Eiroa 1990, MA, Azrou, Oum-er-Rbia, Ifrane; Oosterbroek and Theowald 1992, HA, Setti Fatma; Oosterbroek 1994a, Rif, Dardara, MA, Ifrane, Aghbalou, HA, Oukaimeden, Setti Fatma, Imlil, Tizi-N’Tichka, Tadmant, Asni, Tachdirt; Dakki 1997; Adghir et al. 2018, Rif, Ras el Ma (Chefchaouen), Jebel Tissouka, Bab Taza, Dardara, 4 km SE Ketama, Aïn El Manzela, Aïn Bab Tariouante, Dayat Aïn Afersiw, Oued El Kanar, Dayat Afrate, Douar Kitane, Wilaya (Tétouan), 15 km from Fifi, Aïn Sidi Brahim Ben Arrif, Oued Nakhla, Âounsar Aheramen, Oued Boumerouil, Douar Zaouya, Oued Tizekhte, Oued Samsa, Dayat Aïn Jdioui, Douar Ouled Laghmari-Rmal, maison forestière Tazia, Hejar Nehal, Lot Hemmadi, tributary of Oued Ouara, Oued Sidi Mohamed Saâda, Oued Amsemlil, Tourbière Amesmlil, Beni Salah, Oued Jnane Niche, Oued Maggou, Souk Lhed Beni Darkoul, Oued Imassouden, Aïn Helouma, Source Zarka, Aïn Kchour; Oosterbroek 2020 (CCW) – MISR

Trichoceridoidea

TRICHOCERIDAE

K. Kettani, E. Krzemińska

Number of species: 8. Expected: 10

Faunistic knowledge of the family in Morocco: good

Trichocerinae

Trichocera Meigen, 1803

Trichocera (Trichocera) hiemalis (DeGeer, 1776)

Pierre 1922b (AP, Rabat); Dakki 1997; Mouna 1998; AP (Rabat) – MISR

Trichocera (Trichocera) marocana Driauach, Krzemińska & Belqat, 2015

Driauach et al. 2015, Rif, Oued Akrir

Trichocera (Saltrichocera) annulata Meigen, 1818

Driauach et al. 2015, Rif, affluent Oued Akrir, Dayat Fifi, Oued Taria, Oued Kelâa, Oued Ouara, Oued à 15 km de Fifi, Aïn Mâaze, Oued Maggou, Aïn Quanquben, Oued Tizekhte, Forêt Jebel Bouhachem, EM, Grotte du Chameau, Oued Zegzel, Aïn Sidi Yahia, Cascade Grotte des pigeons, Oued Tafoughalt

Trichocera (Saltrichocera) pappi Krzemińska, 2003

Driauach et al. 2015, Rif, Dayat Fifi, affluent Oued Akrir, Oued Amsemlil, Ruisseau Agoummir, maison forestière, EM, Cascade Grotte des Pigeons, MA, Seguia El Hajeb

Trichocera (Saltrichocera) saltator (Harris, 1776)

Driauach et al. 2015, Rif, Dayat Fifi, Oued à 15 km de Fifi, Forêt Jebel Bouhachem, maison forestière, EM, Cascade Grotte des pigeons

Trichocera (Saltrichocera) sardiniensis Petrašiūnas, 2009

Driauach et al. 2015, Rif, Oued à 15 km de Fifi, affluent Oued Akrir, Oued Amsemlil, maison forestière

Trichocera (Saltrichocera) regelationis (Linnaeus, 1758)

Driauach et al. 2015, Rif, Oued Ouara, Aïn el Ma Bared, Oued à 15 km de Fifi, Cascade Chrafate, Aïn Quanquben

Trichocera (Saltrichocera) rufescens Edwards, 1921

Driauach et al. 2015, Rif, affluent Oued Akrir, Dayat Fifi, Oued Tassikeste, Oued Farda, Aïn Mâaze

Psychodoidea

PSYCHODIDAE

K. Kettani, R. Wagner

Number of species: 51. Expected: 70

Faunistic knowledge of the family in Morocco: moderate

Phlebotominae

Phlebotomus Loew, 1845

Phlebotomus (Larroussius) ariasi Tonnoir, 1921

Gaud 1947a; Gaud and Laurent 1952, AP, Rabat; Bailly-Choumara et al. 1971, AP, MA, HA; Rioux et al. 1974; Mouna 1998; Guernaoui et al. 2005, MA, HA; Bounamous 2010; Boussaa et al. 2005; Boussaa et al. 2010

Phlebotomus (Larroussius) chadlii Rioux, Juminer & Gibily, 1966

Rioux et al. 1974, HA; Rioux et al. 1975; Croset et al. 1978; Mouna 1998; Bounamous 2010

Phlebotomus (Larroussius) langeroni Nitzulescu, 1930

Rislorcelli 1941, HA; Bailly-Choumara et al. 1971, AP, EM; Croset et al. 1978; Mouna 1998

Phlebotomus (Larroussius) longicuspis Nitzulescu, 1930

Rislorcelli 1941, HA; Gaud and Laurent 1952, AP, Rabat; Bailly-Choumara et al. 1971, Rif, EM, AP, MA, HA, AA; Rioux et al. 1974, HA; Croset et al. 1978; Mouna 1998; Guernaoui et al. 2005, Rif, Chefchaouen, HA, AA, Agadir; Boussaa et al. 2005; Boussaa 2008; Boussaa et al. 2008; Boussaa et al. 2010; Bounamous 2010; Zarrouk et al. 2016

Phlebotomus (Larroussius) mariae Rioux, Croset, Léger and Bailly-Choumara, 1974

Hervy et al. 1994; Rioux et al. 1974, HA; Mouna 1998; Guernaoui et al. 2005, MA, HA

Phlebotomus (Larroussius) perfiliewi Parrot, 1930

Rioux et al. 1977; Croset et al. 1978; Mouna 1998

Phlebotomus (Larroussius) perniciosus Newstead, 1911

Séguy 1930a; Gaud and Laurent 1952, AP, Rabat; Bailly-Choumara et al. 1971, AP, EM, MA, HA, AA; Mouna 1998; Guernaoui et al. 2005, Rif, Chefchaouen, HA; Boussaa et al. 2008; Bounamous 2010; Boussaa et al. 2010; Zarrouk et al. 2016

Phlebotomus (Paraphlebotomus) alexandri Sinton, 1928

Bailly-Choumara et al. 1971, AA; Abonnenc 1972; Rioux et al. 1974, HA; Croset et al. 1978; Mouna 1998; Colacicco-Mayhugh et al. 2010, EM; Boussaa et al. 2010; Bounamous 2010

Phlebotomus (Paraphlebotomus) chabaudii Croset, Abonnenc & Rioux, 1970

Rioux et al. 1974, HA; Rioux et al. 1975; Mouna 1998

Phlebotmus (Paraphlebotomus) kazeruni Theodor & Mesghali, 1964

Mouna 1998

Phlebotomus (Paraphlebotomus) riouxi Depaquit, Killick-Kendrick & Léger, 1998

Bounamous 2010

Phlebotomus (Paraphlebotomus) sergenti Parrot, 1917

Séguy 1930a; Rislorcelli 1941; Rislorcelli 1947; Gaud and Laurent 1952, AP, Rabat; Bailly-Choumara et al. 1971, Rif, AP, EM, MA, HA, AA; Abonnenc 1972; Rioux et al. 1974, HA; Mouna 1998; Boussaa et al. 2009; Bounamous 2010

Phlebotomus (Phlebotomus) bergeroti Parrot, 1934

Rioux et al. 1975, HA; Mouna 1998; Bounamous 2010

Phlebotomus (Phlebotomus) papatasi (Scopoli, 1786)

Séguy 1930a; Rislorcelli 1941, Rislorcelli 1947, HA; Bailly-Choumara et al. 1971, AP, EM, MA, HA, AA; Abonnenc 1972; Rioux et al. 1974, HA; Croset et al. 1978; Mouna 1998; Boussaa et al. 2005; Boussaa 2008; Colacicco-Mayhugh et al. 2010, Mediterranean region; Bounamous 2010; Boussaa et al. 2010; Prudhomme et al. 2012

Phlebotomus clydei Sinton, 1928

Bailly-Choumara et al. 1971, EM; Mouna 1998

Phlebotomus lewisi Parrot, 1948

Bailly-Choumara et al. 1971, South AP; Mouna 1998

Sergentomyia França & Parrot, 1920

Sergentomyia (Grassomyia) dreyfussi (Parrot, 1933)

Rislorcelli 1941, Rislorcelli 1947, HA; Bailly-Choumara et al. 1971, AP, EM, MA, HA; Abonnenc 1972; Rioux et al. 1974, HA; Croset et al. 1978; Mouna 1998; Boumezzough et al. 2009, HA, Marrakech; Bounamous 2010

Sergentomyia (Parrotomyia) africana (Newstead, 1912)

= Phlebotomus (Parrotomyia) africana (Newstead), in Rislorcelli 1941: 522, Bailly-Choumara et al. 1971: 454; Rioux et al. 1974: 99

Séguy 1930a (as subspecies of minutus Rondani: 43), HA, Marrakech; Rislorcelli 1941 (as subspecies of minutus Rondani: 528), AA, Ksar es Souk; Gaud and Laurent 1952 (as subspecies of minutus Rondani: 75), AP, Rabat; Bailly-Choumara et al. 1971 (as subspecies of minutus Rondani: 438), AP, AA; Rioux et al. 1974, HA; Croset et al. 1978 (as subspecies of minutus Rondani: 722); Mouna 1998; Boussaa et al. 2005; Boumezzough et al. 2009, HA, Marrakech; Bounamous 2010; Boussaa et al. 2010

Sergentomyia (Sergentomyia) antennata (Newstead, 1912)

= Phlebotomus cinctus Parrot & Martin, 1944, in Mouna 1998: 86

= Phlebotomus signatipennis Newstead, 1920, in Mouna 1998: 86

Bailly-Choumara et al. 1971, EM; Rioux et al. 1974, HA; Léger et al. 1974, AA (south of Morocco); Mouna 1998

Sergentomyia (Sergentomyia) fallax (Parrot, 1921)

Rislorcelli 1947; Gaud 1954; Bailly-Choumara et al. 1971, AP, EM, MA, AA; Rioux et al. 1974, HA; Mouna 1998; Guernaoui et al. 2005; Boussaa et al. 2005; Boussaa et al. 2007; Boumezzough et al. 2009, HA, Marrakech; Boussaa et al. 2010; Bounamous 2010

Sergentomyia (Sergentomyia) minuta (Rondani, 1843)

= Phlebotomus minutus Rondani, in Gaud and Laurent 1952: 75, Mouna 1998: 86

= Phlebotomus (Sergentomyia) parroti (Adler and Theodor), in Rislorcelli 1941: 526, Rislorcelli 1947: 487, Bailly-Choumara et al. 1971: 450, Rioux et al. 1974, 99

Séguy 1930a, HA, Marrakech; Gaud and Laurent 1952, AP, Rabat; Rislorcelli 1941; Rislorcelli 1947, HA; Bailly-Choumara et al. 1971, Rif, AP, EM, MA, HA; Rioux et al. 1974, HA; Croset et al. 1978 (from the mediterranean region to the Sahara); Mouna 1998; Boussaa et al. 2005; Boumezzough et al. 2009, HA, Marrakech; Bounamous 2010; Boussaa et al. 2010; Depaquit et al. 2015, Rif, Chefchaouen

Sergentomyia (Sergentomyia) schwetzi Adier, Theodor & Parrot, 1929

Bailly-Choumara and Léger 1976, SA, Aouinet-Torkoz; Mouna 1998

Sergentomyia (Sintonius) christophersi (Sinton, 1927)

Rioux et al. 1974, HA; Rioux et al. 1975; Croset et al. 1978; Mouna 1998

Psychodinae

Maruinini

Tonnoiriella Vaillant, 1982

Tonnoiriella paveli Ježek, 1999

Ježek 1999, HA, AA; Afzan and Belqat 2016

Tonnoiriella pulchra (Eaton, 1893) (?) [probably mis-identification]

Wagner 1990; Ježek and Hájek 2007; Afzan and Belqat 2016

Mormiini

Mormia Enderlein, 1935

Mormia tenebricosa Vaillant, 1954

= Telmatoscopus (Mormia) tenebricosus Vaillant, in Vaillant 1956b: 244

Vaillant 1956b, HA, Imi-N’Ifri; Afzan and Belqat 2016, Rif, Oued Achekrade

Paramormiini

Clogmia Enderlein, 1937

Clogmia albipunctata (Williston, 1893)

Afzan and Belqat 2016, Rif, Douar Kitane, Douar Moukhlata, Oued Mhannech, AP, Douar Aoulad Ali (Central Plateau (Coastal region))

Panimerus Eaton, 1913

Panimerus thienemanni (Vaillant, 1954)

= Panimerus maynei (Tonnoir, 1919), in Dakki 1997: 62

Boumezzough and Vaillant 1986b, HA, Assif Réghaya; Afzan and Belqat 2016

Paramormia Enderlein, 1935

Paramormia ustulata (Walker, 1856)

Vaillant 1956b, HA; Mouna 1998; Ježek and Yağcı 2005; Omelková and Ježek 2012a; Afzan and Belqat 2016, Rif, Seguia Barrage Dar Chaoui, Douar Kitane, Oued Jnane Niche

Pericomini

Bazarella Vaillant, 1964

Bazarella atra (Vaillant, 1955)

= Pericoma atra Vaillant, in Vaillant 1956b: 234, 238, 242

Vaillant 1956b, HA, Assif Tasouat (M’Goum), Siroua, Imi-N’Ifri, Aguelmous, Sidi Chamarouch, Lac Tamhda (Anremer), Oukaimeden; Boumezzough and Thomas 1987, HA, Oued Réghaya (Imlil, 1750 m); Dakki and Himmi 2008; Afzan and Belqat 2016, Rif, Oued Inesmane, Oued Madissouka, Aïn Quanquben

Pericoma Walker, 1856

Pericoma (Pachypericoma) blandula Eaton, 1893

Boumezzough and Vaillant 1986b, HA; Dakki 1997; Ježek 2004, Rif; Ježek and Hájek 2007; Dakki and Himmi 2008, MA, Oued Sebou; Omelková and Ježek 2012a; Afzan and Belqat 2016, Rif, Oued Taida, Âounsar Aherman, Oued Beni Ouachekradi, Oued Aâyaden, Cascade Ras el Ma

Pericoma (Pericoma) barbarica Vaillant, 1955

Vaillant 1956b, HA, M’Goum; Afzan and Belqat 2016, Rif, Oued Taida, Douar Taria, Cascade Grotte des pigeons

Pericoma (Pericoma) granadica Vaillant, 1978

Boumezzough and Vaillant 1986b, HA; Vaillant and Moubayed 1987; Dakki 1997; Dakki and Himmi 2008, MA, Oued Sebou; Afzan and Belqat 2016, Rif, Oued Taida, Ametrasse, Oued Farda, Oued Aâyaden, Oued Ras el Ma, MA, Aïn Vittel, HA, Cascade sur sol cuivreux, Oued Réghaya

Pericoma (Pericoma) diversa Tonnoir, 1920

Vaillant 1978, HA; Afzan and Belqat 2016, Rif, Cascade Chrafate

Pericoma (Pericoma) exquisita Eaton, 1893

Ježek 2004, Rif, HA; Afzan and Belqat 2016

Pericoma (Pericoma) latina Sarà, 1954

Vaillant 1955, HA; Afzan and Belqat 2016, Rif, Cascade Chrafate, Oued Maggou, Nord Village Maggou, Oued Kelâa, Oued Talembote, Oued associé à Dayat Fifi, Oued Tiffert, Oued à 20 km de Fifi, Oued El Kanar, Beni Fenzar

Pericoma (Pericoma) maroccana Vaillant, 1955

= Pericoma numidica var. marocana Vaillant, 1955

Boumezzough and Vaillant 1986b, HA, Tissaout; Dakki 1997; Afzan and Belqat 2016, Rif, Cascade Chrafate, ruisseau maison forestière; Dakki and Himmi 2008, MA, Oued Sebou

Pericoma (Pericoma) modesta Tonnoir, 1922

= Pericoma numidica Vaillant, in Vaillant 1956b: 236, 237, 240

Vaillant 1956b, HA, Assif Tassouat (M’Goum), Lac Tamhda (L’Anremer); Boumezzough and Vaillant 1986b, HA; Dakki 1997; Dakki and Himmi 2008, MA, Oued Sebou; Afzan and Belqat 2016

Pericoma pseudexquisita Tonnoir, 1940

Afzan and Belqat 2016, Rif, Oued Azila

Pneumia Enderlein, 1935

Pneumia nubila (Meigen, 1818)

Afzan and Belqat 2016, Rif, Aïn Mâaze

Pneumia pilularia (Tonnoir, 1940)

Ježek 2004; Ježek and Hájek 2007; Omelková and Ježek 2012a

Pneumia propinqua (Satchell, 1955)

Afzan and Belqat 2016, Rif, Chrafate, Oued Zarka

Pneumia reghayana (Boumezzough & Vaillant, 1986)

= Satchelliella reghayana Boumezzough & Vaillant, 1986, in Boumezzough and Vaillant 1986b: 238

Boumezzough and Vaillant 1986b, HA; Dakki 1997; Dakki and Himmi 2008; Afzan and Belqat 2016

Pneumia toubkalensis Omelková & Ježek, 2012

Omelková and Ježek 2012b, HA, Toubkal; Afzan and Belqat 2016, Rif, Oued Aâyaden, Aïn Ras el Ma

Psychodini

Philosepedon Eaton, 1904

Philosepedon (Philosepedon) humeralis (Meigen, 1818)

Afzan and Belqat 2016, Rif, Oued Hachef, Cascade Ras el Ma, Oued El Kanar, 2 km de Douar Assoul, Oued Aâyaden

Psychoda Latreille, 1796

Psychoda (Logima) albipennis (Zettersdedt, 1850)

= Logima albipennis (Zettersdedt), in Mouna 1998: 86

Mouna 1998

Psychoda (Psycha) grisescens Tonnoir, 1922

Ježek 2004, Rif; Afzan and Belqat 2016, Rif, Douar Kitane, MA, Gîte Aït Ayoub

Psychoda (Psychoda) uniformata Haseman, 1907

Ježek 2004, Rif; Afzan and Belqat 2016

Psychoda (Psychodocha) cinerea Banks, 1894

Boumezzough and Thomas 1987, HA, Azib Oukaimeden (2730 m); Dakki 1997; Dakki and Himmi 2008; Afzan and Belqat 2016, Rif, Oued Tazzarine, Douar Taria, Douar Kitane, Oued Chrafate, Oued Aâyaden, EM, Cascade Grotte des Pigeons (Béni Snassen)

Psychoda (Psychodocha) gemina (Eaton, 1904)

Afzan and Belqat 2016, Rif, Dayat Fifi, Oued Zarka, Douar Kitane, Oued Aâyaden

Psychoda (Tinearia) alternata Say, 1824

Tonnoir 1920, HA, La Maire; Boumezzough and Thomas 1987, HA, Oued Réghaya (1740 m), Imlil; Dakki 1997; Dakki and Himmi 2008; Afzan and Belqat 2016, Rif, Oued Nakhla, Oued Farda, Oued Ouara, Oued Ametrasse, Oued Chrafate, Douar Derâa, Douar Ihermochene, Douar Ikhlafene, Douar Taria, Douar Idrene, Douar Kitane, Oued 2 km de Douar Assoul, Oued Aâyaden, ruisseau maison forestière, Oued Mhannech, Aïn Sidi Yahia, MA, Gîte Aït Ayoub

Scatopsoidea

SCATOPSIDAE3

K. Kettani, J.P. Haenni

Number of species: 13. Expected: 30–40

Faunistic knowledge of the family in Morocco: poor

Ectaetiinae

Ectaetia Enderlein, 1912

Ectaetia clavipes (Loew, 1846)

= Scatopse clavipes Loew, in Mouna 1998: 84

Mouna 1998; Haenni and Kettani 2016, Rif, Amsa – MISR

Psectrosciarinae

Anapausis Enderlein, 1912

Anapausis sp.*

Rif, Source Aïn Tissemlal

Psectrosciara Kieffer, 1911

Psectrosciara sp. 1*

Rif, Adrou

Psectrosciara sp. 2*

Rif, Adrou

Scatopsinae

Coboldia Melander, 1916

Coboldia fuscipes (Meigen, 1830)

Haenni and Kettani 2011, Rif, Kitane, AP, El Jadida, Rabat, AA, Souss; Haenni and Kettani 2016, Rif, Maggou, Aârkob, Jnane Niche; Rif (M’Diq farm) – MISR

Parascatopse Cook, 1955

Parascatopse sp.

