Catalogue |
James O'Hara‡, Pierfilippo Cerretti§
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Corresponding author: James O'Hara ( james.ohara@canada.ca ) Corresponding author: Pierfilippo Cerretti ( pierfilippocerretti@yahoo.it ) Academic editor: Daniel Whitmore
© 2016 James O'Hara, Pierfilippo Cerretti.
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation:
O’Hara JE, Cerretti P (2016) Annotated catalogue of the Tachinidae (Insecta, Diptera) of the Afrotropical Region, with the description of seven new genera. ZooKeys 575: 1–344. https://doi.org/10.3897/zookeys.575.6072
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Abstract
The Tachinidae of the Afrotropical Region are catalogued and seven genera and eight species are newly described. There are 237 genera and 1126 species recognized, of which 101 genera and 1043 species are endemic to the region. The catalogue is based on examination of the primary literature comprising about 525 references as well as numerous name-bearing types and other specimens housed in collections. Taxa are arranged hierarchically and alphabetically under the categories of subfamily, tribe, genus, subgenus (where recognized), species, and rarely subspecies. Nomenclatural information is provided for all genus-group and species-group names, including lists of synonyms (mostly restricted to Afrotropical taxa) and name-bearing type data. Species distributions are recorded by country within the Afrotropical Region and by larger geographical divisions outside the region. Additional information is given in the form of notes, numbering about 300 in the catalogue section. Seven genera and eight species are described as new: Afrophylax Cerretti & O’Hara with type species Sturmia aureiventris Villeneuve, 1910, gen. n. (Exoristinae, Eryciini); Austrosolieria Cerretti & O’Hara with type species Austrosolieria londti Cerretti & O’Hara, gen. n. and sp. n. (South Africa) and Austrosolieria freidbergi Cerretti & O’Hara, sp. n. (Malawi) (Tachininae, Leskiini); Carceliathrix Cerretti & O’Hara with type species Phorocera crassipalpis Villeneuve, 1938, gen. n. (Exoristinae, Eryciini); Filistea Cerretti & O’Hara with type species Viviania aureofasciata Curran, 1927, gen. n. and Filistea verbekei Cerretti & O’Hara, sp. n. (Cameroon, D.R. Congo, Uganda) (Exoristinae, Blondeliini); Mesnilotrix Cerretti & O’Hara with type species Dexiotrix empiformis Mesnil, 1976, gen. n. (Dexiinae, Dexiini); Myxophryxe Cerretti & O’Hara with type species Phorocera longirostris Villeneuve, 1938, gen. n., Myxophryxe murina Cerretti & O’Hara, sp. n. (South Africa), Myxophryxe regalis Cerretti & O’Hara, sp. n. (South Africa), and Myxophryxe satanas Cerretti & O’Hara, sp. n. (South Africa) (Exoristinae, Goniini); and Stiremania Cerretti & O’Hara with type species Stiremania karoo Cerretti & O’Hara, gen. n. and sp. n. (South Africa), and Stiremania robusta Cerretti & O’Hara, sp. n. (South Africa) (Exoristinae, Goniini). Paraclara Bezzi, 1908 is transferred from the Cylindromyiini to the Hermyini, comb. n. Sarrorhina Villeneuve, 1936 is transferred from the Minthoini to the Graphogastrini, comb. n. Three genera are newly recorded from the Afrotropical Region: Madremyia Townsend, 1916 (Eryciini); Paratrixa Brauer & Bergenstamm, 1891 (Blondeliini); and Simoma Aldrich, 1926 (Goniini). Three genera previously recorded from the Afrotropical Region are no longer recognized from the region: Calozenillia Townsend, 1927 (Palaearctic, Oriental and Australasian regions); Eurysthaea Robineau-Desvoidy, 1863 (Palaearctic, Oriental and Australasian regions); and Trixa Meigen, 1824 (Palaearctic and Oriental regions). Two species are newly recorded from the Afrotropical Region: Amnonia carmelitana Kugler, 1971 (Ethiopia, Kenya); and Simoma grahami Aldrich, 1926 (Namibia). Three species previously recorded from the Afrotropical Region are no longer recognized from the region: Euthera peringueyi Bezzi, 1925 (Oriental Region); Hamaxia incongrua Walker, 1860 (Palaearctic, Oriental and Australasian regions); Leucostoma tetraptera (Meigen, 1824) (Palaearctic Region). New replacement names are proposed for five preoccupied names of Afrotropical species: Billaea rubida O’Hara & Cerretti for Phorostoma rutilans Villeneuve, 1916, preoccupied in the genus Billaea Robineau-Desvoidy, 1830 by Musca rutilans Fabricius, 1781, nom. n.; Cylindromyia braueri O’Hara & Cerretti for Ocyptera nigra Villeneuve, 1918, preoccupied in the genus Cylindromyia Meigen, 1803 by Glossidionophora nigra Bigot, 1885, nom. n.; Cylindromyia rufohumera O’Hara & Cerretti for Ocyptera scapularis Villeneuve, 1944, preoccupied in the genus Cylindromyia Meigen, 1803 by Ocyptera scapularis Loew, 1845, nom. n.; Phytomyptera longiarista O’Hara & Cerretti for Phytomyzoneura aristalis Villeneuve, 1936, preoccupied in the genus Phytomyptera Rondani, 1845 by Phasiostoma aristalis Townsend, 1915, nom. n.; and Siphona (Siphona) pretoriana O’Hara & Cerretti for Siphona laticornis Curran, 1941, preoccupied in the genus Siphona Meigen, 1803 by Actia laticornis Malloch, 1930, nom. n. New type species fixations are made under the provisions of Article 70.3.2 of the ICZNCode for two genus-group names: Lydellina Villeneuve, 1916, type species newly fixed as Lydellina villeneuvei Townsend, 1933 (valid genus name); and Sericophoromyia Austen, 1909, type species newly fixed as Tachina quadrata Wiedemann, 1830 (synonym of Winthemia Robineau-Desvoidy, 1830). Lectotypes are designated for the following nine nominal species based on examination of one or more syntypes of each: Degeeria crocea Villeneuve, 1950; Degeeria semirufa Villeneuve, 1950; Erycia brunnescens Villeneuve, 1934; Exorista oculata Villeneuve, 1910; Kiniatilla tricincta Villeneuve, 1938; Myxarchiclops caffer Villeneuve, 1916; Ocyptera linearis Villeneuve, 1936; Peristasisea luteola Villeneuve, 1934; and Phorocera crassipalpis Villeneuve, 1938. The following four genus-group names that were previously treated as junior synonyms or subgenera are recognized as valid generic names: Bogosiella Villeneuve, 1923, status revived; Dyshypostena Villeneuve, 1939, status revived; Perlucidina Mesnil, 1952, status revived; and Thelymyiops Mesnil, 1950, status n. The following six species-group names that were previously treated as junior synonyms are recognized as valid species names: Besseria fossulata Bezzi, 1908, status revived; Degeeria cinctella Villeneuve, 1950, status revived (as Medina cinctella (Villeneuve)); Nemoraea miranda intacta Villeneuve, 1916, status revived (as Nemoraea intacta Villeneuve); Succingulum exiguum Villeneuve, 1935, status revived (as Trigonospila exigua (Villeneuve)); Wagneria rufitibia abbreviata Mesnil, 1950, status n. (as Periscepsia abbreviata (Mesnil)); and Wagneria rufitibia nudinerva Mesnil, 1950, status n. (as Periscepsia nudinerva (Mesnil)). The following 25 new or revived combinations are proposed: Afrophylax aureiventris (Villeneuve, 1910), comb. n.; Blepharella orbitalis (Curran, 1927), comb. n.; Bogosiella pomeroyi Villeneuve, 1923, comb. revived; Brachychaetoides violacea (Curran, 1927), comb. n.; Carceliathrix crassipalpis (Villeneuve, 1938), comb. n.; Charitella whitmorei (Cerretti, 2012), comb. n.; Dyshypostena edwardsi (van Emden, 1960), comb. n.; Dyshypostena tarsalis Villeneuve, 1939, comb. revived; Estheria buccata (van Emden, 1947), comb. n.; Estheria surda (Curran, 1933), comb. n.; Filistea aureofasciata (Curran, 1927), comb. n.; Madremyia setinervis (Mesnil, 1968), comb. n.; Mesnilotrix empiformis (Mesnil, 1976), comb. n.; Myxophryxe longirostris (Villeneuve, 1938), comb. n.; Nealsomyia chloronitens (Mesnil, 1977), comb. n.; Nealsomyia clausa (Curran, 1940), comb. n.; Nilea longicauda (Mesnil, 1970), comb. n.; Paratrixa aethiopica Mesnil, 1952, comb. revived; Paratrixa stammeri Mesnil, 1952, comb. revived; Perlucidina africana (Jaennicke, 1867), comb. n.; Perlucidina perlucida (Karsch, 1886), comb. revived; Prolophosia retroflexa (Villeneuve, 1944), comb. n.; Sturmia profana (Karsch, 1888), comb. n.; additionally, Ceromasia rufiventris Curran, 1927 is treated as an unplaced species of Goniini, comb. n. and Hemiwinthemia stuckenbergi Verbeke, 1973 is treated as an unplaced species of Leskiini, comb. n. New or revived generic and specific synonymies are proposed for the following nine names: Afrosturmia Curran, 1927 with Blepharella Macquart, 1851, syn. n.; Archiphania van Emden, 1945 with Catharosia Rondani, 1868, syn. revived; Besseria longicornis Zeegers, 2007 with Besseria fossulata Bezzi, 1908 (current name Besseria fossulata), syn. n.; Dexiomera Curran, 1933 with Estheria Robineau-Desvoidy, 1830, syn. n.; Hemiwinthemia francoisi Verbeke, 1973 with Nemoraea capensis Schiner, 1868 (current name Smidtia capensis), syn. n.; Kinangopana van Emden, 1960 with Dyshypostena Villeneuve, 1939, syn. n.; Metadrinomyia Shima, 1980 with Charitella Mesnil, 1957, syn. n.; Phorocera majestica Curran, 1940 with Phorocera longirostris Villeneuve, 1938 (current name Myxophryxe longirostris), syn. n.; and Podomyia discalis Curran, 1939 with Antistasea fimbriata Bischof, 1904 (current name Antistasea fimbriata), syn. n.
Keywords
Afrotropical Region, parasitoids, classification, distribution, zoological nomenclature, systematics, new taxa
Introduction
The Tachinidae are a large cosmopolitan family of flies that are parasitoids of other arthropods, primarily other insects (
Crosskey’s exemplary skills as a taxonomist, nomenclaturalist and bibliographer ensured that his Afrotropical catalogue and keys were virtually free of errors in their presentation of factual information. His higher classification of the Tachinidae, however, was little changed from his earlier conspecti and in this respect was not progressive. Nevertheless, it suited Crosskey’s desire to construct keys that would first separate tribes and then genera within tribes. His classification was already at odds with the advances being made in tachinid relationships by Mesnil, Herting and Verbeke (
The main impetus for preparing this catalogue was the announcement in 2010 during the 7th International Congress of Dipterology in San José, Costa Rica, of an international effort to publish a Manual of Afrotropical Diptera (A.H. Kirk-Spriggs and B.J. Sinclair, editors, in prep.). The Tachinidae are by far the largest family of Afrotropical Diptera in terms of genera and the Manual chapter detailing this diversity is recognized as a considerable challenge by the authors (P. Cerretti, J.E. O’Hara, J.O. Stireman and D.M. Wood, in prep.). This catalogue is intended as both a companion volume to the Manual chapter and a resource for the chapter authors as they prepare a key to genera and evaluate the diversity, biology and biogeography of the tachinid fauna.
The geographic limits of the Afrotropical Region for the purposes of this catalogue have been changed slightly from those of
Numerous specimens of Afrotropical Tachinidae were examined during the preparation of this catalogue. This has led to taxonomic changes within the catalogue and also revealed numerous new species and a smaller number of new genera. Described herein are seven new genera that are well characterized and worthy of formal recognition in this catalogue and by such treatment will be available for inclusion in the key to tachinid genera in the Manual.
The Catalogue of the Diptera of the Afrotropical Region recognized 95 families, 2020 genera and 16,550 species (
The number of Afrotropical tachinid genera and species has risen modestly over the past 35 years due to taxonomic activity, an expansion of the region’s boundaries, and the new taxa described herein. The present catalogue records 237 genera, of which 101 (43%) are endemic to the region. Of the 1126 species recorded, a total of 1043 (93%) are endemic. The current numbers represent an increase since 1980 of 29 genera and 130 species. Despite these advances, the tachinid fauna of the region remains understudied and many new taxa await discovery and description.
Materials and methods
Format
This catalogue is arranged in a similar manner to the one on the Tachinidae of China by
General
This catalogue cites all nominal species in their original combinations, provides details about name-bearing types, gives known distributions, and is based on the examination of all but a very few of the approximately 525 publications listed in the References.
Valid names are arranged hierarchically and alphabetically according to the categories of subfamily, tribe, genus, subgenus, species, and subspecies. Synonyms are given for valid names of genera, subgenera, and species and are listed chronologically. Synonymic lists comprise taxa described from the Afrotropical Region, synonyms that have been used as valid names in the literature on Afrotropical Tachinidae, and (where known) misidentifications (given last in synonymic lists).
Each genus-group name is listed with the following information: genus name in italics and capital letters (and additionally in bold if valid, unless misidentified from the Afrotropics), author, year (with letter if applicable), page, note in parentheses if applicable (e.g., junior homonym, proposed as subgenus), type species with author and date, form of type fixation, and region of origin of type species in square brackets if not the Afrotropics. Each type species is cited in its original binomen (Recommendation 67B of the Code, ICZN 1999), and if that name is a synonym then it is followed by the valid name of the species in parentheses. We have invoked Article 70.3.2 of the Code (ICZN 1999) to fix the intended species as the type species for generic names that were based on misidentified type species. This maintains the concepts of these generic names as currently accepted and in prevailing usage. The genera so affected are listed below under “Summary of new taxonomic and nomenclatural changes”.
Type species were fixed by original designation, monotypy, subsequent designation, or in a few instances subsequent monotypy, except for type species newly fixed here for nominal genera based on misidentified type species. Fixation by original designation requires an explicit designation of a type species (Article 68.2 of the Code, ICZN 1999), so a new genus “proposed for” or “erected for” a single species has its type species fixed by monotypy. A new genus proposed before 1931 for a single species and accompanied by the expression “gen. n., sp. n.” or an equivalent also has its type species fixed by monotypy (Article 68.2.1). If, on the other hand, the new genus is proposed for more than one new species and the expression “gen. n., sp. n.” or an equivalent is applied to only one of the new species, then that species is fixed as type species by original designation (Article 68.2.1).
Species are listed by valid name followed by the available name(s) associated with it; i.e., the available name of the valid name plus synonyms. The valid name is represented by the valid specific epithet in bold and italics (in italics only if questionably recorded or misidentified from the Afrotropics) followed by the author, date (no letter), and known distribution. Author and date are enclosed in parentheses if the species has moved from its original genus. The distribution is given first for the Afrotropical Region and then for other regions as explained under “Geographic divisions” and “Distributional data”. Each available name is given in italics in its original combination and spelling followed by author, year (with letter if applicable to match a publication listed in the References), page, and a note in parentheses if applicable (e.g., junior homonym, subsequent spelling). A questionable synonym is preceded by a question mark (e.g., “? Ocyptera cribrata Villeneuve”). Given next is name-bearing type information consisting of status (holotype, lectotype, neotype, or syntypes), sex (of single type, or number and sex of syntypes), type depository (in parentheses), and type locality. If a neotype or lectotype was designated then a citation is given to the designation. Additional information may be given in parentheses with the type depository to cite the number and sex of syntypes existing in a collection if that number is different from the information given in the original description, or if the original description did not provide details about the type series; also, a reference may be cited wherein information can be found about the name-bearing type.
A subsequent spelling of a generic or specific name can be an incorrect subsequent spelling (which is not an available name) or an unjustified emendation (which is an available name with its own author and date). Incorrect subsequent spellings encountered during this study are cited but there are certainly others that escaped our notice. In a departure from the catalogue of
The following acronyms are used in this work:
Code
ICZN International Commission on Zoological Nomenclature
JEOH James E. O’Hara
PC Pierfilippo Cerretti
Name-bearing types
We follow the same method developed by
Type(s), male: One or more males. This citation is used for a species described from the male sex without indication of whether a single male (i.e., a holotype) or more than one male (i.e., syntypes) comprised the type series.
Type(s), female: One or more females. See “Type(s), male”.
Type(s), unspecified sex: One or more specimens with no indication of sex.
Syntypes, [number] male[s] and [number] female[s] (e.g., “Syntypes, 3 males and 2 females”): Species described from an indicated number of males and females.
Syntypes, males and females: Species described from both sexes but the number of each sex was not given.
Syntypes, males: Species described from more than one male but without indication of the number of males.
Syntypes, females: Species described from more than one female but without indication of the number of females.
Syntypes, unspecified number and sex: Species described from more than one specimen but without indication of sex or number of specimens.
Avoidance of assumption of holotype
In following the foregoing format we have complied with Recommendation 73F of the Code (ICZN 1999), “Avoidance of assumption of holotype”, which states: “Where no holotype or syntype was fixed for a nominal species-group taxon established before 2000, and when it is possible that the nominal species-group taxon was based on more than one specimen, an author should proceed as though syntypes may exist and, where appropriate, should designate a lectotype rather than assume a holotype (see also Article 74.6)”. See
By following Recommendation 73F of the Code, assumed holotypes take on the status of syntypes. The recommendation favors “where appropriate” the designation of lectotypes. We have combined the spirit of Recommendation 73F and the provisions of Article 74.5 of the Code (ICZN 1999) to recognize certain published statements (as discussed in next section) about assumed holotypes as lectotype fixations. This follows
Lectotypifications
There are two types of lectotypification in zoological nomenclature, explicit and implicit. In the former, a single syntype in a type series is designated as lectotype; in the latter, there is some form of statement that can be construed as the selection of a single name-bearing type. We follow
“In a lectotype designation made before 2000, either the term ‘lectotype’, or an exact translation or equivalent expression (e.g. ‘the type’), must have been used or the author must have unambiguously selected a particular syntype to act as the unique name-bearing type of the taxon. When the original work reveals that the taxon had been based on more than one specimen, a subsequent use of the term ‘holotype’ does not constitute a valid lectotype designation unless the author, when wrongly using that term, explicitly indicated that he or she was selecting from the type series that particular specimen to serve as the name-bearing type”.
What constitutes a valid lectotypification (or lectotype fixation in our terminology) in the foregoing is largely dependent on how one interprets the passage about an author explicitly indicating “that he or she was selecting from the type series that particular specimen to serve as the name-bearing type”. At one end of the spectrum is the mere mention of a “holotype” or “type” by a subsequent author when the original type series clearly consisted of two or more syntypes. This statement does not constitute a lectotype fixation because the “holotype” is not distinguishable from other syntypes. At the other end of the spectrum is the mention of a “holotype” or “type” with accompanying details about its labelling, features, damage, etc. that clearly distinguishes that specimen from other syntypes; or perhaps there is only one type specimen in a collection and it is an “assumed holotype” (see section above) for a species described from an unspecified number of specimens. We considered these latter statements about a single type to qualify as lectotype fixations under Article 74.5 because they contain an explicit indication that an author accepted the cited “holotype” as the name-bearing type and restricted the term to a single recognizable specimen in a collection. We encountered many “holotype” statements that were not so easily interpretable as the aforementioned ones. For these, we adopted the criteria that there had to be reasonable grounds to believe the information provided would permit the “holotype” or “type” to be recognized in a collection, and we generally required some additional data beyond the mere mention of a “holotype” or “type”, for a statement to qualify as a lectotype fixation.
Type localities
Type localities are cited first by country and then by location within that country from larger to smaller geographic area or place. Spellings of geographic areas and places largely follow The Times Comprehensive Atlas of the World (
The type localities of almost 30 nominal species were published as the Rwenzori (often published as “Ruwenzori”) Range on the border of D.R. Congo and Uganda, frequently with additional data.
Criteria for citing type localities from Sweden, and for nominal species described by Meigen, are explained in
Collections housing name-bearing types
The location of the name-bearing type (holotype, lectotype, neotype, or syntypes) is cited for each nominal species, where known. The collections housing these name-bearing types are listed below with the acronyms used in the text. We largely accepted as accurate the statements about the deposition of name-bearing types given in the original literature unless we had reason to doubt the information given (e.g., types known to have been relocated or are presumed lost). We personally examined many of the types cited in
The acronyms of collections cited in this work are as follows:
BMNH
FMNHH
HUJI
JOS Private collection of J.O. Stireman, Dayton, Ohio, USA
MZF Museo Zoologico “La Specola”, Firenze [Florence], Italy
NMDA
Department of
SDEI
ZMHB
ZMUK
Zoologisches
Geographic divisions
The known distribution of each tachinid species recorded from the Afrotropical Region is given next to the valid name in the following order: Afrotropical Region, Palaearctic Region, Oriental Region, Australasian and Oceanian regions [cited as Australasian for brevity], Nearctic Region, and Neotropical Region. Each of these regions is subdivided according to the scheme explained below. Areas close to the Afrotropical Region are subdivided more finely than those that are distant from it. Spellings of countries and areas within countries follow, with few exceptions, The Times Comprehensive Atlas of the World (
Afrotropical Region (Fig. 1 )
The geographic limits of the Afrotropical Region follow
The names of countries and islands listed below and shown in Fig.
Angola.
Ascension (an island dependency of the United Kingdom Overseas Territory of Saint Helena).
Benin.
Botswana.
Burkina [Burkina Faso] (as Upper Volta in
Burundi.
Cameroon (as Cameroun in
Cape Verde [Cape Verde Islands].
C.A. Republic [Central African Republic].
Chad.
Comoros [Comoros Islands].
Congo.
Côte d’Ivoire [or Ivory Coast].
Djibouti.
D.R. Congo [Democratic Republic of the Congo] (as Zaire in
Eq. Guinea [Equatorial Guinea] (including Annobón and Bioco [as “Fernando Póo”] islands of
Eritrea (new country since
Ethiopia.
Gabon.
Gambia [The Gambia].
Ghana.
Guinea.
Guinea-Bissau.
Kenya.
Lesotho.
Liberia.
Madagascar.
Malawi.
Mali.
Mauritania.
Mauritius (including Cargados Carajos and Rodrigues islands of
Mozambique.
Namibia.
Niger.
Nigeria.
Oman (not included in Afrotropical Region of
Réunion (France).
Rwanda.
Saint Helena (United Kingdom Overseas Territory).
São Tomé & Príncipe (treated separately in
Senegal.