Haenni and Kettani 2011, AA, Ouarzazate ­– MHNN

Quateiella Cook, 1975

Quateiella inexpectata Haenni, 1988

Haenni and Kettani 2016, Rif, Afertane

Reichertella Enderlein, 1912

Reichertella geniculata (Zetterstedt, 1850)

Haenni and Kettani 2016, Rif, Jnane Niche

Reichertella maroccana Haenni, 2011

Haenni and Kettani 2011, HA, Oukaimeden – RMNH

Rhegmoclemina Enderlein, 1936

Rhegmoclemina lunensis Haenni & Godfrey, 2009

Haenni and Kettani 2011, Rif, Boujdad

Rhexoza Enderlein, 1936

Rhexoza freyi (Duda, 1936)

Haenni and Kettani 2011, AA, Agadir – MHNN

Scatopse Geoffroy, 1762

Scatopse notata (Linnaeus, 1758)

Mouna 1998; Haenni and Kettani 2016, Rif, Onsar Lile – MISR

Swammerdamella Enderlein, 1912

Swammerdamella brevicornis (Meigen, 1830)

Haenni and Kettani 2011, Rif, Kitane, Oued Laou, Ketama, AP, El Jadida, Essaouira, HA, Ouarzazate; Haenni and Kettani 2016, Rif, Amsa, Jnane Niche, Ametrasse – MISR

New records for Morocco

An undescribed species of Anapausis has been collected in the Rif mountains in 2014 by K. Kettani and will be described elsewhere.

Anapausis sp.

Rif: Forêt Azilane (NPT), Source Aïn Tissemlal, 1255 m, 35°11.67N, 5°15.20W, 7.vi.2014, Sapinière à Abies maroccana et Pinus nigra, 1♂.

Two undescribed species of Psectrosciara have been collected in the Rif mountains in 2013 by K. Kettani and will be described elsewhere.

Psectrosciara sp. 1

Rif: Taghzout (PNPB), Adrou, 556 m, 35°22.39N, 05°32.28W, chênes-liège, 14.vii–15.viii.2013, 6♂♂.

Psectrosciara sp. 2

Rif: Taghzout (PNPB), Adrou, 556m, 35°22.39N, 05°32.28W, chênes-liège, 14.vii–15.viii.2013, 3♂♂.

PTYCHOPTERIDAE

K. Kettani, R. Wagner

Number of species: 5. Expected: 8

Faunistic knowledge of the family in Morocco: moderate

Ptychopterinae

Ptychoptera Meigen, 1803

Ptychoptera albimana (Fabricius, 1787)

MISR (No locality given)

Ptychoptera contaminata (Linnaeus, 1758)

MISR (No locality given)

Ptychoptera lacustris Meigen, 1830

MISR (No locality given)

Ptychoptera paludosa Meigen, 1804

MISR (No locality given)

Ptychoptera scutellaris Meigen, 1818

MISR (No locality given)

Culicoidea

CHAOBORIDAE

K. Kettani, R. Wagner

Number of species: 2. Expected: 4

Faunistic knowledge of the family in Morocco: poor

Chaoborinae

Chaoborus Lichtenstein, 1800

Chaoborus crystallinus (De Geer, 1776)

Dakki 1997: 60

Mochlonyx Loew, 1844

Mochlonyx culiciformis (De Geer, 1776)

Dakki 1997: 60

CULICIDAE

K. Kettani, B. Trari, O. Himmi, M. Dakki

Number of species: 43. Expected: 60

Faunistic knowledge of the family in Morocco: good

Anophelinae

Anopheles Meigen, 1818

Anopheles (Anopheles) algeriensis Theobald, 1903

Viallate 1922, AP, Kénitra; Séguy 1930a; Bonjean 1947, EM, MA; Gaud 1957a, HA, north of High Atlas; Guy 1959a, MA, Béni Mellal, HA; Guy 1959b, HA; Guy 1959a, MA, Béni Mellal; Bailly-Choumara 1967a, MA, Ghorm El Alem; Benmansour et al. 1972, MA, Barrage Bin El Ouidane; Bailly-Choumara 1973b, AP, Sidi Yahia du Gharb; Metge 1986, AP, Casablanca; Trari and Himmi 1987, AP; Himmi et al. 1995; Louah 1995, Rif, Tahaddart, Schroda; Dakki 1997; Ramdani 1997, AP, Skhirat, Casablanca; Trari et al. 2002; Trari et al. 2004b; Himmi 2007, Rif, Chefchaouen; El Ouali Lalami et al. 2010 a,b, MA, Fès, Boulmane; El Ouali Lalami 2012, MA, Fès; Trari 2017, Rif, Chefchaouen, AA, Tiznit; Trari and Dakki 2017a, Rif, Chefchaouen, AA, Tiznit; Trari and Dakki 2017b, AA, Tiznit; Trari et al. 2017

Anopheles (Anopheles) claviger (Meigen, 1804)

Vialatte 1923, AP, Kénitra; Séguy 1930a, AP, Rabat; Langeron 1938, HA, Tounfit, Massou, Anefgou, Tirghist, Tighermine, Louggouargh; Callot 1940, HA, Anefgou, Tirghist; Bonjean 1947, EM, MA; Gaud 1947c, MA, Sefrou, Meknès; Gaud 1948, AP, Rabat, MA, El Hajeb, AA, Errachidia, Tadla; Gaud et al. 1948, AP, Skhirat; Guy 1963, Rif, Taounate, MA, Meknès, Ifrane; Bailly-Choumara 1967a, MA, piste Tafechna-Taoujgelt, maison forestière Ouiouane, piste Tafechna-Senoual-Itzer, piste Ksiba-Naour, piste Naour-Arbala, Zaouia Cheikh, Oued Sarif (environs El Ksiba), Dayet Aoua (environs Ifrane), Boulemane; Bailly-Choumara 1967b, EM, 8 km N Itzer; Bailly-Choumara 1967c, Rif, piste Ketama-Mt Tiguidin; Guy 1967, HA, Marrakech, AA, Tafilalt; Guy and Holstein 1968, SA; Metge 1986, AP, Casablanca; Himmi et al. 1995; Louah 1995, Rif, Haidra, Marina Smir; Dakki 1997; Ramdani 1997, AP, Skhirat, Tamaris drains, Tamaris merja; Trari et al. 2002; Trari et al. 2004b; Himmi 2007, Rif, Bab Berred, Tanakoub; Faraj et al. 2008b, Rif, Assoul, Mizgane; El Ouali Lalami et al. 2010a, MA, Boulmane; Larhbali et al. 2010, MA, Zhiliga, Boukachmir, Aït Ichou; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Anopheles (Anopheles) labranchiae Falleroni, 1926

d’Anfreville 1916, AP, Salé; Delanoe 1917, AP, El Jadida; Viallate 1922, AP, Rabat, Boulhaut, Bouznika, Gharb, MA, Sidi Kacem, Tiflet, Fès, Taza; Charrier 1924, Rif, Tanger; Séguy 1930a, MA; Roubaud 1935, AP, Rabat; Sicault et al. 1935, AP, Merja Ras Eddaoura, Merja Zerga, dayas entre Sebou et Maâmora, Dar bel Hamri (entre barrage El Kansera et Sidi Slimane), à proximité de Merja Zerga, Douar Anabsa; Langeron 1938, AA; Callot 1940, AA; Ristorcelli 1946a, b, HA, Oued Tensift, Oued Issil; Bonjean 1947, AP, Gharb; Gaud 1947c, MA, Oulmès, HA, Marrakech; Gaud et al. 1948, AP, Merja Ras Eddaoura; Gaud et al. 1949, AP, Salé, SA, Foum Zguid, Tagounit; Gaud 1953a, AP, from Tanger to El Jadida, MA, Tissa, Timhadit, Bekrit, Meknès, Ifrane (1700 m), HA, Sidi Aissa, Tizi-n’Tichka; Guy 1958, HA, Oued N’fis; Sacca and Guy 1960b, Rif, Tétouan, AP, Skhirat, Sidi Yahia, Sidi Bettache, Mazagan, Braila (près Sidi Allal Tazi), Aït Lahsen, MA, Meknès, HA, Marrakech; Guy 1962, Rif, Taounate, AP, Gharb, HA, Marrakech (ville et banlieu); Guy 1963, Rif, Tanger, Tétouan, EM, Berkane, Debdou, Oujda, AP, Kénitra, Souk Larba, Settat, Chamaîa, Safi, Casablanca, Rabat, Essaouira, MA, Meknès, Fès, Azrou, Oued Zem, Béni Mellal, HA, Kelaâ of Sraghna; Bailly-Choumara 1967a, MA, piste Tafechna-Znan Imes, piste Tafechna-Taoujgelt, piste Tafechna-Assoul, Aguelmane Azigza, piste Aguelmane Azigza-Aïn Leuh, maison forestière Ouiouane, route Khénifra-Tafechna, piste Tafechna-Senoual-Itzer, Ajdir, Itzer, Bords Moulouya, piste Itzer-Boumia, RN P.33 Boumia-El Kbab, Ouaoumana, piste Ksiba-Naour, piste Naour-Arbala, piste Arbala-El Kbab, Kafensour, Oued Sarif (environs El Ksiba), Dayet Aoua, Ifrane, Imouzer Marmoucha; Bailly-Choumara 1967b, EM, route Itzer-Midelt, Douar Sherba, Douar Aïn Shebbak, Douar Aïn Zabia, Douar Madarh, Douar Sidi Hashas, Mechraa safsaf; Bailly-Choumara 1967c, Rif, route Chaouen-Bab Taza, piste Bab Taza-Fifi, piste Bab Taza-Talassemtane, piste Bab Taza-Béni Ahmed, piste Bab Taza-Bab Berred, Anasar, piste Bab Berred-Tamorote, Ketama, route Ketama-Targuist, route nationale Jebha, Al Hoceima Club Med, Béni Bouayache, Targuist et environs; Guy 1967, HA, Marrakech, AA, Tafilalt; Bailly-Choumara 1968b, AP, Larache; Guy and Holstein 1968, AA, Ouarzazate; Bailly-Choumara 1970, Rif, Tétouan, EM, Berkane, AP, Larache, Sidi Yahia du Gharb, HA, Marrakech; Benmansour et al. 1972, AA, Agadir; Bailly-Choumara 1972a, AP, Merja Sheishat; Bailly-Choumara 1972b, AP, Merja Sheishat; Bailly-Choumara 1973a, AP, Merja de l’Oued Smir; Bailly-Choumara 1973b, Rif, Tétouan, EM, Berkane, AP, Merja Sheishat, HA, Souk des Oudaias; de Zulueta et al. 1983, Rif; Ibn Jilali 1984, AP, Maâmora; Metge 1986, AP, Entre Oulad Dlim et Al Ara’ra; Trari and Himmi 1987, AP, Gharb; Himmi 1991, AP, Kénitra, Maâmora; Metge 1991, AP, Sidi Bettache; Trari 1991, Rif, Chefchaouen, Tanger, Taounate, EM, Oujda, AP, Sidi Amira, Sidi Boughaba, Merja Zerga, Sidi Allal Tazi, Aïn Chouk, Oued Loukous, Merja Oulad Skhar, MA, Khémissat, Béni Mellal, Khouribga, HA, El Kelaâ des Sraghrna, AA, Ouarzazate; El Bermaki 1993, AP, entre l’aeroport Anfa et l’aménagement d’El Oulfa, Sidi Maârouf, El Oulfa; Chlaida and Bouzidi 1995, AP, Aïn Blal, Oued Sidi Messoud; Louah 1995, Rif, Tres piedras, Marina Smir, Bouzerlal, Oued Maleh, Zekri, Lajour, Azla, Tahaddart, Skroda, Stehat, Moulay Bouchta, Kebbache, Talembote, Loubart, Chefchaouen, Oued Maggou, Bab Berred, Sidi Kankoch, Oued kbir, Oued Jebel Lehbib; Louah et al. 1995; Chlaida 1997, AP, Aïn Blal, Oued Sidi Messoud, Douar Chlihat, Barrage Al Massira; Moussalim 1997, AP, Sidi Allal Tazi, Maâmora; Ramdani 1997, AP, Tamaris, Skhirat; Faraj et al. 1997, AP, Ouled Moussa; Handaq 1998, HA, Oukaimeden, Amizmiz, Tiguenziouine; Himmi et al. 1998, AP, Dayat d’El Menzeh (north-east of Kénitra), Sidi Boughaba; Alaoui Slimani et al. 1999, AP, Bou-Regreg Salé, Sidi Bouguettaya, Quartier industriel Takaddoum, Marjane; Alaoui Slimani 2002, AP, Rabat-Salé; Trari et al. 2002; Faraj et al. 2003, Rif, Azib Bouflou, Azib Jrou, Imzouren, Amezzaourou, Tizi-Tamalout, MA, Aït Abdelsalam, Aït Lamfadel; Faraj et al. 2004, Rif, Azib Jrou, Tanghaya Akarkar, Amezzaourou, Ouled Nsar, AP, Fedalate, MA, Talaa Chougaga, Aïn Smen (Fès), Aït Lamfadel; Trari et al. 2004a, Rif, Larache; Trari et al. 2004b, Rif, Ketama, Gzenaya, Taounate, Bouzaghlal, Oued Laou, Smir (Merja), Béni Hassane, Azib Jrou, Azib Bouflou, AP, Laaouamra, Aarabat Sidi Abdelaziz, Moukaouama, Dar Belamri, Laksibia, Mgadid, Beggara, Rabat (Chellah), MA, Adouz, Rommani (Khémisset), Aït Abderrahmane, Aït Ishaq, Oulad Messoud, Ouled Fennane, El karma, Oulad Zguida, Oulad Abbou; Aouinty et al. 2006, AP, Mohammedia; Himmi 2007, Rif, Bab Berred, Bab Taza, Tanakoub, AP, Skhirate, Maâmora, Oulja, Bouknadel, Sidi Azzouz, Tamaris, MA, Khémisset; Faraj et al. 2008a, AP, Laouamra, Boucharen, MA, Béni Khlef, Talaa Chougaga; Faraj et al. 2008b, Rif, Assoul, Mizgane; El Ouali Lalami et al. 2010b, MA, Fès; Faraj et al. 2010, AP, Begara, Boucharen, Ben Slimane, Skhirat, Rabat, Sehoul, MA, Sidi Allal Msader, Aïn Aghbal, Aïn Elouali, Sidi Kacem; Larhbali et al. 2010, MA, Aït Haddou Said; Adlaoui et al. 2011, AP, Larache; Larhbali et al. 2011, MA, Oulmès, Aït Yadin, Sfassif, Mâaziz, Rommani, Laghoualem, Ezzhiliga, Sidi Allal Bahraoui, Boukachmine, Aït Malek, Sidi Boukhalkhal, Bni Ounzar, Ganzra, Aït Siberne, Sidi Allal Msader, El Ghandour; El Ouali Lalami 2012, MA, Pont Diamant vert, Sidi Harazem, Oued El Himmer, Moulay Yakoub, Oued Sebou, Aïn Kansara, Oued Aïn Chkef, Sefrou, Boulemane; Laboudi et al. 2012, AP, Larache; Hadji et al. 2013, AP, Sidi Slimane; El Joubari et al. 2014, Rif, Smir lagoon; Laboudi et al. 2014, Rif, Tétouan, Tanger, Chefchaouen, Al Hoceima, AP, Larache, Salé, MA, Taza, Khémissat; El Joubari et al. 2015a, Rif, Smir lagoon; El Joubari et al. 2015b, Rif, Smir lagoon; Marc et al. 2016, AP, Kénitra; Trari 2017, Rif, Chefchaouen, Tétouan, AP, Larache, Rabat, Settat, EM, Oujda, MA, Khémisset, Meknès, Khouribga, HA, Marrakech, AA, Tiznit, Ouarzazate; Trari and Dakki 2017a, Rif, Chefchaouen, Tétouan, AP, Larache, Rabat, Settat, EM, Oujda, MA, Khémisset, Meknès, Khouribga, HA, Marrakech, AA, Tiznit, Ouarzazate; Trari and Dakki 2017b; Trari et al. 2017

Anopheles (Anopheles) marteri Senevet & Prunnelle, 1927

Gaud 1945b, MA, El Hajeb, Khénifra, HA, Tizi-n’test, Tillougite; Gaud et al. 1949, HA, Tizi-n’test; Gaud 1953b, HA, Tizi-n’test; Bailly-Choumara 1967b, EM, Grotte du Zegzel; Bailly-Choumara 1967c, Rif, route Chaouen-Bab Taza, piste Bab Taza-Béni Ahmed, route Bab Taza-Bab Berred, piste Bab Taza-Asifane, piste Bab Berred-Tamorote, Ketama, route Ketama-Targuist, route nationale Jebha, Boured; Benmansour et al. 1972, MA, Taza; Trari 1991, Rif, Taounate; Himmi et al. 1995; Dakki 1997; Trari et al. 2002; Himmi 2007; Trari 2017, Rif, Chefchaouen, AP, Settat; Trari and Dakki 2017a, Rif, Chefchaouen, AP, Settat; Trari and Dakki 2017b; Trari et al. 2017

Anopheles (Anopheles) ziemanni Grünberg, 1902

Senevet 1935, MA; Gaud et al. 1949, HA, plain of south and north of the Occidental Atlas; Gaud et al. 1950, HA, plain of south and north of the Occidental Atlas; Guy 1958, HA, Marrakech; Guy et al. 1958, HA, Oued N’fis; Guy 1967, EM, Oujda, AP, Rabat, HA, Marrakech, AA, Tafilelt; Bailly-Choumara 1970, HA, Marrakech; Benmansour et al. 1972, MA, Taza, HA, Haouz, Tadla Azilal; Bailly-Choumara 1973b, HA, Souk des Oudaias (Souk Tnine des Oudaias, 470 m); Trari 1991, MA, Tissa; Moussalim 1997, AP, 7.5 km de Sidi Allal Tazi; Trari et al. 2002; Trari et al. 2004b; Himmi 2007; Trari 2017, Rif, Tétouan, AA, Tiznit; Trari and Dakki 2017a, Rif, Tétouan, AA, Tiznit; Trari and Dakki 2017b; Trari et al. 2017

Anopheles (Cellia) cinereus Theobald, 1901

Viallate 1922, MA, Sefrou, Sidi Lamine, HA, Mtougui; Sicault et al. 1935, AP, Souk Larba of Gharb; Senevet 1935, AP, Souk Larba of Zemmour; Langeron 1938, HA, Anefgou, Tirghist, Valley of Sidi Yahia Ouyoussef, Tighermine, Louggouargh, Massou; Callot 1940, HA, Anefgou, Tirghist; Gaud 1945a, Rif, Meridional Rif, EM, Moulouya, HA, Marrakech, AA, Tansikht, valley de Sous; Gaud and Duthu 1945, HA, Marrakech; Viamonte and Ramirez 1945, Rif; Viamonte and Ramirez 1946, Rif; Ristorcelli 1946a, HA, Oued Tensift, Oued Issil; Ristorcelli 1946b, HA, Oued Tensift, Oued Issil; Gaud 1945b, AA, Tansikht; Gaud et al. 1949, Rif, Meridional Rif, EM, Moulouya, HA, Marrakech, AA, Tansikht, valley de Sous; Gaud et al. 1950, Rif, Meridional Rif, EM, Moulouya, HA, Marrakech, AA, Tansikht, valley de Sous; Gaud 1953a; Gaud 1958, HA, Marrakech; Guy 1962, Rif, Taounate, HA, Marrakech; Guy 1963, MA, Midelt, AA, Hamada of Draa; Bailly-Choumara 1965, EM, Aman d’Aït Oussa, Tiglit, El Megrinat, Taskala, Aïn Aït delouine, Oued mesdourt, Talmesdourt, Assa, AA, Aït Melloul, Oued Teima, Issen, Taroudant, Talaint, Tiznit, Oued Assaka, Anezi, Pont de la route Agadir-Tiznit, valley of low Draa, Tafraoute, Tacharicht, Bou Izakarn, Jemâa N’tirhirte, Aït Erkha,Tazert, Barrage Taourirt, AA, Goulmima, SA, Aouinet Torkoz, Tirh Mzoun; Bailly-Choumara 1966, AA, piste Foum Zguid-Lac Iriqui, Agadir-Tissint, Akka-Iguiren, Tirhem, Taoujgelt, Souk El Khémis Dades, Akka, Aït Ouabelli, Foum-el-Hassan, Tarhjicht, Aït melloul, Taliouine, Tazenakht (Rocade of Draa); Bailly-Choumara 1967a, MA, route Azrou-Khénifra, Jnane Imasse, piste Tafechna-Taoujgelt, piste Tafechna-Assoul, Sources Oum-er-Rbia, Itzer, piste Itzer-Boumia, Ouaoumana, piste Ghorm El Alem-El Ksiba, piste Naour-Arbala, Zaouia Cheich, kafensour, Ifrane, Imouzzer Marmoucha; Bailly-Choumara 1967b, EM, 7 km N Itzer, 8 km N Itzer, route Itzer-Midelt, Aïn Srouna, Gouttitir, Cascade Oued Za, Grotte du Zegzel, Douar Aïn Soultane, Mechraa Safsaf, Oujda, Berguent (valley of Moulouya), Figuig; Bailly-Choumara 1967c, Rif, Chaouen ville, route Chaouen-Bab Taza, Bab Taza, piste Bab Taza-Fifi, piste Bab Taza-Talassemtane, piste Bab Taza-Béni Ahmed, route Bab Taza-Bab Berred, piste Bab Berred-Assifane, Anasar, piste Bab berred-Tamorote, Ketama, route Ketama-Targuist, route nationale Jebha, route Al Hoceima-Arba Taourirt, Arba Taourirt, Targuist et environs, route Targuist Al Hoceima, Jebha, Oued Ouergha, route Aknoul-Al Hoceima; Guy 1967, AP, Rabat, EM, Oujda, HA, Marrakech, AA, Tafilalet; Bailly-Choumara 1970, HA, Marrakech; Bailly-Choumara 1973b, HA, Souk des Oudaias; Trari 1991, Rif, Al Hoceima, Chefchaouen, Taounate, EM, Nador, Oujda, Figuig, AP, Larache, Settat, Ben Slimane, MA, Khénifra, Taza, Khouribga, HA, Kelaâ of Sraghna, AA, Ouarzazate, Goulmima; Bouallam 1992, HA, Oued N’fis; Louah 1995, Rif, Marina Smir, Bouzerlal, Tahaddart, Stehat, Moulay Bouchta, Kebbache, Loubart, 0ued Maggou; Louah et al. 1995; Bouallam et al. 1997b, HA, Marrakech; Handaq 1998, HA, Amizmiz, Tiguenziouine; Bouallam 2001, HA, Oued N’fis; Trari et al. 2002; Trari et al. 2004b, Rif, Ketama, Azib Jrou, Sidi Mokhfi, MA, Taghzirt, Aghbala, Aït Shak, Smaala, Mlalih, Ouled Fennane, Béni Khlef, Tachrafte; Faraj et al. 2007, Rif, Assoul, Mizgane; Himmi 2007, Rif, Bab Berred, Bab Taza, Stehat, Tanakoub; Faraj et al. 2008, Rif, Assoul, Mizgane; El Ouali Lalami et al. 2010a, MA, Fès; El Ouali Lalami et al. 2010b, MA, Oued El Himmer; Larhbali et al. 2010, MA, Roumani Aïn Sbite, Jamâa M. B., Ghoualem, Zhiliga, Oulmès, Tarmilate, Boukachmir, Mrirte, Aït Ichou, Mâaziz, Tiddas, Bni Ounzar, Ganzra; El Ouali Lalami 2012, MA, Oued El Himmer; Trari 2017, Rif, Chefchaouen, Tétouan, MA, Khouribga, AA, Tiznit; Trari and Dakki 2017a, Rif, Chefchaouen, Tétouan, MA, Khouribga, AA, Tiznit; Trari and Dakki 2017b; Trari et al. 2017; Mouatassem et al. 2019, MA, Fès

Anopheles (Cellia) dthali Patton, 1905

Saccà 1960, AA, Aoufous, Meski, Erfoud, Agdz, Zogora, Tagounit, Tamsrruth; Guy 1961, HA, Sud de Zagora (en bordure Hamada du Draa); Guy 1963, AA, Zagora and south of the High Atlas (at edge of Hamada of Draa), Oued Ziz; Bailly-Choumara 1965c, AA, Tiznit, EM, Aïn Aït Delouine, Aouinet Torkoz, Rich Tamlougout (eastern borders of Jebel Bani); Bailly-Choumara 1966, AA, piste Foum Zguid au Lac Iriqui, Agadir-Tissint, Akka-Iguiren, Souk El Khémis Dades, Akka et Environs, Aït Ouabelli, Tarhjicht, piste Tazenakhte à Foum Zguid (Rocade du Draa); Bailly-Choumara 1967b, EM, valley of Moulouya; Guy 1967, HA, Marrakech, AA, Tafilalet; Guy and Holstein 1968, EM, N Outat El Haj (valley of Moulouya), Gouttitir (environs de Taourirt); Bailly-Choumara 1970, AA, Foum Zquid; Bailly-Choumara 1973b, AA, Foum Zquid; Himmi et al. 1995; Dakki 1997; Trari et al. 2002; Trari et al. 2004b; Faraj et al. 2007, Rif, Assoul, Mizgane (SE of Bab Berred); Faraj et al. 2008, Rif, Assoul, Mizgane (SE Bab Berred); Himmi 2007; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Anopheles (Cellia) multicolor Cambouliu, 1902

Messerlin and Treillard 1938, HA, Marrakech; Viamonte and Ramirez 1945, Rif; Guy 1963, SA; Bailly-Choumara 1965c, EM, Aït Oussa, Aman d’Aït Oussa, Aïn Aït Delouine, Aouinet Torkoz, Rich tamlougout (Confins orientaux du Jebel Bani), AA, Tafnidilt, Guelta Zerga, Aïn Temda (valley of low Draa), Tirhmert (Goulmima), SA, Vallée et embouchure de l’Oued Assaka, Tantan ville, Tirh Mzoun; Bailly-Choumara 1966, AA, Rocade of Draa; Bailly-Choumara 1967b, EM, 40 km N de Outat El Haj, Gouttitir, Cascade Oued Za, Douar Aïn Shebbak (valley of Moulouya); Guy 1967, HA, Marrakech, AA, Tafilelt; Guy and Holstein 1968, HA, south of Atlas, AP, plain located between Marrakech and the Atlantic from Tanger along the length of the Mediterranean; Bailly-Choumara 1970, AA, Foum Zquid; Bailly-Choumara 1973b, AA, Foum Zquid; Metge 1986, AP, Casablanca; Trari 1991, Rif, Al Hoceima, Taounate, Oued Maleh, Bab Berred, AP, Tamaris Merja, AA, Ouarzazate; Himmi et al. 1995; Dakki 1997; Trari et al. 2002; Trari et al. 2004b; El Joubari et al. 2014, Rif, Smir lagoon; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Anopheles (Cellia) sergentii (Theobald, 1907)

Séguy 1930a; Messerlin and Treillard 1938, HA, Tamelelt; Langeron 1938, Rif, Targhist; Callot 1940, AA, Taghjicht; Gaud 1947c, AA, Wadi Draa; Gaud et al. 1949, Rif, Zoumi, HA, Zaouia Sidi Hamza, Tizi-n’test (1700 m), Tillougit (1800 m); Gaud et al. 1950, Rif, Zoumi, MA; Guy et al. 1958, HA, Oued N’fis; Guy 1961, Rif, AP, south of Casablanca, AA, Sud de Zagora; Guy 1962, Rif, AP, south of Casablanca, HA, Marrakech; Guy 1963, Rif, Tanger, EM, Berkane, AP, south of Casablanca, MA, Béni Mellal, HA, Oued Tensift, Oued Ziz, Marrakech, Chichaoua, AA, Oued Draâ, Oued Dades, Goulmima, sud de Zagora, SA, Foum Zguid; Bailly-Choumara 1965c, EM, iglit, Aman d’Aït Oussa, Oued Isker, El Megrinat, Aïn Aït Delouine, Oued Mesdourt, almesdourt, Aouinet Torkoz, Bouanama, Rich Tamlougout, Assa, AA, Tiznit, Tafraoute, Oued Izi, Bou Izakarn, Abeino (region of Goulmima), SA, Aouzeroual, Tirhmert, Tacharicht, Jebel Bani, Tantan; Bailly-Choumara 1966, AA, Rocade of Draa, Taliouine; Bailly-Choumara 1967a, Rif, AP, south of Casablanca, MA, Itzer, Ghorm El Alem, Zaouia Cheikh, Kafensour, HA, north of High Atlas; Bailly-Choumara 1967b, EM, Aïn Srouna, Grotte du Zegzel, Douar Mardarh, Douar Aïn Soultane, Merja Boubker, Selouane, Driouch (valley of Moulouya); Bailly-Choumara 1967c, Rif, route nationale Jebha, Targuist-Béni Boufrah, Al Hoceima, Béni Bouayache, Marchica, Had El Rouadi, Pont du Srah; Guy 1967, HA, Marrakech, AA, Tafilelt; Guy and Holstein 1968, AP, Casablanca; Bailly-Choumara 1970, EM, Berkane, HA, Marrakech; Bailly-Choumara 1973b, EM, Berkane, HA, Marrakech; Metge 1986, AP, Casablanca; Trari 1991, Rif, Al Hoceima, Taounate, AP, Larache, AA, Ouarzazate; Himmi et al. 1995; Dakki 1997; Trari et al. 2002; Faraj et al. 2003, Rif, Al Hoceima, Chefchaouen, Taounate, HA, Khouribga; Trari et al. 2004b, Rif, Ketama, Sidi Mokhfi; Faraj et al. 2007, Rif, Assoul, Mizgane; Faraj et al. 2008, Rif, Assoul, Mizgane; El Ouali Lalami et al. 2010a, MA, Fès; Larhbaliet al. 2010, MA, Oulmès, Ganzra; Trari 2017, Rif, Chefchaouen, Tétouan, EM, Oujda, MA, Khouribga; Trari and Dakki 2017a, Rif, Chefchaouen, Tétouan, EM, Oujda, MA, Khouribga; Trari and Dakki 2017b; Trari et al. 2017; Benabdelkrim Filali et al. 2018; Mouatassem et al. 2019, MA, Fès

Culicinae

Aedini

Aedes Meigen, 1818

Aedes (Acartomyia) mariae (Sergent & Sergent, 1903)

Séguy 1930b; Messerlin 1938, AP, Rabat; Séguy 1930a, Rif, littoral méditerranéen; Bailly-Choumara 1967b, Rif, Al Hoceima; Bailly-Choumara 1967c, Rif, Al Hoceima; Bailly-Choumara 1968a, Rif, Al Hoceima, AP, Larache, Sidi Yahia, Sidi Allal Tazi, Rabat, HA, Marrakech, AA, Tiznit; Himmi et al. 1995; Mestari 1997, AP, Mohammedia; Moussalim 1997, AP, Sidi Allal Tazi; Trari et al. 2002; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Aedes (Aedimorphus) vexans (Meigen, 1830)

Gaud 1947c, AP, Sidi Allal Tazi, MA, Khémisset; Metge 1986, AP, Littoral Casablanca; Himmi et al. 1995; Handaq 1998, AP, Gharbia; Dakki 1997; Trari et al. 2002; El Ouali Lalami et al. 2010a, MA, Fès; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Aedes (Dahliana) echinus (Edwards, 1920)

Séguy 1924; Séguy 1930a; Gaud 1953a, AP, Rabat, Sidi Yahia, MA, Moulay Bouazza, Taza, Fès, Meknès, Ifrane; Bailly-Choumara 1965b, AP, Maâmora; Bailly-Choumara 1967c, Rif, piste Bab Taza-Talassemtane; Himmi et al. 1995; Trari et al. 2002; Nikookar et al. 2010; El Joubari et al. 2014; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Aedes (Dahliana) geniculatus (Olivier, 1791)

Séguy 1924, MA; Séguy 1930a; Metge and El Alaoui 1987, AP, Subéraies de Béni Abid-Benslimane (Casablanca); Metge and Belakoul 1989, AP, Sidi Bettache; Himmi et al. 1995; El Ouali Lalami et al. 2010a, MA, Fès Boulmane; Trari et al. 2002; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Aedes (Ochlerotatus) berlandi (Séguy, 1921)

Séguy 1930a, AP, Rabat; Gaud 1953a, MA, Fès; Bailly-Choumara 1967a, MA, Jnane Imasse, piste Tafechna-Taoujgelt; Bailly-Choumara 1967c, Rif, piste Bab Taza-Béni Ahmed; Belakoul 1985, AP, Benslimane, Sidi Bettache; Metge and El Alaoui 1987, AP, Casablanca; Metge and Belakoul 1989, AP, Benslimane, Sidi Bettache; Trari et al. 2002; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Aedes (Ochlerotatus) caspius (Pallas, 1771)

Séguy 1930b; Viamonte et Ramirez 1946, AP, Larache; Gaud 1952, AP, Rabat, Casablanca; Gaud 1953a, AP, Rabat, Casablanca; Senevet and Andarelli 1954, EM, Embouchure de la Moulouya, Figuig, AP, Jorf Lasfar, Mohammedia, Rabat, MA, Meknès, Fès, Taza, HA, Marrakech, Midelt, AA, Tiznit; Bailly-Choumara 1966, AA, environs de Tiznit; Bailly-Choumara 1967a, EM, Bords Moulouya (près Itzer), Cherarba; Bailly-Choumara 1967b, EM, Cherarba, Aïn Shebbak, Saidia, Berguent; Bailly-Choumara 1967c, Rif, piste Al Hoceima-Arba Taourirt; Bailly-Choumara 1972a, AP, Merja Sheishat; Bailly-Choumara 1973b, Rif, Merja de l’Oued Smir; El Kaim 1972, AP, Bou-Regreg; Rioux et al. 1975, AP, Rabat-Salé; Metge 1986, AP, Littoral casablancais; Himmi 1991, AP, Sidi Boughaba; Trari 1991, AP, Sidi Boughaba, Merja Zerga, Oued Loukous; Himmi et al. 1995; Mestari 1997, AP, Mohammedia; Moussalim 1997, AP, Sidi Allal Tazi, Kénitra; Ramdani 1997, AP, Tamaris Merja; Handaq 1998, AP, Zemamra, B. Iffou (Entre El Oualidia et Youssoufia), MA, Béni Mellal, HA, Marrakech, Zaouiet Ben Sassi, Bengrir; Himmi et al. 1998, AP, Sidi Boughaba; Trari et al. 2002; Himmi 2007, AP, Sidi Boughaba; Alaoui Slimani 2002, AP, Rabat; Aouinty et al. 2006, AP, Mohammedia; El Joubari et al. 2014, Rif, Smir lagoon; El Joubari et al. 2015a, Rif, Smir lagoon; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Aedes (Ochlerotatus) coluzzii Rioux, Guilvard & Pasteur, 1998 et Aedes (Ochlerotatus) detritus (Haliday, 1833) [Complexe detritus]

Charrier 1924, Rif, Tanger; Séguy 1930a; Gaud 1953a, EM, Saidia, AP, Kénitra, Rabat, Bouznika, El jadida, Oualidia, HA, Marrakech, AA, Agadir, Tafnidilt; Bailly-Choumara 1965c, EM, Aïn Aït delouine, SA, Tirhmert; Bailly-Choumara 1970, Rif, Tétouan; Knight 1971, AP, Kénitra; El Kaim 1972, AP, Bou-Regreg; Bailly-Choumara 1973b, Rif, Merja de l’Oued Smir; Rioux et al. 1975, AP, Rabat; Pasteur et al. 1978, AP, Bou-Regreg; Metge 1986, AP, Littoral casablancais; Himmi 1991, AP, Sidi Boughaba; Trari 1991, AP, Sidi Boughaba, Merja Zerga, Oued Loukous; Louah 1995, Rif, Tres piedras, Cabo Negro, Lajour, Azla, Tahaddart; Himmi et al. 1995; Louah et al. 1995; Mestari 1997, AP, Mohammedia; Moussalim 1997, AP, Sidi Allal Tazi, Rabat; Ramdani 1997, AP, Tamaris Merja; Handaq 1998, AP, Essaouira, Zima-Chemaîa; Himmi et al. 1998, AP, Sidi Boughaba; Himmi 2007, AP, Sidi Boughaba; Alaoui Slimani 2002, AP, Rabat; Trari et al. 2002; El Joubari et al. 2014, Rif, Smir lagoon; El Joubari et al. 2015a, Rif, Smir lagoon; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Aedes (Ochlerotatus) pulchritarsis (Rondani, 1872)

Gaud 1953a, AP, Benslimane, Sidi Yahia, Rabat, MA, Oued Zem, Khénifra, Fès; Metge and El Alaoui 1987, AP, Benslimane; Himmi et al. 1995; Trari et al. 2002; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Aedes (Rusticoidus) rusticus (Rossi, 1790)

Viamonte and Ramirez 1946, Rif, Tétouan, AP, Larache; Gaud 1953a, MA, Taza; Himmi et al. 1995; Handaq 1998, HA, Bengrir; Trari et al. 2002; El Ouali Lalami et al. 2010a, MA, Fès; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Aedes (Stegomyia) aegypti (Linnaeus in Hasselquist, 1762)

d’Anfreville 1916, AP, Salé; Vialatte 1923, AP, Rabat, Casablanca, HA, Marrakech; Charrier 1924, Rif, Tanger; Gaud 1953a, AP, Salé, HA, Marrakech; Himmi et al. 1995; Dakki 1997; Handaq 1998, HA, Bengrir; Trari et al. 2002; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Aedes (Stegomyia) albopictus (Skuse, 1895)

Bennouna et al. 2016, AP, Agdal (Rabat); Trari 2017; Trari and Dakki 2017b; Trari et al. 2017; Faraj et al. 2018; Amraoui et al. 2019

Culicini

Culex Linnaeus, 1758

Culex (Barraudius) modestus Ficalbi, 1889

Séguy 1930a; Bailly-Choumara 1968a, AP, Larache; Trari 1991, AP, Gharb; Himmi et al. 1995; Dakki 1997; Handaq 1998, HA, Bengrir; Trari et al. 2002; Himmi 2007, Rif, Bab Berred, AP, Maâmora; Hadji et al. 2013, AP, Sidi yahia du Gharb, Kcebia, Sidi Hagouch (Sidi Slimane); Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Culex (Culex) brumpti Galliard, 1931

Bailly-Choumara 1968a, AP, Merja Bokka, Larache, HA, Marrakech; Bailly-Choumara 1972a, AP, Merja Sheishat; Himmi et al. 1995; Dakki 1997; Himmi 2007; Trari et al. 2002; El Ouali Lalami et al. 2010a, MA, Fès; El Joubari et al. 2014, Rif, Smir lagoon; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Culex (Culex) laticintus Edwards, 1913

Charrier 1924, Rif, Tanger; Callot 1940, AA, Goulmima (mares); Gaud 1953a, HA, Marrakech, AA, Agadir; Gaud 1957a, EM, Nador; Bailly-Choumara 1965c, EM, Oued Isker, Aïn Aït delouine, Talmesdourt, AA, Ouled Teima, ounaamane, Bou Izakarn, Agunil Khnufa, Akka-guiren; Bailly-Choumara 1966, AA, Akka-Iguiren, Tirherm, Taoujgelt, Aït Ouabelli, Anamere-Smougue, Aït melloul, Tiznit (Rocade de Draa); Bailly-Choumara 1967b, Rif, Béni Bouayache, Targuist et environs, EM, Grotte du Zegzel; Himmi et al. 1995; Dakki 1997; Trari et al. 2002; Himmi 2007, AP, Skhirat; Faraj et al. 2008b, AP, Louamra; Hadji et al. 2013, AP, Sidi Hagouch (Sidi Slimane); Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Culex (Culex) mimeticus Noè, 1899

Séguy 1930a; Viamonte and Ramirez 1946, Rif, Béni Ider, Fnideq, Khemis Anjra, Ketama, Oued Amsa, Oued Krikra, Oued Martil, Oued Laou; Gaud 1953a, Rif, Ouezzane, Ghafsai, EM, Berkane, Martinpray (près Berkane), El Aïoun Sidi Mellouk, AP, Tamri, MA, Meknès, Fès, Ifrane, Taza, Béni Mellal, HA, Midelt, Marrakech, Azilal, AA, Tinghir, Tichka; Guy et al. 1958, HA, Oued N’fis; Bailly-Choumara 1965c, AA, Oued Noun, Anezi, Tafraoute; Bailly-Choumara 1966, AA, Agadir; Bailly-Choumara 1967a, MA, route Azrou-Khénifra, piste Tafechna-Taoujgelt, Source Oumerrbia, Ghorm El Alem, piste Ksiba-Naour, piste Naour-Arbala, Zaouia Cheikh, Oued Sarif; Bailly-Choumara 1967b, EM, 7 km N d’Itzer, 8 km d’Itzer; Bailly-Choumara 1967c, Rif, route Chaouen-Bab Taza, Bab Taza, piste Bab Taza-Fifi, piste Bab Taza-Talassemtane, piste Bab Taza-Béni Ahmed, route Bab Taza-Bab Berred, piste Bab Taza-Asifane, piste Bab Berred-Tamorote, Ketama, piste Ketama-Jebel Tidighine, route Ketama-Targuist, route nationale Jebha, Al Hoceima, Béni Bouayache, Marchica, Targuist et environs, Jebha, Boured; Trari 1991, AP, Sidi Yahia du Gharb; Himmi et al. 1995; Louah 1995, Rif, Riffien, Tres piedras, Marina Smir, Bouzaghlal, M’diq, Oued Maleh, Azla, Tahaddart, Moulay Bouchta, Schroda, Kebbache, Talambote, Oued Maggou; Louah et al. 1995; Chlaida 1997, AP, Oued Sidi Messoud, Aïn Blal, Douar Chlihat, Barrage Al Massira; Chlaida and Bouzidi 1995, AP, Barrage El Massira; Dakki 1997; Ramdani 1997, AP, Skhirat, Tamaris Merja; Handaq 1998, HA, Oukaimeden, Amizmiz, Tiguenziouine (près Oued N’fis); Trari et al. 2002; Himmi 2007, Rif, Bab Berred, AP, Bouknadel, Douar jdid (Skhirat); El Ouali Lalami et al. 2010a, MA, Fès; Larhbali et al. 2010, MA, Oulmès; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Culex (Culex) perexiguus Theobald, 1903