Seychelles (including Aldabra, Amirante, Astove, Coëtivy, and Cosmoledo islands of
Sierra Leone.
Somalia.
South Africa.
South Sudan (see note for Sudan; new country since
Sudan (including, for distributional purposes, South Sudan).
Swaziland.
Tanzania.
Togo.
Tristan da Cunha (an island dependency of the United Kingdom Overseas Territory of Saint Helena).
Tromelin (disputed island territory of France).
U.A. Emirates [United Arab Emirates] (not included in Afrotropical Region of
Uganda.
Yemen (including South Yemen and Suquţrá [as Socotra] of
Zambia.
Zimbabwe (as Rhodesia in
Palaearctic Region (Fig. 2 )
The traditional limits of the Palaearctic Region are recognized except that Oman and United Arab Emirates are assigned to the Afrotropical Region to conform with the upcoming Manual of Afrotropical Diptera and the boundary with the Oriental Region through China is as newly defined under Oriental China (area 12). The subdivisions of the Palaearctic Region are explained below and are shown in Fig.
1. Europe.
a. British Is. [British Isles].—United Kingdom and Republic of Ireland.
b. Scand. [Scandinavia].—Iceland, Denmark (excluding Greenland), Norway, Sweden, and Finland.
c. W. Eur. [Western Europe].—Austria, Belgium, Channel Islands, France (excluding Corse), Germany, Liechtenstein, Luxembourg, Netherlands, and Switzerland.
d. E. Eur. [Eastern Europe].—Belarus, Czech Republic, Estonia, Hungary, Kaliningradskaya [or Kaliningrad] Oblast’ (Russia), Latvia, Lithuania, Moldova, Poland, Romania, Slovakia, and Ukraine.
e. SW. Eur. [Southwestern Europe].—Andorra, Portugal (including Azores, excluding Madeira), and Spain (excluding Canary Islands).
f. SC. Eur. [Southcentral Europe].—Corse (France), Italy, Malta, Monaco, and San Marino.
g. SE. Eur. [Southeastern Europe].—Albania, Bosnia and Herzegovina, Bulgaria, Croatia, Greece, Montenegro, Macedonia, Serbia, and Slovenia.
h. Turkey.—Cyprus and Turkey.
2. N. Africa [North Africa].
a. Canary Is. [Canary Islands].—Canary Islands (Spain).
b. Madeira.—Madeira (Portugal).
c. NW. Africa [Northwestern Africa].—Algeria, Morocco, Tunisia, and Western Sahara.
d. NE. Africa [Northeastern Africa].—Egypt and Libya.
3. M. East [Middle East].
a. Israel (treated as a separate division because the Tachinidae are significantly better known from Israel than from the other countries of the Middle East).
b. M. East [Middle East] (excluding Israel).—Afghanistan, Bahrain, Iran, Iraq, Jordan, Kuwait, Lebanon, Oman, [Occupied] Palestinian territories, Qatar, Saudi Arabia, and Syria.
4. Transcaucasia.—Armenia, Azerbaijan, and Georgia.
5. C. Asia [Central Asia].—Kyrgyzstan, Tajikistan, Turkmenistan, and Uzbekistan.
6. Kazakhstan.
7. Russia [or Russian Federation].
a. W. Russia [Western Russia, excluding Kaliningradskaya Oblast’].—Bordering Scandinavia and Eastern Europe to the west, Transcaucasia to the south, Ural Mountains to the east, and Kazakhstan to the southeast.
b. W. Siberia [Western Siberia].—Bordering Western Russia to the west, Kazakhstan and Mongolia to the south, and Yenisey River to the east.
c. E. Siberia [Eastern Siberia].—Bordering Western Siberia to the west, Mongolia and China to the south, and Russian administrative divisions of Chukotskiy [or Chukotka] Avtonomnyy Okrug, Magadanskaya [or Magadan] Oblast’, Khabarovskiy [or Khabarovsk] Kray, and Amurskaya [or Amur] Oblast’ to the east.
d-e. Far East [Russian Far East].—Bordering Eastern Siberia to the west, China and North Korea to the south, and Japan to the southeast.
d. N. Far East [Northern Russian Far East].—Russian administrative divisions of Chukotskiy Avtonomnyy Okrug, Magadanskaya Oblast’, and Kamchatskiy [or Kamchatka] Kray.
e. S. Far East [Southern Russian Far East].—Russian administrative divisions of Khabarovskiy Kray, Amurskaya Oblast’, Yevreyskaya [or Jewish] Avtonomnaya Oblast’, and Sakhalinskaya [or Sakhalin] Oblast’ (including Kuril Islands).
8. Mongolia.
9. Korea.—North and South Korea. Cited as Korea when more detailed distributional data is not available.
a. N. Korea [North Korea].
b. S. Korea [South Korea].
10. Japan (excluding Ryukyu I.).
11. Pal. China [Palaearctic China]. North of the dotted line in Fig.
Oriental Region (Fig. 2 )
The Oriental Region is bounded on the south by Weber’s Line (following Evenhuis 1989) and on the north and west by the Palaearctic Region. The subdivisions of the Oriental Region are explained below and are shown on Fig.
12. Orien. China [Oriental China]. The Oriental portion of China is newly defined here as comprising the southern half of Chongqing, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hunan, Jiangxi, Macau, Shanghai, southern half of Sichuan, most of Yunnan except for the extreme northwest portion, and Zhejiang. A species recorded from Palaearctic China and additionally Sichuan and/or Yunnan, with no other records from Oriental China, is recorded only from Palaearctic China; e.g., Periscepsia carbonaria (Panzer).
13. Maldives etc.—Maldives, Lakshadweep (India), British Indian Ocean Territory [or Chagos Archipelago] (United Kingdom Overseas Territory).
14. Pakistan.
15. India.
16. Sri Lanka.
17. Nepal.
18. Bhutan.
19. Bangladesh.
20. Myanmar [or Burma].
21. Laos.
22. Vietnam.
23. Cambodia.
24. Thailand.
25. Andaman & Nicobar Is.—Andaman and Nicobar Islands (India).
26. Malaysia.
27. Singapore.
28. Brunei.
29. Indonesia (Oriental part as delimited by Weber’s Line; mainly Borneo, Jawa [or Java], Lesser Sunda Islands, Sulawesi [or Celebes], and Sumatera [or Sumatra]).
30. Christmas & Cocos Is.—Territories of Christmas Island and Cocos [or Keeling] Islands (Australia).
31. Philippines.
32. Taiwan.
33. Ryukyu Is.—Ryukyu Islands [or Nansei-shotō] (Japan).
Australasian and Oceanian regions
These regions are combined under the title of Australasian Region for the purposes of this catalogue. The combined region is bounded on the north by the Oriental Region (Weber’s Line) and is subdivided as follows.
N. Australasian.—Indonesia (Australasian part as delimited by Weber’s Line; mainly Maluku [or Moluccas] Islands, Western New Guinea [or Irian Jaya], and Papua New Guinea (including Bismarck Archipelago).
Australia.
Hawaii.—Hawaiian Islands (USA).
Melanesia.—Melanesia (excluding Papua New Guinea and Bismarck Archipelago, listed as part of N. Australasian), principally Fiji, New Caledonia (France), Solomon Islands, and Vanuatu.
Micronesia.—Federated States of Micronesia, principally Guam (USA), Kiribati, Marshall Islands, Nauru, Northern Mariana Islands (USA), and Palau.
New Zealand.
Polynesia.—Polynesia (excluding New Zealand and Hawaii, each listed separately), principally American Samoa (USA), Cook Islands (New Zealand), Easter Island (Chile), French Polynesia (France), Niue (New Zealand), Pitcairn Islands (United Kingdom), Samoa, Tokelau (New Zealand), Tonga, Tuvalu, and Wallis and Futuna (France).
Nearctic Region
The Nearctic Region is arbitrarily defined as America north of Mexico for the purposes of this catalogue, including Greenland (Denmark) and Bermuda (United Kingdom Overseas Territory) but not Hawaii (USA) and the West Indies (following
Neotropical Region
This region is bounded on the north by the Nearctic Region. There are only three species recorded from the region in this catalogue: Leucostoma simplex (Fallén), Trichopoda giacomellii (Blanchard) (introduced into South Africa and establishment unknown), and Voria ruralis (Fallén).
Sample distribution
A species recorded from all regions and subdivisions recognized here would be cited with the following distribution:
Afrotropical: Angola, Ascension, Benin, Botswana, Burkina, Burundi, Cameroon, Cape Verde, C.A. Republic, Chad, Comoros, Congo, Côte d’Ivoire, Djibouti, D.R. Congo, Eq. Guinea, Eritrea, Ethiopia, Gabon, Gambia, Ghana, Guinea, Guinea-Bissau, Kenya, Lesotho, Liberia, Madagascar, Malawi, Mali, Mauritania, Mauritius, Mozambique, Namibia, Niger, Nigeria, Oman, Réunion, Rwanda, Saint Helena, São Tomé & Príncipe, Senegal, Seychelles, Sierra Leone, Somalia, South Africa, Sudan, Swaziland, Tanzania, Togo, Tristan da Cunha, Tromelin, U.A. Emirates, Uganda, Yemen, Zambia, Zimbabwe. Palaearctic: C. Asia, Europe (British Is., Scand., W. Eur., E. Eur., SW. Eur., SC. Eur., SE. Eur., Turkey) [or Europe (all), if recorded from all subdivisions], Japan, Kazakhstan, Korea (N. Korea, S. Korea), M. East (Israel, M. East) [or M. East (all)], Mongolia, N. Africa (Canary Is., Madeira, NW. Africa, NE. Africa) [or N. Africa (all)], Pal. China, Russia (W. Russia, W. Siberia, E. Siberia, N. Far East, S. Far East) [or Russia (all)], Transcaucasia. Oriental: Andaman & Nicobar Is., Bangladesh, Bhutan, Brunei, Cambodia, Christmas & Cocos Is., India, Indonesia, Laos, Malaysia, Maldives etc., Myanmar, Nepal, Orien. China, Pakistan, Philippines, Ryukyu Is., Singapore, Sri Lanka, Taiwan, Thailand, Vietnam. Australasian: Australia, Hawaii, Melanesia, Micronesia, N. Australasian, New Zealand, Polynesia. Nearctic: [individual distribution]. Neotropical: [individual distribution].
Distributional data
Distributions within the Afrotropical Region
Distributions are cited for each species based on published records, examination of specimens in collections, and material collected by the authors (primarily PC) or made available to us by colleagues (see Acknowledgements). The principle source for published records was
Distributions outside the Afrotropical Region
The primary sources for extralimital distributions were
Classification
The classification adopted here recognizes the usual four subfamilies—Dexiinae, Exoristinae, Phasiinae and Tachininae—a classificatory scheme that has been generally accepted since the time of
At the tribal level, the greatest difference between the classification of
A few tribes have been moved to other subfamilies since
The Campylochetini, Thelairiini, Voriini and Wagneriini were recognized as distinct tribes within the Tachininae by
The Eloceriini, Linnaemyini, and Loewiini were recognized as tribes (within the Tachininae) by
The Eutherini, a small tribe with one of its two genera (Euthera Loew) present in the Afrotropics, have the distinction of being one of only two tribes in recent decades to have been treated in the Phasiinae by some authors (e.g.,
The Imitomyiini have also been treated in the Phasiinae by some authors (e.g.,
Another tribe of enigmatic placement, the Strongygastrini, is newly recorded from the Afrotropical Region. The tribe has typically been considered an unusual member of the Phasiinae (because it is ovolarviparous and not restricted to parasitizing heteropterans) (e.g.,
Within the Phasiinae, the Cinochirini of
With respect to the priority of family-group names,
Review of generic changes since Crosskey’s Afrotropical catalogue
There has been no dramatic reappraisal of the Afrotropical genera of Tachinidae since
Three lists are given in this section. In the first list that follows are the genera and subgenera that have been described or recorded from the Afrotropical Region since
Acemya Robineau-Desvoidy, 1830 (
Amnonia Kugler, 1971 (
Anomalostomyia Cerretti & Barraclough, 2007.
Apomorphomyia Crosskey, 1984.
Brachychaetoides Mesnil, 1970 (treated as a synonym of Chlorolydella Townsend, 1933 by
Calliethilla Shima, 1979 (
Calyptromyia Villeneuve, 1915 (
Campylocheta Rondani, 1859 (as Elpe Robineau-Desvoidy, 1863 in
Chryserycia Mesnil, 1977 (described from Madagascar by
Clairvilliops Mesnil, 1959 (treated as a synonym of Dionaea Robineau-Desvoidy, 1830 by
Clausicella Rondani, 1856 (as Istoglossa Rondani, 1856 in
Conopomima Mesnil, 1978 (published too late to be included in
Crassicornia Kugler, 1980.
Dionomelia Kugler, 1978 (
Estheria Robineau-Desvoidy, 1830 (as Dolichodexia Brauer & Bergenstamm, 1889 in
Eugaedioxenis Cerretti, O’Hara & Stireman, 2015 (
Exoristella Herting, 1984 as subgenus of Exorista Meigen, 1803.
Istocheta Rondani, 1859 (as Prosopofrontina Townsend, 1926 in
Kaiseriola Mesnil, 1970 (treated as a synonym of Diaprochaeta Mesnil, 1970 by
Kuwanimyia Townsend, 1916 (
Lydella Robineau-Desvoidy, 1830 (as Metoposisyrops Townsend, 1916 in
Mediosetiger Barraclough, 1983.
Meigenia Robineau-Desvoidy, 1830 (
Minthosoma Zeegers, 2007.
Montanothalma Barraclough, 1996.
Myxogaedia Mesnil, 1956 (treated as a synonym of Pretoriana Curran, 1938 by
Nardia Cerretti, 2009.
Nealsomyia Mesnil, 1939 (
Neophryxe Townsend, 1916 (
Nilea Robineau-Desvoidy, 1863 (recorded from Madagascar by
Ossidingia Townsend, 1919 (treated as a synonym of Nemorilla Rondani, 1856 by
Paraclara Bezzi, 1908 (treated as a synonym of Clara Brauer & Bergenstamm, 1889 by
Phasia Latreille, 1804 (as Alophora Robineau-Desvoidy, 1830 in
Piligenoides Barraclough, 1985.
Pseudalsomyia Mesnil, 1968 (
Ptilotachina Brauer & Bergenstamm, 1891 as subgenus of Exorista Meigen, 1803.
Ramonella Kugler, 1980 (
Rhinophoroides Barraclough, 2005.
Rhynchogonia Brauer & Bergenstamm, 1893 (
Rossimyiops Mesnil, 1953 (
Schembria Rondani, 1861 (
Senometopia Macquart, 1834 was treated as a subgenus of Carcelia Robineau-Desvoidy, 1830 by
Smidtia Robineau-Desvoidy, 1830 (as Timavia Robineau-Desvoidy, 1863 in
Spixomyia Crosskey, 1967 as subgenus of Exorista Meigen, 1803.
Stomina Robineau-Desvoidy, 1830 (undetermined species noted by
Stylocarcelia Zeegers, 2007.
Thrixion Brauer & Bergenstamm, 1889 (
Trichopoda Berthold, 1827 (two species introduced into South Africa in the 1990s but no confirmation of establishment).
In the second list below are given genus-group names that have changed status since
Alophorella Townsend, 1912 was treated as a subgenus of Alophora Robineau-Desvoidy, 1830 by
Asiphona Mesnil, 1954 was treated as a genus by
Carcelita Mesnil, 1975 was treated as a nomen nudum by
Caricelia Mesnil, 1975 was treated as a subgenus of Carcelia Robineau-Desvoidy, 1830 by
Ceranthia Robineau-Desvoidy, 1830 was treated as a genus by
Cuphocera Macquart, 1845 was treated as a genus by
Elfia Robineau-Desvoidy, 1850 was treated as a genus by
Mapolomyia Verbeke, 1960 was treated as a genus by
Mormonomyia Brauer & Bergenstamm, 1891 was treated as a subgenus of Alophora Robineau-Desvoidy, 1830 by
Palexorista Townsend, 1921 was treated as a genus by
Phaniola Mesnil, 1978 was listed as a genus by Crosskey (1981a) in the Appendix to the Afrotropical catalogue but was placed in synonymy with Catapariprosopa Townsend, 1927 by
Podotachina Brauer & Bergenstamm, 1891 was treated as a synonym of Exorista Meigen, 1803 by
Stomatomyia Brauer & Bergenstamm, 1889 was treated as a genus by
Tricoliga Rondani, 1856 was treated as a synonym of Exorista Meigen, 1803 by
Trypherosoma Verbeke, 1962 was treated as a genus by
Zelindomyia Verbeke, 1962 was treated as a genus by
Ziminiola Mesnil, 1978 was treated as a genus by
Zygobothria Mik, 1891 was treated as a genus by
In the third list below are given the genus-group names that were treated as valid by
Clairvillia Robineau-Desvoidy, 1830 was treated as a genus by
Dexiotrix Villeneuve, 1936 was treated as a genus by
Dionaea Robineau-Desvoidy, 1830 was treated as a genus by
Gymnophryxe Villeneuve, 1922 was treated as a genus by
Recent family reassignment of copal inclusions from East Africa
Two copal inclusions from East Africa were believed to be Baltic amber fossils of Tachinidae until
Summary of new taxonomic and nomenclatural changes
New genera and species
Afrophylax Cerretti & O’Hara. Type species: Sturmia aureiventris Villeneuve, 1910, by designation herein. Gen. n.
Austrosolieria Cerretti & O’Hara. Type species: Austrosolieria londti Cerretti & O'Hara, sp. n., by designation herein. Gen. n.
Austrosolieria freidbergi Cerretti & O’Hara. Sp. n. (Malawi).
Austrosolieria londti Cerretti & O’Hara. Sp. n. (South Africa).
Carceliathrix Cerretti & O’Hara. Type species: Phorocera crassipalpis Villeneuve, 1938, by designation herein. Gen. n.
Filistea Cerretti & O’Hara. Type species: Viviania aureofasciata Curran, 1927, by designation herein. Gen. n.
Filistea verbekei Cerretti & O’Hara. Sp. n. (Cameroon, D.R. Congo, Uganda).
Mesnilotrix Cerretti & O’Hara. Type species: Dexiotrix empiformis Mesnil, 1976, by designation herein. Gen. n.
Myxophryxe Cerretti & O’Hara. Type species: Phorocera longirostris Villeneuve, 1938, by designation herein. Gen. n.
Myxophryxe murina Cerretti & O’Hara. Sp. n. (South Africa).
Myxophryxe regalis Cerretti & O’Hara. Sp. n. (South Africa).
Myxophryxe satanas Cerretti & O’Hara. Sp. n. (South Africa).
Stiremania Cerretti & O’Hara. Type species: Stiremania karoo Cerretti & O’Hara, sp. n., by designation herein. Gen. n.
Stiremania karoo Cerretti & O’Hara. Sp. n. (South Africa).
Stiremania robusta Cerretti & O’Hara. Sp. n. (South Africa).
Genera newly recorded from the Afrotropical Region
The following genera are newly recorded from the Afrotropical Region based on species that were placed in other genera by
Madremyia Townsend, 1916 (one species placed in Phryxe Robineau-Desvoidy, 1830 by
Paratrixa Brauer & Bergenstamm, 1891 (two species placed in Medina Robineau-Desvoidy, 1830 by
The following genus is newly recorded from the Afrotropical Region based on a described species not previously reported from the region.
Simoma Aldrich, 1926 (based on new record of Simoma grahami Aldrich). New record.
Genera no longer recognized from the Afrotropical Region
The following genera, which are currently recorded from the Afrotropical Region in the literature (e.g.,
Calozenillia Townsend, 1927 [Oriental; also Australasian and Palaearctic]. The two species placed under Calozenillia by
Eurysthaea Robineau-Desvoidy, 1863 [Palaearctic; also Oriental and Australasian]. The single species recognized under Eurysthaea by
Trixa Meigen, 1824 [Palaearctic; also Oriental]. Dexiotrix empiformis Mesnil, 1976 from Madagascar was transferred to Trixa by
Species newly recorded from the Afrotropical Region
The following species are newly recorded from the Afrotropical Region. New country records for Afrotropical species are noted in the Catalogue section.
Amnonia carmelitana Kugler, 1971 (Ethiopia, Kenya).
Simoma grahami Aldrich, 1926 (Namibia).
Species misidentified or misrecorded from the Afrotropical Region
Species that are newly recognized as misidentified or misrecorded from the Afrotropical Region are listed here.
Euthera peringueyi Bezzi, 1925 [Oriental]. The type locality was originally given as “Chabra, Congo” and on this basis E. peringueyi was recorded from “Congo: Chabra” by van
Hamaxia incongrua Walker, 1860 [Australasian; also Oriental and Palaearctic]. Recorded from Tanzania by
Leucostoma tetraptera (Meigen, 1824) [Palaearctic]. Recorded from Botswana, Nigeria and South Africa by
New replacement names
Five new names are proposed for preoccupied names of Afrotropical species. Preoccupied names that are currently recognized as junior synonyms are not renamed in this work.
Billaea rubida O’Hara & Cerretti is proposed as a nomen novum for Phorostoma rutilans Villeneuve, 1916, a name preoccupied in the genus Billaea Robineau-Desvoidy, 1830 by Musca rutilans Fabricius, 1781 [Nearctic]. Nom. n.
Cylindromyia braueri O’Hara & Cerretti is proposed as a nomen novum for Ocyptera nigra Villeneuve, 1918, a name preoccupied in the genus Cylindromyia Meigen, 1803 by Glossidionophora nigra Bigot, 1885 [Neotropical]. Nom. n.
Cylindromyia rufohumera O’Hara & Cerretti is proposed as a nomen novum for Ocyptera scapularis Villeneuve, 1944, a junior primary homonym of Ocyptera scapularis Loew, 1845 [Palaearctic]. Nom. n.
Phytomyptera longiarista O’Hara & Cerretti is proposed as a nomen novum for Phytomyzoneura aristalis Villeneuve, 1936, a name preoccupied in the genus Phytomyptera Rondani, 1845 by Phasiostoma aristalis Townsend, 1915 [Nearctic]. Nom. n.
Siphona (Siphona) pretoriana O’Hara & Cerretti is proposed as a nomen novum for Siphona laticornis Curran, 1941, a name preoccupied in the genus Siphona Meigen, 1803 by Actia laticornis Malloch, 1930 [Oriental]. Nom. n.
New type species fixations
Article 70.3.2 of the Code (ICZN 1999) allows the type species of a nominal genus to be fixed as the species intended by the original author if the type species designated by that author was misidentified. We have invoked Article 70.3.2 for the two instances of misidentified type species in this catalogue that had not been dealt with previously (e.g.,
Lydellina Villeneuve, 1916c: 490. Type species newly fixed as Lydellina villeneuvei Townsend, 1933. Valid generic name.