Séguy 1930a; Callot 1940, AA, Assa; Senevet and Andarelli 1959a, EM, Oujda, Taourirt, AP, Aïn el Aouda, Arbaoua, Had Kourt, Oued Beht, Rabat, Allal Tazi, Oued Sahli, Zaouia Ech cheikh, Taghzirt, MA, Béni Mellal, Foum Zabel, Ifrane, Meknès, Aït Atta du Rteb, Fès, Sidi Mokhfi, Tahala, HA, Tazert, Midelt, AA, akka, Tamri; Bailly-Choumara 1965c, EM, Assa; Bailly-Choumara 1966, AA, Souk El Khémis Dades, Aït Ouabelli, Tarhjicht, Tirherm, Taoujgelt, Aït melloul; Bailly-Choumara 1967a, MA, piste Tafechna-Taoujgelt, piste Tafechna-Assoul, Ajdir; Bailly-Choumara 1967b, EM, Madagh, Merja Boubker, Aïn Béni Mathar; Bailly-Choumara 1967c, Rif, piste Bab Taza-Fifi, piste Bab Taza-Talassemtane, piste Bab Taza-Béni Ahmed, route Bab Taza-Bab Berred, Anasar, piste Bab Berred-Tamorote, Ketama, route Ketama-Targuist, route nationale Jebha, Béni Bouayache, Targusit et environs; Bailly-Choumara 1972a, AP, Merja Sheishat; Louah 1995, Rif, Riffien, Tres piedras, Marina Smir, Bouzeghlal, Oued Maleh, Tahaddart, Talembote, Schroda, Tanger; Louah et al. 1995; Mestari 1997, AP, Mohammedia; Moussalim 1997, AP, Sidi Allal Tazi, El Oulja, Fouarate; Ramdani 1997, AP, Skhirat, Tamaris; Handaq 1998, HA, Marrakech, entre Oued N’fis et Chichaoua, Kelaâ Sraghna; Alaoui Slimani 2002, AP, Rabat, Salé; Trari et al. 2002; Himmi 2007, Rif, Bab Berred, Bab Taza; Faraj et al. 2008c, AP, Larache Louamra; El Ouali Lalami et al. 2010a, MA, Fès; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017; Mouatassem et al. 2019, MA, Fès

Culex (Culex) pipiens Linnaeus, 1758

d’Anfreville 1916, AP, Salé; Charrier 1924, Rif, Tanger; Séguy 1930a; Callot 1940, SA, Goulimine; Viamonte and Ramirez 1946, Rif, Boudinar, Dar Benkarrich, Tétouan, Tanger, Asilah, Ksar El Kébir, Chefchaouen, Ketama, Nador; Gaud 1952, AP, Gharb; Séguy 1953a, SA, Tindouf; Guy 1958, HA, Oued N’fis; Guy et al. 1958, HA, Oued N’fis; Bailly-Choumara 1965c, EM, Aouinet Aït Oussa, Aïn Isker, Aïn Aït Delouine, Oued Mesdourt, Talmesdourt, Toudi, AA, Aït Onmar, Oulad Teima, Taroudant, Tiznit ville, Talaint, Hassi Tafnidilt, Zaouiat Cheikh, Aïn Guerzim, Tafraout ville, SA, Goulimine ville, Vallée de l’Oued Assaka, Ouaroun, Zriouila, Labyar, Tighmert, Abeino, Tantan ville, Zag; Bailly-Choumara 1966, AA, piste d’Akka au Draa, Akka et environs, Aït Ouabelli, Anamere-Smougue, Tarhjicht, Aït Melloul, Tiznit et environs; Bailly-Choumara 1967a, MA, route Azrou-Khénifra, piste Tafechna-Taoujgelt, piste Tafechna-Assoul, maison forestière Ouiouane, piste Tafechna-Senoual-Itzer, Ajdir, Itzer, piste Itzer-Boumia, Ghorm El Alem, Ghorm El Alem-El Ksiba, El Ksiba, piste Ksiba-Naour, Zaouia Cheikh, Oued Sarif, Dayet Aoua, Pont Tarmilate, Ifrane, Imilchil; Bailly-Choumara 1967b, EM, route Itzer-Midelt, Gaada de Debdou, Guercif ville, Cascade Oued Za, Grotte du Zegzel, Environs Saidia, Douar Aïn Shebbak, Douar Aïn Zabia, Douar Mardarh, Douar Sidi Hashas, Saidia, Merja Boubker, Berguent, Tendrara, Figuig; Bailly-Choumara 1967c, Rif, Chaouen ville, route Chaouen-Bab Taza, piste Bab Taza-Talassemtane, piste Bab Taza-Béni Ahmed, route Bab Taza-Bab Berred, Anasar, piste Bab Berred-Tamorote, Ketama, route Ketama-Targuist, route nationale Jebha, Al Hoceima, Al Hoceima Club Med, Béni Bouayache, Marchica, Targuist et environs, Jebha, Ghafsai; Bailly-Choumara 1972a, AP, Merja Sheishat; Bailly-Choumara 1973b, AP, Merja Bokka, Merja Qodiya; Metge and Belakoul 1989, AP, Benslimane, Sidi Bettache; Himmi 1991, AP, Sidi Boughaba, Sidi Amira; Trari 1991, AP, Maâmora, El Menzeh, Sidi Boughaba, Chkaïfien, Sidi Yahia du Gharb, Bokka, Merja Zerga, Sidi Allal Tazi, Oued Loukous, Aïn Chouk, Merja Bargha, Merja Oulad Skhar; Bouallam and Ramdani 1992, HA, Marrakech; El Bermaki 1993, AP, Sidi Maârouf; Himmi et al. 1995; Louah 1995, Rif, Fnideq, Riffien, Tres piedras, Marina Smir, M’diq, Cabo Negro, Zekri, Azla, Tahaddart, Schroda, Kebbache, Ouadras, Punta cirres, Sidi Kankoch, Oued Kbir; Louah et al. 1995; Bouallam et al. 1997b, HA, Marrakech; Dakki 1997; Mestari 1997, AP, Mohammedia; Himmi et al. 1998, AP, Sidi Boughaba, Puits de Sidi Amira (forest of Maâmora); Moussalim 1997, AP, El Oulja, Maâmora, Fouarate, Sidi Boughaba; Ramdani 1997, AP, meseta côtière (Témara-Casablanca); Faraj et al. 1997, AP, Kénitra; Handaq 1998, AP, Zemamra, MA, Oued Zem, HA, Marrakech, Zaouiet Ben Sassi, Sidi Bou Othmane, Kelaâ Sraghna, Bengrir; Bouallam 2001, HA, Marrakech; Alaoui Slimani 2002, AP, Rabat; Trari et al. 2002; Aouinty et al. 2006, AP, Mohammedia; Faraj et al. 2006, AP, Salé; Himmi 2007, Rif, Bab Berred, Bab Taza, Stehat, AP, Skhirate, forest of Maâmora, forest of Hilton, Sidi Boughaba, El Oulja, Ouled Salem, Ouled dlim, Douar Ould Yahia Ben Ali, Larouaza, Douar Elarja, Douar Jdid, Douar Jnaja; Faraj et al. 2008b, AP, Louamra; El Ouali Lalami et al. 2010a, MA, Fès; Larhbali et al. 2010, MA, Oulmès, Tarmilate, Bni Ounzar, Ganzra;Louali Lalami et al. 2010b, MA, Oued El Himmer; Amraoui 2012, HA, Marrakech; Amraoui et al. 2012, Rif, Tanger, AP, Mohammedia, Casablanca, HA, Marrakech; Amraoui et al. 2012, AP, Mohammedia, Casablanca; El Ouali lalami 2012, MA, route de Sidi Harazem; Hadji et al. 2013, AP, Sidi Slimane; El Joubari et al. 2014, Rif, Smir lagoon; El Joubari et al. 2015a, Rif, Smir lagoon; Marc et al. 2016, AP, Kénitra; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017; Bkhache et al. 2018, Rif, Tanger, AP, Rabat, Mohammedia, HA, Marrakech; Tmimi et al. 2018, AP, Mohammedia; Mouatassem et al. 2019, MA, Fès

Culex (Culex) simpsoni Theobald, 1905

Callot 1940, AA, Taghicht; Senevet et al. 1949, AA, Oued Noun, Tafnidelt, Akka; Gaud 1953a, AA, Imsouane, Aït Melloul, Akka, O’Noun, Taghjicht, Assa, Tafnidilt; Bailly-Choumara 1965c, EM, Aman d’Aït Oussa, El Megrinat, AA, Oued Izi, Oued Massa-Pont de la route Agadir-Tiznit, Guelta Zerga, Tafnidilt, SA, Poste militaire de Boujrif, Tirhmert, Taourirt-Barrage, Aouinet Torkoz; Bailly-Choumara 1966, AA, Tirherm, Taoujgelt, Taghjicht, Aït melloul, Tiznit; Chlaida and Bouzidi 1995, AP, Sidi M’barek, Oued Sidi Messoud, Mechrâa, Sidi Boulâarais, Douar Chlihat (south of Settat); Chlaida 1997, AP, Sidi M’barek, Mechrâa Settir, Sidi Boulâarais, Douar Chlihat (south of Settat); Dakki 1997; Handaq 1998, HA, Sidi Bou Othmane (Marrakech); Trari et al. 2002; Himmi 2007, Rif, Bab Berred, MA, Khémisset, AA, sud Anti Atlas; El Ouali Lalami et al. 2010a, MA, Fès; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Culex (Culex) theileri Theobald, 1903

Callot 1940, AA, Taghjicht; Viamonte and Ramirez 1946, Rif, Dar Benkarrich, Boudinar, Ketama, Tanger, Tétouan, Chefchaouen, AP, Larache; Guy 1958, HA, Oued N’fis; Guy et al. 1958, HA, Oued N’fis; Bailly-Choumara 1965c, EM, Aouinet Aït Oussa, Aïn Oumesdour, Aouinet Torkoz, SA, Tirh Mzoun; Bailly-Choumara 1966, AA, Souk El Khémis Dades, Akka et environs, Aït Melloul, Tazenakhte, piste Tiznit-Tafraout, SA, Goulimine; Bailly-Choumara 1967a, MA, piste Tafechna-Taoujgelt, piste Tafechna-Assoul, Aguelmane Azigza, piste Aguelmane Azigza-Aïn Leuh, maison forestière Ouiouane, Khénifra, piste Tafechna-Senoual-Itzer, Ajdir, Itzer, Col du Zad, piste Itzer-Boumia, Ghorm El Alem, piste Ghorm El Alem-El Ksiba, El Ksiba, piste Ksiba-Naour, Aguelmane Moulay Yakoub, Zaouia Cheich, Kafensour, Dayet Aoua, Pont Tarmilate; Bailly-Choumara 1967b, EM, route Itzer-Midelt, 40 km N Outat El Haj, Taourirt, Driouch, Figuig; Bailly-Choumara 1967c, Rif, piste Bab Taza-Fifi, piste Bab Taza-Talassemtane, route Bab Taza-Bab Berred, Anasar, piste Bab Berred-Tamorote, Ketama, route Ketama-Targuist, ArbaaTaourirt, Al Hoceima, Targuist et environs, Ghafsai; Bailly-Choumara 1972a, AP, Merja Sheishat; Bailly-Choumara 1973b, AP, Merja Sheishat, Merja Bokka; Himmi 1991, AP, Sidi Boughaba, El menzeh, Sidi Amira; Trari 1991, AP, Maâmora, Oued Sebou, Bordure Oued Loukous, Sidi Yahia du Gharb, Bokka, Entre Moulay Bousselham et Larache, Merja Zerga, Larache; El Bermaki 1993, AP, Sidi Maârouf, El Oulfa; Louah 1995, Rif, Bouzeghlal, M’Diq, Oued Maleh, Zekri, Lajour, Tahaddart, Loubart, Chefchaouen, Tanger; Himmi et al. 1995; Louah et al. 1995; Dakki 1997; Mestari 1997, AP, Mohammedia; Moussalim 1997, AP, Sidi Allal Tazi, Maâmora, Rabat, Kénitra; Ramdani 1997, AP, Skhirat, Tamaris; Handaq 1998, AP, Zemamra, MA, Béni Mellal, HA, Oukaimeden, Amizmiz, Tiguenziouine, Marrakech, Sidi Bou Othmane, Seguarta-Kelaâ, Had Mhara, Aîn Äounate, Bengrir; Himmi et al. 1998, AP, Sidi Boughaba, Dayat d’El Menzeh (north east of Kénitra), Gharb; Alaoui Slimani 2002, AP, Rabat; Trari et al. 2002; Himmi 2007, Rif, Bab Berred, Tanakoub, AP, Skhirate, forest of Maâmora, Chiahna, Sidi Amira, Sidi Boughaba, Bouknadel, Sidi Azzouz, Ehssaïne, Bettana, Sidi Yahia, Sebbah, Larouaza, Douar Elarja, Douar Jdid, Douar Jnaja; Faraj et al. 2008b, AP, Larache Louamra; El Ouali Lalami et al. 2010a, AP, Boucharen;Larhbali et al. 2010, MA, Roumani, Aïn Sbite, Ezzhiliga, Oulmès, Tarmilate, M’rirt, Bni Ounzar, Ganzra (Khémisset); Hadji et al. 2013, AP, Sidi yahia du Gharb, Kcebia, Sidi Hagouch, Dar Belamri (Sidi Slimane); El Joubari et al. 2014, Rif, Smir lagoon; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017; Mouatassem et al. 2019, MA, Fès

Culex (Maillotia) deserticola Kirkpatrick, 1925

Gaud 1947c, AA, Tansikht; Gaud 1953a, EM, Figuig, Berguent, Anoual, Boudnib, Aïn Chair, Aoufous, Tarhit, HA, Tazenakht, Tichka, Tizi-n’Telghemt, AA, Ouarzazate, Zagora; Bailly-Choumara 1965c, EM, Oued Isker, El Megrinat, Taskala, Aïn Aït Delouine, Talmesdourt, Aouinet Torkoz, Bouanama, Rich Tamlougout, Assa, AA, Tafraout ville, Oued Jemâa Idaousmaal, Aït abdallah, Issedrim Igmur Igues, SA, Vallée de l’Oued Assaka, Tacharicht, Bou Izakarn, Aïn Erkha; Chlaida and Bouzidi 1995, AP, Sud de Settat; Himmi et al. 1995; Chlaida 1997, AP, Barrage Al Massira; Dakki 1997; Trari et al. 2002; Himmi 2007; El Ouali Lalami et al. 2010a, MA, Fès; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Culex (Maillotia) hortensis Ficalbi, 1889

Charrier 1924, Rif, Tanger; Séguy 1930a; Langeron 1938, HA, Tounfite; Callot 1940, HA, Anefgou (2500 m), Tirghist (2500 m), Tighermine (2500 m); Gaud 1953a, HA, Tizi-n’Telghemt, Tizi-n’Tichka; Bailly-Choumara 1967a, MA, piste tafechna-Znan Imes, Khénifra, piste Tafechna-Senoual-Itzer, Ajdir, Itzer, Ghorm El Alem, El Ksiba, piste Ksiba-Naour, Piste Naour-Arbala, piste Arbala-El Kbab, Zouia Cheikh, Oued Sarif, Ifrane, Boulmane, Imilchil; Bailly-Choumara 1967b, EM, Tafraout, Grotte du Zegzel; Bailly-Choumara 1967c, Rif, Chaouen ville, route Chaouen-Bab Taza, Bab Taza, piste Bab Taza-Fifi, piste Bab Taza-Talassemtane, piste Bab Taza-Béni Ahmed, route Bab Taza-Bab Berred, piste Bab Berred-Tamorote, route Bab Berred-Ketama, Ketama, piste Ketama-Mt Tiguidin, route nationale Jebha, route Targuist-Béni Boufrah,Trari 1991, AP, Gharb; El Bermaki 1993, AP, Casablanca; Himmi et al. 1995; Louah 1995, Rif, Riffien, Marina Smir, M’diq, Oued Maleh, Lajour, Tahaddart, Schroda, Talembote, Bab Berred, Punta Cirres, Sidi Kankoch, Oued kbir, Oued Jebel Lehbib; Louah et al. 1995; Chlaida and Bouzidi 1995, AP, Barrage Al Massira; Chlaida 1997, AP, Barrage Al Massira; Dakki 1997; Ramdani 1997, AP, Skhirat, Tamaris; Handaq 1998, HA, Amizmiz, Sidi Bou Othmane, Had Mhara, Bengrir; Trari et al. 2002; Himmi 2007, Rif, Bab Berred, Bab Taza, AP, Skhirate; El Ouali Lalami et al. 2010b, MA, Oued Fès, Oued El Himmer, Camping Sidi Harazem; Larhbali et al. 2010, MA, Merchouch, Ghoualem, Ezzhiliga, Oulmès, Tarmilate, M’rirt, Bni Ounzar, Ganzra (Khémisset); El Ouali Lalami 2012, MA, Oued Fès; Oued El Himmer, route de Sidi Harazem, Camping Sidi Harazem; Hadji et al. 2013, AP, Sidi Yahia du Gharb, Kcebia, Sidi Hagouch, Dar Belamri, Lalla Itto, Soualem (Sidi Slimane); Trari 2017; Trari and Dakki 2017b; Trari et al. 2017; Mouatassem et al. 2019, MA, Fès

Culex (Neoculex) impudicus Ficalbi, 1890

Charrier 1924, Rif, Tanger; Séguy 1930a; Bailly-Choumara 1966, AA, Taliouine; Bailly-Choumara 1967a, MA, route Azrou-Khénifra, piste Tafechna-Senoual-Itzer, Itzer, piste Naour-Arbala, Zaouia Cheikh, Pont Tarmilate; Bailly-Choumara 1967b, EM, route Itzer-Midelt, Cascade Oued Za, Grotte du Zegzel; Bailly-Choumara 1967c, Rif, Chaouen ville, route Chaouen-Bab Taza, piste Bab Taza-Fifi, piste Bab Taza-Talassemtane, piste Bab Taza-Béni Ahmed, route Bab Taza-Bab Berred, piste Bab Berred-Asifane, piste Bab Berred-Tamorote, Ketama, route nationale Jebha, Al Hoceima Club Med, Targuist et environs; Trari 1991, AP, Sidi Amira, El Menzeh, Oued Sebou, Sidi Yahia du Gharb, Moulay Bousselham, Larache, Bordure Oued Loukous; Dakki 1997; Himmi et al. 1998, AP, Sidi Boughaba, Dayat d’El Menzeh (north east of Kénitra), Puits de Sidi Amira (forest of Maâmora); Trari et al. 2002; Himmi 1991, AP, Sidi Boughaba, El Menzeh, Sidi Amira; Himmi et al. 1995; Louah 1995, Rif, Haidra, Marina Smir, Tahaddart, Tanger; Louah et al. 1995; Ramdani 1997, AP, Tamaris; Alaoui Slimani 2002, AP, Rabat; Himmi 2007, Rif, Bab Berred, Bab Taza, AP, forest Maâmora; El Ouali Lalami et al. 2010a, MA, Fès; El Joubari et al. 2014, Rif, Smir lagoon; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Culex (Neoculex) martinii Medschid, 1930

Bailly-Choumara 1968a, Rif, Al Hoceima, AP, Sidi Yahia du Gharb, Sidi Allal Tazi, Larache, Rabat, HA, Marrakech, AA, Tiznit; Himmi et al. 1995; Louah 1995, Rif, Haidra, Marina Smir, Cabo Negro, Oued Maleh; Louah et al. 1995; Dakki 1997; Trari et al. 2002; Himmi 2007; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Culisetini

Culiseta Felt, 1904

Culiseta (Allotheobaldia) longiareolata (Macquart, 1838)

d’Anfreville 1916, AP, Salé; Séguy 1930a; Gaud 1947c, AA, Tansikht; Gaud 1953a, HA, Marrakech; Bailly-Choumara 1965c, EM, El Aïoun du Draa, Aïn Aït Delouine, Talmesdourt, Aouinet Torkoz, Rich Tamlougout, AA, Aït Onmar, Ouled Teima, Tiznit ville, Bounaamane, Id Baha, Tafnidilt, Guelta Zerga, Aïn Kerma, Tafraout ville, Oued Jemâa Idaousmaal, Toudi, Aït Abdallah, Igherm, Issedrim Igmur Igues, SA, Goulimine ville, Poste militaire de Boujrif, Embouchure de l’Oued Assaka, Ouaroun, Labyar, Asrir, Tacharicht, Bou Izakarn ville, Jemâa N’Tirhirte, Aït Erkha, Tantan ville; Bailly-Choumara 1966, AA, Aït melloul, Tiznit; Bailly-Choumara 1967a, MA, Jnane Imasse, Khénifra, piste Tafechna-Senoual-Itzer, Itzer, Ghorm El Alem, El ksiba, piste Naour-Arbala, Ifrane, Pont Tarmilate, Imouzzer Marmoucha; Bailly-Choumara 1967b, EM, Tafraout, Guercif ville, Grotte du Zegzel, Berguent, Tendrara; Bailly-Choumara 1967c, Rif, Chaouen ville, route Chaouen-Bab Taza, Bab Taza, route Bab Taza-Bab Berred, route Ketama-Targuist, route nationale Jebha, Targuist et environs, Aïn Hamra; Bailly-Choumara 1968a, AP, Meja Bokka, AA, Tiznit; Himmi 1991, AP, Sidi Boughaba, El Menzeh; Trari 1991, AP, El Menzeh, Gharb; El Bermaki 1993, AP, Sidi Maârouf; Himmi et al. 1995; Louah 1995, Rif, Fnideq, Riffien, Marina Smir, M’diq, Cabo Negro, Lajour, Ouadras, Punta Cirres, Ksar Sghir, Tanger; Louah et al. 1995; Mestari 1997, AP, Mohammedia; Moussalim 1997, AP, Maâmora, Fouarate; Ramdani 1997, AP, Témara, Skhirat, Tamaris; Dakki 1997; Handaq 1998, HA, Marrakech, Kelaâ Sraghna, Bengrir Himmi et al. 1998, AP, Sidi Boughaba, Dayat d’El Menzeh (north east of Kénitra); Bouallam 2001, HA, Bordure Oued N’fis; Alaoui Slimani 2002, AP, Rabat; Trari et al. 2002; Aouinty et al. 2006, AP, Mohammedia; Himmi 2007, Rif, Bab Berred, AP, Skhirate, Maâmora, forest of Hilton; Koçak and Kemal 2013; El Joubari et al. 2014, Rif, Smir lagoon; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017; Mouatassem et al. 2019, MA, Fès