Sericophoromyia Austen, 1909: 95. Type species newly fixed as Tachina quadrata Wiedemann, 1830. Synonym of Winthemia Robineau-Desvoidy, 1830.
Lectotype designations
Lectotypes are designated for the following nominal species (see Lectotype Designations section).
Degeeria crocea Villeneuve, 1950. This is a valid name in the genus Medina Robineau-Desvoidy, 1830, as Medina crocea (Villeneuve).
Degeeria semirufa Villeneuve, 1950. This is a valid name in the genus Medina Robineau-Desvoidy, 1830, as Medina semirufa (Villeneuve).
Erycia brunnescens Villeneuve, 1934. This is a valid name in the genus Thelairosoma Villeneuve, 1916, as Thelairosoma brunnescens (Villeneuve).
Exorista oculata Villeneuve, 1910. This is a valid name in the genus Carcelia Robineau-Desvoidy, 1830 (subgenus Carcelita Mesnil, 1975), as Carcelia (Carcelita) oculata (Villeneuve).
Kiniatilla tricincta Villeneuve, 1938. This is a valid name in the genus Kiniatilla Villeneuve, 1938.
Myxarchiclops caffer Villeneuve, 1916. This is a valid name in the genus Myxarchiclops Villeneuve, 1916.
Ocyptera linearis Villeneuve, 1936. This is a junior synonym in the genus Cylindromyia Meigen, 1803. The valid name of the species is Cylindromyia soror (Wiedemann, 1830).
Peristasisea luteola Villeneuve, 1934. This is a valid name in the genus Peristasisea Villeneuve, 1934.
Phorocera crassipalpis Villeneuve, 1938. This valid name is designated as the type species of Carceliathrix Cerretti & O’Hara, gen. n.
New and revived status
Changes to genus-group names
Bogosiella Villeneuve, 1923, which was synonymized with Phasia Latreille, 1804 by
Dyshypostena Villeneuve, 1939, which was treated as a junior synonym of Sumpigaster Macquart, 1855 by
Perlucidina Mesnil, 1952, which was synonymized with Calozenillia Townsend, 1927 by
Thelymyiops Mesnil, 1950, which was originally proposed as a subgenus of Carcelia Robineau-Desvoidy, 1830 and was treated as such by
Changes to species-group names
Besseria fossulata Bezzi, 1908, which was treated as a junior synonym of Actia zonaria Loew, 1847 in the genus Besseria Robineau-Desvoidy by
Degeeria cinctella Villeneuve, 1950, which was treated as a junior synonym of Degeeria lateralis Villeneuve, 1950 in the genus Medina Robineau-Desvoidy by
Nemoraea miranda intacta Villeneuve, 1916, which was treated as a valid name by
Succingulum exiguum Villeneuve, 1935, which was treated as a junior synonym of Succingulum mista Villeneuve, 1913 in the genus Trigonospila Pokorny by
Wagneria rufitibia abbreviata Mesnil, 1950, which was treated as a junior synonym of Wagneria rufitibia Villeneuve, 1938 in the genus Periscepsia Gistl by
Wagneria rufitibia nudinerva Mesnil, 1950, which was treated as a junior synonym of Wagneria rufitibia Villeneuve, 1938 in the genus Periscepsia Gistl by
New and revived combinations
New and revived combinations proposed in this work are listed below. These are based on the study of type material, authoritatively identified specimens, and/or descriptions and figures in the literature by PC.
Afrosturmia orbitalis Curran, 1927 (type species of Afrosturmia Curran) is moved from its original placement in Afrosturmia to Blepharella Macquart (with Afrosturmia in synonymy). Comb. n.
Alsomyia chloronitens Mesnil, 1977, which was published too late to be included in
Bogosiella pomeroyi Villeneuve, 1923 (type species of Bogosiella Villeneuve) is returned to Bogosiella from its placement in Phasia Latreille by
Campylochaeta violacea Curran, 1927 is moved to Brachychaetoides Mesnil from its placement in Chlorolydella Townsend by
Ceromasia rufiventris Curran, 1927 is moved to Goniini, and treated as an unplaced species within the tribe, from its placement in Eurysthaea Robineau-Desvoidy by
Degeeria profana Karsch, 1888 is moved to Sturmia Robineau-Desvoidy from its placement in “Unplaced species of Goniinae” by
Dexia buccata van Emden, 1947 is moved to Estheria Robineau-Desvoidy from its treatment as a “species of uncertain generic affiliation” by
Dexiomera surda Curran, 1933 (type species of Dexiomera Curran) is moved from its original placement in Dexiomera to Estheria Robineau-Desvoidy (with Dexiomera in synonymy). Comb. n.
Dexiotrix empiformis Mesnil, 1976 is moved to Mesnilotrix gen. n. from its placement in Trixa Meigen by
Dyshypostena tarsalis Villeneuve, 1939 (type species of Dyshypostena Villeneuve) is returned to Dyshypostena Villeneuve from its placement in Sumpigaster Macquart by
Exorista africana Jaennicke, 1867 is moved to Perlucidina Mesnil from its placement in Calozenillia Townsend by
Exorista perlucida Karsch, 1886 (type species of Perlucidina Mesnil) is returned to Perlucidina from its placement in Calozenillia Townsend by
Hemiwinthemia stuckenbergi Verbeke, 1973 is moved to Leskiini, and treated as an unplaced species within the tribe, from its original placement in Hemiwinthemia Verbeke. Comb. n.
Kinangopana edwardsi van Emden, 1960 (type species of Kinangopana van Emden) is moved from its original placement in Kinangopana to Dyshypostena Villeneuve (with Kinangopana in synonymy). Comb. n.
Metadrinomyia whitmorei Cerretti, 2012 is moved to Charitella Mesnil from its original placement in Metadrinomyia Shima. Comb. n.
Ocyptera retroflexa Villeneuve, 1944 is moved to Prolophosia Townsend from its placement in Cylindromyia Meigen by
Paratrixa aethiopica Mesnil, 1952 is returned to Paratrixa Brauer & Bergenstamm from its placement in Medina Robineau-Desvoidy by
Paratrixa stammeri Mesnil, 1952 is returned to Paratrixa Brauer & Bergenstamm from its placement in Medina Robineau-Desvoidy by
Phorocera clausa Curran, 1940 is moved to Nealsomyia Mesnil from its placement in “Unplaced species of Goniinae” by
Phorocera crassipalpis Villeneuve, 1938 is moved to Carceliathrix gen. n. (and designated as its type species) from its placement in “Unplaced species of Carceliini” by
Phorocera longirostris Villeneuve, 1938 is moved to Myxophryxe gen. n. (and designated as its type species) from its placement in Pretoriana Curran, 1938 by
Phryxe setinervis Mesnil, 1968 is moved to Madremyia Townsend from its original placement in Phryxe Robineau-Desvoidy. Comb. n.
Sturmia aureiventris Villeneuve, 1910 is moved to Afrophylax gen. n. (and designated as its type species) from its placement in “Unplaced species of Carceliini” by
Sturmia longicauda Mesnil, 1970 is moved to Nilea Robineau-Desvoidy from its original placement in Sturmia Robineau-Desvoidy. Comb. n.
Viviania aureofasciata Curran, 1927 is moved to Filistea gen. n. (and designated as its type species) from its placement in “Unplaced species of Tachinidae” by
New and revived synonymies
New and revived generic and specific synonymies are proposed for the names below. As with the new and revived combinations listed above, they result from the study of type material, authoritatively identified specimens, and/or descriptions and figures in the literature by PC.
Afrosturmia Curran, 1927, which was treated as a genus by
Archiphania van Emden, 1945 was treated as a genus by
Besseria longicornis Zeegers, 2007 is synonymized with Besseria fossulata Bezzi, 1908. The current combination is Besseria fossulata Bezzi. Syn. n.
Dexiomera Curran, 1933, which was treated as a genus by
Hemiwinthemia francoisi Verbeke, 1973, which was overlooked by
Kinangopana van Emden, 1960, which was treated as a genus by
Metadrinomyia Shima, 1980 is synonymized with Charitella Mesnil, 1957. Syn. n.
Phorocera majestica Curran, 1940 is synonymized with Phorocera longirostris Villeneuve, 1938. The current combination is Myxophryxe longirostris (Villeneuve). Syn. n.
Podomyia discalis Curran, 1939 is synonymized with Antistasea fimbriata Bischof, 1904. The current combination is Antistasea fimbriata Bischof. Syn. n.
Catalogue
Subfamily DEXIINAE (Fig. 3 )
Tribe DEXIINI
Genus BILLAEA Robineau-Desvoidy, 1830
BILLAEA Robineau-Desvoidy, 1830: 328. Type species: Billaea grisea Robineau-Desvoidy, 1830 (= Dexia pectinata Meigen, 1826), by monotypy [Palaearctic].
OMALOGASTER Macquart, 1834: 51 [also 1834: 187]. Type species: Billaea grisea Robineau-Desvoidy, 1830 (= Dexia pectinata Meigen, 1826), by subsequent designation of
GIGAMYIA Macquart, 1844: 115 [also 1844: 272]. Type species: Stomoxys gigantea Wiedemann, 1824, by original designation.
HOMALOGASTER Agassiz, 1846b: 184. Unjustified emendation of Omalogaster Macquart, 1834.
PARAPROSENA Brauer & Bergenstamm, 1889: 127 [also 1890: 59]. Type species: Paraprosena waltlii Brauer & Bergenstamm, 1889 (= Dexia marmorata Meigen, 1838), by monotypy [Palaearctic].
GYMNODEXIA Brauer & Bergenstamm, 1891: 364 [also 1891: 60]. Type species: Dexia triangulifera Zetterstedt, 1844, by subsequent designation of
AMPHIBOLIOPSIS Townsend, 1926b: 538. Type species: Gymnostylia minor Villeneuve, 1913, by original designation.
CHAETOBILLAEA Mesnil, 1976: 44 (as subgenus of Billaea Robineau-Desvoidy, 1830). Type species: Billaea (Chaetobillaea) communis Mesnil, 1976, by original designation.
africana (Villeneuve, 1935).—Afrotropical: D.R. Congo, Ethiopia, Kenya, South Africa, Tanzania.
Paraprosena marmorata africana Villeneuve, 1935a: 138. Syntypes, 5 males (possibly 1 male in
Billaea neavei van Emden, 1947: 643. Holotype male (BMNH). Type locality: Kenya, Marsabit [as “Marsabit, Rendili Njoro, N. Frontier District”].
Note:
capensis van Emden, 1947.—Afrotropical: South Africa.
Billaea capensis van Emden, 1947: 645. Holotype male (BMNH). Type locality: South Africa, Western Cape, 40 miles from Cape Town, Viljoen’s Pass [as “Viljoeus Pass”, ca. 34°5′S 19°3′E].
communis Mesnil, 1976.—Afrotropical: Madagascar.
Billaea (Chaetobillaea) communis Mesnil, 1976: 45. Holotype male (
decisa (Curran, 1927).—Afrotropical: D.R. Congo.
Gymnodexia decisa Curran, 1927a: 7. Holotype male (
edwardsi (van Emden, 1947).—Afrotropical: Uganda.
Paraprosena edwardsi van Emden, 1947: 658. Holotype female (BMNH). Type locality: Uganda, Rwenzori Range [as “Ruwenzori”], Mobuku Valley, 7300ft.
gigantea (Wiedemann, 1824).—Afrotropical: South Africa.
Stomoxys gigantea Wiedemann, 1824: 41. Type(s), female (1 syntype in
grandis Mesnil, 1976.—Afrotropical: Madagascar.
Billaea (Chaetobillaea) grandis Mesnil, 1976: 46. Holotype male (
interrupta (Curran, 1927).—Afrotropical: D.R. Congo.
Gymnodexia interrupta Curran, 1927a: 8. Holotype male (
Note: Billaea interrupta (Curran, 1927) is a senior secondary homonym of B. interrupta (Curran, 1929), a name currently treated as valid in the Nearctic Region (
lateralis (Curran, 1927).—Afrotropical: D.R. Congo.
Gymnodexia lateralis Curran, 1927a: 6. Holotype male (
lativentris van Emden, 1947.—Afrotropical: Kenya.
Billaea lativentris van Emden, 1947: 646. Holotype male (BMNH). Type locality: Kenya, Mt. Elgon, 10,500–11,500ft.
minor (Villeneuve, 1913).—Afrotropical: D.R. Congo, Ethiopia, Kenya, South Africa, Uganda.
Gymnostylia minor Villeneuve, 1913c: 37. Lectotype male (
Note: Gymnostylia minor Villeneuve, 1913 was described from two males, one from “Natal” (South Africa) and the other from Kericho (Kenya) and dated “27-XI-1911”.
orbitalis van Emden, 1947.—Afrotropical: South Africa.
Billaea orbitalis van Emden, 1947: 644. Holotype male (BMNH). Type locality: South Africa, Western Cape, Malgas [as “Malagas”].
ovata Mesnil, 1976.—Afrotropical: Madagascar.
Billaea (Chaetobillaea) ovata Mesnil, 1976: 45. Holotype male (
rhingiaeformis van Emden, 1959.—Afrotropical: Ethiopia.
Billaea rhingiaeformis van Emden, 1959: 186. Holotype male (BMNH). Type locality: Ethiopia, Simien Mountains, Lori, 11,500ft [ca. 13°17′N 38°12′E, see map in
rubida O’Hara & Cerretti, nom. n.—Afrotropical: South Africa.
Phorostoma rutilans Villeneuve, 1916c: 504 (junior secondary homonym of Musca rutilans Fabricius, 1781). Syntypes, males (1 male in
Billaea rubida O’Hara & Cerretti, nom. n. for Phorostoma rutilans Villeneuve, 1916.
Note: Phorostoma rutilans Villeneuve, 1916 is a junior secondary homonym of Musca rutilans Fabricius, 1781, the valid name of a Nearctic species of Billaea (
setosa (Macquart, 1844).—Afrotropical: South Africa.
Gymnostylia setosa Macquart, 1844: 88 [also 1844: 245]. Syntypes, males and females (lost,
sjostedti Speiser, 1910.—Afrotropical: Ethiopia, Kenya, Tanzania, Uganda.
Billaea sjostedti Speiser, 1910: 146 (as “sjöstedti”). Lectotype male (
Note: Billaea sjostedti Speiser, 1910 was described from two males from the area of “Kilimandjaro”, with one male further restricted to “Kibonoto” [now Kibongoto] at 1000m.
solivaga Mesnil, 1976.—Afrotropical: Madagascar.
Billaea (Chaetobillaea) solivaga Mesnil, 1976: 46. Holotype male (
vanemdeni Fennah, 1959.—Afrotropical: Ghana.
Billaea vanemdeni Fennah, 1959: 682. Holotype male (BMNH). Type locality: Ghana, Tafo, West African Cacao Research Institute.
velutina Mesnil, 1976.—Afrotropical: Madagascar.
Billaea velutina Mesnil, 1976: 42. Holotype male (
versicolor (Curran, 1927).—Afrotropical: D.R. Congo.
Gymnodexia versicolor Curran, 1927a: 7. Holotype male (
villeneuvei (Curran, 1927).—Afrotropical: D.R. Congo.
Gymnodexia villeneuvei Curran, 1927a: 5. Holotype male (
vitripennis Mesnil, 1950.—Afrotropical: Zimbabwe.
Billaea (Homalogaster) vitripennis Mesnil, 1950d: 116. Syntypes, males and females (“Plusieurs exemplaires”) (1 male in
Genus CHAETODEXIA Mesnil, 1976
CHAETODEXIA Mesnil, 1976: 49. Type species: Chaetodexia keiseri Mesnil, 1976, by original designation.
keiseri Mesnil, 1976.—Afrotropical: Madagascar.
Chaetodexia keiseri Mesnil, 1976: 50. Holotype male (
nigrescens Mesnil, 1976.—Afrotropical: Madagascar.
Chaetodexia keiseri nigrescens Mesnil, 1976: 50. Holotype male (
pallida Mesnil, 1976.—Afrotropical: Madagascar.
Chaetodexia pallida Mesnil, 1976: 50. Holotype male (
trilineata Mesnil, 1976.—Afrotropical: Madagascar.
Chaetodexia trilineata Mesnil, 1976: 51. Holotype male (
Genus DEXIA Meigen, 1826
DEXIA Meigen, 1826: 33. Type species: Musca rustica Fabricius, 1775, by designation under the Plenary Powers of ICZN (1988: 74) [Palaearctic].
DEXILLA Westwood, 1840: 140. Type species: Musca rustica Fabricius, 1775, by original designation [Palaearctic].
aurohumera van Emden, 1947.—Afrotropical: Mozambique.
Dexia aurohumera van Emden, 1947: 634. Holotype male (BMNH). Type locality: Mozambique, Maputo [as “Lorenzo Marques”].
capensis Robineau-Desvoidy, 1830.—Afrotropical: Kenya, South Africa, Tanzania.
Dexia capensis Robineau-Desvoidy, 1830: 314. Type(s), unspecified sex (
Dexia afra Curran, 1927f: 104. Holotype male (BMNH). Type locality: South Africa, KwaZulu-Natal, Durban.
cuthbertsoni (Curran, 1941).—Afrotropical: Kenya, Liberia, Nigeria, Sierra Leone, Zimbabwe.
Dexilla cuthbertsoni Curran, 1941: 1. Holotype female (
Dexilla bequaerti Curran, 1941: 2. Holotype male (
Note: The relative priority of Dexilla cuthbertsoni Curran, 1941 and Dexilla bequaerti Curran, 1941, when the two are treated as synonyms, was established by van
inappendiculata Austen, 1909.—Afrotropical: D.R. Congo, Uganda.
Dexia inappendiculata Austen, 1909: 97. Syntypes, 2 males (BMNH). Type locality: Uganda, Rwenzori Range [as “Ruwenzori”], 7000–8000ft.
Dexia monticola Villeneuve, 1935a: 137. Holotype male (
orphne Curran, 1927.—Afrotropical: Kenya.
Dexia orphne Curran, 1927f: 105. Holotype male (BMNH). Type locality: Kenya, Amboseli National Park [as “Southern Masai Reserve”].
pollinosa Villeneuve, 1943.—Afrotropical: Nigeria, Tanzania.
Dexia pollinosa Villeneuve, 1943b: 94. Syntypes, 2 males (1 male in
rhodesia (Curran, 1941).—Afrotropical: Ghana, Mozambique, Tanzania, Zimbabwe.
Dexilla rhodesia Curran, 1941: 2. Holotype female (
torneutopoda (Speiser, 1914).—Afrotropical: Cameroon, Nigeria.
Dolichodexia torneutopoda Speiser, 1914: 10. Syntypes, 2 males (1 syntype in SDEI,
Dexia venusta Curran, 1927f: 105. Holotype male (SDEI). Type locality: southern Nigeria [as “N. Cameroons”; Northern Cameroons became part of Nigeria in 1961].
uelensis van Emden, 1954.—Afrotropical: D.R. Congo.
Dexia uelensis van Emden, 1954: 551. Holotype male (
uniseta Curran, 1927.—Afrotropical: Kenya, Malawi, South Africa, Tanzania, Uganda.
Dexia uniseta Curran, 1927f: 105. Holotype female (BMNH). Type locality: South Africa, KwaZulu-Natal, Weenen [ca. 28°51′S 30°4′E].
varivittata Curran, 1927.—Afrotropical: Cameroon, Kenya, Tanzania.
Dexia varivittata Curran, 1927f: 106. Holotype male (SDEI). Type locality: Cameroon (not Nigeria as published,
Genus DINERA Robineau-Desvoidy, 1830
DINERA Robineau-Desvoidy, 1830: 307. Type species: Dinera grisea Robineau-Desvoidy, 1830 (= Musca carinifrons Fallén, 1817), by subsequent designation of
PHOROSTOMA Robineau-Desvoidy, 1830: 326. Type species: Phorostoma subrotunda Robineau-Desvoidy, 1830 (= Musca ferina Fallén, 1817), by monotypy [Palaearctic].
MYOCERA Robineau-Desvoidy, 1830: 328. Type species: Myocera longipes Robineau-Desvoidy, 1830 (= Musca ferina Fallén, 1817), by subsequent designation of
MYIOCERA Rondani, 1868b: 597. Unjustified emendation of Myocera Robineau-Desvoidy, 1830 (see
MYOCEROPS Townsend, 1916c: 178. Type species: Musca carinifrons Fallén, 1816, by original designation [Palaearctic].
AFRICODEXIA Townsend, 1933: 462. Type species: Dexia lugens Wiedemann, 1830, by original designation.
Note: We have not determined who, as the First Reviser (Article 24.2.2 of the Code, ICZN 1999), established the relative priority of Dinera Robineau-Desvoidy, 1830, Phorostoma Robineau-Desvoidy, 1830 and Myocera Robineau-Desvoidy, 1830 when the three are treated as synonyms.
femoralis (van Emden, 1947).—Afrotropical: Ethiopia, Kenya.
Paraprosena femoralis van Emden, 1947: 659. Holotype male (BMNH). Type locality: Kenya, Lake Naivasha [as “Lake Naivasha, Masai Reserve”], 6000ft.
fulvotestacea (Villeneuve, 1943).—Afrotropical: South Africa.
Myiocera fulvotestacea Villeneuve, 1943b: 95 (as “fulvo-testacea”). Holotype male (not located). Type locality: South Africa, KwaZulu-Natal, Durban.
latigena (van Emden, 1947).—Afrotropical: Malawi.
Paraprosena latigena van Emden, 1947: 663. Holotype male (BMNH). Type locality: Malawi, plateau on Mt. Mulanje [as “Mlanje Mt.”], 6000–7000ft.
lugens (Wiedemann, 1830).—Afrotropical: Kenya, South Africa, Zimbabwe.
Dexia lugens Wiedemann, 1830: 374. Type(s), male (not located). Type locality: South Africa, Western Cape, Cape of Good Hope [as “Kap”].
palliventris (van Emden, 1947).—Afrotropical: Kenya, Uganda.
Paraprosena palliventris van Emden, 1947: 661. Holotype male (BMNH). Type locality: Uganda, Rwenzori Range [as “Ruwenzori”], Kilembe, 4500ft.
spinosa (Walker, 1858).—Afrotropical: South Africa.
Dexia spinosa Walker, 1858: 204. Type(s), male (BMNH). Type locality: South Africa, KwaZulu-Natal, Durban [as “Port Natal”].
suffulva (Villeneuve, 1943).—Afrotropical: D.R. Congo, Zimbabwe.