Culiseta (Culicella) fumipennis (Stephens, 1825)

Gaud 1947c, AP, Rabat, Sidi Allal Tazi, MA, Khémisset; Senevet and Andarelli 1959, AP, Rabat, Casablanca, Bouznika, MA, Fès; Bailly-Choumara 1967a, MA, Jnane Imasse; Bailly-Choumara 1967c, Rif, route Bab Taza-Bab Berred; Himmi et al. 1995; Dakki 1997; Trari et al. 2002; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Culiseta (Culicella) litorea (Shute, 1928)

Metge 1986, AP, Casablanca; Dakki 1997; Carles-Tolrá 2002; Trari et al. 2002; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Culiseta (Culiseta) annulata (Schrank, 1776)

d’Anfreville 1916, AP, Salé; Charrier 1924, Rif, Tanger; Séguy 1930a, Rif, Tanger; Viamonte and Ramirez 1946, Rif, Ben Karrich, Asilah, Ketama, Malaliene, Tétouan, AP, Larache; Gaud 1952, AP, Souk El Hadd (Gharb); Gaud 1953a, Rif, Tanger, Tétouan, EM, Saidia, Talsint, AP, Casablanca, Rabat, Kénitra, MA, Meknès, Fès, Ifrane, Taza, HA, Marrakech, Aït Bouguemez, Midelt; Gaud 1957b, Rif, Tétouan, Tanger, AP, Sidi Allal Tazi, Rabat, Casablanca, MA, Meknès, Ifrane; Bailly-Choumara 1967a, MA, Jnane Imasse, maison forestière Ouiouane, piste Tafechna-Senoual-Itzer, Ghorm El Alem, Zaouia Cheikh, Pont Tarmilate, Ifrane; Bailly-Choumara 1967b, EM, Tafraout, route Itzer-Midelt; Bailly-Choumara 1967c, Rif, piste Bab Taza-Talassemtane; Bailly-Choumara 1972a, AP, Merja Sheishat; Himmi et al. 1995; Louah 1995, Rif, Riffien, M’diq, Tanger; Louah et al. 1995; Chlaida 1997, AP, Barrage Al Massira; Dakki 1997; Mestari 1997, AP, Mohammedia; Moussalim 1997, AP, Rabat; Ramdani 1997, AP, meseta côtière (Casablanca-Rabat); Himmi et al. 1998, AP, Sidi Boughaba; Himmi 1991, AP, Sidi Boughaba, El Menzeh; Trari 1991, AP, Sidi Boughaba, El Menzeh; Trari et al. 2002; Himmi 2007, AP, Skhirate, Maâmora, forest of Hilton, Oulja; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Culiseta (Culiseta) subochrea (Edwards, 1921)

Gaud 1952, AP, Gharb; Gaud 1957b, Rif, Tanger, Chefchaouen, EM, Oujda, AP, Sidi Allal Tazi, Casablanca; Bailly-Choumara 1967a, MA, Imilchil; Bailly-Choumara 1967c, Rif, Ketama; Bailly-Choumara 1972a, AP, Merja Sheishat; Himmi 1991, AP, Sidi Boughaba; Trari 1991, AP, Sidi Boughaba; Louah 1995, Rif, Riffien, Tres piedras, M’diq, Cabo Negro, Oued Maleh, Tanger; Louah et al. 1995; Dakki 1997; Mestari 1997, AP, Mohammedia; Moussalim 1997, AP, Kénitra; Ramdani 1997, AP, Témara, Skhirat, Casablanca; Handaq 1998, AP, El Jadida, HA, Bengrir; Himmi et al. 1998, AP, Sidi Boughaba; Trari et al. 2002; Himmi 2007, AP, Skhirat, Dayet Aïn Chems, Maâmora, forest of Hilton; El Joubari et al. 2014, Rif, Smir lagoon; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Mansoniini

Coquillettidia Dyar, 1905

Coquillettidia (Coquillettidia) buxtoni (Edwards, 1923)

Bailly-Choumara 1965a, AP, Merja Bokka; Bailly-Choumara 1970, AP, Merja Bokka; Bailly-Choumara 1972a, AP, Merja Sheishat (Larache); Bailly-Choumara 1973b, AP, Merja Bokka, Larache; Himmi et al. 1995; Trari et al. 2002; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Coquillettidia (Coquillettidia) richiardii (Ficalbi, 1889)

Bailly-Choumara 1965a, AP, Merja Bokka; Bailly-Choumara 1967a, MA, Zaouia Cheikh; Bailly-Choumara 1970, AP, Merja Bokka; Bailly-Choumara 1972a, AP, Merja Sheishat (Larache); Bailly-Choumara 1973b, AP, Merja Bokka; Trari 1991, AP, Sidi Yahia du Gharb, Aïn Chouk, Merja Bargha; Himmi et al. 1995; Dakki 1997; Moussalim 1997, AP, Fouarate sur les bordures de la Merja; Trari et al. 2002; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Orthopodomyiini

Orthopodomyia Theobald, 1904

Orthopodomyia pulcripalpis (Rondani, 1872)

Bailly-Choumara 1965b, AP, Maâmora; Himmi et al. 1995; Dakki 1997; Trari et al. 2002; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017

Uranotaeniini

Uranotaenia Lynch Arribálzaga, 1891

Uranotaenia (Pseudoficalbia) unguiculata Edwards, 1913

Séguy 1930a; Gaud 1953a, HA, Marrakech; Senevet and Andarelli 1959a, EM, Oujda, Berguent, Guercif, AP, Sidi Allal Tazi, Bouznika, Casablanca, Béni Moussa, MA, Oulad Massine, Meknès, Moulay Yacoub, Béni Mellal, AA, Ksar Mzizel, Aït Melloul; Bailly-Choumara 1965c, EM, Aïn Aït Delouine, Talmesdourt, Rich Tamlougout; Bailly-Choumara 1966, AA, Aït Ouabelli; Bailly-Choumara 1967a, MA, Pont Tarmilate; Bailly-Choumara 1967b, EM, environs de Taourirt, Madagh, Merja Boubker; Bailly-Choumara 1967c, Rif, route Bab Taza-Bab Berred, piste Ketama-Jebel Tidighine; Bailly-Choumara 1968a, AP, Oued Loukous, Merja Bokka, Kénitra, Oued Bou-Regreg; Himmi 1991, AP, Sidi Boughaba; Trari 1991, AP, Oued Sebou; El Bermaki 1993, AP, El Oulfa (Casablanca); Himmi et al. 1995; Dakki 1997; Ramdani 1997, AP, Tamaris; Handaq 1998, AP, Sidi Bennour, HA, Zima Chemaîa (Bengrir); Trari et al. 2002; Himmi 2007, AP, Sidi Boughaba; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017; Mouatassem et al. 2019, MA, Fès

Uranotaenia (Uranotaenia) balfouri Theobald, 1904

Bailly-Choumara 1968a, AP, Merja Bokka, Oued Loukous, Sidi Yahia, Oued Sebou (Kénitra), Oued Bou-Regreg; Himmi et al. 1995; Dakki 1997; Moussalim 1997, AP, Sidi Allal Tazi, Fouarate, Sidi Boughaba; Trari et al. 2002; Himmi 2007, AP, Sidi Boughaba; El Ouali Lalami et al. 2010a, MA, Fès Boulmane; Trari 2017; Trari and Dakki 2017b; Trari et al. 2017; Mouatassem et al. 2019, MA, Fès

DIXIDAE

K. Kettani, R. Wagner

Number of species: 12. Expected: 15

Faunistic knowledge of the family in Morocco: moderate

Dixa Meigen, 1818

Dixa caudatula Séguy, 1928

Séguy 1930a, HA, Arround, Skoutana (2400 m); Vaillant 1959; Dakki 1997; Mouna 1998

Dixa dilatata Strobl, 1900

= Dixa riparia Vaillant, in Vaillant 1959: 180

Vaillant 1959, HA, Source de M’Goum (2500 m), Gorges d’Imi-N’Ifri (1050 m); Vaillant 1965; Dakki 1997

Dixa maculata Meigen, 1818

Séguy 1930a; Dakki 1997; Mouna 1998

Dixa mera Séguy, 1930

Séguy 1930a, MA, forest of Timelilt (1900 m); Vaillant 1959; Dakki 1997; Mouna 1998

Dixa nebulosa Meigen, 1830

Séguy 1930a, HA; Dakki 1997; Mouna 1998

Dixa perexilis Séguy, 1928

Séguy 1930a, HA, riverside of Oued Imminen (Tachdirt, 2400 m); Vaillant 1959; Dakki 1997; Mouna 1998

Dixa puberula Loew, 1849

Vaillant 1959, HA, headwaters of Asif M’Goum (2500 m); Vaillant 1965; Dakki 1997; Mouna 1998

Dixa submaculata Edwards, 1920

Séguy 1930a, MA, Sidi Yahia, Talzent (1800 m); Dakki 1997; Mouna 1998

Dixella Dyar & Shannon, 1924

Dixella aestivalis (Meigen, 1818)

Séguy 1930a, AP, Merja Boughaba; Vaillant 1965; Ramdani 1981; Ramdani and Tourenq 1982; Mouna 1998

Dixella attica (Pandazis, 1933)

= Dixella numidica (Sicart, 1955)

Ebejer et al. 2019, Rif, Cabo Negro (indoors: 10 m)

Dixella martinii (Peus, 1934)

Ebejer et al. 2019, Rif, Moulay Abdelsalam (965 m)

Dixella serotina (Meigen, 1818)

= Dixa serotina Wied, in Mouna 1998: 85

Séguy 1930a, AP, Casablanca (between Kénitra and Oued Beth); Dakki 1997; Mouna 1998

Chironomoidea

CERATOPOGONIDAE

K. Kettani, B. Mathieu

Number of species: 62. Expected: 80

Faunistic knowledge of the family in Morocco: moderate

Ceratopogoninae

Culicoidini

Culicoides Latreille, 1809

Culicoides (Avaritia) imicola Kieffer, 1913

Kremer et al. 1971, MA, Fès-Meknès, SA, Guelmim-Oued Noun; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, SA, Guelmim-Oued Noun; Chaker et al. 1980, MA, Fès, Rhafsai, AA, Torkoz, Tarhjisht; Remm 1988a; Dakki 1997; Bouayoune et al. 1998, Rif, Tanger-Tétouan-Al Hoceima, EM, Oriental, AP, Rabat-Salé-Kénitra, Safi, MA, Fès-Meknès, Béni Mellal-Khénifra, HA, Marrakech, AA, Draa-Tafilalet, Souss-Massa, SA, Guelmim-Oued Noun; Cêtre-Sossah and Baldet 2004, AP, Rabat-Salé-Kénitra; Lhor et al. 2015, Rif, Sahel Chamali, MA, Sidi Hammadi, Benioukil, Aïn Leuh, Ait Siberne, Meknès, AA, Errachidia, Sidi Dahmane, SA, Foum El Oued; Bourquia et al. 2019, AP, Rabat

Culicoides (Avaritia) montanus Shakirzjanova, 1962

Kremer et al. 1971, AP, Rabat-Salé-Kénitra, Safi, HA, Marrakech, SA, Guelmim-Oued Noun; Kremer et al. 1975; Kremer et al. 1979, Rif, Tanger-Tétouan-Al Hoceima, AP, Safi, HA, Marrakech; Chaker et al. 1980, Rif, Al Hoceima, AP, Oued Cherrat, HA, Souk Tnine de Oudaias (Haouz), Marrakech, AA, Torkoz; Remm 1988a; Dakki 1997; Bourquia et al. 2019, AP, Rabat

Culicoides (Avaritia) obsoletus (Meigen, 1818)

Callot et al. 1968, Rif, Al Hoceima, AP, Merja Bokka, Sidi Yahia du Gharb, Sidi-Bettache (Zaeir), Rabat-Salé-Kénitra, HA, El Harcha (plateau central); Bailly-Choumara and Kremer 1970, AP, estuaire de Bou-Regreg; Kremer et al. 1971, AP, Rabat-Salé-Kénitra, Safi, MA, Béni Mellal-Khénifra, HA, Marrakech; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, Rif, Tanger-Tétouan-Al Hoceima, AP, Rabat-Salé-Kénitra, Casablanca, Settat, Safi, HA, Marrakech; Chaker et al. 1980, Rif, Al Hoceima, AP, Zaers, Sidi Bettache, HA, El Harcha, Talet Inaouane (Haouz); Remm 1988a; Dakki 1997; Bouayoune et al. 1998, Rif, Tanger-Tétouan-Al Hoceima, AP, Rabat-Salé-Kénitra, Casablanca, Settat, Safi, MA, Fès-Meknès, Béni Mellal-Khénifra, HA, Marrakech, AA, Souss-Massa, SA, Guelmim-Oued Noun; Cêtre-Sossah and Baldet 2004; Lhor et al. 2015, Rif, Sahel Chamali, MA, Ait Siberne; Bourquia et al. 2019, AP, Rabat

Culicoides (Avaritia) scoticus Downes & Kettle, 1952

Kremer et al. 1971, AP, Safi, MA, Béni Mellal-Khénifra, HA, El Harcha, Talet Inaouane (Haouz), Marrakech; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, AP, Casablanca-Settat, Safi, HA, Marrakech; Remm 1988a; Dakki 1997; Bourquia et al. 2019, AP, Rabat

Culicoides (Beltranmyia) circumscriptus Kieffer, 1918

Callot et al. 1968; Bailly-Choumara and Kremer 1970, Rif, Smir lagoon, Oued Negro, EM, Merja Boubker (Berkane), Gouttitir (NE Guercif), AP, Merja Sheishat (Larache), Aïn Muelha (near Oued Sidi Allal Tazi, estuaire Oued Bou-Regreg, Dayat Qoudiya (Sidi Yahia Gharb); Kremer et al. 1971, AP, Rabat-Salé-Kénitra, Casablanca, Settat, Safi, MA, Fès-Meknès, HA, Marrakech, SA, Guelmim-Oued Noun; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, Rif, Tanger-Tétouan-Al Hoceima, AP, Rabat-Salé-Kénitra, Casablanca, Settat, Safi, MA, Fès-Meknès, AA, Souss Massa; Chaker et al. 1980, Rif, Al Hoceima, AP, Merja Qoudiya, Merja Bokka, Sidi Yahia du Gharb, MA, Aïn Karma (Saiss), Oulmès, HA, Setti Fatma, AA, Aït Melloul (Souss); Dakki 1997; Cêtre-Sossah and Baldet 2004, AP, Rabat-Salé-Kénitra; Lhor et al. 2015, Rif, Sahel Chamali, MA, Sidi Hammadi, Benioukil, Meknès, AA, Errachidia, SA, Foum El Oued; Bourquia et al. 2019, AP, Rabat

Culicoides (Culicoides) fagineus Edwards, in Edwards et al. 1939

Kremer et al. 1971, AP, Rabat-Salé-Kénitra, Safi, MA, Béni Mellal-Khénifra, HA, Marrakech; Kremer et al. 1975; Chaker et al. 1979, AP, Rabat-Salé-Kénitra; Kremer et al. 1979; Chaker et al. 1980, AP, Rabat, Sidi Bettache, MA, Khemisset, HA, Marrakech; Remm 1988a; Dakki 1997; Bourquia et al. 2019, AP, Rabat

Culicoides (Culicoides) newsteadi Austen, 1921

= Culicoides (Culicoides) halophilus Kieffer, in Callot et al. 1968: 886, Bailly-Choumara and Kremer 1970: 386, Dakki 1997: 60

Callot et al. 1968, Rif, Cabo Negro (Ferma), Tétouan, Talerhza, Tanger-Tétouan-Al Hoceima, AP, Larache, Merja Bokka (Gharb), Aïn Chok, HA, Talet-Inaouan (Haouz); Bailly-Choumara and Kremer 1970, Rif, Smir lagoon, Oued Negro, AP, Merja Sheishat (Larache), Aïn Muelha (near Oued Sidi Allal Tazi, estuaire Oued Bou-Regreg, Dayat Qoudiya (Sidi Yahia du Gharb), EM, Merja Boubker (Berkane), Ksabi (NE Midelt), HA, Souk Tnine des Oudaias (bordure Oued N’fis), AA, Aïn Sefra (south Foum Zquid); Kremer et al. 1971, AP, Rabat-Salé-Kénitra, Casablanca, Settat, MA, Fès-Meknès, Béni Mellal-Khénifra, SA, Guelmim-Oued Noun; Kremer et al. 1975; Chaker et al. 1979, Rif, Tanger-Tétouan-Al Hoceima, AP, Rabat-Salé-Kénitra, Safi, Casablanca, Settat, MA, Béni Mellal- Khénifra, HA, Marrakech; Kremer et al. 1979; Baylis et al. 1997, Rif, Tanger, HA, Marrakech; Remm 1988a; Dakki 1997; Cêtre-Sossah and Baldet 2004, AP, Rabat-Salé-Kénitra; Lhor et al. 2015, MA, Aïn Leuh, Ait Siberne, Meknès, SA, Foum El Oued; Bourquia et al. 2019, AP, Rabat

Culicoides (Culicoides) pulicaris (Linnaeus, 1758)

Kremer et al. 1971, AP, Rabat-Salé-Kénitra, MA, Fès-Meknès, Béni Mellal-Khénifra, SA, Guelmim-Oued Noun; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, AP, Rabat-Salé-Kénitra, Safi, HA, Marrakech, SA, Guelmim-Oued Noun; Chaker et al. 1980, MA, Lalla Outka, Khénifra, Oulmès, HA, Talet Inaouan (Haouz), AA, Aouinet-Torkoz, Tarhjicht; Remm 1988a; Dakki 1997; Bouayoune et al. 1998, Rif, Tanger-Tétouan-Al Hoceima, EM, Oriental, AP, Rabat-Salé-Kénitra, Casablanca, Settat, Safi, MA, Fès-Meknès, Béni Mellal-Khénifra, HA, Marrakech, AA, Drâa-Tafilalet, Souss-Massa; Cêtre-Sossah and Baldet 2004; Lhor et al. 2015, Rif, Sahel Chamali, MA, Aïn Leuh, Aït Siberne, Meknès; Bourquia et al. 2019

Culicoides (Culicoides) punctatus (Meigen, 1804)

Callot et al. 1968, Rif, Tanger-Tétouan-Al Hoceima; Bailly-Choumara and Kremer 1970, Rif, Merja Smir, Oued Negro, AP, Merja Sheishat (Larache), estuaire Bou-Regreg, EM, Merja Boubker (Berkane); Kremer et al. 1971, Rif, Cabo Negro (Ferma), Tétouan, EM, Berkane, AP, Rabat-Salé-Kénitra, MA, Fès-Meknès, AA, Foum Zguid; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, AP, Casablanca-Settat; Dakki 1997; Lhor et al. 2015, Rif, Sahel Chamali, MA, Sidi Hammadi, Benioukil, Aïn Leuh, Meknès; Bourquia et al. 2019

Culicoides (Culicoides) subfagineus Delécolle & Ortega, 1998

Bourquia et al. 2019, AP, Rabat

Culicoides (Monoculicoides) parroti Kieffer, 1922

Bailly-Choumara and Kremer 1970, HA, Dar Saâda (Haouz); Kremer et al. 1971, AP, Rabat-Salé-Kénitra; Kremer et al. 1975; Kremer et al. 1979, AP, Safi, HA, Marrakech; Chakeret al. 1980, AP, Rabat, HA, Marrakech, Souk Tnine des Oudaias (Haouz); Remm 1988a; Dakki 1997; Bourquia et al. 2019

Culicoides (Monoculicoides) puncticollis (Becker, 1903)

Callot et al. 1968, AP, Merja Qoudiya, Sidi Yahia du Gharb, Romani (Zaers), Rabat-Salé-Kénitra, MA, Aïn Karma (Saiss), HA, Souk Tnine des Oudaias (Haouz); Bailly-Choumara and Kremer 1970 (reported as C. riethi and corrected by Kremer et al. 1971), Rif, Merja Smir, AP, Merja Sheishat (Larache), estuaire Bou-Regreg, Dayat Qoudiya (Sidi Yahia du Gharb), HA, Souk Tnine des Oudaias (bordure de l’Oued N’fis), Dar Saâda (Haouz), Talet Inouane (bordure marécageuse du lac du Barrage Lalla Taguergoust); Kremer et al. 1971, AP, Rabat-Salé-Kénitra, MA, Fès-Meknès; Kremer et al. 1975; Kremer et al. 1979, AP, Rabat-Salé-Kénitra, Safi, MA, Fès-Meknès, HA, Marrakech; Chaker et al. 1980, AP, Aïn Karma, Zaers, Roumani, HA, Souk-Tnine des Oudaias; Remm 1988a; Cêtre-Sossah and Baldet 2004, AP, Rabat-Salé-Kénitra; Bourquia et al. 2019, AP, Rabat

Culicoides (Oecacta) azerbajdzhanicus Dhzafarov, 1962

Bailly-Choumara and Kremer 1970, HA, Souk Tnine des Oudaias (bordure Oued N’fis); Kremer et al. 1971, AP, Safi, MA, Fès-Meknès, Beni Mellal-Khénifra, HA, Marrakech, SA, Guelmim-Oued Noun; Kremer et al. 1975; Kremer et al. 1979, AA, Souss-Massa SA, Guelmim-Oued Noun; Chaker et al. 1980, MA, Kkénifra, Rhafsai, HA, Marrakech, AA, Torkoz, Tarhjisht; Dakki 1997; Bourquia et al. 2019

Culicoides (Oecacta) longipennis Khalaf, 1957

Kremer et al. 1971, EM, Berkane AP, Safi, MA, Fès-Meknès, HA, Marrakech; Kremer et al. 1975; Remm 1988a; Dakki 1997; Bourquia et al. 2019, AP, Rabat

Culicoides (Oecacta) marcleti Callot, Kremer & Basset, 1968

Kremer et al. 1971, MA, Rhafsai, Fès-Meknès, SA, Guelmim-Oued Noun; Kremer et al. 1975; Remm 1988a; Dakki 1997; Bourquia et al. 2019

Culicoides (Oecacta) pallidus Khalaf, 1957

= Culicoides stackelbergi Dhzafarov, in Kremer et al. 1971: 662, Dakki 1997: 61

Kremer et al. 1971, AA, Torkoz, SA, Guelmim-Oued Noun; Kremer et al. 1971; Remm 1988a; Dakki 1997; Bourquia et al. 2019

Culicoides (Oecacta) ravus De Meillon, 1936

= Culicoides (Synhelea) subravus Cornet and Château, in Kremer et al. 1971: 664, Chaker et al. 1980: 85, Dakki 1997: 61

Kremer et al. 1971, AP, Safi, HA, Marrakech, SA, Guelmim-Oued Noun; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, AA, Souss-Massa, SA, Guelmim-Oued Noun; Chaker et al. 1980, HA, Marrakech, AA, Torkoz, Tarhjicht, Aït Ouaballi (Draa); Remm 1988a; Dakki 1997; Bourquia et al. 2019

Culicoides (Oecacta) sahariensis Kieffer, 1923

= Culicoides colluzzii Callot, Kremer and Bailly-Choumara, in Bailly-Choumara and Kremer 1970: 386, Chaker et al. 1980: 83, Dakki 1997: 61

Bailly-Choumara and Kremer 1970, AP, Merja Sheishat (Larache), EM, Merja Boubker (Berkane), HA, Souk Tnine des Oudaias (bordure Oued N’fis); Callot et al. 1970, AP, Larache, HA, Marrakech; Kremer et al. 1971, AP, Rabat-Salé-Kénitra, MA, Fès-Meknès, SA, Guelmim-Oued Noun; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, AP, Rabat-Salé-Kénitra, SA, Guelmim-Oued Noun; Chaker et al. 1980, AP, Merja Bokka (Gharb), Rabat (Zaers), EM, Berkane, MA, Fès, Khémisset, AA, Tarhjisht; Baylis et al. 1997; Dakki 1997; Bouayoune et al. 1998; Cêtre-Sossah and Baldet 2004; Bourquia et al. 2019, AP, Rabat

Culicoides (Oecacta) santonicus Callot, Kremer, Rault & Bach, 1966

Bailly-Choumara and Kremer 1970, AP, Merja Sheishat (Larache); Kremer et al. 1971, AP, Rabat-Salé-Kénitra; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, AP, Rabat-Salé-Kénitra; Bailly-Choumara et al. 1980, AP, Larache, Sidi Bettache, MA, Oulmès, EM, El-Harcha; Remm 1988a; Dakki 1997; Bourquia et al. 2019

Culicoides (Oecacta) semimaculatus Clastrier, 1958

Kremer et al. 1975, Rif, Tanger-Tétouan-Al Hoceima, AP, Larache, Casablanca-Settat, MA, Plateau Central (Khatouate); Chaker et al. 1979; Remm 1988a; Dakki 1997; Szadziewski and Dominiak 2006; Bourquia et al. 2019

Culicoides (Oecacta) sergenti Kieffer, 1921

= Culicoides (Oecacta) mosulensis Khalaf, in Chaker et al. 1980: 84, Dakki 1997: 60

Kremer et al. 1979, EM, Oriental, SA, Guelmim-Oued Noun; Chaker et al. 1980, AA, Tarhjicht, SA, Bou-Arfa; Dakki 1997; Bourquia et al. 2019

Culicoides (Oecacta) similis Carter, Ingram & Macfie, 1920

Kremer et al. 1971, AA, Torkoz, SA, Guelmim-Oued Noun; Kremer et al. 1975; Remm 1988a; Dakki 1997; Bourquia et al. 2019

Culicoides (Oecacta) truncorum Edwards, 1939

= Culicoides (Oecacta) sylvarum Callot and Kremer, in Kremer et al. 1971: 662, Remm 1988a: 65, Dakki 1997: 61

Kremer et al. 1971, AP, Rabat-Salé-Kénitra, MA, Béni Mellal- Khénifra; Kremer et al. 1975; Remm 1988a; Dakki 1997; Bourquia et al. 2019

Culicoides (Pontoculicoides) saevus Kieffer, 1922

Callot et al. 1968, AP, Sidi Yahia du Gharb, Rabat-Salé-Kénitra, Safi, MA, Aïn Karma (Saiss), HA, Talet Inouane, Marrakech, Souk Tnine des Oudaias (Haouz), AA, Aït Melloul (Souss), Ksar er Souk (Tafilalt), Tarhjicht, SA, Bou-Arfa; Bailly-Choumara and Kremer 1970, AP, Merja Sheishat (Larache), HA, Souk Tnine des Oudaias (bordure de l’Oued N’fis); Kremer et al. 1971, AP, Rabat-Salé-Kénitra, MA, Fès-Meknès SA, Guelmim-Oued Noun; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, EM, Oriental, AP, Safi, MA, Fès-Meknès, HA, Marrakech, AA, Drâa-Tafilalet, Souss-Massa, SA, Guelmim-Oued Noun; Bailly-Choumara and Kremer 1980; Dakki 1997; Cêtre-Sossah and Baldet 2004, AP, Rabat-Salé-Kénitra; Bourquia et al. 2019

Culicoides (Pontoculicoides) sejfadinei Dzhafarov, 1958

Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, EM, Oriental, AA, Drâa-Tafilalet; Bourquia et al. 2019

Culicoides (Remmia) kingi Austen, 1912

Bailly-Choumara and Kremer 1970, AA, Mrimima (Oued de Foum Zquid), Souss-Massa; Chaker et al. 1980, MA, Meknès, AA, Tarhjisht; Cornet and Brunhes 1994; Bourquia et al. 2019, AP, Rabat

Culicoides (Remmia) schultzei (Enderlein, 1908)

Callot et al. 1968, AP, Safi, HA, Talet Inouane, Marrakech; Dakki 1997; Bourquia et al. 2019

Culicoides (Sensiculicoides) badooshensis Khalaf, 1961

Bailly-Choumara and Kremer 1970, AP, Merja Sheishat (Larache), HA, Marrakech, Souk Tnine des Oudaias; Kremer et al. 1971, AP, Rabat-Salé-Kénitra, Safi, MA, Fès-Meknès; Kremer et al. 1975; Kremer et al. 1979, AP, Rabat-Salé-Kénitra, Safi, HA, Marrakech; Dakki 1997; Chaker et al. 1980, AP, Oued Cherrat, Bousselham, Sidi Yahia, MA, Fès, HA, Marrakech; Bourquia et al. 2019

Culicoides (Sensiculicoides) cataneii Clastrier, 1957

Kremer et al. 1971, AP, Rabat-Salé-Kénitra; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, Rif, Tanger-Tétouan-Al Hoceima, AP, Rabat-Salé-Kénitra; Chaker et al. 1980, AP, Oued Cherrat, Rabat; Remm 1988a; Dakki 1997; Bourquia et al. 2019, AP, Rabat

Culicoides (Sensiculicoides) clastrieri Callot, Kremer & Deduit, 1962

Bourquia et al. 2019, MA, Fès-Meknès

Culicoides (Sensiculicoides) derisor Callot & Kremer, 1965

Chaker et al. 1980, AP, Rabat; Kremer et al. 1971, AP, Rabat-Salé-Kénitra, MA, Fès-Meknès; Kremer et al. 1975; Chaker et al. 1979, AP, Rabat-Salé-Kénitra; Kremer et al. 1979; Remm 1988a; Dakki 1997; Bourquia et al. 2019

Culicoides (Sensiculicoides) duddingstoni Kettle & Lawson, 1955

Bourquia et al. 2019, MA, Fès-Meknès

Culicoides (Sensiculicoides) dzhafarovi Remm, 1967

Kremer et al. 1971, AP, Rabat-Salé-Kénitra; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, SA, Guelmim-Oued Noun; Chaker et al. 1980, AP, Oued Cherrat, AA, Tarhjisht; Remm 1988a; Dakki 1997; Bourquia et al. 2019

Culicoides (Sensiculicoides) festivipennis Kieffer, 1914

= Culicoides (Oecacta) odibilis Austen, in Bailly-Choumara and Kremer 1970: 387, Chaker et al. 1980: 84, Dakki 1997: 61

Callot et al. 1968, Rif, Tanger-Tétouan-Al Hoceima; Bailly-Choumara and Kremer 1970, Rif, Merja Smir, HA, Souk Tnine des Oudaias (bordure Oued N‘fis); Kremer et al. 1971, AP, Rabat-Salé-Kénitra; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, Rif, Tanger-Tétouan-Al Hoceima, AP, Rabat-Salé-Kénitra, Safi, Casablanca-Settat, HA, Marrakech; Chaker et al. 1980, Rif, Tétouan, EM, El-Harcha, AP, Aïn Chok, Rabat, Larache, MA, Oulmès, HA, Marrakech, Talet-Inaouan (Haouz); Dakki 1997; Cêtre-Sossah and Baldet 2004, AP, Rabat-Salé-Kénitra; Bourquia et al. 2019, AP, Rabat

Culicoides (Sensiculicoides) heteroclitus Kremer and Callot, in Callot & Kremer, 1965

Kremer et al. 1975, AP, Safi, HA, Marrakech, AA, Tafraout, Tiznit, Souss-Massa; Chaker et al. 1979; HA, Haouz; Remm 1988a; Dakki 1997; Bourquia et al. 2019

Culicoides (Sensiculicoides) jumineri Callot & Kremer, 1969

Kremer et al. 1971, AP, Rabat-Salé-Kénitra, MA, Fès-Meknès, SA, Guelmim-Oued Noun; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, EM, Oriental, AP, Rabat-Salé-Kénitra, Safi, HA, Marrakech, AA, Souss-Massa, SA, Guelmim-Oued Noun; Chaket et al. 1980, AP, Oued Cherrat, Merja Bokka, Rabat, MA, Fès, HA, Talet-Inaouan (Haouz), AA, Torkoz, Tarhjisht, Aït Oubelli, SA, Bou-Arfa; Remm 1988a; Dakki 1997; Cêtre-Sossah and Baldet 2004, AP, Rabat-Salé-Kénitra; Bourquia et al. 2019, AP, Rabat

Culicoides (Sensiculicoides) kibunensis Tokunaga, 1937

= Culicoides (Oecata) cubitalis Edwards, in Kremer et al. 1975: 205, Dakki 1997: 60

Kremer et al. 1975, AP, Safi, MA, Ifrane, Imouzzer-du-Kander, Fès-Meknès, HA, Haouz, Marrakech; Chaker et al. 1979; Dakki 1997; Bourquia et al. 2019

Culicoides (Sensiculicoides) kurensis Dzhafarov in Gutsevich, 1960

Remm 1988a; Cêtre-Sossah and Baldet 2004, AP, Rabat-Salé-Kénitra; Bourquia et al. 2019

Culicoides (Sensiculicoides) landauae Kremer, Rebholtz-Hirtzel & Bailly-Choumara, 1975

Kremer et al. 1975, MA, Imouzzer-du-Kander, Sefrou, Fès-Meknès; Chaker et al. 1979; Hervy et al. 1994; Borkent and Wirth 1997; Remm 1988a; Dakki 1997; Chilasse and Dakki 2004, MA; Borkent 2012; Bourquia et al. 2019

Culicoides (Sensiculicoides) langeroni Kieffer, 1921

Bailly-Choumara and Kremer 1970, HA, Tnine des Oudaias (bordure de Oued N‘fis); Kremer et al. 1971, AP, Rabat-Salé-Kénitra; Kremer et al. 1975; Kremer et al. 1979; Bailly-Choumara and Kremer 1980, MA, Khénifra, AA, Tarhjicht, Torkoz (Draa); Dakki 1997; Bourquia et al. 2019

Culicoides (Sensiculicoides) maritimus Kieffer, 1924

Bailly-Choumara and Kremer 1970, AP, estuaire de Bou-Regreg; Kremer et al. 1971, Rif, Tanger-Tétouan-Al Hoceima, AP, Rabat-Salé-Kénitra; Kremer et al. 1975; Kremer et al. 1979; Bailly-Choumara and Kremer 1980, Rif, Tétouan, AP, Larache, Rabat, Sidi Bettache, HA, Talet-Inaouan (Haouz), Souk Tnine des Oudaias (Haouz); Remm 1988; Dakki 1997; Bourquia et al. 2019

Culicoides (Sensiculicoides) odiatus Austen, 1921

= Culicoides lailae Khalaf, in Bailly-Choumara and Kremer 1970: 387, Dakki 1997: 60

= Culicoides indistinctus Khalaf, in Kremer et al. 1975: 206, Dakki 1997: 60

Bailly-Choumara and Kremer 1970, HA, Tnine des Oudaias (bordure de Oued N‘fis); Kremer et al. 1971, AP, Rabat-Salé-Kénitra, MA, Fès-Meknès, AA, Torkoz, SA, Guelmim-Oued Noun; Kremer et al. 1975, AA, Tafraout, Souss-Massa; Kremer et al. 1979; Chaker et al. 1979, AP, Rabat-Salé-Kénitra, Safi, HA, Marrakech, SA, Guelmim-Oued Noun; Chaker et al. 1980, AP, Kénitra, Merja Bokka (Gharb), HA, Souk Tnine des Oudaias (Haouz), AA, Tarhjicht; Remm 1988a; Dakki 1997; Baylis et al. 1997; Bouayoune et al. 1998; Cêtre-Sossah and Baldet 2004, AP, Rabat-Salé-Kénitra; Sarvašová et al. 2014; Bourquia et al. 2019

Culicoides (Sensiculicoides) paolae Boorman, 1996

Bourquia et al. 2019, AP, Rabat

Culicoides (Sensiculicoides) pictipennis (Staeger, 1839)

Bailly-Choumara and Kremer 1970, AP, estuaire de Bou-Regreg; Kremer et al. 1971; Remm 1988a; Bourquia et al. 2019, AP, Rabat

Culicoides (Sensiculicoides) pseudopallidus Khalaf, 1961

Bailly-Choumara and Kremer 1970, HA, Souk Tnine des Oudaias (bordure Oued N’fis); Kremer et al. 1971, AP, Safi, MA, Rhafsai, Fès-Meknès, HA, Marrakech; Kremer et al. 1975; Remm 1988a; Dakki 1997; Bourquia et al. 2019

Culicoides (Sensiculicoides) shaklawensis Khalaf, 1957

Kremer et al. 1975, AP, Safi, MA, Sefrou, Fès-Meknès, HA, Setti Fatma, Marrakech; Chaker et al. 1979; Remm 1988a; Dakki 1997; Bourquia et al. 2019

Culicoides (Sensiculicoides) simulator Edwards, 1939

Kremer et al. 1975, MA, Ifrane, Fès-Meknès, HA, Setti Fatma; Chaker et al. 1979; Dakki 1997; Bourquia et al. 2019

Culicoides (Sensiculicoides) univittatus Vimmer, 1932

= Culicoides agathensis Callot, Kremer and Rioux, in Bailly-Choumara and Kremer 1970: 386, Kremer et al. 1971: 663, Chaker et al. 1980: 82, Dakki 1997: 60

Bailly-Choumara and Kremer 1970, AP, estuaire Bou-Regreg; Kremer et al. 1971, AP, Rabat-Salé-Kénitra, Rif, Tanger-Tétouan-Al Hoceima; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, Rif, Tanger-Tétouan-Al Hoceima, AP, Rabat-Salé-Kénitra, Safi, MA, Fès-Meknès, HA, Marrakech; Chaker et al. 1980, Rif, Tétouan, AP, Larache, Sidi Bettache MA, Oulmès; Dakki 1997; Bourquia et al. 2019

Culicoides (Sensiculicoides) vidourlensis Callot, Kremer, Molet & Bach, 1968

Bailly-Choumara and Kremer 1970, AP, Merja Sheishat (Larache), estuaire de Oued Bou-Regreg, HA, Souk Tnine des Oudaias (bordure de Oued N’fis); Kremer et al. 1971; Remm 1988a; Bourquia et al. 2019

Culicoides (Silvaticulicoides) pallidicornis Kieffer, 1919

Bailly-Choumara and Kremer 1970, HA, Souk Tnine des Oudaias (bordure Oued N’fis); Kremer et al. 1971, AA, Torkoz, SA, Guelmim-Oued Noun; Dakki 1997; Balenghien et al. 2014, SA, Guelmim-Oued Noun; Bourquia et al. 2019

Culicoides (Silvaticulicoides) picturatus Kremer & Deduit, 1961

Bailly-Choumara and Kremer 1970, AP, Merja Sheishat (Larache); Kremer et al. 1971, Rif, Talerhza, EM, El-Harcha, AP, Bousselham, Rabat-Salé-Kénitra, MA, Oulmès; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, AP, Casablanca-Settat, MA, Béni Mellal-Khénifra; Remm 1988a; Dakki 1997; Sarvašová et al. 2014; Bourquia et al. 2019

Culicoides (Silvaticulicoides) subfasciipennis Kieffer, 1919

Kremer et al. 1971, AP, Rabat-Salé-Kénitra, MA, Fès-Meknès, Béni Mellal-Khénifra; Kremer et al. 1975; Kremer et al. 1979; Chaker et al. 1979, Rif, Tanger-Tétouan-Al Hoceima; Bailly-Choumara and Kremer 1980, AP, Larache, Zaers, Rabat, Aïn Chok, MA, Sefrou; Remm 1988a; Hervy et al. 1994; Dakki 1997; Cêtre-Sossah and Baldet 2004, AP, Rabat-Salé-Kénitra; Bourquia et al. 2019

Culicoides (Wirthomyia) faghihi Navai, 1971

Kremer et al. 1975, AA, Tafraout, Souss-Massa; Chaker et al. 1979; Remm 1988a; Hervy et al. 1994; Dakki 1997; Bourquia et al. 2019

Culicoides (Wirthomyia) minutissimus (Zetterstedt, 1855)

Referred as C. pumilus

Culicoides (Wirthomyia) pumilus (Winnertz, 1852)

Kremer et al. 1975, AP, Safi, MA, Ifrane, Imouzzer-du-Kander, Fès-Meknès, HA, Setti Fatma, Marrakech; Chaker et al. 1979; Dakki 1997; Bourquia et al. 2019

Culicoides calloti Kremer, Delécolle, Bailly-Choumara & Chaker, 1979

Chaker et al. 1979, AA, Souss Massa, SA, Guelmim-Oued Noun; Kremer et al. 1979, AA, Tarhjigt, Aït Ouaballi, Souss Massa, SA, Guelmim-Oued Noun; Chaker et al. 1980; Remm 1988a; Hervy et al. 1994; Borkent and Wirth 1997; Baylis et al. 1997; Dakki 1997; Koçak and Kemal 2010; Borkent 2012; Bourquia et al. 2019

Ceratopogonini

Alluaudomyia Kieffer, 1913

Alluaudomyia hygropetrica Vaillant, 1954

Vaillant 1956b, HA, Sidi Chamarouch

Palpomyiini

Bezzia Kieffer, 1899

Bezzia (Bezzia) nigritula (Zetterstedt, 1838)