Myiocera suffulva Villeneuve, 1943b: 96. Syntypes, 3 males (1 male in
Genus ESTHERIA Robineau-Desvoidy, 1830
ESTHERIA Robineau-Desvoidy, 1830: 305. Type species: Estheria imperatoriae Robineau-Desvoidy, 1830 (= Dexia cristata Meigen, 1826), by subsequent designation of
DEXIMORPHA Rondani, 1856: 84. Type species: Deximorpha marittima Rondani, 1856 (as “Dexia marittima Macq:”) (= Dexia picta Meigen, 1826), by original designation (see
DOLICHODEXIA Brauer & Bergenstamm, 1889: 118 [also 1890: 50]. Type species: Dolichodexia rufipes Brauer & Bergenstamm, 1889 (= Dinera pallicornis Loew, 1873), by original designation [Palaearctic].
DEXIOMERA Curran, 1933: 164. Type species: Dexiomera surda Curran, 1933, by original designation. Syn. n.
buccata (van Emden, 1947).—Afrotropical: Mozambique. Comb. n.
Dexia buccata van Emden, 1947: 633. Holotype female (BMNH). Type locality: Mozambique, Maputo [as “Lorenzo Marques”].
Note:
capensis (Brauer & Bergenstamm, 1891).
Deximorpha capensis Brauer & Bergenstamm, 1891: 417 [also 1891: 113] (as “capensis S. litt. Cap. [Cape of Good Hope]”). Nomen nudum.
Note: Although Deximorpha capensis Brauer & Bergenstamm, 1891 is an unavailable name, there are seven specimens labelled as capensis from “Cap” [= Cape of Good Hope] and “Coll. Winthem” in
notopleuralis (van Emden, 1947).—Afrotropical: South Africa.
Dexiomera notopleuralis van Emden, 1947: 639. Holotype male (BMNH). Type locality: South Africa, KwaZulu-Natal, Willow Grange.
surda (Curran, 1933).—Afrotropical: South Africa. Comb. n.
Dexiomera surda Curran, 1933: 165. Holotype male (formerly in
Note: Dexiomera surda Curran, 1933 is the type species of Dexiomera Curran, 1933.
turneri (van Emden, 1947).—Afrotropical: South Africa.
Dexiomera turneri van Emden, 1947: 638. Holotype male (BMNH). Type locality: South Africa, Eastern Cape, Somerset East.
Genus EUPODODEXIA Villeneuve, 1915
EUPODODEXIA Villeneuve, 1915b: 200. Type species: Eupododexia festiva Villeneuve, 1915, by subsequent designation of
HOMOTRIXODES Townsend, 1926b: 529. Type species: Eupododexia diaphana Villeneuve, 1915, by original designation.
amoena Mesnil, 1976.—Afrotropical: Madagascar.
Eupododexia amoena Mesnil, 1976: 42. Holotype male (
diaphana Villeneuve, 1915.—Afrotropical: Madagascar.
Eupododexia diaphana Villeneuve, 1915b: 202. Holotype male (
festiva Villeneuve, 1915.—Afrotropical: Madagascar.
Eupododexia festiva Villeneuve, 1915b: 201. Lectotype male (
gigantea Mesnil, 1976.—Afrotropical: Madagascar.
Eupododexia gigantea Mesnil, 1976: 41. Holotype male (
picta Mesnil, 1976.—Afrotropical: Madagascar.
Eupododexia picta Mesnil, 1976: 40. Holotype female (
Genus FRONTODEXIA Mesnil, 1976
FRONTODEXIA Mesnil, 1976: 51. Type species: Frontodexia lutea Mesnil, 1976, by original designation.
lutea Mesnil, 1976.—Afrotropical: Madagascar.
Frontodexia lutea Mesnil, 1976: 51. Holotype male (
Genus MESNILOTRIX Cerretti & O’Hara, gen. n
MESNILOTRIX Cerretti & O’Hara, gen. n. Type species: Dexiotrix empiformis Mesnil, 1976, by present designation.
Note: This new genus is described in the New Taxa of Afrotropical Tachinidae section.
empiformis (Mesnil, 1976).—Afrotropical: Madagascar. Comb. n.
Dexiotrix empiformis Mesnil, 1976: 48. Holotype male (
Note: Dexiotrix empiformis Mesnil, 1976 was treated in the genus Dexiotrix Villeneuve, 1936 by
Genus PILIGENA van Emden, 1947
PILIGENA van Emden, 1947: 666. Type species: Piligena mackieae van Emden, 1947, by monotypy.
mackieae van Emden, 1947.—Afrotropical: South Africa, Zimbabwe (new record,
Piligena mackieae van Emden, 1947: 667. Holotype male (BMNH). Type locality: South Africa, Western Cape, Bot River.
Undescribed sp.: South Africa (Limpopo Province) (
Genus PILIGENOIDES Barraclough, 1985
PILIGENOIDES Barraclough, 1985b: 268. Type species: Piligenoides vittata Barraclough, 1985, by original designation.
vittata Barraclough, 1985.—Afrotropical: South Africa.
Piligenoides vittata Barraclough, 1985b: 269. Holotype male (NMDA). Type locality: South Africa, KwaZulu-Natal, St Lucia Nature Reserve.
Genus PLATYDEXIA van Emden, 1954
PLATYDEXIA van Emden, 1954: 550. Type species: Platydexia maynei van Emden, 1954, by original designation.
maynei van Emden, 1954.—Afrotropical: D.R. Congo.
Platydexia maynei van Emden, 1954: 551 (as “maynéi”). Holotype male (
Genus PODODEXIA Brauer & Bergenstamm, 1889
PODODEXIA Brauer & Bergenstamm, 1889: 117 [also 1890: 49]. Type species: Pododexia arachna Brauer & Bergenstamm, 1889, by monotypy.
arachna Brauer & Bergenstamm, 1889.—Afrotropical: Madagascar.
Pododexia arachna Brauer & Bergenstamm, 1889: 117, 166 [also 1890: 49, 98]. Type(s), published as male (7 males and 4 females in
Note: Pododexia arachna Brauer & Bergenstamm, 1889 was described from an unspecified number of males from Madagascar. There are seven males and four females in
hirtipleura Mesnil, 1976.—Afrotropical: Madagascar.
Pododexia hirtipleura Mesnil, 1976: 39. Holotype male (
similis Mesnil, 1976.—Afrotropical: Madagascar.
Pododexia similis Mesnil, 1976: 39. Holotype male (
Genus PRETORIAMYIA Curran, 1927
PRETORIAMYIA Curran, 1927f: 106. Type species: Pretoriamyia munroi Curran, 1927, by original designation.
anacrostichalis van Emden, 1947.—Afrotropical: Kenya.
Pretoriamyia anacrostichalis van Emden, 1947: 653. Holotype female (BMNH). Type locality: Kenya, Mt. Elgon, 8500ft.
munroi Curran, 1927.—Afrotropical: D.R. Congo (new record,
Pretoriamyia munroi Curran, 1927f: 107. Holotype male (
ogilviei van Emden, 1947.—Afrotropical: South Africa.
Pretoriamyia ogilviei van Emden, 1947: 650. Holotype male (BMNH). Type locality: South Africa, Free State, Norvalspont [as “Norvals Pont”], “North Bank Halt” [not located but presumably north of the Orange River in Free State, across the river from Norvalspont in Northern Cape].
plumicornis van Emden, 1947.—Afrotropical: South Africa.
Pretoriamyia plumicornis van Emden, 1947: 651. Holotype female (BMNH). Type locality: South Africa, Eastern Cape, Graaf-Reinet.
sellifera van Emden, 1947.—Afrotropical: South Africa.
Pretoriamyia sellifera van Emden, 1947: 652. Holotype female (BMNH). Type locality: South Africa, Western Cape, Doring [as “Doorn”] River.
somereni van Emden, 1947.—Afrotropical: D.R. Congo (new record,
Pretoriamyia somereni van Emden, 1947: 655. Holotype female (BMNH). Type locality: Uganda, Semliki National Park [as “Bwamba Valley”, ca. 0°49′N 30°3′E].
Genus PROSENA Lepeletier & Serville, 1828
CALIRRHOE Meigen, 1800: 39. Name suppressed by ICZN (1963: 339).
PROSENA Lepeletier & Serville in
siberita (Fabricius, 1775).—Afrotropical: Mozambique. Palaearctic: C. Asia, Europe (all except Turkey), Japan, Mongolia, Pal. China, Russia (W. Russia, W. Siberia, E. Siberia, S. Far East), Transcaucasia. Oriental: India, Indonesia, Malaysia, Myanmar, Nepal, Philippines, Ryukyu Is., Sri Lanka, Taiwan. Australasian: Australia, ?Melanesia. Nearctic: introduced and established in United States.
Stomoxys siberita Fabricius, 1775: 798. Type(s), unspecified sex (
Stomoxys flavipennis Wiedemann, 1819: 20. Lectotype male (
Prosena longirostris Egger, 1860: 798. Syntypes, males and females (
Prosena sybarita Rondani, 1861a: 280. Unjustified emendation of Stomoxys siberita Fabricius, 1775.
Calirrhoe malayana Townsend, 1926c: 25. Lectotype male (
Prosena brevirostris van Emden, 1947: 630. Holotype male (BMNH). Type locality: Mozambique, Maputo [as “Lorenzo Marques”].
sibirita. Incorrect subsequent spelling of siberita Fabricius, 1775 (e.g.,
Note:
There are likely syntypes of Prosena longirostris Egger, 1860 among the specimens of Prosena siberita (Fabricius) in
Genus PROSENOIDES Brauer & Bergenstamm, 1891
PROSENOIDES Brauer & Bergenstamm, 1891: 370 [also 1891: 66]. Type species: Prosenoides papilio Brauer & Bergenstamm, 1891 (as “Prosena papilio S. litt.”) (= Prosena curvirostris Bigot, 1889), by monotypy [Neotropical].
NEOPROSENA Townsend, 1927a: 221. Type species: Neoprosena haustellata Townsend, 1927, by original designation [Neotropical].
PERIPROSENA Villeneuve, 1938c: 14. Type species: Periprosena dispar Villeneuve, 1938, by monotypy.
cytorus (Walker, 1849).—Afrotropical: South Africa, “West Africa”.
Stomyxys cytorus Walker, 1849: 1160 (as “Stomyxys ? cytorus”, with “Stomyxys” as an error for Stomoxys). Type(s), unspecified sex (1 male in BMNH according to BMNH database). Type locality: “West Africa”.
dispar (Villeneuve, 1938).—Afrotropical: D.R. Congo.
Periprosena dispar Villeneuve, 1938c: 14. Holotype female (
longilingua (Villeneuve, 1943).—Afrotropical: D.R. Congo.
Myiocera longilingua Villeneuve, 1943b: 95. Holotype male (not located). Type locality: D.R. Congo, Nord-Kivu, Kibati [ca. 1°36′S 29°16′E].
Note:
tenuipes (van Emden, 1947).—Afrotropical: Uganda.
Paraprosena tenuipes van Emden, 1947: 665. Holotype male (BMNH). Type locality: Uganda, Rwenzori Range [as “Ruwenzori”], Namwamba Valley, 6500ft.
Genus PSEUDODINERA Brauer & Bergenstamm, 1891
PSEUDODINERA Brauer & Bergenstamm, 1891: 378 [also 1891: 74]. Type species: Pseudodinera nigripes Brauer & Bergenstamm, 1891, by monotypy.
nigripes Brauer & Bergenstamm, 1891.—Afrotropical: South Africa.
Pseudodinera nigripes Brauer & Bergenstamm, 1891: 379 [also 1891: 75] (as “nigripes Wd. Coll. Winth. litt.”). Type(s), male (2 males in
Note: Pseudodinera nigripes Brauer & Bergenstamm, 1891 was described from an unspecified number of males. There are two male syntypes in
spinigera (Thomson, 1869).—Afrotropical: South Africa.
Dinera spinigera Thomson, 1869: 531. Type(s), male (
Genus ZELIOMIMA Mesnil, 1976
ZELIOMIMA Mesnil, 1976: 37. Type species: Zeliomima caudata Mesnil, 1976, by original designation.
caudata Mesnil, 1976.—Afrotropical: Madagascar.
Zeliomima caudata Mesnil, 1976: 39. Holotype male (
chaetosa Mesnil, 1976.—Afrotropical: Madagascar.
Zeliomima chaetosa Mesnil, 1976: 39. Holotype male (
Genus ZEUXIOTRIX Mesnil, 1976
ZEUXIOTRIX Mesnil, 1976: 46. Type species: Zeuxiotrix atra Mesnil, 1976, by original designation.
atra Mesnil, 1976.—Afrotropical: Madagascar.
Zeuxiotrix atra Mesnil, 1976: 48. Holotype male (
cinerosa Mesnil, 1976.—Afrotropical: Madagascar.
Zeuxiotrix cinerosa Mesnil, 1976: 47. Holotype male (
Unplaced species of Dexiini
brunnicornis Macquart, 1844.—Afrotropical: Réunion.
Dexia brunnicornis Macquart, 1844: 86 [also 1844: 243]. Lectotype male (
crassipalpis Mesnil, 1950.—Afrotropical: Zimbabwe.
Dinera crassipalpis Mesnil, 1950d: 115. Syntypes, 3 females (not located). Type locality: Zimbabwe, Hurungwe [as “Urungwe”], Gota Gota.
Tribe DUFOURIINI
Genus CHETOPTILIA Rondani, 1862
CHETOPTILIA Rondani, 1862: 166. Type species: Ptilops puella Rondani, 1862, by monotypy [Palaearctic].
CHAETOPTILIA Bezzi & Stein, 1907: 402. Unjustified emendation of Chetoptilia Rondani, 1862 (see
PARAPTILOPS Mesnil, 1975a: 1358. Type species: Chaetoptilia angustifrons Mesnil, 1953, by original designation [Oriental].
cyanea Mesnil, 1968.—Afrotropical: Madagascar.
Chaetoptilia cyanea Mesnil, 1968a: 53. Holotype male (BMNH). Type locality: Madagascar, Toamasina, Toamasina [as “Tamatave”].
metallica Mesnil, 1968.—Afrotropical: Madagascar.
Chaetoptilia metallica Mesnil, 1968a: 54. Holotype male (
plumicornis Villeneuve, 1942.—Afrotropical: Uganda.
Chaetoptilia plumicornis Villeneuve, 1942a: 53. Holotype male (not located). Type locality: Uganda, Kampala.
Genus MESNILANA van Emden, 1945
MESNILANA van Emden, 1945: 413. Type species: Mesnilana bevisi van Emden, 1945, by monotypy.
Note: We follow van
bevisi van Emden, 1945.—Afrotropical: South Africa.
Mesnilana bevisi van Emden, 1945: 414. Holotype female (BMNH). Type locality: South Africa, KwaZulu-Natal, Greenwood Park [suburb of Durban].
Genus PANDELLEIA Villeneuve, 1907
PANDELLEIA Villeneuve, 1907: 392. Type species: Etheria sexpunctata Pandellé, 1896, by monotypy [Palaearctic].
AFROPHASIA Curran, 1939: 1. Type species: Afrophasia dimorphia Curran, 1939, by original designation.
dimorphia (Curran, 1939).—Afrotropical: Burundi, D.R. Congo, Kenya, Lesotho, South Africa, Tanzania, Uganda.
Afrophasia dimorphia Curran, 1939: 1. Holotype male (
Pandelleia francoisi Mesnil, 1952a: 2 (as “françoisi”). Holotype male (
translucens (Mesnil, 1959).—Afrotropical: Tanzania.
Rondania translucens Mesnil, 1959: 27. Holotype female (
Genus RHINOPHOROIDES Barraclough, 2005
RHINOPHOROIDES Barraclough, 2005: 381. Type species: Rhinophoroides minutus Barraclough, 2005, by original designation.
Note: Rhinophoroides Barraclough, 2005 is possibly a junior synonym of Mesnilana van Emden, 1945.
minutus Barraclough, 2005.—Afrotropical: South Africa.
Rhinophoroides minutus Barraclough, 2005: 382. Holotype male (NMDA). Type locality: South Africa, KwaZulu-Natal, Merrivale, Tshwalabenyoni, 1000m (29°31′S 20°15′E).
Tribe EUTHERINI
Genus EUTHERA Loew, 1866
EUTHERA Loew, 1866: 46, 47. Type species: Euthera tentatrix Loew, 1866, by monotypy [Nearctic].
EUTHEROPSIS Townsend, 1916c: 178. Type species: Euthera mannii Mik, 1889 (= Ocyptera fascipennis Loew, 1854), by original designation.
PREUTHERA Townsend, 1933: 452. Type species: Euthera (Eutheropsis) peringueyi Bezzi, 1925, by original designation [Oriental].
fascipennis (Loew, 1854).—Afrotropical: Malawi, Tanzania, Yemen. Palaearctic: C. Asia, Europe (SW. Eur., SC. Eur., SE. Eur., Turkey). Oriental: India, Taiwan.
Ocyptera fascipennis Loew, 1854: 20. Type(s), male (1 male in ZMHB). Type locality: Greece, Crete [or Kriti], Heraklion [as “Candia”].
Euthera mannii Mik, 1889: 132. Lectotype female (
Euthera burtti van Emden, 1960: 383. Holotype male (BMNH). Type locality: Tanzania, Old Shinyanga.
manni. Incorrect subsequent spelling of mannii Mik, 1889 (e.g.,
peringueyi Bezzi, 1925.—Not Afrotropical [Oriental].
Euthera (Eutheropsis) peringueyi Bezzi, 1925a: 280 (as “péringueyi”).
Note:
tuckeri Bezzi, 1925.—Afrotropical: Botswana (new record, NMDA [PC]), Ghana, Kenya (new record,
Euthera (Eutheropsis) tuckeri Bezzi, 1925a: 279. Holotype male (
Tribe VORIINI
Genus ALLOTHELAIRA Villeneuve, 1915
ALLOTHELAIRA Villeneuve, 1915c: 226. Type species: Allothelaira diaphana Villeneuve, 1915, by monotypy.
diaphana Villeneuve, 1915.—Afrotropical: Cameroon, D.R. Congo, Ghana, Nigeria, Sierra Leone, Tanzania.
Allothelaira diaphana Villeneuve, 1915c: 226. Lectotype male (BMNH), by designation of van
Note: Allothelaira diaphana Villeneuve, 1915 was described from five males and two females, including two males from Aburi (Ghana).
Genus CAMPYLOCHETA Rondani, 1859
CAMPYLOCHETA Rondani, 1859: 157, 169. Type species: Tachina praecox Meigen, 1824, by fixation of
ELPE Robineau-Desvoidy, 1863a: 488. Type species: Tachina inepta Meigen, 1824, by original designation [Palaearctic].
MYXACTIA Villeneuve, 1915b: 197. Type species: Myxactia inclinata Villeneuve, 1915, by monotypy.
CAMPYLOCHAETA Bezzi & Stein, 1907: 305. Unjustified emendation of Campylocheta Rondani, 1859 (see
CHAETOPHLEPSIS Townsend, 1915b: 422. Type species: Chaetophlepsis tarsalis Townsend, 1915, by original designation [Neotropical].
inclinata (Villeneuve, 1915).—Afrotropical: Madagascar.
Myxactia inclinata Villeneuve, 1915b: 197. Holotype male (
Note:
keiseri Mesnil, 1978.—Afrotropical: Madagascar.
Campylochaeta keiseri Mesnil, 1978b: 284. Holotype male (
plumbea (Mesnil, 1952).—Afrotropical: D.R. Congo, Rwanda (new record,
Frivaldszkia plumbea Mesnil, 1952a: 8. Holotype male (not located). Type locality: D.R. Congo, Nord-Kivu, Bweza, Tshamugussa, 2250m [ca. 1°20′S 29°31′E].
risbeci (Mesnil, 1944).—Afrotropical: Mali, Nigeria, Senegal, Uganda.
Frivaldzkia risbeci Mesnil, 1944: 16. Type(s), unspecified sex (
vansomereni van Emden, 1960.—Afrotropical: Kenya.
Campylochaeta vansomereni van Emden, 1960: 352. Holotype male (BMNH). Type locality: Kenya, Meru.
Genus CYRTOPHLOEBA Rondani, 1856
CYRTOPHLOEBA Rondani, 1856: 207. Type species: Tachina ruricola Meigen, 1824, by original designation [Palaearctic].
CYRTHOPLAEBA Rondani, 1857: 13. Unjustified emendation of Cyrtophloeba Rondani, 1856 (see
STACKELBERGULA Richter, 1967: 478. Type species: Stackelbergula eremophila Richter, 1967, by original designation.
CYRTHOPHLAEBA. Incorrect subsequent spelling of Cyrtophloeba Rondani, 1856 (
CYRTHOPHLEBA. Incorrect subsequent spelling of Cyrtophloeba Rondani, 1856 (
CYRTOPHLEBA. Incorrect original spelling of Cyrtophloeba Rondani, 1856 (
arabica Zeegers, 2007.—Afrotropical: Yemen.
Cyrtophleba (Stackelbergula) arabica Zeegers, 2007: 374. Holotype male (
eremophila (Richter, 1967).—Afrotropical: U.A. Emirates. Palaearctic: C. Asia, Mongolia.
Stackelbergula eremophila Richter, 1967: 479. Holotype male (
Undescribed spp.: Kenya (
Genus HYLEORUS Aldrich, 1926
HYLEORUS Aldrich, 1926a: 16. Type species: Hyleorus furcatus Aldrich, 1926, by monotypy [Australasian].
STEINIOMYIA Townsend, 1932: 54. Type species: Plagia elata Meigen, 1838, by monotypy [Palaearctic].
NEUROPLAGIA Townsend, 1933: 479. Type species: Plagia elata nudinerva Villeneuve, 1920, by original designation.
AFROPLAGIA Curran, 1938: 6. Type species: Afroplagia fasciata Curran, 1938, by original designation.
fasciatus (Curran, 1938).—Afrotropical: Ghana, South Africa, Uganda.
Afroplagia fasciata Curran, 1938: 6. Holotype male (
nudinerva (Villeneuve, 1920).—Afrotropical: Yemen. Palaearctic: Europe (SW. Eur.), M. East (Israel).
Plagia elata nudinerva Villeneuve, 1920b: 200. Holotype, unspecified sex [female, examined by PC] (
Genus HYSTRICOVORIA Townsend, 1928
HYSTRICOVORIA Townsend, 1928: 395. Type species: Hystricovoria bakeri Townsend, 1928, by original designation.
AFROVORIA Curran, 1938: 5. Type species: Afrovoria munroi Curran, 1938 (= Hystricovoria bakeri Townsend, 1928), by original designation.
ANAVORIA Mesnil, 1953b: 170 (as subgenus of Voria Robineau-Desvoidy, 1830). Type species: Voria (Anavoria) indica Mesnil, 1953 (= Hystricovoria bakeri Townsend, 1928), by monotypy.
bakeri Townsend, 1928.—Afrotropical: Botswana, Ghana, Kenya, South Africa, Yemen. Australasian: ?Australia. Oriental: India, Orien. China, Philippines.