= Palpomyia tenebricosa Goetghebuer, 1912, in Vaillant 1956b: 241

Vaillant 1956b, HA, Tamesrit

Palpomyia Meigen, 1818

Palpomyia helviscutellata Borkent, in Borkent & Wirth 1997

= Dasyhelea flavoscutellata (Zetterstedt, 1850), in Vaillant 1956b: 244

Vaillant 1956b, HA, Tahanaout

Forcipomyiinae

Dasyheleini

Dasyhelea Kieffer, 1911

Dasyhelea (Prokempia) flaviventris (Goetghebuer, 1910)

Dominiak 2012

Dasyhelea (Pseudoculicoides) turficola Kieffer, 1925

Dominiak 2012

Leptoconopinae

Leptoconops Skuse, 1889

Leptoconops (Holoconops) laurae (Weiss, 1912)

Remm 1988b

CHIRONOMIDAE

K. Kettani

Number of species: 412. Expected: 600

Faunistic knowledge of the family in Morocco: good

Buchonomyiinae

Buchonomyia Fittkau, 1955

Buchonomyia thienemanni Fittkau, 1955

Ashe and O’Connor 2009; Kettani et al. 2010, Rif, Oued Kelaâ (Akoumi, 400 m); Kettani and Langton 2012; Ashe et al. 2015, Rif, Oued Kelaâ (Akoumi, 400 m); Kettani and Moubayed-Breil 2018, Rif

Podonominae

Paraboreochlus Thienemann, 1939

Paraboreochlus minutissimus (Strobl, 1895)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Tanypodinae

Macropelopiini

Apsectrotanypus Fittkau, 1962

Apsectrotanypus trifascipennis (Zetterstedt, 1838)

Kettani et al. 2010, Rif, Aïn Abou Hayane (Tiouertiouane, 880 m), Oued Maggou (Maggou village, 777 m), Oued Kanar (Gorges Kanar, 280 m); Kettani and Langton 2012

Macropelopia Thienemann, 1916

Macropelopia adaucta Kieffer, 1916

Kettani and Langton 2011, Rif, Fifi, Issaguen; Kettani and Langton 2012

Macropelopia nebulosa (Meigen, 1804)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1994, Rif, Oued Siflaou; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Psectrotanypus Kieffer, 1909

Psectrotanypus varius (Fabricius, 1787)

Kettani et al. 1996; Kettani et al. 1997, Rif, Ras el Ma (Chefchaouen); Kettani et al. 2001; Ashe and O’Connor 2009; Kettani et al. 2010, Rif, Oued Tassikeste (Afechtal, 240 m); Kettani and Langton 2012

Pentaneurini

Ablabesmyia Johannsen, 1905

Ablabesmyia (Ablabesmyia) ebbae Lehmann, 1981

Lehmann 1981; Azzouzi and Laville 1987; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Ablabesmyia (Ablabesmyia) longistyla Fittkau, 1962

El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, aval Barrage Talembote, aval Oued Laou; Kettani et al. 1995, Rif, aval Oued El Kbir, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou); Kettani et al. 2001; Ashe and O’Connor 2009; Kettani et al. 2010, Rif, source Maggou (Maggou, 1300 m), Oued Talembote; Kettani and Langton 2012

Ablabesmyia (Ablabesmyia) monilis (Linnaeus, 1758)

Reiss 1977, Rif, Tétouan, HA, kranichsee (Dra-Tal); Azzouzi and Laville 1987, Rif, retenue El Makhazine; El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Naya 1988, MA, Haut Sebou; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Conchapelopia Fittkau, 1957

Conchapelopia (Conchapelopia) melanops (Meigen, 1818)

Kettani et al. 1996; Kettani et al. 1997, Rif, Ras el Ma (Chefchaouen); Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Conchapelopia (Conchapelopia) pallidula (Meigen, 1818)

Kettani and Moubayed-Breil 2018, Rif

Conchapelopia (Conchapelopia) viator (Kieffer, 1911)

= Conchapelopia Pe 1 Langton 1991 in Kettani et al. 1994: 28, Kettani et al. 1995: 256

Azzouzi et al. 1992, HA, Gorges de Dadès (Imdiazen, 1900 m); Kettani et al. 1994; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Ashe and O’Connor 2009; Kettani and Langton 2012

Larsia Fittkau, 1962

Larsia atrocincta (Goetghebuer, 1942)

Azzouzi et al. 1992, HA, Gorges de Dadès (Imdiazen, 1900 m); Kettani et al. 1994, Rif, Oued Moulay Bouchta; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Larsia curticalcar (Kieffer, 1918)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani et al. 2010, Rif, Nord Maggou village (Maggou, 905 m); Kettani and Langton 2012

Nilotanypus Kieffer, 1923

Nilotanypus dubius (Meigen, 1804)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1994, Rif, Oued Siflaou, Oued Moulay Bouchta, aval Oued Tassikeste; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani et al. 2010, Rif, Oued Kanar (Gorges Kanar, 280 m), Oued Tassikeste (Afechtal, 240 m), Oued Talembote (Usine électrique, 120 m), Oued Laou (Afertane, 55 m); Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Paramerina Fittkau, 1962

Paramerina cingulata (Walker, 1856)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Ashe and O’Connor 2009; Kettani et al. 2010, Rif, Oued Talembote (aval Barrage Talembote, 245 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Paramerina divisa (Walker, 1856)

Kettani et al. 1996; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani and Langton 2012

Paramerina mauretanica Fittkau, 1962

Fittkau 1962, Atlas (850 m), SA; Azzouzi and Laville 1987; Ashe and Cranston 1990, EM, Figuig; Kettani et al. 2001; Ashe and O’Connor 2009, EM, Figuig; Kettani et al. 2010, Rif, Oued Tassikeste (Afechtal, 240 m); Kettani and Langton 2012

Paramerina spec. Greichenland (Fittkau, 1962)

Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Laville and Langton 2002; Kettani et al. 2010, Rif, Oued Chrafat (Armotah, 900 m), Oued Talembote (aval Barrage Talembote, 245 m); Kettani and Moubayed-Breil 2018, Rif

Pentaneurella Fittkau & Murray, 1983

Pentaneurella sp. Ourika

Azzouzi et al. 1992, HA; Kettani et al. 2001

Rheopelopia Fittkau, 1962

Rheopelopia maculipennis (Zetterstedt, 1838)

Naya 1988, MA, Haut et Moyen Sebou; Azzouzi and Laville 1987, MA, Oum-er-Rbia, HA, Tensift; Kettani et al. 1994, Rif, Oued Siflaou, Oued Moulay Bouchta; Kettani et al. 1995, Rif, aval Oued El Kbir, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Dakki et al. 2008, MA, Oued Sebou; Ashe and O’Connor 2009; Kettani et al. 2010, Rif, Ruisselet maison forestière (Talassemtane, 1683 m), Source Maggou (Maggou, 1300 m), Oued Talembote (avant village Talembote, 320 m), Oued Tassikeste (Afechtal, 240 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Rheopelopia murrayi Dowling, 1983

Dowling 1983, AA, Tata (Moyen Draa); Ashe and Cranston 1990; Kettani et al. 2001; Ashe and O’Connor 2009; 2012; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Rheopelopia ornata (Meigen, 1838)

Azzouzi and Laville 1987, MA, Oued Fès; Kettani et al. 1995, Rif, Oued El Kbir, amont Oued Nakhla, aval Oued Khemis; Kettani et al. 1996; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Dakki et al. 2008, MA, Oued Sebou; Ashe and O’Connor 2009; Kettani et al. 2010, Rif, Oued Talembote (avant village Talembote, 320 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Telopelopia Roback, 1971

Telopelopia fascigera (Verneaux, 1970)

= Telopelopia maroccana Murray, 1980, in Reiss 1977: 91, Murray 1980: 151, Azzouzi and Laville 1987: 218, Ashe and Cranston 1990: 133

Reiss 1977, AP, Larache, HA, Dra-Tal; Murray 1980, AP, Larache, HA, Dra-Tal; Azzouzi and Laville 1987; Ashe and Cranston 1990; Azzouzi and Laville 1987, MA, Oum-er-Rbia; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Telmatopelopia Fittkau, 1962

Telmatopelopia nemorum (Goetghebuer, 1921)

Kettani et al. 1996, Rif, Oued Khizana (Oued Laou); Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Thienemannimyia Fittkau, 1957

Thienemannimyia (Thienemannimyia) berkanea Dowling, 1987

Dowling 1987, EM, Berkane; Azzouzi et al. 1992, EM, Environs de Berkane, HA, Ouarzazate (1160 m), Oasis Meski (1160 m), Aït Saoun; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Thienemannimyia (Thienemannimyia) carnea (Fabricius, 1805)

Kettani and Langton 2012, Rif

Thienemannimyia (Thienemannimyia) choumara Dowling, 1983

Dowling 1983, EM, Environ de Berkane (Monts de Bni Snassen), HA, Souk des Judais (Marrakech); Azzouzi et al. 1987, HA, Dra-Tal; Ashe and Cranston 1990; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Thienemannimyia (Thienemannimyia) geijskesi (Goetghebuer, 1934)

Kettani and Langton 2012, Rif, Oued Zarka

Thienemannimyia (Thienemannimyia) laeta (Meigen, 1818)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Thienemannimyia (Thienemannimyia) lentiginosa (Fries, 1823)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Thienemannimyia (Thienemannimyia) northumbrica (Edwards, 1929)

Fittkau 1962; Azzouzi and Laville 1987, MA, Oum-er-Rbia; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Ashe and O’Connor 2009; Kettani and Langton 2012

Trissopelopia Kieffer, 1923

Trissopelopia longimana (Staeger, 1839)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Xenopelopia Fittkau, 1962

Xenopelopia falcigera (Kieffer, 1911)

Kettani and Langton 2011, Rif, Anasser, Fifi, AP, marais de Loukous; Kettani and Langton 2012

Xenopelopia nigricans (Goetghebuer, 1927)

Kettani et al. 1994, Rif, aval Oued Talembote (usine éléctrique); Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani et al. 2010, Rif, Oued Talembote (aval affluent Talembote, 155 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Zavrelimyia Fittkau, 1962

Zavrelimyia (Zavrelimyia) barbatipes (Kieffer, 1911)

Naya 1988, MA, Moyen Sebou; Kettani et al. 2001; Ashe and O’Connor 2009 (?); Kettani et al. 2010, Rif, Oued Tiffert (Tiffert Talassemtane, 1230 m), Aïn Abou Hayane (Tiouertiouane, 880 m), Oued Abiyati (Ifansa, 140 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Zavrelimyia (Zavrelimyia) berberi Fittkau, 1962

Azzouzi and Laville 1987; Ashe and Cranston 1990, HA, Tamhda; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Zavrelimyia (Zavrelimyia) hirtimana (Kieffer, 1918)

Kettani and Langton 2012

Zavrelimyia (Zavrelimyia) melanura (Meigen, 1804)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Zavrelimyia (Zavrelimyia) nubila (Meigen, 1830)

Kettani and Langton 2011, Rif, marais de Lemtahane (PNPB), Dayat Aïn Rami, Dayat Amlay; Kettani and Langton 2012

Procladiini

Procladius Skuse, 1889

Procladius (Holotanypus) brevipetiolatus (Goetghebuer, 1935)

Azzouzi et al. 1992, HA, Oued Meski (1160 m), Khettaras de Marrakech; Kettani et al. 2001; Kettani and Langton 2012

Procladius (Holotanypus) choreus (Meigen, 1804)

Ramdani and Tourenq 1982, AP, Merja Sidi Boughaba; El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Azzouzi and Laville 1987, AP, Merja Sidi Boughaba; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote; Kettani et al. 1995, Rif, amont Oued Nakhla, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 2010, Rif, Aïn Talassemtane (Talassemtane, 1700 m), Oued Talembote (aval Barrage Talembote, 245 m), Oued Tassikeste (Afechtal, 240 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Procladius (Holotanypus) culiciformis (Linnaeus, 1767)

Kettani and Moubayed-Breil 2018, Rif

Procladius (Holotanypus) noctivagus (Kieffer, 1910)

Azzouzi et al. 1992, HA, Ouarzazate (1160 m); Kettani et al. 2001; Kettani and Langton 2012

Procladius (Holotanypus) sagittalis (Kieffer, 1909)

Kettani et al. 1996; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Procladius (Psilotanypus) anomalus Kieffer, 1906

Nomen dubium in Ashe and O’Connor 2009: 213

Naya 1988, MA; Kettani et al. 2001; Kettani and Langton 2012

Procladius Pe 3 Langton 1991

Kettani et al. 1994, Kettani et al. 1995, Rif, Oued Mhajrat; Kettani et al. 1996; Kettani et al. 2001; Dakki 1997

Tanypodini

Tanypus Meigen, 1803

Tanypus (Tanypus) brevipalpis (Kieffer, 1923)

Reiss 1977, EM, Berkane; Ashe and O’Connor 2009 (?); Kettani and Langton 2012

Tanypus (Tanypus) kraatzi (Kieffer, 1912)

Azzouzi et al. 1992, HA, Oasis Meski; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Tanypus (Tanypus) punctipennis Meigen, 1818

Reiss 1977, EM, Berkane; El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Azzouzi and Laville 1987, HA, Oued Tensift; Kettani et al. 1996; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Diamesinae

Boreoheptagyiini

Boreoheptagyia Brundin, 1966

Boreoheptagyia legeri (Goetghebuer, 1933)

= Boreoheptagyia punctulata (Goetghebuer, 1934), in Kettani et al. 2001: 327

Ashe and Cranston 1990; Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Diamesini

Diamesa Meigen, 1835

Diamesa aberrata Lundbeck, 1898

Saether 1968; Serra-Tosio 1973; Fittkau and Reiss 1987; Serra-Tosio 1983; Azzouzi and Laville 1987, HA (2500–3350 m); Ashe and Cranston 1990; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Diamesa bertrami Edwards, 1935

Serra-Tosio 1983, HA, Gorges de Todra (2500 m); Azzouzi and Laville 1987, HA, Gorges Todra; Ashe and Cranston 1990; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Diamesa hamaticornis Kieffer, 1924

Reiss 1977; Serra-Tosio 1983, HA, M’Goum; Azzouzi and Laville 1987, HA, M’Goum; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Diamesa insignipes Kieffer, 1908

Serra-Tosio 1983, HA (2500 m); Azzouzi and Laville 1987; Naya 1988, MA, Haut and Moyen Sebou; Ashe and Cranston 1990; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Diamesa latitarsis (Goetghebuer, 1921)

Vaillant 1955b; Vaillant 1956b, HA, Asif Tessaout (M’Goum), Lac Tamhda (Anremer); Serra-Tosio 1967; Serra-Tosio 1967; Saether 1968; Serra-Tosio 1973; Azzouzi and Laville 1987, HA; Ashe and Cranston 1990; Kettani et al. 2001, Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Diamesa steinboecki Goetghebuer, 1933

Vaillant 1956b, HA, Cascade Siroua, Oukaimeden, Sidi Chamarouch

Diamesa tonsa (Haliday in Walker, 1856)

= Diamesa thienemanni Kieffer, 1909

Naya 1988, MA, Haut Sebou (Arhbalou Yahya, Oued Arbi, Pont Aït hamza); Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2011, Rif, Oued Ketama, Oued Sgara; Kettani and Langton 2012

Diamesa vaillanti Serra-Tosio, 1972

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Diamesa veletensis Serra-Tosio, 1971

Serra-Tosio 1983, HA (2500 m); Azzouzi and Laville 1987, HA; Ashe and Cranston 1990; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Diamesa zernyi Edwards, 1933

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Potthastia Kieffer, 1922

Potthastia gaedii (Meigen, 1838)

Azzouzi and Laville 1987, MA, oued Boufekrane, Oued Fès, Oued Oum-er-Rbia; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Oued Talembote (usine éléctrique), Oued Afertane, aval Oued Laou; Kettani et al. 1995, Rif, aval Oued El Kbir, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani et al. 2010, Rif, Oued Laou, Oued Afertane; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Potthastia pastoris (Edwards, 1933)

Kettani and Moubayed-Breil 2018, Rif

Pseudodiamesa Goetghebuer, 1939

Pseudodiamesa (Pseudodiamesa) branickii (Nowicki, 1873)

Naya 1988, MA, Haut Sebou; Ashe and Cranston 1990; Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2001; Kettani and Langton 2012

Pseudodiamesa (Pseudodiamesa) nivosa (Goetghebuer, 1928)

Naya 1988, MA, Moyen Sebou; Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Ashe and O’Connor 2009; Kettani and Langton 2012

Sympothastia Pagast, 1947

Sympothastia zavreli Pagast, 1947

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1994, Rif, Haut Laou; Kettani et al. 1995, Rif, aval Oued Krikra; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Syndiamesa Kieffer, 1918

Syndiamesa hygropterica (Kieffer, 1909)

Naya 1988, MA, Moyen Sebou (Sidi Abdellah, Dar El Arsa, Pont Oulad Slimane, Pont Portugais); Ashe and Cranston 1990; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Protanypini

Protanypus Kieffer, 1906

Protanypus morio (Zetterstedt, 1838)

Naya 1988, MA, Moyen Sebou; Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Prodiamesinae

Odontomesa Pagast, 1947

Odontomesa fulva (Kieffer, 1919)

Azzouzi and Laville 1987, MA, Oued Oum-er-Rbia; Kettani et al. 2001; Ashe and O’Connor 2009; Kettani and Langton 2012

Prodiamesa Kieffer, 1906

Prodiamesa olivacea (Meigen, 1818)

Naya 1988, MA, Haut Sebou (Haut Guigou); Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1994, Rif, Haut Laou; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Ras el Ma (Chefchaouen); Kettani et al. 2001; Ashe and O’Connor 2009; Kettani et al. 2010, Rif, Maggou village, Ifansa; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Orthocladiinae

Orthocladiini

Acricotopus Kieffer, 1921

Acricotopus lucens (Zetterstedt, 1850)

Kettani and Moubayed-Breil 2018, Rif

Brilla Kieffer, 1913

Brillia bifida (Kieffer, 1909)

= Brilla modesta (Meigen, 1830)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1996; Kettani et al. 1997, Rif, Ras el Ma (Chefchaouen); Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2001; Kettani et al. 2010, Rif, Oued Tamaridine (Zaouiet El Habtiyine, 819 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Brillia flavifrons (Johannsen, 1905)

Kettani and Langton 2012

Brilla longifurca Kieffer, 1921

Azzouzi and Laville 1987, MA, Oued Boufekrane, Oued Sebou, Oued Oum-er-Rbia; Kettani et al. 1995, Rif, amont Oued Nakhla, Oued Mhajrat; Kettani et al. 1996; Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2010, Rif, Oued Talembote (avant village Talembote, 320 m), Oued Talembote (Usine électrique, 120 m); Kettani and Langton 2012; Ashe and O’Connor 2012

Bryophaenocladius Thienemann, 1934

Bryophaenocladius aestivus (Brundin, 1947)

Kettani and Moubayed-Breil 2018, Rif; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Bryophaenocladius flexidens (Brundin, 1947)

Kettani and Moubayed-Breil 2018, Rif

Bryophaenocladius cf. furcatus Thienemann & Strenzke, 1940

Kettani and Moubayed-Breil 2018, Rif

Bryophaenocladius illimbatus (Edwards, 1929)

Kettani and Moubayed-Breil 2018, Rif; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Bryophaenocladius muscicola (Kieffer, 1906)

Kettani and Moubayed-Breil 2018, Rif

Bryophaenocladius nidorum (Edwards, 1929)

Kettani and Moubayed-Breil 2018, Rif

Bryophaenocladius subvernalis (Edwards, 1929)

Azzouzi and Laville 1987, MA, Oued Boufekrane; Kettani et al. 2001; Kettani and Langton 2011, Rif, Oued Taida (Moulay Abdelsalam); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Bryophaenocladius tuberculatus (Edwards 1929)

Kettani and Moubayed-Breil 2018, Rif

Camptocladius Wulp, 1874

Camptocladius stercorarius (De Geer, 1976)

Kettani and Moubayed-Breil 2018, Rif

Cardiocladius Kieffer, 1912

Cardiocladius capucinus (Zetterstedt, 1850)

Azzouzi and Laville 1987, MA, Oued Oum-er-Rbia; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote, Oued Afertane, aval Oued Tassikeste; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Oued Talembote (aval Barrage Talembote, 245 m), Oued Tassikeste (Afechtal, 240 m), Oued Laou (Afertane, 55 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cardiocladius fuscus Kieffer, 1924

Azzouzi and Laville 1987, MA, Oued Boufekrane, Oued Oum-er-Rbia; Naya 1988, MA, Haut Sebou (Amont de Aïn Tadout, Skhounate, amont confluence avec Oued Atchane, Pont Aït Hamza); Kettani et al. 1994, Rif, Oued Siflaou; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Chaetocladius Kieffer, 1911

Chaetocladius (Chaetocladius) acuticornis (Kieffer in Potthast, 1914)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Chaetocladius dentiforceps (Edwards, 1929)

Kettani and Moubayed-Breil 2018, Rif; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Chaetocladius dissipatus (Edwards, 1929)

Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Chaetocladius (Chaetocladius) melaleucus (Meigen, 1818)

Kettani and Langton 2011, Rif, Oued Sgara, Bab Tariouant, Bouztata; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Chaetocladius piger (Goetghebuer, 1913)