Hystricovoria bakeri Townsend, 1928: 395. Holotype male (
Afrovoria munroi Curran, 1938: 6. Holotype male (
Voria (Anavoria) indica Mesnil, 1953b: 170. Holotype female (BMNH). Type locality: India, Uttarakhand, Dehra Dun.
Genus NARDIA Cerretti, 2009
NARDIA Cerretti, 2009a: 108. Type species: Plagiomima rufolateralis Crosskey, 1984, by original designation.
rufolateralis (Crosskey, 1984).—Afrotropical: Botswana, Namibia.
Plagiomima rufolateralis Crosskey, 1984: 302. Holotype male (BMNH). Type locality: Botswana, South-East, Sebele [as “Bakgatla, Sebele”; 24°34′S 25°58′E according to
tsavo Cerretti, 2009.—Afrotropical: Kenya.
Nardia tsavo Cerretti, 2009a: 114. Holotype female (
Genus PERISCEPSIA Gistel, 1848
SCOPOLIA Robineau-Desvoidy, 1830: 268 (junior homonym of Scopolia Hübner, 1825). Type species: Musca carbonaria Panzer, 1798, by subsequent designation of
PERISCEPSIA Gistel, 1848: x (unnecessary nomen novum for Scopolia Robineau-Desvoidy, 1830) (see
PHORICHETA Rondani, 1861b: 8 (nomen novum for Scopolia Robineau-Desvoidy, 1830).
RAMONDA Robineau-Desvoidy, 1863a: 790. Type species: Ramonda fasciata Robineau-Desvoidy, 1863 (= Tachina spathulata Fallén, 1820), by original designation [Palaearctic].
PHORICHAETA Brauer & Bergenstamm, 1889: 106 [also 1890: 38]. Unjustified emendation of Phoricheta Rondani, 1861 (see
WAGNERIA of authors (e.g.,
Note: Subgenera of Periscepsia Gistel, 1848 are not recognized here because the subgeneric placements of the Afrotropical species require more study.
abbreviata (Mesnil, 1950).—Afrotropical: D.R. Congo. Status n.
Wagneria rufitibia abbreviata Mesnil, 1950a: 1. Holotype, unspecified sex [male, examined by PC] (
Note: Wagneria rufitibia abbreviata Mesnil, 1950 was treated as a synonym of Wagneria rufitibia Villeneuve, 1938 by
amicula (Mesnil, 1950).—Afrotropical: D.R. Congo, South Africa.
Wagneria amicula Mesnil, 1950a: 1. Holotype male (
canina (Mesnil, 1950).—Afrotropical: D.R. Congo, Ethiopia, Rwanda, South Africa.
Wagneria canina Mesnil, 1950a: 2. Holotype, unspecified sex (
carbonaria (Panzer, 1798).—Afrotropical: “widespread n.-e. Afr. to sthn Afr.” (
Musca carbonaria Panzer, 1798: 15 (and coloured figure on unnumbered facing plate). Type(s), unspecified sex [sex cannot be determined from the figure] (lost). Type locality: Austria (
Dexia nigrans Meigen, 1826: 40. Syntypes, published as females (male(s) in
Note: Periscepsia carbonaria (Panzer, 1798) of current authors is likely a species complex but is treated here as a single species pending further study.
caviceps (van Emden, 1960).—Afrotropical: Zimbabwe.
Wagneria caviceps van Emden, 1960: 336. Holotype male (BMNH). Type locality: Zimbabwe, Harare [as “Salisbury”].
decolor (van Emden, 1960).—Afrotropical: Ethiopia, Kenya, South Africa, Uganda.
Wagneria decolor van Emden, 1960: 347. Holotype male (BMNH). Type locality: Uganda, Nyakasura [ca. 0°40′N 30°13′E].
fratella (Villeneuve, 1938).—Afrotropical: D.R. Congo, Kenya, Uganda.
Wagneria fratella Villeneuve, 1938a: 5. Holotype, unspecified sex (
glossinicornis (van Emden, 1960).—Afrotropical: Kenya, South Africa.
Wagneria glossinicornis van Emden, 1960: 337. Holotype male (BMNH). Type locality: Kenya, Chyulu Hills, 6000ft.
guttipennis (van Emden, 1960).—Afrotropical: Kenya.
Wagneria guttipennis van Emden, 1960: 345. Holotype male (BMNH). Type locality: Kenya, Naivasha.
kirbyiformis (van Emden, 1960).—Afrotropical: D.R. Congo.
Wagneria kirbyiformis van Emden, 1960: 344. Holotype male (
lindneri (Mesnil, 1959).—Afrotropical: Tanzania.
Wagneria lindneri Mesnil, 1959: 25. Holotype male (
natalica (van Emden, 1960).—Afrotropical: Ethiopia, Kenya, South Africa.
Wagneria natalica van Emden, 1960: 339. Holotype male (BMNH). Type locality: South Africa, KwaZulu-Natal, “Winzinto River” [not located].
Wagneria laniventris van Emden, 1960: 339. Holotype male (BMNH). Type locality: Kenya, Ngong.
Wagneria nubilipennis van Emden, 1960: 341. Holotype female (BMNH). Type locality: Kenya, Meru.
Wagneria z-fuscum van Emden, 1960: 340. Holotype female (BMNH). Type locality: South Africa, KwaZulu-Natal, Weenen [ca. 28°51′S 30°4′E].
Note:
nudinerva (Mesnil, 1950).—Afrotropical: D.R. Congo. Status n.
Wagneria rufitibia nudinerva Mesnil, 1950a: 1. Holotype, unspecified sex [female, examined by PC] (
Note: Wagneria rufitibia nudinerva Mesnil, 1950 was treated as a synonym of Wagneria rufitibia Villeneuve, 1938 by
pallidipennis (van Emden, 1960).—Afrotropical: D.R. Congo, Kenya.
Wagneria pallidipennis van Emden, 1960: 349. Holotype male (BMNH). Type locality: Kenya, Naivasha.
propleuralis (van Emden, 1960).—Afrotropical: South Africa, Uganda.
Wagneria propleuralis van Emden, 1960: 343. Holotype female (BMNH). Type locality: Uganda, Semliki National Park [as “Bwamba Valley, Ruwenzori”, ca. 0°49′N 30°3′E].
rufitibia (Villeneuve, 1938).—Afrotropical: D.R. Congo, Kenya, South Africa, Tanzania, Uganda.
Wagneria rufitibia Villeneuve, 1938a: 4. Holotype, unspecified sex [male, see van
salti (van Emden, 1960).—Afrotropical: Tanzania.
Wagneria salti van Emden, 1960: 348. Holotype male (BMNH). Type locality: Tanzania, Mt. Kilimanjaro, Shira Plateau, 12,450ft [ca. 3°0′S 37°14′E].
vidua (Mesnil, 1950).—Afrotropical: Kenya, Rwanda, Uganda.
Wagneria vidua Mesnil, 1950a: 3. Holotype, unspecified sex (
Genus PROSHELIOMYIA Brauer & Bergenstamm, 1891
PROSHELIOMYIA Brauer & Bergenstamm, 1891: 375 [also 1891: 71]. Type species: Prosheliomyia nietneri Brauer & Bergenstamm, 1891, by monotypy [Oriental].
Subgenus THRIXIONELLUS Mesnil, 1968
THRIXIONELLUS Mesnil, 1968a: 45 (as subgenus of Prosheliomyia Brauer & Bergenstamm, 1891). Type species: Prosheliomyia (Thrixionellus) mirabilis Mesnil, 1968, by original designation.
mirabilis Mesnil, 1968.—Afrotropical: Madagascar.
Prosheliomyia (Thrixionellus) mirabilis Mesnil, 1968a: 45. Holotype male (
nigricornis Mesnil, 1968.—Afrotropical: Madagascar.
Prosheliomyia (Thrixionellus) nigricornis Mesnil, 1968a: 47. Holotype male (
pallida Mesnil, 1968.—Afrotropical: Madagascar.
Prosheliomyia (Thrixionellus) pallida Mesnil, 1968a: 48. Holotype male (
Genus REICHARDIA Karsch, 1886
REICHARDIA Karsch, 1886a: 137. Type species: Reichardia insignis Karsch, 1886, by monotypy.
insignis Karsch, 1886.—Afrotropical: Tanzania.
Reichardia insignis Karsch, 1886a: 137. Type(s), unspecified sex (1 male in ZMHB). Type locality: Tanzania, east of Lake Tanganyika, “Kawende” [not located].
Undescribed sp.: Ethiopia (
Genus STOMINA Robineau-Desvoidy, 1830
STOMINA Robineau-Desvoidy, 1830: 411. Type species: Stomina rubricornis Robineau-Desvoidy, 1830 (= Musca tachinoides Fallén, 1817), by monotypy [Palaearctic].
Undetermined sp(p).—Afrotropical: Malawi (
Note: Undetermined specimens of this genus were recorded from the Afrotropical Region by
Genus SUBFISCHERIA Villeneuve, 1937
SUBFISCHERIA Villeneuve, 1937a: 210. Type species: Subfischeria flavogrisea Villeneuve, 1937, by monotypy.
flavogrisea Villeneuve, 1937.—Afrotropical: Botswana, Malawi, Namibia, South Africa.
Subfischeria flavogrisea Villeneuve, 1937a: 211 (as “flavo-grisea”). Holotype female (
Genus THELAIRA Robineau-Desvoidy, 1830
THELAIRA Robineau-Desvoidy, 1830: 214 (as “Thelaïra”). Type species: Thelaira abdominalis Robineau-Desvoidy, 1830 (= Musca solivagus Harris, 1780), by subsequent designation of
THELAIRIA. Incorrect subsequent spelling of Thelaira Robineau-Desvoidy, 1830 (
altoplani Speiser, 1914.—Afrotropical: Angola, Cameroon, D.R. Congo, Eritrea, Ghana, Lesotho, Madagascar, Malawi, Mozambique, Nigeria, Sierra Leone, South Africa, Sudan, Tanzania, Uganda, Zimbabwe.
Thelaira altoplani Speiser, 1914: 12. Holotype male (not located). Type locality: Cameroon, Dschang.
Thelaira palliventris Curran, 1928b: 378. Holotype male (
Musca nigripes of authors (e.g.,
aurofasciata van Emden, 1960.—Afrotropical: Ghana, Nigeria.
Thelaira aurofasciata van Emden, 1960: 374. Holotype male (BMNH). Type locality: Ghana, Obuasi, Ashanti.
luteiventris van Emden, 1960.—Afrotropical: Nigeria, Sudan.
Thelaira luteiventris van Emden, 1960: 376. Holotype male (BMNH). Type locality: Nigeria, Azare.
madecassa Mesnil, 1978.—Afrotropical: Madagascar.
Thelaira madecassa Mesnil, 1978b: 285. Holotype male (
Genus VORIA Robineau-Desvoidy, 1830
VORIA Robineau-Desvoidy, 1830: 195. Type species: Voria latifrons Robineau-Desvoidy, 1830 (= Tachina ruralis Fallén, 1810), by monotypy [Palaearctic].
PLAGIA Meigen, 1838: 201. Type species: Tachina verticalis Meigen, 1824 (= Tachina ruralis Fallén, 1810), by subsequent designation of
capensis Villeneuve, 1935.—Afrotropical: Ghana, Kenya, Mozambique (new record,
Plagia setosa Brauer & Bergenstamm, 1891: 409, 439 [also 1891: 105, 135] (as “setosa Wd. litt. Cap. [Cape of Good Hope]”). Nomen nudum.
Voria capensis Villeneuve, 1935a: 138. Holotype male (not located). Type locality: South Africa.
ruralis (Fallén, 1810).—Afrotropical: “Kenya to South Africa, South Yemen [part of present-day Yemen]” (
Tachina ruralis Fallén, 1810: 265. Lectotype male (
Subfamily EXORISTINAE (Fig. 4 )
Tribe ACEMYINI
Genus ACEMYA Robineau-Desvoidy, 1830
ACEMYA Robineau-Desvoidy, 1830: 202. Type species: Acemya oblonga Robineau-Desvoidy, 1830 (= Tachina acuticornis Meigen, 1824), by subsequent designation of Desmarest in d’
ACOMYIA Agassiz, 1846b: 3, 5. Unjustified emendation of Acemya Robineau-Desvoidy, 1830 (see
ACEMYIA Schiner, 1861: 472. Unjustified emendation of Acemya Robineau-Desvoidy, 1830 (see
fishelsoni Kugler, 1968.—Afrotropical: Yemen. Palaearctic: M. East (Israel), Mongolia, Pal. China.
Acemyia fishelsoni Kugler, 1968: 65. Holotype female (
pyrrhocera Villeneuve, 1922.—Afrotropical: U.A. Emirates. Palaearctic: C. Asia, Europe (W. Eur., SW. Eur., SC. Eur.), Mongolia, Russia (E. Siberia), Transcaucasia.
Acomyia pyrrhocera Villeneuve, 1922c: 342. Syntypes, 1 male and 2 females (not located). Type localities: France, Digne-les-Bains [as “Digne”] and “sud de la France”.
Genus ATLANTOMYIA Crosskey, 1977
ATLANTOMYIA Crosskey, 1977: 145. Type species: Atlantomyia nitida Crosskey, 1977, by original designation.
nitida Crosskey, 1977.—Afrotropical: Saint Helena.
Atlantomyia nitida Crosskey, 1977: 147. Holotype male (
Genus CERACIA Rondani, 1865
CERACIA Rondani, 1865: 221. Type species: Ceracia mucronifera Rondani, 1865, by monotypy [Palaearctic].
MYOTHYRIA van der Wulp, 1890: 208. Type species: Myothyria majorina van der Wulp, 1890, by subsequent designation of
MYIOTHYRIA. Incorrect subsequent spelling of Myothyria van der Wulp, 1890 (e.g.,
Note:
africana (Mesnil, 1959).—Afrotropical: Nigeria, South Africa, Tanzania, Uganda.
Myothyria africana Mesnil, 1959: 19. Holotype male (
Ceracia burtti van Emden, 1960: 370. Holotype female (BMNH). Type locality: Tanzania, Old Shinyanga.
freyi (Herting, 1958).—Afrotropical: Cape Verde.
Myiothyria freyi Herting, 1958: 4. Holotype male (FMNHH). Type locality: Cape Verde Islands, São Nicolau, Ribeira da Pulga [as “S. Nicolau: Rib. Pulga”].
mucronifera Rondani, 1865.—Afrotropical: Yemen. Palaearctic: C. Asia, Europe (W. Eur., SW. Eur., SC. Eur., SE. Eur.), M. East (Israel), N. Africa (Canary Is., NW. Africa), Transcaucasia. Oriental: Orien. China [Hunan].
Ceracia mucronifera Rondani, 1865: 222. Syntypes, 2 males (MZF). Type locality: Italy, Apennines, near Parma.
murina Mesnil, 1977.—Afrotropical: Madagascar.
Ceracia murina Mesnil, 1977d: 326. Holotype female (
Genus METACEMYIA Herting, 1969
METACEMYIA Herting, 1969: 197. Type species: Acemyia calloti Séguy, 1936, by original designation.
CERACIA of
aartseni Zeegers, 2007.—Afrotropical: U.A. Emirates, Yemen. Palaearctic: M. East (Israel).
Metacemyia aartseni Zeegers, 2007: 388. Holotype female (
calloti (Séguy, 1936).—Afrotropical: Senegal, Tanzania, U.A. Emirates, Yemen, Zambia, Zimbabwe. Palaearctic: Europe (W. Eur., SW. Eur., SC. Eur., Turkey), M. East (Israel), N. Africa (NW. Africa)
Acemyia calloti Séguy, 1936: 324. Holotype female (not located). Type locality: Tunisia, El Aouina.
Ceracia nomadacridis van Emden, 1960: 369. Holotype male (BMNH). Type locality: Tanzania, Rukwa District, Nkamba-Kati.
Ceracia mucronifera of authors (e.g.,
setosa Crosskey, 1973.—Afrotropical: Malawi.
Metacemyia setosa Crosskey, 1973a: 376. Holotype male (BMNH). Type locality: Malawi, Southern Region, Chambe Plateau.
uncinata (Thomson, 1869).—Afrotropical: Botswana, D.R. Congo, South Africa.
Myobia uncinata Thomson, 1869: 526. Lectotype male (
Note: A record of Myobia uncinata Thomson, 1869 from Israel by
Tribe ANACAMPTOMYIINI
Genus ANACAMPTOMYIA Bischof, 1904
ANACAMPTOMYIA Bischof, 1904: 79. Type species: Anacamptomyia africana Bischof, 1904, by monotypy.
ROUBAUDIA Villeneuve, 1910a: 249. Type species: Roubaudia rufescens Villeneuve, 1910, by monotypy (not by original designation as given by
PARAROUBAUDIA Roubaud & Villeneuve, 1914: 122, 124 (as subgenus of Roubaudia Villeneuve, 1910). Type species: Roubaudia (Pararoubaudia) bisetosa Roubaud & Villeneuve, 1914, by monotypy.
Note: A key to the African species of Anacamptomyia Bischof, 1904 was published by
africana Bischof, 1904.—Afrotropical: D.R. Congo, Kenya, ?Madagascar, Mozambique, Nigeria, Senegal, South Africa, Tanzania.
Anacamptomyia africana Bischof, 1904: 81. Lectotype female (
aurifrons Zeegers, 2014.—Afrotropical: Madagascar.
Anacamptomyia aurifrons Zeegers, 2014: 97. Holotype male (
bisetosa (Roubaud & Villeneuve, 1914).—Afrotropical: Benin, Cameroon, D.R. Congo, Ghana, Nigeria, Senegal, Sierra Leone, Zimbabwe.
Roubaudia (Pararoubaudia) bisetosa Roubaud & Villeneuve, 1914: 125. Syntypes, males and females (1 female in
Note: Roubaudia bisetosa Roubaud & Villeneuve, 1914 was described from an unspecified number of males and females. The only specific locality mentioned was Dakar (the locality where the syntype in
blommersi Zeegers, 2014.—Afrotropical: Madagascar.
Anacamptomyia blommersi Zeegers, 2014: 99. Holotype male (
gymnops Zeegers, 2007.—Afrotropical: Yemen.
Anacamptomyia gymnops Zeegers, 2007: 376. Holotype female (
obscurella Mesnil, 1950.—Afrotropical: “toute Afrique tropicale et australe” (
Anacamptomyia pallida obscurella Mesnil, 1950b: 24. Syntypes, males and females (1 male and possibly other unrecognized syntypes in
pallida (Roubaud & Villeneuve, 1914).—Afrotropical: Benin, Cameroon, D.R. Congo, Ghana, Malawi, Nigeria, Senegal, Sudan, Tanzania, Zambia, Zimbabwe.
Roubaudia rufescens pallida Roubaud & Villeneuve, 1914: 124. Syntypes, only the male sex specifically mentioned (2 females in
Note: A male in
pruinosa (Roubaud & Villeneuve, 1914).—Afrotropical: Nigeria, Senegal, Uganda, Zimbabwe.
Roubaudia pruinosa Roubaud & Villeneuve, 1914: 123. Syntypes, male(s) and female(s) (1 female in
rufescens (Villeneuve, 1910).—Afrotropical: Benin, Nigeria.
Roubaudia rufescens Villeneuve, 1910a: 249. Lectotype male (
Genus LEUCOCARCELIA Villeneuve, 1921
LEUCOCARCELIA Villeneuve, 1921: 30. Type species: Leucocarcelia argyrata Villeneuve, 1921, by monotypy.
argyrata Villeneuve, 1921.—Afrotropical: Malawi.
Leucocarcelia argyrata Villeneuve, 1921: 30. Holotype male (BMNH). Type locality: Malawi, Mt. Mulanje [as “Mont Mlanje”].
Undescribed spp.: D.R. Congo (
Genus PARAPALES Mesnil, 1950
PARAPALES Mesnil, 1949b: 102 (as subgenus of Ctenophorocera Brauer & Bergenstamm, 1891). Nomen nudum (proposed after 1930 without designation of type species; no included species) (see
PARAPALES Mesnil, 1950c: 122 (as subgenus of Ctenophorocera Brauer & Bergenstamm, 1891). Type species: Ctenophorocera (Parapales) pallidula Mesnil, 1950, by original designation (see
brevicornis Mesnil, 1977.—Afrotropical: Madagascar.
Parapales brevicornis Mesnil, 1977b: 192. Holotype male (
brunnea Mesnil, 1977.—Afrotropical: Madagascar.
Parapales brunnea Mesnil, 1977b: 192. Holotype female (
luteicornis Mesnil, 1977.—Afrotropical: Madagascar.
Parapales luteicornis Mesnil, 1977b: 192. Holotype female (
micronychia Mesnil, 1977.—Afrotropical: Madagascar.
Parapales micronychia Mesnil, 1977b: 191. Holotype male (
pallidula (Mesnil, 1950).—Afrotropical: Madagascar.
Ctenophorocera (Parapales) pallidula Mesnil, 1950c: 123. Holotype male (
pectinipes Mesnil, 1977.—Afrotropical: Madagascar.
Parapales pectinipes Mesnil, 1977b: 192. Holotype male (
Tribe BLONDELIINI
Genus AFROLIXA Curran, 1939
AFROLIXA Curran, 1939: 4. Type species: Afrolixa macula Curran, 1939, by original designation.
macula Curran, 1939.—Afrotropical: Malawi, Mozambique, South Africa.
Afrolixa macula Curran, 1939: 4. Holotype male (
Undescribed sp.: Côte d’Ivoire, Sudan, Uganda (BMNH,
Genus ANOMALOSTOMYIA Cerretti & Barraclough, 2007
ANOMALOSTOMYIA Cerretti & Barraclough, 2007: 102. Type species: Anomalostomyia namibica Cerretti & Barraclough, 2007, by original designation.
Note:
namibica Cerretti & Barraclough, 2007.—Afrotropical: Namibia.
Anomalostomyia namibica Cerretti & Barraclough, 2007: 103. Holotype male (
Genus BLONDELIA Robineau-Desvoidy, 1830
BLONDELIA Robineau-Desvoidy, 1830: 122. Type species: Blondelia nitida Robineau-Desvoidy, 1830 (= Tachina nigripes Fallén, 1810), by subsequent designation of Duponchel in d’
tibialis Mesnil, 1962.—Afrotropical: Burundi (new record,
Blondelia tibialis Mesnil, 1962: 753. Holotype male (
Genus CHARITELLA Mesnil, 1957
CHARITELLA Mesnil, 1957: 31. Type species: Charitella gracilis Mesnil, 1957, by monotypy [Oriental].
METADRINOMYIA Shima, 1980: 259. Type species: Metadrinomyia proclinata Shima, 1980, by original designation [Palaearctic]. Syn. n.
Note: Metadrinomyia Shima, 1980 was first recognized from the Afrotropical Region by
nigrescens Mesnil, 1977.—Afrotropical: ?Madagascar, Malawi.
Charitella nigrescens Mesnil, 1977d: 325. Holotype female (
whitmorei (Cerretti, 2012).—Afrotropical: Burundi, D.R. Congo. Comb. n.
Metadrinomyia whitmorei Cerretti, 2012: 325. Holotype male (
Note: The recently described Metadrinomyia whitmorei Cerretti, 2012 is moved here to Charitella Mesnil, 1957.
Undescribed sp. 1: Madagascar (
Undescribed sp. 2: Comoros (
Genus COMPSILURA Bouché, 1834
COMPSILURA Bouché, 1834: 58. Type species: Tachina concinnata Meigen, 1824, by subsequent designation of
concinnata (Meigen, 1824).—Afrotropical: “widespread W. Afr. n.-e. Afr., E. Afr. & sthn Afr.” (
Tachina concinnata Meigen, 1824: 412. Holotype female (
Phorocera selecta Curran, 1940: 6. Holotype male (
solitaria (Curran, 1940).—Afrotropical: Zimbabwe.
Phorocera solitaria Curran, 1940: 6. Holotype male (
Undescribed sp.: Madagascar (
Undetermined sp.: Burundi (
Genus DOLICHOTARSINA Mesnil, 1977
DOLICHOTARSINA Mesnil, 1977d: 324. Type species: Dolichotarsina gracilis Mesnil, 1977, by original designation.
gracilis Mesnil, 1977.—Afrotropical: Madagascar.
Dolichotarsina gracilis Mesnil, 1977d: 325. Holotype female (
Genus EOMEDINA Mesnil, 1960
EOMEDINA Mesnil, 1960b: 652. Type species: Eomedina grisescens Mesnil, 1960 (= Degeeria apicalis Curran, 1927), by original designation.
Note: See
apicalis (Curran, 1927).—Afrotropical: D.R. Congo, Kenya, Nigeria, Sierra Leone, Tanzania (new record,
Degeeria apicalis Curran, 1927c: 8. Holotype male (
Eomedina grisescens Mesnil, 1960b: 651. Holotype female (BMNH). Type locality: D.R. Congo [as “Südafrika”, in error], Katanga, Bukama.
hamoyensis Cerretti & Wyatt, 2006.—Afrotropical: Namibia.
Eomedina hamoyensis Cerretti & Wyatt, 2006: 64. Holotype female (
Genus EOPHYLLOPHILA Townsend, 1926
EOPHYLLOPHILA Townsend, 1926c: 19. Type species: Eophyllophila elegans Townsend, 1926, by original designation [Oriental].
africana Villeneuve, 1935.—Afrotropical: Angola, Burundi, Cameroon, Kenya, Malawi, Nigeria, Sierra Leone, Tanzania, Uganda.
Eophyllophila africana Villeneuve, 1935a: 136. Syntypes, 1 male and 1 female (not located). Type localities: Nigeria (Oshogbo) and Uganda (west Rwenzori Range [as “W. Ruwenzori”], 1800m).
Undescribed spp.: Kenya, Malawi, Uganda (all in
Genus ERYNNIOLA Mesnil, 1977
ERYNNIOLA Mesnil, 1977c: 179. Type species: Erynniola atricolor Mesnil, 1977, by original designation.
atricolor Mesnil, 1977.—Afrotropical: Madagascar.
Erynniola atricolor Mesnil, 1977c: 181. Holotype male (
russipes Mesnil, 1977.—Afrotropical: Madagascar.
Erynniola russipes Mesnil, 1977c: 181. Holotype female (
Genus FILISTEA Cerretti & O’Hara, gen. n
FILISTEA Cerretti & O’Hara, gen. n. Type species: Viviania aureofasciata Curran, 1927, by present designation.
Note: This new genus is described in the New Taxa of Afrotropical Tachinidae section.
aureofasciata (Curran, 1927).—Afrotropical: Cameroon (new record, ZMHB [PC]), D.R. Congo, Nigeria, Uganda. Comb. n.
Viviania aureofasciata Curran, 1927c: 8. Holotype male (
verbekei Cerretti & O’Hara, sp. n.—Afrotropical: Cameroon, D.R. Congo, Nigeria, Uganda.
Filistea verbekei Cerretti & O’Hara, sp. n. Holotype male (ZMHB). Type locality: Cameroon, Kumba [as “Johann-Albrechtshöhe”] (4°38′N 9°28′E).
Note: This new species is described in the New Taxa of Afrotropical Tachinidae section.
Genus ISTOCHETA Rondani, 1859
FALLENIA Meigen, 1838: 265 (junior homonym of Fallenia Meigen, 1820). Type species: Tachina longicornis Fallén, 1810, by subsequent designation of
ISTOCHETA Rondani, 1859: 157, 171. Type species: Istocheta frontosa Rondani, 1859 (as “Sp. Typ. novaFrontalis Mihi”, incorrect original spelling, see
ISTOCHAETA Marschall, 1873: 334. Unjustified emendation of Istocheta Rondani, 1859 (see
HISTOCHAETA Brauer & Bergenstamm, 1891: 445 [also 1891: 141]. Unjustified emendation of Istocheta Rondani, 1859 (see
PROSOPOFRONTINA Townsend, 1926c: 33. Type species: Prosopofrontina pulchra Townsend, 1926, by original designation [Oriental].
UROPHYLLINA Villeneuve, 1937c: 5 (as subgenus of Urophylloides Brauer & Bergenstamm, 1893). Type species: Urophylloides (Urophyllina) rufipes Villeneuve, 1937, by monotypy [Oriental].
ANUROPHYLLINA Mesnil, 1961: 693 (as subgenus of Urophyllina Villeneuve, 1937). Nomen nudum (proposed after 1930 without designation of type species from four included species) (see note below and
ANUROPHYLLINA Mesnil, 1977d: 322 (as subgenus of Urophyllina Villeneuve, 1937). Type species: Urophylloides bicolor Villeneuve, 1937, by original designation [Oriental].
Note:
cerina (Mesnil, 1977).—Afrotropical: Madagascar.
Urophyllina (Anurophyllina) cerina Mesnil, 1977d: 322. Holotype female (
conifrons (Villeneuve, 1950).—Afrotropical: Uganda.
Degeeria conifrons Villeneuve, 1950: 2. Holotype male (
crucigera (Mesnil, 1977).—Afrotropical: Madagascar.
Urophyllina (Anurophyllina) crucigera Mesnil, 1977d: 322. Holotype male (
flava (Curran, 1927).—Afrotropical: Kenya, Nigeria, Sierra Leone.
Viviania flava Curran, 1927f: 108. Holotype male (BMNH). Type locality: Sierra Leone, Njala [ca. 8°14′N 12°1'W].
Degeeria frontosa Villeneuve, 1950: 3. Holotype female (
Genus KINIATILIOPS Mesnil, 1955
KINIATILIOPS Mesnil, 1955: 365. Type species: Kiniatiliops elegans Mesnil, 1955 (= Lomatacantha nigrapex Mesnil, 1952), by monotypy.
bilineatus (Mesnil, 1952).—Afrotropical: D.R. Congo.
Lomatacantha bilineata Mesnil, 1952a: 11. Holotype female (not located). Type locality: D.R. Congo, Nord-Kivu, Kamatembe, 2100m [ca. 1°19′S 29°6′E].
nigrapex (Mesnil, 1952).—Afrotropical: D.R. Congo, Ethiopia, Kenya, Rwanda, Tanzania, Zambia.
Lomatacantha nigrapex Mesnil, 1952a: 13. Holotype male (
Kiniatiliops elegans Mesnil, 1955: 365. Holotype male (
trispina Mesnil, 1959.—Afrotropical: Kenya.
Kiniatiliops trispina Mesnil, 1959: 14. Holotype female (
Genus KINIATILLA Villeneuve, 1938
KINIATILLA Villeneuve, 1938c: 10. Type species: Kiniatilla tricincta Villeneuve, 1938, by original designation.
KINIATILIA. Incorrect subsequent spelling of Kiniatilla Villeneuve, 1938 (
brevipalpis Mesnil, 1952.—Afrotropical: Burundi, D.R. Congo.
Kiniatilia brevipalpis Mesnil, 1952a: 14. Holotype male (
tricincta Villeneuve, 1938.—Afrotropical: Burundi, D.R. Congo, Rwanda, Uganda.
Kiniatilla tricincta Villeneuve, 1938c: 11. Lectotype female (
Genus LATIGINELLA Villeneuve, 1936
LATIGINELLA Villeneuve, 1936a: 4. Type species: Latiginella rufogrisea Villeneuve, 1936, by monotypy.
handeni Verbeke, 1963.—Afrotropical: Malawi (new record, NMDA [PC]), Mozambique, Tanzania.
Latiginella handeni Verbeke, 1963: 176. Holotype female (
rufogrisea Villeneuve, 1936.—Afrotropical: D.R. Congo, Kenya, Nigeria.
Latiginella rufogrisea Villeneuve, 1936a: 4. Holotype female (
Genus LINDNERIOLA Mesnil, 1959
LINDNERIOLA Mesnil, 1959: 17. Type species: Lindneriola paradoxa Mesnil, 1959, by monotypy.
paradoxa Mesnil, 1959.—Afrotropical: Tanzania, Uganda.
Lindneriola paradoxa Mesnil, 1959: 17. Holotype female (
Undescribed sp. 1: South Africa (
Undescribed sp. 2: Tanzania (
Genus MAURITIODORIA Townsend, 1932
MAURITIODORIA Townsend, 1932: 52. Type species: Medoria spinicosta Thomson, 1869, by original designation.
GASTROLEPTINA Villeneuve, 1938c: 6. Type species: Gastroleptina discolor Villeneuve, 1938 (= Medoria spinicosta Thomson, 1869), by monotypy.
spinicosta (Thomson, 1869).—Afrotropical: Mauritius, Réunion.
Medoria spinicosta Thomson, 1869: 522. Lectotype male (
Clytia spinicosta Thomson, 1869: 523 (junior secondary homonym of Medoria spinicosta Thomson, 1869). Type(s), male (
Gastroleptina discolor Villeneuve, 1938c: 7. Syntypes, 1 male and 1 female (BMNH). Type locality: Mauritius.
Note: The relative priority of Medoria spinicosta Thomson, 1869 and Clytia spinicosta Thomson, 1869, when both are placed in the same genus, was established by
Genus MEDINA Robineau-Desvoidy, 1830
MEDINA Robineau-Desvoidy, 1830: 138. Type species: Medina cylindrica Robineau-Desvoidy, 1830 (= Tachina collaris Fallén, 1820), by subsequent designation of
DEGEERIA Meigen, 1838: 249. Type species: Tachina collaris Fallén, 1820, by subsequent designation of
carbonata Mesnil, 1968.—Afrotropical: Madagascar, South Africa, Tanzania.
Medina carbonata Mesnil, 1968b: 8. Holotype male (
cinctella (Villeneuve, 1950).—Afrotropical: Malawi. Status revived.
Degeeria cinctella Villeneuve, 1950: 7. Holotype male (
Note: Degeeria cinctella Villeneuve, 1950 was treated as a synonym of Medina lateralis (Villeneuve, 1950) by
crocea (Villeneuve, 1950).—Afrotropical: Kenya, Malawi.
Degeeria crocea Villeneuve, 1950: 3. Lectotype male (
decellei Verbeke, 1964.—Afrotropical: Côte d’Ivoire.
Medina decellei Verbeke, 1964: 169. Holotype male (
denticulata (Villeneuve, 1950).—Afrotropical: Madagascar, Nigeria.
Degeeria denticulata Villeneuve, 1950: 6. Holotype female (
egregia (Villeneuve, 1950).—Afrotropical: Nigeria, Zambia, Zimbabwe.
Degeeria egregia Villeneuve, 1950: 4. Holotype male (
lateralis (Villeneuve, 1950).—Afrotropical: Burundi, D.R. Congo (new record,
Degeeria lateralis Villeneuve, 1950: 7. Holotype male (
mira Mesnil, 1977.—Afrotropical: Madagascar.
Medina mira Mesnil, 1977c: 185. Holotype male (
nigra Mesnil, 1968.—Afrotropical: Angola, Madagascar, South Africa.
Medina nigra Mesnil, 1968b: 8. Holotype male (
pectinifera Mesnil, 1977.—Afrotropical: Madagascar.
Medina pectinifera Mesnil, 1977c: 187. Holotype female (
rubricosa (Villeneuve, 1913).—Afrotropical: Nigeria.
Lydella rubricosa Villeneuve, 1913c: 30. Holotype female (BMNH). Type locality: Nigeria, Oshogbo.
Note:
semirufa (Villeneuve, 1950).—Afrotropical: Kenya, Malawi.
Degeeria semirufa Villeneuve, 1950: 6. Lectotype female (
setosella (Villeneuve, 1950).—Afrotropical: Burundi (new record,
Degeeria setosella Villeneuve, 1950: 5. Holotype male (
sopha Mesnil, 1977.—Afrotropical: Madagascar.
Medina sopha Mesnil, 1977c: 184. Holotype male (
spinulifera Mesnil, 1968.—Afrotropical: Tanzania.
Medina spinulifera Mesnil, 1968b: 9. Holotype female (
succuba Mesnil, 1977.—Afrotropical: Madagascar.
Medina succuba Mesnil, 1977c: 186. Holotype male (
vidua Mesnil, 1977.—Afrotropical: Madagascar.
Medina vidua Mesnil, 1977c: 187. Holotype female (
Possibly undescribed sp.: Nigeria (
Genus MEIGENIA Robineau-Desvoidy, 1830
MEIGENIA Robineau-Desvoidy, 1830: 198. Type species: Meigenia cylindrica Robineau-Desvoidy, 1830, by subsequent designation of Desmarest in d’
Note: Meigenia cylindrica Robineau-Desvoidy, 1830 is accepted as the type species of Meigenia Robineau-Desvoidy, 1830, following
Undetermined sp.: Yemen (
Genus MEDINOSPILA Mesnil, 1977
MEDINOSPILA Mesnil, 1977d: 322. Type species: Medinospila nigella Mesnil, 1977, by original designation.
nigella Mesnil, 1977.—Afrotropical: Madagascar.
Medinospila nigella Mesnil, 1977d: 323. Holotype male (
Genus PARARONDANIA Villeneuve, 1916
PARARONDANIA Villeneuve, 1916c: 498. Type species: Pararondania multipunctata Villeneuve, 1916, by monotypy.
multipunctata Villeneuve, 1916.—Afrotropical: South Africa.
Pararondania multipunctata Villeneuve, 1916c: 498. Holotype female (
Genus PARATRIXA Brauer & Bergenstamm, 1891
PARATRIXA Brauer & Bergenstamm, 1891: 357 [also 1891: 53]. Type species: Paratrixa polonica Brauer & Bergenstamm, 1891, by monotypy [Palaearctic]. New record.
Note:
aethiopica Mesnil, 1952.—Afrotropical: D.R. Congo, Rwanda, South Africa. Comb. revived.
Paratrixa aethiopica Mesnil, 1952a: 10. Holotype female (not located). Type locality: Rwanda, Ruhengeri [1°30′S 29°38′E], “sources Kirii” [not located], 1800–1825m.
stammeri Mesnil, 1952.—Afrotropical: D.R. Congo, South Africa (new record,
Paratrixa stammeri Mesnil, 1952a: 9. Holotype male (not located). Type locality: D.R. Congo, Nord-Kivu, Rutshuru, 1285m.
Genus PELASHYRIA Villeneuve, 1935
PELASHYRIA Villeneuve, 1935a: 138. Type species: Pelashyria grisescens Villeneuve, 1935, by monotypy.
grisescens Villeneuve, 1935.—Afrotropical: D.R. Congo.
Pelashyria grisescens Villeneuve, 1935a: 139. Syntypes, 1 male and 1 female (
Genus PRODEGEERIA Brauer & Bergenstamm, 1894
PRODEGEERIA Brauer & Bergenstamm, 1894: 617 [also 1895: 81]. Type species: Prodegeeria javana Brauer & Bergenstamm, 1894, by monotypy [Oriental].
MYXHYPOSTENA Villeneuve, 1939: 6. Type species: Myxhypostena consobrina Villeneuve, 1939, by original designation.
Note:
consobrina (Villeneuve, 1939).—Afrotropical: D.R. Congo, Ghana, Nigeria.
Myxhypostena consobrina Villeneuve, 1939: 6. Syntypes, 1 male and 1 female (
straeleni Mesnil, 1952.—Afrotropical: D.R. Congo, Uganda.
Prodegeeria straeleni Mesnil, 1952a: 14. Holotype male (
Genus PROSUCCINGULUM Mesnil, 1959
PROSUCCINGULUM Mesnil, 1959: 16. Type species: Prosuccingulum aberrans Mesnil, 1959, by monotypy.
aberrans Mesnil, 1959.—Afrotropical: Tanzania.
Prosuccingulum aberrans Mesnil, 1959: 16. Holotype female (
Undescribed sp.: Malawi (
Genus RIOTERIA Herting, 1973
RIOTERIA Herting, 1973: 3. Type species: Rioteria submacula Herting, 1973, by monotypy [Palaearctic].
flava Zeegers, 2007.—Afrotropical: Yemen.
Rioteria flava Zeegers, 2007: 395. Holotype male (
rufitibia (Mesnil, 1959).—Afrotropical: Nigeria, Tanzania.
Tachinophytopsis rufitibia Mesnil, 1959: 14. Holotype male (
Undescribed sp. 1: South Africa (
Undescribed sp. 2: Burkina (
Genus TRIGONOSPILA Pokorny, 1886
TRIGONOSPILA Pokorny, 1886: 191. Type species: Trigonospila picta Pokorny, 1886 (= Tachina ludio Zetterstedt, 1849), by monotypy [Palaearctic].
SUCCINGULUM Pandellé, 1894: 52. Type species: Succingulum transvittatum Pandellé, 1896, by subsequent monotypy of
bimaculata (Villeneuve, 1935).—Afrotropical: Ghana, Malawi, Mozambique, Nigeria, Sudan, Uganda.
Succingulum bimaculatum Villeneuve, 1935a: 142. Holotype female (
Note:
exigua (Villeneuve, 1935).—Afrotropical: South Africa. Status revived.
Succingulum exiguum Villeneuve, 1935a: 142. Holotype male (
Note: Succingulum exiguum Villeneuve, 1935 was treated as a synonym of Trigonospila mista (Villeneuve, 1913) by
integra (Villeneuve, 1935).—Afrotropical: “Afrique”. Oriental: India, Myanmar.
Succingulum integrum Villeneuve, 1935a: 142. Holotype male (possibly lost,
mista (Villeneuve, 1913).—Afrotropical: Angola, D.R. Congo, Kenya, Malawi, ?South Africa, Tanzania, Uganda.
Succingulum mista Villeneuve, 1913c: 39. Holotype female (
prasius Mesnil, 1977.
prasiusprasius Mesnil, 1977.—Afrotropical: Madagascar.
Trigonospila prasius prasius Mesnil, 1977c: 181, 183. Holotype male (
prasius trifida Mesnil, 1977.—Afrotropical: Madagascar.
Trigonospila prasius trifidus Mesnil, 1977c: 183. Holotype male (
Unplaced species of Blondeliini
triquetra Macquart, 1844.—Afrotropical: Réunion.
Dexia triquetra Macquart, 1844: 86 [also 1844: 243]. Lectotype male (
Tribe ERYCIINI
Genus AFROPHYLAX Cerretti & O’Hara, gen. n
AFROPHYLAX Cerretti & O’Hara, gen. n. Type species: Sturmia aureiventris Villeneuve, 1910, by present designation.
Note: This new genus is described in the New Taxa of Afrotropical Tachinidae section.
aureiventris (Villeneuve, 1910).—Afrotropical: Cameroon (new record, ZMHB [PC]), D.R. Congo, Nigeria, Sierra Leone, Tanzania, Uganda. Comb. n.
Sturmia aureiventris Villeneuve, 1910a: 252. Holotype male (
Note:
Genus ANTISTASEA Bischof, 1904
ANTISTASEA Bischof, 1904: 82. Type species: Antistasea fimbriata Bischof, 1904, by monotypy.
fimbriata Bischof, 1904.—Afrotropical: Kenya (new record,
Antistasea fimbriata Bischof, 1904: 83. Lectotype male (
Podomyia discalis Curran, 1939: 2. Holotype male (
Note:
mutans Mesnil, 1970.—Afrotropical: Botswana, South Africa.
Antistasea mutans Mesnil, 1970b: 106. Holotype male (
Genus APLOMYA Robineau-Desvoidy, 1830
APLOMYA Robineau-Desvoidy, 1830: 184. Type species: Aplomya zonata Robineau-Desvoidy, 1830 (= Tachina confinis Fallén, 1820), by subsequent designation of
APLOMYIA Agassiz, 1846a: 3. Unjustified emendation of Aplomya Robineau-Desvoidy, 1830 (see
HAPLOMYIA Agassiz, 1846b: 172. Unjustified emendation of Aplomya Robineau-Desvoidy, 1830 (see
PROZENILLIA Villeneuve, 1916c: 487. Type species: Prozenillia distans Villeneuve, 1916, by monotypy.
WIEDEMANNIOMYIA Townsend, 1933: 469. Type species: Tachina metallica Wiedemann, 1824, by original designation.
APLOMYIELLA Mesnil, 1939d: 31. Type species: Tricholyga impexa Villeneuve, 1916 (= Tachina metallica Wiedemann, 1824), by original designation.
ATRICHOLYGA Villeneuve, 1939: 9. Type species: Tricholyga impexa Villeneuve, 1916 (= Tachina metallica Wiedemann, 1824), by monotypy.
confinis (Fallén, 1820).—Afrotropical: ?Malawi, Yemen. Palaearctic: C. Asia, Europe (all), Japan, M. East (all), Mongolia, N. Africa (Canary Is., Madeira), Pal. China, Russia (W. Russia, W. Siberia, E. Siberia, S. Far East), Transcaucasia. Oriental: Orien. China.
Tachina confinis Fallén, 1820: 32. Syntypes, males and females (
Note: Tachina confinis Fallén, 1820 was recorded from Malawi by
distans (Villeneuve, 1916).—Afrotropical: Nigeria, South Africa, Sudan, Uganda.
Prozenillia distans Villeneuve, 1916c: 488. Lectotype male (
latimana Villeneuve, 1934.—Afrotropical: D.R. Congo, Kenya, Uganda.