Kettani and Moubayed-Breil 2018, Rif

Chaetocladius (Chaetocladius) perennis (Meigen, 1830)

Kettani and Langton 2011, Rif, Oued Hamma; Kettani and Langton 2012

Chaetocladius (Chaetocladius) vitellinus (Kieffer in Kieffer & Thienemann, 1908)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012

Corynoneura Winnertz, 1846

Corynoneura carriana Edwards, 1924

Naya 1988, MA, Haut Sebou (Haut Guigou, Aïn Nokra); Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Corynoneura celtica Edwards, 1924

Kettani and Langton 2011, Rif, Oued Hamma; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Corynoneura coronata Edwards, 1924

Kettani and Langton 2011, Rif, Oued Hamma; Kettani and Langton 2012; Ashe and O’Connor 2012

Corynoneura edwardsi Brundin, 1949

Kettani and Langton 2012

Corynoneura lacustris Edwards, 1924

El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Corynoneura lobata Edwards, 1924

Kettani et al. 1996, Rif, Oued Nakhla; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Oued Kelaâ (Akoumi, 400 m); Azzouzi et al. 1992, HA; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Corynoneura scutellata Winnertz, 1846

Kettani and Moubayed-Breil 2018, Rif

Corynoneura Pe 2 Langton 1991

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012

Corynoneurella Brundin, 1949

Corynoneurella paludosa Brundin, 1949

Kettani et al. 2010, Rif, Nord Maggou village (Maggou, 905 m), Oued Kelaâ (Akoumi, 400 m), Oued Talembote (avant village Talembote, 320 m), Oued Laou (Afertane, 55 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus van der Wulp, 1874

Cricotopus (Cricotopus) albiforceps (Kieffer in Thienemann and Kieffer 1916)

Kettani et al. 1996; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Cricotopus) annulator Goetghebuer, 1927

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani et al. 2010, Rif, Oued Talembote (avant village Talembote, 320 m), Oued Tassikeste (Afechtal, 240 m), Oued Laou (Afertane, 55 m); Kettani and Langton 2011, Rif, Oued Sgara; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Cricotopus) beckeri Hirvenoja, 1973

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1995, Rif, aval Oued El Kbir; Kettani et al. 1996; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou); Kettani et al. 2001; Langton and Laville 2002; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Cricotopus) bicinctus (Meigen, 1818)

Azzouzi and Laville 1987, MA, Oued Boufekrane; Naya 1988, MA, Haut et Moyen Sebou; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote, Oued Afertane, aval Oued Laou; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis, Oued Martil (Tamuda); Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2010, Rif, Source Maggou (Maggou, 1300 m), Oued Talembote (aval Barrage Talembote, 245 m), Oued Laou (Afertane, 55 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Cricotopus) caducus Hirvenoja, 1973

Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Cricotopus) ephippium (Zetterstedt, 1838)

Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Cricotopus) levantinus Moubayed & Hirvenoja, 1986

Kettani et al. 1996, Rif, Haut Maggou; Kettani et al. 1997, Rif, Maggou (Oued Laou); Kettani et al. 2001; Laville and Langton 2002; Kettani et al. 2010, Rif, Source Maggou (Maggou, 1300 m), Oued Inesmane (Adeldal, 1173 m); Kettani and Langton 2012; Ashe and O’Connor 2012

Cricotopus (Cricotopus) pallidipes Edwards, 1929

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1995, Rif, Oued Martil (Tamuda); Kettani et al. 1996; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Cricotopus) pulchripes Verrall, 1912

Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Cricotopus) similis Goetgnebuer, 1921

Azzouzi and Laville 1987, MA, Oued Oum-er-Rbia; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Afertane, aval Oued Talembote (usine éléctrique), aval Oued Tassikeste; Kettani et al. 1995, Rif, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani et al. 2010, Rif, Oued Talembote (aval Barrage Talembote, 245 m), Oued Talembote (aval affluent Talembote, 155 m), Oued Laou (Afertane, 55 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Cricotopus) tremulus (Linnaeus, 1758)

Kettani et al. 2010, Rif, Oued Maggou (Maggou village, 777 m); Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Cricotopus) triannulatus (Macquart, 1826)

Kettani et al. 1994, Rif, Haut Laou, Oued Moulay Bouchta, Oued Afertane; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Oued Talembote (aval affluent Talembote, 155 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Cricotopus) trifascia Edwards, 1929

Azzouzi and Laville 1987, MA, Oued Boufekrane, Oued Fès, Oued Oum-er-Rbia; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote, aval Oued Talembote (usine éléctrique), aval Oued Tassikeste; Kettani et al. 1995, Rif, aval Oued Krikra, amont Oued Nakhla, Oued Mhajrat; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2010, Rif, Oued Maggou (Maggou village, 777 m), Oued Talembote (aval Barrage Talembote, 245 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Cricotopus) vierriensis Goetghebuer, 1935

El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Azzouzi and Laville 1987, MA, Oued Boufekrane, Oued Fès, Oued Sebou, Oued Oum-er-Rbia, HA, Oued Tensift; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote, aval Oued Talembote (usine éléctrique), Oued Afertane, aval Oued Tassikeste, aval Oued Laou; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2010, Rif, Oued Talembote (aval Barrage Talembote, 245 m), Oued Talembote (aval affluent Talembote, 155 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Isocladius) brevipalpis Kieffer, 1909

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012

Cricotopus (Isocladius) laetus Hirvenoja, 1973

Kettani et al. 1994, Rif, Oued Siflaou; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012

Cricotopus (Isocladius) ornatus (Meigen, 1818)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012

Cricotopus (Isocladius) sylvestris (Fabricius, 1794)

Fittkau and Reiss 1978; Ramdani and Tourenq 1982, AP, Merja Sidi Boughaba; El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Azzouzi and Laville 1987, MA, Oued Boufekrane, Oued Fès, HA, Oued Tensift; Kettani et al. 1994, Rif, Haut Laou; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Isocladius) tricinctus (Meigen, 1818)

Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Paratrichocladius) micans (Kieffer, 1918)

Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote, aval Oued Talembote (usine éléctrique), Oued Afertane, aval Oued Tassikeste; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Kettani et al. 1997, Rif, Maggou (Oued Laou); Dakki 1997; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Haut Maggou (1300 m), Oued Talembote, Oued Laou (Afertane, 56 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Paratrichocladius) osellai Rossaro, 1990

Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Paratrichocladius) rufiventris (Meigen, 1830)

El Mezdi and Giudicelli 1985, HA, Khettaras de Marrakech; Azzouzi and Laville 1987, MA, Oued Fès, Oued Boufekrane, Oued Sebou, Oued Oum-er-Rbia, HA, Oued Tensift; (Dar El Arsa, Pont oulad Slimane, Pont portugais); Naya 1988, MA, Haut et Moyen Sebou; Kettani et al. 1995, Rif, Oued Mhajrat; Kettani et al. 1996; Kettani et al. 2001; Kettani et al. 2010, Rif, Oued Kelaâ (Akoumi, 400 m), Oued Talembote; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Cricotopus (Paratrichocladius) skirwithensis (Edwards, 1929)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Eukieferiella Thienemann, 1926

Eukieferiella ancyla Svensson, 1986

Kettani and Langton 2011, Rif, Oued Tkarae; Kettani and Langton 2012; Ashe and O’Connor 2012

Eukiefferiella bedmari Vilchez-Quero & Laville, 1988

Azzouzi and Laville 1987, MA, Oued Fès, Oued Oum-er-Rbia; Kettani et al. 1994, Rif, Haut Laou, aval Oued Talembote (usine éléctrique), Oued Afertane, aval Oued Tassikeste; Kettani et al. 1995, Rif, aval Oued Krikra, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Laville and Langton 2002; Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2010, Rif, Oued Kelaâ (Akoumi, 400 m), Oued Talembote (aval Barrage Talembote, 245 m), Oued Talembote (aval affluent Talembote, 155 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Eukiefferiella brehmi Gowin, 1943

Kettani et al. 1996; Kettani et al. 1997, Rif, Maggou (Oued Laou); Kettani et al. 2001; Kettani et al. 2010, Rif, Source Maggou (Maggou, 1300 m), Oued Kelaâ (Akoumi, 400 m), Oued Talembote (avant village Talembote, 320 m), Oued Talembote (Usine électrique, 120 m)

Eukiefferiella brevicalcar (Kieffer, 1911)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani et al. 2010, Rif, Nord Maggou village (Maggou, 905 m), Oued Ametrasse (Ametrasse, 820 m); Kettani and Langton 2011, Rif, Oued Issaguen, Oued Ketama, Oued Sgara; Bab Tariouant, Bouztate (Fifi); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Eukiefferiella claripennis (Lundbeck, 1898)

Fittkau and Reiss 1978; Naya 1988, MA, Moyen Sebou (Dar Cheik Harazem); Azzouzi and Laville 1987, MA, Oued Oum-er-Rbia; Kettani et al. 1994, Rif, Oued Siflaou; Kettani et al. 1996; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Eukieffeiella clypeata (Thienemann, 1919)

Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Oued Talembote (usine éléctrique); Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat; Kettani et al. 1996; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Oued Laou (Afertane, 55 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Eukiefferiella coerulescens (Kieffer in Zavřel, 1926)

Azzouzi and Laville 1987, MA, Oued Oum-er-Rbia; Naya 1988, MA, Haut Sebou (Skhounate, Arhbalou Aberchane); Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, Oued Moulay Bouchta, aval Barrage Talembote, aval Oued Talembote (usine éléctrique), aval Oued Tassikeste, aval Oued Laou; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2010, Rif, Source Maggou (Maggou, 1300 m), Nord Maggou village (Maggou, 905 m), Oued Tamaridine (Zaouiet El Habtiyine, 819 m), Oued Maggou (Maggou village, 777 m), Oued Kelaâ (Akoumi, 400 m), Oued Talembote (aval Barrage Talembote, 245 m), Oued Talembote (aval affluent Talembote, 155 m), Oued Laou (Afertane, 55 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Eukiefferiella cyanea Thienemann, 1936

Vaillant 1955b, HA; Fittkau and Reiss 1978; Azzouzi and Laville 1987, HA; Kettani et al. 1996, Rif, Oued Nakhla; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Eukiefferiella devonica (Edwards, 1929)

Azzouzi and Laville 1987, MA, Oued Oum-er-Rbia; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Kettani et al. 1997, Rif, Oued Khizana (Oued Laou); Kettani et al. 2001; Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Eukiefferiella dittmari Lehmann, 1972

Kettani and Langton 2011, Rif, Oued Boujdad, Fifi; Kettani and Langton 2012, Rif, Oued Zarka; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Eukiefferiella fittkaui Lehmann, 1972

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012

Eukiefferiella fuldensis Lehmann, 1972

Azzouzi and Laville 1987, MA, Oued Oum-er-Rbia; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Eukiefferiella gracei (Edwards, 1929)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1994, Rif, aval Oued Tassikeste; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Oued Talembote (aval Barrage Talembote, 245 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Eukiefferiella ilkleyensis (Edwards, 1929)

Azzouzi and Laville 1987, MA, Oued Oum-er-Rbia; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Eukiefferiella lobifera Goetghebuer, 1934

Azzouzi and Laville 1987, MA, Oued Fès, Oued Oum-er-Rbia; Kettani et al. 2001; Kettani and Langton 2011, Rif, Oued Ketama, Oued Sgara; Kettani and Langton 2012; Ashe and O’Connor 2012

Eukiefferiella minor (Edwards, 1929)

Vaillant 1955b, HA (1050 m); Vaillant 1956b, HA, Imi-N’Ifri; Azzouzi and Laville 1987, HA; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012

Eukiefferiella pseudomontana Goetghebuer, 1935

Kettani et al. 2010, Rif, Oued Madissouka (Talassemtane, 1530 m), Oued Dchar d’Amran (Béni M’Hamed, 1180 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Eukiefferiella similis Goetghebuer, 1939

Azzouzi and Laville 1987, MA, Oued Boufekrane, Oued Fès, Oued Sebou, Oued Oum-er-Rbia; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Eukiefferiella tirolensis Goetghebuer, 1938

Kettani et al. 1994, Rif, Oued Afertane; Kettani et al. 1996; Kettani et al. 2001; Kettani et al. 2010, Rif, Oued Talembote (aval affluent Talembote, 155 m); Azzouzi et al. 1992, HA, Oued Tensift; Dakki 1997; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Eukiefferiella Pe 2 Langton 1991

Kettani et al. 2010, Rif, Oued Kelaâ (Akoumi, 400 m); Kettani and Langton 2012

Halocladius Hirvenoja, 1973

Halocladius (Halocladius) varians (Staeger, 1839)

Ramdani and Tourenq 1982, AP, Merja Sidi Boughaba; Azzouzi and Laville 1987; Kettani et al. 1996; Kettani et al. 1997, Rif, Maggou (Oued Laou); Kettani et al. 2001; Kettani et al. 2010, Rif, Source Maggou (Maggou, 1300 m); Ashe and Cranston 1990; Kettani and Langton 2012; Ashe and O’Connor 2012

Heleniella Gowin, 1943

Heleniella dorieri Serra-Tosio, 1967

Kettani and Langton 2012

Heleniella ornaticollis (Edwards, 1929)

Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, amont Oued Nakhla, Oued Mhajrat; Kettani et al. 1996; Kettani et al. 1997, Rif, Maggou (Oued Laou); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Source Maggou (Maggou, 1300 m), Oued Kelaâ (Akoumi, 400 m); Kettani and Langton 2012; Ashe and O’Connor 2012

Heleniella serratosioi Ringe, 1976

Kettani and Langton 2011, Rif, Oued Hamma, Bouztate (Fifi); Kettani and Langton 2012; Ashe and O’Connor 2012

Heterotrissocladius Spärck, 1923

Heterotrissocladius marcidus (Walker, 1856)

Naya 1988, MA, Moyen Sebou; Kettani et al. 2001; Kettani and Langton 2012; Dakki et al. 2008, MA, Oued Sebou; Ashe and O’Connor 2012

Hydrobaenus Fries, 1830

Hydrobaenus conformis (Holmgren, 1869)

Kettani and Moubayed-Breil 2018, Rif

Hydrosmittia Ferrington & Sæther, 2011

Hydrosmittia oxoniana (Edwards, 1929)

= Pseudosmittia recta (Edwards, 1929), in Azzouzi and Laville 1987: 218, Kettani et al. 2001: 330, Kettani and Langton 2012: 422

Azzouzi and Laville 1987, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012; Kettani and Moubayed-Breil 2018, Rif

Hydrosmittia ruttneri (Strenzke & Thienemann, 1942)

Kettani and Moubayed-Breil 2018, Rif

Krenosmittia Thienemann & Krüger, 1939

Krenosmittia boreoalpina (Goetghebuer, 1944)

Kettani et al. 1996, Rif, Oued Nakhla; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani and Langton 2012; Ashe and O’Connor 2012

Krenosmittia camptophleps (Edwards, 1929)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2011, Rif, Oued Ketama, Oued Sgara; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Krenosmittia halvorseni (Cranston & Sæther, 1986)

Kettani et al. 1995, Rif, aval Oued El Kbir; Kettani et al. 1996; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012

Krenosmittia hispanica Wülker, 1957

Kettani et al. 2001; Laville and Langton 2002; Ashe and O’Connor 2012

Limnophyes Eaton, 1875

Limnophyes difficilis Brunidin, 1947

Kettani and Moubayed-Breil 2018, Rif

Limnophyes gelasinus Saether, 1990

Kettani and Moubayed-Breil 2018, Rif

Limnophyes habilis (Walker, 1856)

Kettani and Moubayed-Breil 2018, Rif

Limnophyes madeirae Sæther, 1985

Kettani and Moubayed-Breil 2018, Rif

Limnophyes minimus (Meigen, 1818)

Ramdani and Tourenq 1982, AP, Merja Sidi Boughaba; Azzouzi and Laville 1987, MA, Lac Aguelmane Azigza; Kettani et al. 2001; Kettani and Langton 2011; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Limnophyes natalensis (Kieffer, 1914)

Kettani and Moubayed-Breil 2018, Rif

Limnophyes ninae Sæther, 1975

Kettani et al. 1996, Rif, Oued Nakhla; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Limnophyes pentaplastus (Kieffer, 1921)

Kettani and Moubayed-Breil 2018, Rif

Limnophyes pumilio (Holmgren, 1869)

Kettani and Moubayed-Breil 2018, Rif

Limnophyes punctipennis (Goetghebuer, 1919)

Kettani and Langton 2012

Limnophyes Pe 1a Langton 1991

Kettani et al. 2010, Rif, Oued Talembote (Usine électrique, 120 m); Kettani and Langton 2012

Metriocnemus van der Wulp, 1874

Metriocnemus (Metriocnemus) albolineatus Meigen, 1818

Kettani and Moubayed-Breil 2018, Rif

Metriocnemus (Metriocnemus) eurynotus (Holmgren, 1883)

= Metriocnemus hygropetricus Kieffer, 1912, in Ashe and O’Connor 2012: 372

= Metriocnemus (Metriocnemus) obscuripes (Holmgren, 1869), in Azzouzi et al. 1992: 229, Kettani et al. 2001: 329, Kettani and Langton 2012: 421

Boumezzough and Thomas 1987, HA, Oued Réghaya (1740 m), Imlil; Azzouzi and Laville 1987, HA, Oued Tensift; Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif; Moubayed-Breil and Kettani 2019, Rif, Chrafate, Challal Sghir (Akchour)

Metriocnemus (Metriocnemus) fuscipes (Meigen, 1818)

Azzouzi and Laville 1987, HA, Oued Tensift; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Metriocnemus (Metriocnemus) hirticollis (Staeger, 1839)

Kettani and Moubayed-Breil 2018, Rif

Metriocnemus (Metriocnemus) ursinus Holmgren, 1869

Kettani and Moubayed-Breil 2018, Rif

Nanocladius Kieffer, 1913

Nanocladius (Nanocladius) balticus (Palmén, 1959)

Kettani et al. 1995, Rif, Oued El Kbir, amont Oued Nakhla; Kettani et al. 1996; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Nanocladius (Nanocladius) dichromus (Kieffer 1906)

Kettani and Moubayed-Breil 2018, Rif

Nanocladius (Nanocladius) parvulus (Kieffer 1909)

Kettani and Moubayed-Breil 2018, Rif

Nanocladius (Nanocladius) rectinervis (Kieffer, 1911)

Azzouzi and Laville 1987, MA, Oued Boufekrane, Oued Oum-er-Rbia; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, aval Oued Tassikeste; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Oued Khizana (Oued Laou); Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Kettani et al. 2010, Rif, Source Maggou (Maggou, 1300 m), Oued Inesmane (Adeldal, 1173 m), Oued Talembote (aval Barrage Talembote, 245 m); Dakki et al. 2008, MA, Oued Sebou; Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Orthocladius van der Wulp, 1874

Orthocladius (Eudactylocladius) fuscimanus (Kieffer, 1908)

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1994, Rif, Haut Laou, Oued Siflaou, aval Oued Talembote (usine éléctrique); Kettani et al. 1995, Rif, aval Oued Krikra, Oued Mhajrat; Kettani et al. 1996; Dakki 1997; Kettani et al. 1997, Rif, Maggou (Oued Laou), Ras el Ma (Chefchaouen); Kettani et al. 2001; Kettani et al. 2010, Rif, source Maggou (Maggou, 1300 m), Oued Chrafat (Armotah, 900 m), Oued Tamaridine (Zaouiet El Habtiyine, 819 m), Oued Talembote (Usine électrique, 120 m), Oued Laou (Afertane, 55 m); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Orthocladius (Euorthocladius) ashei Soponis, 1990

= Orthocladius luteipes Goetghebuer, in Azzouzi and Laville 1987: 218

= Orthocladius rivicola Kieffer, in Azzouzi and Laville 1987: 218

Azzouzi and Laville 1987, MA, Oued Fès, Oued boufekrane, Oued Sebou, Oued Oum-er-Rbia; Kettani et al. 1994, Rif, aval Oued Talembote (usine éléctrique), aval Oued Tassikeste, aval Oued Laou; Kettani et al. 1995, Rif, aval Oued El Kbir, aval Oued Krikra, Oued El Kbir, amont Oued Nakhla, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Kettani and El Ouazzani 2005, Rif, amont Oued Nakhla; Dakki et al. 2008, MA, Oued Sebou; Kettani et al. 2010, Rif, Oued Talembote (avant village Talembote, 320 m), Oued Kanar (Gorges Kanar, 280 m), Oued Talembote (aval Barrage Talembote, 245 m), Oued Talembote (Usine électrique, 120 m), Oued Laou (Afertane, 55 m); Kettani and Langton 2011, Rif, Bouztate (Fifi); Kettani and Langton 2012; Ashe and O’Connor 2012; Kettani and Moubayed-Breil 2018, Rif

Orthocladius (Euorthocladius) rivulorum Kieffer, 1909

Azzouzi et al. 1992, HA, Oued Tensift; Kettani et al. 1994, Rif, Haut Laou; Kettani et al. 1995, Rif, Oued Mhajrat, aval Oued Khemis; Kettani et al. 1996; Dakki 1997; Kettani et al. 2001; Kettani and Langton 2012; Ashe and O’Connor; 2012 Kettani and Moubayed-Breil 2018, Rif

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