Aplomyia latimana Villeneuve, 1934c: 409. Holotype female (
lycaena (Curran, 1927).—Afrotropical: Ethiopia, Senegal, South Africa.
Zenillia lycaena Curran, 1927d: 333. Holotype male (
metallica (Wiedemann, 1824).—Afrotropical: “W. Afr. to n.-e. Afr., E. Afr. & sthn Afr.” (
Tachina metallica Wiedemann, 1824: 46. Lectotype male (
Tachina nigriventris Wiedemann, 1824: 43. Lectotype male (
Tachina notata Wiedemann, 1830: 653. Type(s), male (
Tachina socia Wiedemann, 1830: 654. Type(s), female (
Phorocera eucalypta Loew, 1852: 659 [also 1862: 19, full description]. Type(s), unspecified sex (1 male in ZMHB). Type locality: Mozambique (Tete [as “Tette”] according to
Parexorista laeviventris van der Wulp, 1893: 173. Lectotype male (
Tricholyga impexa Villeneuve, 1916c: 494. Syntypes, 2 males (1 male in
Note: The relative priority of Tachina metallica Wiedemann, 1824 and Tachina nigriventris Wiedemann, 1824, when the two are treated as synonyms, was established by
The male syntype of Tricholyga impexa Villeneuve, 1916 in
poultoni (Villeneuve, 1922).—Afrotropical: Kenya, Nigeria, South Africa.
Exorista poultoni Villeneuve, 1922a: 518. Holotype male (not located). Type locality: Nigeria, near Ibadan, Moor Plantation.
seyrigi Mesnil, 1954.—Afrotropical: Madagascar.
Aplomyia (Aplomyiella) seyrigi Mesnil, 1954: 330. Holotype male (
versicolor (Curran, 1927).—Afrotropical: South Africa, Uganda.
Zenillia versicolor Curran, 1927d: 334. Holotype male (
Genus CADURCIELLA Villeneuve, 1927
CADURCIELLA Villeneuve, 1927: 120. Type species: Cadurciella rufipalpis Villeneuve, 1927, by monotypy.
rufipalpis Villeneuve, 1927.—Afrotropical: Namibia, South Africa, Zimbabwe. Palaearctic: M. East (Israel).
Cadurciella rufipalpis Villeneuve, 1927: 120. Lectotype male (not located), by fixation of
uniseta (Curran, 1933).—Afrotropical: South Africa, Zimbabwe.
Zenillia uniseta Curran, 1933: 166. Holotype male (BMNH). Type locality: Zimbabwe.
Undetermined sp.: U.A. Emirates, as “cf. Cadurciella spec.” (
Genus CARCELIA Robineau-Desvoidy, 1830
Subgenus CARCELIA Robineau-Desvoidy, 1830
CARCELIA Robineau-Desvoidy, 1830: 176. Type species: Carcelia bombylans Robineau-Desvoidy, 1830, by subsequent designation of
CARCELLIA. Incorrect subsequent spelling of Carcelia Robineau-Desvoidy, 1830 (
nudioculata Villeneuve, 1938.—Afrotropical: D.R. Congo, Rwanda, Uganda.
Carcelia nudioculata Villeneuve, 1938c: 4. Holotype male (not located). Type locality: D.R. Congo, Maniema, Lubutu.
Subgenus CARCELITA Mesnil, 1975
CARCELITA Mesnil, 1975a: 1384. Type species: Carcelia peraequalis Mesnil, 1950, by monotypy.
CARICELIA Mesnil, 1975a: 1384. Nomen nudum (proposed after 1930 without designation of type species; no included species).
CARICELIA Mesnil, 1975b: 1388. Type species: Carcelia obliterata Mesnil, 1950, by original designation.
Note: See
abrelicta Mesnil, 1950.—Afrotropical: Burundi, D.R. Congo, South Africa, Tanzania, Uganda.
Carcelia abrelicta Mesnil, 1950b: 16. Syntypes, males and females (1 female in
aequalis Villeneuve, 1939.—Afrotropical: Nigeria, South Africa, Tanzania, Zimbabwe.
Carcelia aequalis Villeneuve, 1939: 1. Syntypes, males (“plusieurs individus”) (1 male in
Note: One male of Carcelia aequalis Villeneuve, 1939 in
angulicornis Villeneuve, 1916.—Afrotropical: Ghana (new record,
Carcelia angulicornis Villeneuve, 1916c: 481. Syntypes, males and females (BMNH,
argyriceps (Curran, 1927).—Afrotropical: Uganda.
Zenillia argyriceps Curran, 1927d: 328. Holotype male (BMNH). Type locality: Uganda, [Kanungu District in southwestern Uganda], Kinkizi County, “Kizazi” [not located].
Zenillia hargreavesi Curran, 1928a: 238. Holotype male (BMNH). Type locality: Uganda, Kampala.
atricans Mesnil, 1955.—Afrotropical: Burundi (new record,
Carcelia atricans Mesnil, 1955: 362. Holotype male (
bigoti (Jaennicke, 1867).—Afrotropical: Ethiopia.
Exorista bigoti Jaennicke, 1867: 384 [also 1868: 76]. Type(s), female (
forcipata Mesnil, 1977.—Afrotropical: Madagascar.
Carcelia (Carcelita) forcipata Mesnil, 1977b: 178. Holotype male (
inusta Mesnil, 1950.—Afrotropical: Malawi (new record,
Carcelia inusta Mesnil, 1950b: 11. Syntypes, males and females (1 male in
keiseri Mesnil, 1977.—Afrotropical: Madagascar.
Carcelia (Carcelita) keiseri Mesnil, 1977b: 176. Holotype male (
lindneri Mesnil, 1959.—Afrotropical: South Africa (new record,
Carcelia lindneri Mesnil, 1959: 2. Holotype male (
lucidula Villeneuve, 1941.—Afrotropical: C.A. Republic (new record,
Carcelia lucidula Villeneuve, 1941b: 125. Syntypes, 2 males and 1 female (2 males in
normula (Curran, 1927).—Afrotropical: D.R. Congo, Ghana, Nigeria, Tanzania, Uganda.
Zenillia normula Curran, 1927d: 329. Holotype female (BMNH). Type locality: Uganda, “Rosaka” [not located].
oblectanea Mesnil, 1950.—Afrotropical: D.R. Congo, Kenya, South Africa (new record,
Carcelia oblectanea Mesnil, 1950b: 15. Syntypes, males and females (1 female in
oblimata Mesnil, 1950.—Afrotropical: South Africa.
Carcelia oblimata Mesnil, 1950b: 14. Syntypes, males and females (1 female in
obliterata Mesnil, 1950.—Afrotropical: Rwanda, South Africa.
Carcelia obliterata Mesnil, 1950b: 13. Lectotype female (
Note:
oculata (Villeneuve, 1910).—Afrotropical: D.R. Congo.
Exorista oculata Villeneuve, 1910a: 251. Lectotype male (
occulata. Incorrect subsequent spelling of oculata Villeneuve, 1910 (
Note:
orbitalis (Curran, 1927).—Afrotropical: South Africa, Zimbabwe.
Zenillia orbitalis Curran, 1927d: 330. Holotype male (
patellata Mesnil, 1977.—Afrotropical: Madagascar.
Carcelia (Carcelita) patellata Mesnil, 1977b: 177. Holotype female (
pellex Mesnil, 1950.—Afrotropical: Kenya, South Africa, Uganda.
Carcelia pellex Mesnil, 1950b: 13. Type(s), unspecified sex (not located). Type locality: South Africa.
peraequalis Mesnil, 1950.—Afrotropical: D.R. Congo, Kenya, Lesotho, Malawi, Rwanda, South Africa, Tanzania, Uganda, Zimbabwe.
Carcelia peraequalis Mesnil, 1950b: 18. Syntypes, males and females (possibly 1 male in
persimilis Mesnil, 1950.—Afrotropical: Madagascar, South Africa.
Carcelia persimilis Mesnil, 1950b: 17. Lectotype male (
Note:
vaga (Curran, 1927).—Afrotropical: Uganda.
Zenillia vaga Curran, 1927d: 332. Holotype male (BMNH). Type locality: Uganda, Kampala.
vara (Curran, 1927).—Afrotropical: Ghana, Kenya, South Africa, Tanzania.
Zenillia vara Curran, 1927d: 331. Holotype male (BMNH). Type locality: Kenya, Kabete [ca. 1°16′S 36°43′E, near Nairobi].
vexor (Curran, 1927).—Afrotropical: South Africa.
Zenillia vexor Curran, 1927d: 330. Holotype male (
Subgenus EURYCLEA Robineau-Desvoidy, 1863
EURYCLEA Robineau-Desvoidy, 1863a: 290. Type species: Euryclea tibialis Robineau-Desvoidy, 1863, by original designation [Palaearctic].
setifrons Mesnil, 1949.—Afrotropical: D.R. Congo, Nigeria (new record,
Carcelia (Eucarcelia) setifrons Mesnil, 1949a: 90. Holotype male (
Possibly undescribed spp.: Yemen, as “Carcelia (Caricelia) sp. 1 cf. vexor”, “Carcelia (Caricelia) sp. 2”, and “Carcelia (Caricelia) sp. 3” (
Genus CARCELIATHRIX Cerretti & O’Hara, gen. n
CARCELIATHRIX Cerretti & O’Hara, gen. n. Type species: Phorocera crassipalpis Villeneuve, 1938, by present designation.
Note: This new genus is described in the New Taxa of Afrotropical Tachinidae section.
crassipalpis (Villeneuve, 1938).—Afrotropical: D.R. Congo. Comb. n.
Phorocera crassipalpis Villeneuve, 1938c: 2. Lectotype male (
claripalpis. Incorrect subsequent spelling of crassipalpis Villeneuve, 1938 (original usage not found but spelling listed by
Note:
Undescribed sp. 1: Namibia (NNIC, examined by PC).
Undescribed sp. 2: South Africa (
Genus CESTONIA Rondani, 1861
CESTONIA Rondani, 1861b: 105. Type species: Cestonia cineraria Rondani, 1861, by monotypy [Palaearctic].
canariensis Villeneuve, 1936.—Afrotropical: U.A. Emirates. Palaearctic: N. Africa (Canary Is.), M. East (Israel).
Cestonia canariensis Villeneuve in Frey, 1936: 145. Syntypes, 1 male and 1 female (FMNHH). Type locality: Canary Islands, Gran Canaria, Las Palmas de Gran Canaria.
Note: Cestonia canariensis Villeneuve, 1936, was redescribed by
harteni Zeegers, 2007.—Afrotropical: Yemen.
Cestonia harteni Zeegers, 2007: 381. Holotype female (
Note:
rufipes Zeegers, 2007.—Afrotropical: Yemen.
Cestonia rufipes Zeegers, 2007: 382. Holotype male (
rutilans Villeneuve, 1929.—Afrotropical: Senegal, Yemen. Palaearctic: N. Africa (NE. Africa).
Cestonia rutilans Villeneuve, 1929a: 102. Syntypes, 1 male and 1 female (not located). Type locality: Egypt, Al Qāhirah [as “Caire”].
Genus CESTONIONERVA Villeneuve, 1929
CESTONIONERVA Villeneuve, 1929b: 43. Type species: Conogaster petiolata Villeneuve, 1910, by subsequent designation of
Note: Cestonionerva Villeneuve, 1929 was “Formé pour Conogaster petiolata Villen.” (
petiolata (Villeneuve, 1910).—Afrotropical: U.A. Emirates, Yemen. Palaearctic: C. Asia, M. East (Israel), Mongolia, N. Africa (Canary Is., NE. Africa), Pal. China.
Conogaster petiolata Villeneuve in Becker, 1910b: 144 [also 1910b: 14]. Holotype female (
Genus CHRYSERYCIA Mesnil, 1977
CHRYSERYCIA Mesnil, 1977b: 185. Type species: Chryserycia fulviceps Mesnil, 1977, by original designation.
fulviceps Mesnil, 1977.—Afrotropical: Madagascar.
Chryserycia fulviceps Mesnil, 1977b: 186. Holotype female (
Genus DESCAMPSINA Mesnil, 1956
DESCAMPSINA Mesnil, 1956b: 76. Type species: Descampsina sesamiae Mesnil, 1956, by original designation.
sesamiae Mesnil, 1956.—Afrotropical: Cameroon, D.R. Congo (new record,
Descampsina sesamiae Mesnil, 1956b: 76. Holotype, unspecified sex [male, examined by PC] (
Note:
Genus DIAPROCHAETA Mesnil, 1970
DIAPROCHAETA Mesnil, 1970b: 103. Type species: Diaprochaeta (Diaprochaeta) illustris Mesnil, 1970, by original designation.
illustris Mesnil, 1970.—Afrotropical: Zimbabwe.
Diaprochaeta (Diaprochaeta) illustris Mesnil, 1970b: 105. Holotype male (
Genus DRINO Robineau-Desvoidy, 1863
Subgenus DRINO Robineau-Desvoidy, 1863
DRINO Robineau-Desvoidy, 1863a: 250. Type species: Drino volucris Robineau-Desvoidy, 1863 (= Tachina lota Meigen, 1824), by original designation [Palaearctic].
STURMIODORIA Townsend, 1928: 391. Type species: Sturmiodoria facialis Townsend, 1928, by original designation.
cordata (Curran, 1927).—Afrotropical: Burundi, D.R. Congo, Guinea, Malawi, Rwanda.
Sturmia cordata Curran, 1927a: 12. Holotype male (
Note: Sturmia cordata Curran, 1927 is moved here from Drino subgenus Palexorista Townsend, 1921 based on examination of the holotype by PC.
facialis (Townsend, 1928).—Afrotropical: D.R. Congo. Palaearctic: Pal. China. Oriental: India, Indonesia, Malaysia, Orien. China, Philippines, Sri Lanka, Taiwan, Thailand.
Sturmiodoria facialis Townsend, 1928: 392. Holotype female (
Note: Sturmiodoria facialis Townsend, 1928 was recorded from Africa (D.R. Congo) by
lota (Meigen, 1824).—Afrotropical: Tanzania. Palaearctic: Europe (all except SW. Eur., Turkey), Japan, Pal. China [Ningxia], Russia (W. Russia, W. Siberia, S. Far East). Oriental: Orien. China.
Tachina lota Meigen, 1824: 326. Lectotype male (
Note: Tachina lota Meigen, 1824 was recorded from Africa (Tanzania) by
Subgenus PALEXORISTA Townsend, 1921
PALEXORISTA Townsend, 1921: 134. Type species: Tachina succini Giebel, 1862 (as “Tichina succini Giebel”), by monotypy.
PROSTURMIA Townsend, 1927c: 69. Type species: Prosturmia profana Townsend, 1927 (= Masicera solennis Walker, 1858), by original designation [Oriental].
PROSTURMINA Mesnil, 1949b: 103 (as subgenus of Drino Robineau-Desvoidy, 1863). Nomen nudum (proposed after 1930 without designation of type species; no included species).
PROSTURMINA Mesnil, 1949c: 8, 32 (as subgenus of Drino Robineau-Desvoidy, 1863). Nomen nudum (proposed after 1930 without type designation from three included species).
PROSTURMINA Mesnil, 1951: 161 (as subgenus of Drino Robineau-Desvoidy, 1863). Nomen nudum (proposed after 1930 without type designation; no included species).
PROSTURMINA Mesnil, 1970b: 110 (as subgenus of Drino Robineau-Desvoidy, 1863). Type species: Sturmia vigilans Villeneuve, 1933 (= Sturmia pulchra Curran, 1927), by original designation.
Note: The nomenclatural history of Prosturmina Mesnil was discussed by
amicula Mesnil, 1949.—Afrotropical: Cameroon, Ghana, Mozambique, Nigeria, Senegal, Tanzania.
Drino (Prosturmia) amicula Mesnil, 1949c: 30. Syntypes, males and females (1 male in
ampliceps (Karsch, 1886).—Afrotropical: Angola.
Masicera (Blepharipa) ampliceps Karsch, 1886b: 340. Holotype, unspecified sex [female, examined by JEOH] (ZMHB). Type locality: Angola, Pungo Andongo.
aureocincta Mesnil, 1977.—Afrotropical: Madagascar.
Drino (Prosturmia) aureocincta Mesnil, 1977b: 179. Holotype male (
aureola Mesnil, 1970.—Afrotropical: Sierra Leone.
Drino (Prosturmina) aureola Mesnil, 1970b: 110. Holotype male (
aurifera (Villeneuve, 1943).—Afrotropical: D.R. Congo.
Sturmia aurifera Villeneuve, 1943a: 36. Syntypes, males and females (2 males in
crassiseta Mesnil, 1968.—Afrotropical: South Africa.
Drino crassiseta Mesnil, 1968b: 5. Holotype male (
curvipalpis (van der Wulp, 1893).—Misidentification, not Afrotropical [known from Palaearctic, Oriental and Australasian regions].
Note: An unknown species was recorded as “Drino (Prosturmia T.T.) unisetosa Bar.” (originally described as Sturmia (Sturmia) unisetosa Baranov, 1932, currently a synonym of Drino curvipalpis (van der Wulp, 1893)) from Tanzania by
flavicans (Wiedemann, 1819).—Afrotropical: D.R. Congo, Malawi, South Africa, Uganda.
Tachina flavicans Wiedemann, 1819: 24. Type(s), female (not located). Type locality: South Africa, Western Cape, Cape of Good Hope [as “Prom. bon. sp.” = “Promontorium Bonae Spei”].
Sturmia congolensis Villeneuve, 1910a: 253. Syntypes, 3 females (not located). Type locality: D.R. Congo (as “Congo”, p. 249).
Note:
flaviseta (Thomson, 1869).—Afrotropical: Mauritius.
Masicera flaviseta Thomson, 1869: 522. Type(s), unspecified sex (
gilva (Hartig, 1838).—Misidentification, not Afrotropical [known from Palaearctic Region].
Note: An unknown species was recorded as “Sturmia gilva Hartig” (originally described as Tachina gilva Hartig, 1838) from D.R. Congo by
gilvoides (Curran, 1927).—Afrotropical: D.R. Congo, South Africa.
Sturmia gilvoides Curran, 1927f: 117. Holotype male (
idonea (Brauer & Bergenstamm, 1891).—Afrotropical: ?Eritrea, Mozambique, South Africa.
Argyrophylax idonea Brauer & Bergenstamm, 1891: 344 [also 1891: 40]. Type(s), male (
Sturmia partitor Curran, 1927f: 116. Holotype male (
Note:
imberbis (Wiedemann, 1830).—Afrotropical: D.R. Congo, Kenya, Malawi, South Africa, Sudan, Tanzania, U.A. Emirates, Uganda, Yemen. Palaearctic: C. Asia, Europe (SC. Eur., Turkey), M. East (all), N. Africa (Canary Is., NE. Africa), Transcaucasia.
Tachina imberbis Wiedemann, 1830: 317. Syntypes, 2 or more males (lost,
Sturmia zonata Curran, 1927d: 336. Holotype male (BMNH). Type locality: Uganda, Entebbe.
Note: See
inconspicua (Meigen, 1830).—Misidentification, not Afrotropical [known from Palaearctic and Oriental regions].
Note: An unknown species was recorded as “Sturmia (Sturmia) inconspicua” (originally described as Tachina inconspicua Meigen, 1830) from Tanzania by
iterata Mesnil, 1949.—Afrotropical: South Africa, Uganda.
Drino (Prosturmia) iterata Mesnil, 1949c: 31. Syntypes, males and females (1 male in
latigena (Mesnil, 1944).—Afrotropical: Djibouti, U.A. Emirates. Palaearctic: M. East (Israel), N. Africa (NE. Africa).
Phorcida latigena Mesnil, 1944: 15. Holotype male (
Tachina imberbis of authors (e.g.,
lavinia (Curran, 1927).—Afrotropical: D.R. Congo, Uganda.
Sturmia lavinia Curran, 1927c: 14. Holotype male (
laxa (Curran, 1927).—Afrotropical: Botswana, Kenya, Malawi, South Africa, Sudan, Swaziland, Tanzania, Uganda, Zimbabwe. Oriental: India.
Sturmia laxa Curran, 1927d: 335. Holotype male (BMNH). Type locality: Tanzania, Morogoro.
mayneana (Villeneuve, 1930).—Afrotropical: D.R. Congo.
Sturmia mayneana Villeneuve, 1930b: 59. Syntypes, males and females (“plusieurs individus”) (
melancholica Mesnil, 1949.—Afrotropical: Zimbabwe.
Drino (Prosturmia) melancholica Mesnil, 1949c: 16. Syntypes, 1 male and 1 female (
nova Mesnil, 1949.—Afrotropical: Madagascar.
Drino (Prosturmia) nova Mesnil, 1949c: 27. Syntypes, males and females (1 male in
obliterata Mesnil, 1949.—Afrotropical: Malawi, Senegal, South Africa.
Drino (Prosturmia) patruelis obliterata Mesnil, 1949c: 18. Syntypes, males and females (
parachrysops (Bezzi, 1925).—Afrotropical: Ghana, Kenya, Mali, Nigeria, Senegal, Yemen. Palaearctic: M. East (M. East [Saudi Arabia,
Sturmia parachrysops Bezzi, 1925b: 114. Lectotype male (BMNH), by designation of
patruelis Mesnil, 1949.—Afrotropical: Malawi, South Africa, Tanzania, Uganda, Zimbabwe.
Drino (Prosturmia) patruelis Mesnil, 1949c: 17. Syntypes, males and probably females (“nombreux exemplaires”) (1 male and possibly other syntypes in
pulchra (Curran, 1927).—Afrotropical: D.R. Congo, Uganda.
Sturmia pulchra Curran, 1927a: 16. Holotype male (BMNH). Type locality: Uganda, Entebbe.
Sturmia vigilans Villeneuve, 1933: 278. Holotype female (
quadrizonula (Thomson, 1869).—Afrotropical: widespread, including D.R. Congo, Ghana, Kenya, Saint Helena, São Tomé & Principe, Senegal, Seychelles, South Africa, Tanzania, Uganda, Zimbabwe (
Masicera quadrizonula Thomson, 1869: 521. Lectotype female (
Note: Masicera quadrizonula Thomson, 1869 was redescribed by
rufa Zeegers, 2007.—Afrotropical: Yemen.
Drino rufa Zeegers, 2007: 385. Holotype male (
salva (Wiedemann, 1830).—Afrotropical: South Africa.
Tachina salva Wiedemann, 1830: 340. Type(s), female (1 syntype in
subaurata (Walker, 1853).—Afrotropical: Madagascar, South Africa.
Tachina subaurata Walker, 1853: 298. Type(s) female (BMNH). Type locality: South Africa, Western Cape, Cape of Good Hope [as “Cape”].
succini (Giebel, 1862).—Afrotropical: ?Tanzania.
Tachina succini Giebel, 1862: 319. Holotype female (
Note: Tachina succini Giebel, 1862 was described from a copal inclusion originally thought to be an amber fossil (
tenella (Bezzi, 1911).—Afrotropical: South Africa.
Erycia (Bactromyia) tenella Bezzi, 1911: 60. Holotype female (
terrosa Mesnil, 1949.—Afrotropical: Madagascar.
Drino (Prosturmia) terrosa Mesnil, 1949c: 20. Type(s), unspecified sex (
ugandana (Curran, 1927).—Afrotropical: Burundi, D.R. Congo, Malawi, South Africa, Uganda, Zimbabwe.
Sturmia ugandana Curran, 1927c: 16. Holotype male (
Subgenus ZYGOBOTHRIA Mik, 1891
ZYGOBOTHRIA Mik, 1891: 193. Type species: Sturmia atropivora Robineau-Desvoidy, 1830, by original designation.
FORMOSODORIA Townsend, 1933: 475. Type species: Sturmia (Argyrophylax) dilabida Villeneuve, 1916 (= Meigenia ciliata van der Wulp, 1881), by original designation.
atropivora (Robineau-Desvoidy, 1830).—Afrotropical: “widespread Afrotrop Reg.” (
Sturmia atropivora Robineau-Desvoidy, 1830: 171. Syntypes, more than 80 males and females (lost,
Sturmia masakensis Curran, 1927f: 117. Holotype male (BMNH). Type locality: Uganda, Masaka.
masakesnsis. Incorrect subsequent spelling of masakensis Curran, 1927 (
ciliata (van der Wulp, 1881).—Afrotropical: “widespread mainland Afrotrop. Reg.” (
Meigenia ciliata van der Wulp, 1881: 38. Lectotype male (
Sturmia (Argyrophylax) dilabida Villeneuve, 1916c: 479. Type(s), unspecified number and including at least 1 male (
Note: Sturmia dilabida Villeneuve, 1916 was described from one or more specimens, at least one of which was male. The type locality was given as Durban and the depository as
Sturmia munroi Curran, 1927c: 17. Holotype male (
Sturmia (Sturmia) macrophallus Baranov, 1932: 76. Lectotype male (SDEI), by designation of
Formosodoria foeda Villeneuve, 1933: 280 (as “Formosodoria foeda T. T.”). Nomen nudum (proposed in synonymy [with Sturmia dilabida Villeneuve, 1916 and Tachina convergens Wiedemann, 1824, the latter misidentified] and not made available by subsequent usage before 1961).
Tachina convergens of
grandicornis Mesnil, 1977.—Afrotropical: Madagascar.
Drino (Zygobothria) grandicornis Mesnil, 1977b: 178. Holotype male (
Genus EUGAEDIOXENIS Cerretti, O’Hara & Stireman, 2015
EUGAEDIOXENIS Cerretti, O’Hara & Stireman in
haematodes (Villeneuve, 1937).—Afrotropical: South Africa.
Gaedioxenis haematodes Villeneuve, 1937a: 207. Holotype male (
horridus Cerretti, O’Hara & Stireman, 2015.—Afrotropical: South Africa.
Eugaedioxenis horridus Cerretti, O’Hara & Stireman in
Genus HYPERSARA Villeneuve, 1935
HYPERSARA Villeneuve, 1935a: 139. Type species: Hypersara argentata Villeneuve, 1935, by monotypy.
argentata Villeneuve, 1935.—Afrotropical: D.R. Congo, Nigeria.
Hypersara argentata Villeneuve, 1935a: 140. Holotype male (not located). Type locality: D.R. Congo, Nord-Kivu, Walikale [ca. 1°25′S 28°00′E].
Undescribed sp.: Ethiopia (
Genus INTRAPALES Villeneuve, 1938
INTRAPALES Villeneuve, 1938c: 8. Type species: Intrapales remotella Villeneuve, 1938, by monotypy.
hirsuta Mesnil, 1977.—Afrotropical: Madagascar.
Intrapales hirsuta Mesnil, 1977b: 185. Holotype male (
insularis Mesnil, 1977.—Afrotropical: Madagascar.
Intrapales insularis Mesnil, 1977b: 184. Holotype male (
remotella Villeneuve, 1938.—Afrotropical: D.R. Congo, Nigeria (new record,
Intrapales remotella Villeneuve, 1938c: 8. Syntypes, 2 males and 1 female (
Genus KAISERIOLA Mesnil, 1970
KAISERIOLA Mesnil, 1970b: 105 (as subgenus of Diaprochaeta Mesnil, 1970). Type species: Diaprochaeta (Kaiseriola) aperta Mesnil, 1970, by original designation.
Note: Kaiseriola Mesnil, 1970 was treated as a synonym of Diaprochaeta Mesnil, 1970 by
aperta (Mesnil, 1970).—Afrotropical: Mozambique (new record, JOS [PC]), South Africa.
Diaprochaeta (Kaiseriola) aperta Mesnil, 1970b: 105. Holotype male (
obscura (Mesnil, 1970).—Afrotropical: Madagascar.
Diaprochaeta (Kaiseriola) obscura Mesnil, 1970b: 106. Holotype male (
Genus LUBUTANA Villeneuve, 1938
LUBUTANA Villeneuve, 1938c: 10. Type species: Lubutana divaricata Villeneuve, 1938, by original designation.
divaricata Villeneuve, 1938.—Afrotropical: D.R. Congo, Ethiopia, Ghana, Malawi, Nigeria, Sierra Leone, Uganda.
Lubutana divaricata Villeneuve, 1938c: 10. Syntypes, males (
mayeri Mesnil, 1955.—Afrotropical: Nigeria.
Lubutana mayeri Mesnil, 1955: 363. Holotype female (
perplexa Mesnil, 1955.—Afrotropical: D.R. Congo, Rwanda, Uganda.
Lubutana perplexa Mesnil, 1955: 362. Holotype female (
Genus LYDELLA Robineau-Desvoidy, 1830
LYDELLA Robineau-Desvoidy, 1830: 112. Type species: Lydella grisescens Robineau-Desvoidy, 1830, by subsequent designation of
METOPOSISYROPS Townsend, 1916d: 320. Type species: Metoposisyrops oryzae Townsend, 1916, by original designation [Oriental].
Note: Metoposisyrops Townsend, 1916 was synonymized with Lydella Robineau-Desvoidy, 1830 by
sesamiae (Mesnil, 1968).—Afrotropical: D.R. Congo (new record,
Metagonistylum sesamiae Mesnil, 1968b: 4. Holotype female (
Genus MADREMYIA Townsend, 1916
MADREMYIA Townsend, 1916d: 305. Type species: Madremyia parva Townsend, 1916 (= Phorocera saundersii Williston, 1889), by original designation [Neotropical]. New record.
Note: Madremyia Townsend, 1916 is newly recorded from the Afrotropical Region for a species previously placed in Phryxe Robineau-Desvoidy, 1830.
setinervis (Mesnil, 1968).—Afrotropical: Tanzania. Comb. n.
Phryxe setinervis Mesnil, 1968b: 5. Holotype female (
Note: Phryxe setinervis Mesnil, 1968 was treated as a species of Phryxe Robineau-Desvoidy, 1830 by
Genus MYXARCHICLOPS Villeneuve, 1916
MYXARCHICLOPS Villeneuve, 1916c: 494. Type species: Myxarchiclops caffer Villeneuve, 1916, by subsequent designation of
caffer Villeneuve, 1916.—Afrotropical: South Africa.
Myxarchiclops caffer Villeneuve, 1916c: 495. Lectotype male (
major Villeneuve, 1930.—Afrotropical: South Africa.
Myxarchiclops major Villeneuve, 1930a: 353. Syntypes, 2 females (
Genus NEOLYDELLA Mesnil, 1939
NEOLYDELLA Mesnil, 1939a: 209 (as subgenus of Lydella Robineau-Desvoidy, 1830). Type species: Lydella (Neolydella) pruinosa Mesnil, 1939, by monotypy.
pruinosa (Mesnil, 1939).—Afrotropical: Madagascar.
Lydella (Neolydella) pruinosa Mesnil, 1939a: 209. Syntypes, 3 males (
Genus NILEA Robineau-Desvoidy, 1863
NILEA Robineau-Desvoidy, 1863a: 275. Type species: Nilea innoxia Robineau-Desvoidy, 1863, by original designation [Palaearctic].
longicauda (Mesnil, 1970).—Afrotropical: Madagascar. Comb. n.
Sturmia longicauda Mesnil, 1970b: 91. Holotype male (
Note: Sturmia longicauda Mesnil, 1970 was treated as a species of Sturmia Robineau-Desvoidy, 1830 by
perplexa Mesnil, 1977.—Afrotropical: Burundi (new record,
Nilea perplexa Mesnil, 1977b: 188. Holotype male (
Undescribed sp.: Tanzania (
Genus PARADRINO Mesnil, 1949
PARADRINO Mesnil, 1949b: 103 (as subgenus of Drino Robineau-Desvoidy, 1863). Type species: Sturmia halli Curran, 1939 (as “Paradrino Halli Curr.”, p. 100), by monotypy (see
halli (Curran, 1939).—Afrotropical: Botswana, Tanzania, Uganda, Zimbabwe.
Sturmia halli Curran, 1939: 2. Holotype male (
Sturmia rhodesiensis Jones, 1939: 16. Syntypes, males and females (BMNH). Type locality: Zimbabwe, Mazoe.
Note:
Undescribed species of “?Paradrino”: Yemen (
Genus PHRYXE Robineau-Desvoidy, 1830
PHRYXE Robineau-Desvoidy, 1830: 158. Type species: Phryxe athaliae Robineau-Desvoidy, 1830 (= Tachina vulgaris Fallén, 1810), by subsequent designation of
Note: The single species recognized in Phryxe Robineau-Desvoidy, 1830 by
Undescribed sp.: Ethiopia (
Genus PSEUDOPERICHAETA Brauer & Bergenstamm, 1889
PSEUDOPERICHAETA Brauer & Bergenstamm, 1889: 92 [also 1890: 24]. Type species: Pseudoperichaeta major Brauer & Bergenstamm, 1889 (= Phryxe palesioidea Robineau-Desvoidy, 1830), by monotypy [Palaearctic].
ACHAETONEURILLA Mesnil, 1939a: 210 (as subgenus of Pseudoperichaeta Brauer & Bergenstamm, 1889). Type species: Pseudoperichaeta (Achaetoneurilla) madecassa Mesnil, 1939, by monotypy.
laevis Villeneuve, 1932.—Afrotropical: Nigeria, Tanzania, Uganda, Zimbabwe.
Pseudoperichaeta laevis Villeneuve, 1932: 285. Syntypes, males and females (not located). Type locality: Zimbabwe, Harare [as “Salisbury”].
Phorocera bolyodes Curran, 1933: 166. Holotype female (BMNH). Type locality: Zimbabwe, Harare [as “Salisbury”].
bothyodes. Incorrect subsequent spelling of bolyodes Curran, 1933 (original usage not found but spelling listed by
leo (Curran, 1941).—Afrotropical: Zimbabwe.
Phorocera leo Curran, 1941: 10. Holotype female (
Pseudoperichaeta pilosa Villeneuve, 1942a: 52. Syntypes, 2 males (1 male in
madecassa Mesnil, 1939.—Afrotropical: Madagascar.
Pseudoperichaeta (Achaetoneurilla) madecassa Mesnil, 1939a: 210. Syntypes, 12 males and females (3 males and 2 females in
nestor (Curran, 1927).—Afrotropical: D.R. Congo, Nigeria, Tanzania.
Phorocera nestor Curran, 1927c: 12. Holotype male (
pacta Villeneuve, 1932.—Afrotropical: D.R. Congo, Mauritius, South Africa, Zimbabwe.
Pseudoperichaeta pacta Villeneuve, 1932: 285. Holotype female (not located). Type locality: South Africa, Western Cape, “région de Cape-Town”.
sallax (Curran, 1927).—Afrotropical: D.R. Congo.
Phorocera sallax Curran, 1927c: 11. Holotype male (
Genus PTILOCATAGONIA Mesnil, 1956
PTILOCATAGONIA Mesnil, 1956b: 79 (as subgenus of Sisyropa Brauer & Bergenstamm, 1889). Type species: Sisyropa (Ptilocatagonia) viridescens Mesnil, 1956, by monotypy.
viridescens (Mesnil, 1956).—Afrotropical: Sierra Leone, Tanzania, Zambia.
Sisyropa (Ptilocatagonia) viridescens Mesnil, 1956b: 79. Holotype male (
Note:
Genus SENOMETOPIA Macquart, 1834
SENOMETOPIA Macquart, 1834: 160 [also 1834: 296]. Type species: Carcelia aurifrons Robineau-Desvoidy, 1830 (= Tachina excisa Fallén, 1820), by subsequent designation of
STENOMETOPIA Agassiz, 1846b: 351. Unjustified emendation of Senometopia Macquart, 1834.
EOCARCELIA Townsend, 1919b: 582. Type species: Eocarcelia ceylanica Townsend, 1919, by original designation [Oriental].
EOCARCELIOPSIS Townsend, 1928: 392. Type species: Eocarceliopsis bakeri Townsend, 1928, by original designation [Oriental].
EUCARCELIA Baranov, 1934: 393. Type species: Tachina excisa Fallén, 1820, by original designation [Palaearctic].
albatella (Villeneuve, 1941).—Afrotropical: D.R. Congo, Malawi.
Carcelia albatella Villeneuve, 1941b: 125. Syntypes, 1 male and 1 female (
evolans (Wiedemann, 1830).—Afrotropical: Côte d’Ivoire, Senegal, Sierra Leone, ?Yemen.
Tachina evolans Wiedemann, 1830: 321. Type(s), unspecified sex (not located). Type locality: Sierra Leone.
Note: Tachina evolans Wiedemann, 1830 has been misidentified from other places in the Afrotropical Region and from the Palaearctic Region, as noted by
hectica (Speiser, 1910).—Afrotropical: Kenya, Tanzania, Uganda.
Carcelia hectica Speiser, 1910: 141. Holotype male (
illota (Curran, 1927).—Afrotropical: Nigeria, South Africa, Tanzania. Oriental: India, Laos, Orien. China. Australasian: Australia.
Zenillia illota Curran, 1927d: 328. Holotype male (BMNH). Type locality: Tanzania, Morogoro.
judicabilis (Mesnil, 1949).—Afrotropical: D.R. Congo, Malawi, Zimbabwe.
Carcelia (Eucarcelia) evolans judicabilis Mesnil, 1949a: 90. Holotype, unspecified sex [male, examined by PC] (
laetifica (Mesnil, 1949).—Afrotropical: D.R. Congo, Ghana, Nigeria.
Carcelia (Eucarcelia) evolans laetifica Mesnil, 1949a: 89. Holotype male (
norma (Curran, 1927).—Afrotropical: Malawi, Tanzania, Uganda.
Zenillia norma Curran, 1927d: 329. Holotype male (BMNH). Type locality: Uganda, Bugoma Forest [ca. 1°16′N 30°57′E].
Genus SISYROPA Brauer & Bergenstamm, 1889
SISYROPA Brauer & Bergenstamm, 1889: 163 [also 1890: 95]. Type species: Tachina thermophila Wiedemann, 1830, by monotypy [Oriental].
STYLURODORIA Townsend, 1933: 476. Type species: Stylurodoria stylata Townsend, 1933, by original designation.
CTENOPHOROCEROPSIS Baranov, 1938: 408. Type species: Ctenophoroceropsis yerburyi Baranov, 1938, by original designation.
POUJADEA Mesnil, 1949b: 102. Nomen nudum (proposed after 1930 without designation of type species; no included species) (see
EOCATAGONIA Mesnil, 1949b: 103 (as subgenus Sisyropa Brauer & Bergenstamm, 1889). Nomen nudum (proposed after 1930 without designation of type species; no included species) (see
POUJADEA Mesnil, 1950c: 108. Type species: Zenillia insolita Curran, 1927, by monotypy (see
EOCATAGONIA Mesnil, 1950c: 148 (as subgenus Sisyropa Brauer & Bergenstamm, 1889). Type species: Sisyropa (Eocatagonia) argyrata Mesnil, 1950, by monotypy (see
argyrata Mesnil, 1950.—Afrotropical: Senegal.
Sisyropa (Eocatagonia) argyrata Mesnil, 1950c: 148. Holotype male (
boveyi Mesnil, 1958.—Afrotropical: Ghana, Guinea, Kenya, Nigeria, Tanzania.
Sisyropa (Catagonia) boveyi Mesnil, 1958: 252. Holotype male (
insolita (Curran, 1927).—Afrotropical: D.R. Congo.
Zenillia insolita Curran, 1927c: 5. Holotype male (
insoleta. Incorrect subsequent spelling of insolita Curran, 1927 (
madecassa Mesnil, 1944.—Afrotropical: Madagascar.
Sisyropa formosa madecassa Mesnil, 1944: 14. Holotype male (
negator (Curran, 1927).—Afrotropical: D.R. Congo.
Sturmia negator Curran, 1927c: 15. Holotype male (
negastor. Incorrect subsequent spelling of negator Curran, 1927 (
stylata (Townsend, 1933).—Afrotropical: Ghana, Mali, Nigeria, Sierra Leone, Sudan. Oriental: India, Sri Lanka, Taiwan.
Stylurodoria stylata Townsend, 1933: 476. Holotype female (SDEI). Type locality: Taiwan, P’ingtung Hsien, Changkou [as “Kankau”, near Hengch’un].
subdistincta (Villeneuve, 1916).—Afrotropical: Côte d’Ivoire, Ethiopia, Ghana, Senegal, South Africa, Tanzania.
Catagonia subdistincta Villeneuve, 1916c: 484. Syntypes, 2 males (
Sisyropa cinerosa Mesnil, 1944: 15. Holotype male (
yerburyi (Baranov, 1938).—Afrotropical: Yemen.
Ctenophoroceropsis yerburyi Baranov, 1938: 409. Holotype male (BMNH,
yerburi. Incorrect subsequent spelling of yerburyi Baranov, 1938 (original usage not found but spelling listed by
Possibly undescribed spp.: Nigeria (BMNH,
Genus STURMIOPSIS Townsend, 1916
STURMIOPSIS Townsend, 1916d: 313. Type species: Sturmiopsis inferens Townsend, 1916, by original designation.
RHODESINA Curran, 1939: 3 (junior homonym of Rhodesina Malloch, 1921). Type species: Rhodesina parasitica Curran, 1939, by original designation.
CURRANOMYIA Townsend in Cuthbertson & Munro, 1941: 115 (nomen novum for Rhodesina Curran, 1939).
Note:
inferens Townsend, 1916.—Afrotropical: Madagascar (probably introduced,
Sturmiopsis inferens Townsend, 1916d: 313. Holotype female (
parasitica (Curran, 1939).—Afrotropical: Benin, Ghana, Kenya, Nigeria, Senegal, Tanzania, Zimbabwe. Oriental: India (introduced,
Rhodesina parasitica Curran, 1939: 3. Holotype male (
Sturmiopsis angustifrons Mesnil, 1959: 11. Holotype male (
setifrons Mesnil, 1977.—Afrotropical: Madagascar.
Sturmiopsis setifrons Mesnil, 1977b: 187. Holotype male (
Genus STYLOCARCELIA Zeegers, 2007
STYLOCARCELIA Zeegers, 2007: 396. Type species: Stylocarcelia stylata Zeegers, 2007, by original designation.
stylata Zeegers, 2007.—Afrotropical: Yemen.
Stylocarcelia stylata Zeegers, 2007: 396. Holotype male (
Genus THECOCARCELIA Townsend, 1933
THECOCARCELIA Townsend, 1933: 471. Type species: Argyrophylax pelmatoprocta Brauer & Bergenstamm, 1891 (= Masicera acutangulata Macquart, 1851), by original designation [Palaearctic].
THELYCARCELIA Townsend, 1933: 475. Type species: Thelycarcelia thrix Townsend, 1933 (= Sturmia sumatrana Baranov, 1932), by original designation [Oriental].
Note: The relative priority of Thecocarcelia Townsend, 1933 and Thelycarcelia Townsend, 1933, when the two are treated as synonyms, was established by
acutangulata (Macquart, 1851).—Afrotropical: “W. Afr. to E. Afr. & sthn Afr.” (
Masicera acutangulata Macquart, 1851a: 478. Type(s), female (
Masicera incedens Rondani, 1861b: 22. Type(s), female (MZF,
Argyrophylax pelmatoprocta Brauer & Bergenstamm, 1891: 344 [also 1891: 40]. Syntypes, males and females (2 males and 4 females in
Note: Argyrophylax pelmatoprocta Brauer & Bergenstamm, 1891 was described from an unspecified number of males and females from “M.-Europa”.
ebenina Mesnil, 1950.—Afrotropical: D.R. Congo, South Africa.
Thecocarcelia ebenina Mesnil, 1950b: 21. Syntypes, males and possibly females (not located). Type locality: South Africa, KwaZulu-Natal.
flavicosta Zeegers, 2007.—Afrotropical: Yemen.
Thecocarcelia flavicosta Zeegers, 2007: 398. Holotype male (
latifrons Mesnil, 1949.—Afrotropical: Mozambique, South Africa, Uganda, Zimbabwe.
Thecocarcelia latifrons Mesnil, 1949b: 56. Holotype male (
Note:
latimana Mesnil, 1950.—Afrotropical: South Africa.
Thecocarcelia latimana Mesnil, 1950b: 22. Syntypes, males and females (not located). Type locality: South Africa.
pauciseta Mesnil, 1977.—Afrotropical: Madagascar.
Thecocarcelia pauciseta Mesnil, 1977b: 181. Holotype male (
robusta Mesnil, 1950.—Afrotropical: D.R. Congo.
Thecocarcelia robusta Mesnil, 1950b: 22. Syntypes, males (1 male in
trichops Herting, 1967.—Afrotropical: South Africa, Zambia. Palaearctic: Europe (W. Eur., SW. Eur., SC. Eur., SE. Eur.), Japan, Pal. China.
Thecocarcelia trichops Herting, 1967: 4. Holotype male (
Note: Specimens from the Afrotropical Region identified as Thecocarcelia trichops Herting, 1967 should be checked to confirm their identity.
ventralis Mesnil, 1959.—Afrotropical: D.R. Congo, Ghana, Nigeria, Sierra Leone, Tanzania.
Thecocarcelia ventralis Mesnil, 1959: 2. Holotype male (
vibrissata Mesnil, 1977.—Afrotropical: Madagascar.
Thecocarcelia vibrissata Mesnil, 1977b: 181. Holotype male (
Genus THELAIRODRINO Mesnil, 1954
THELAIRODRINO Mesnil, 1954c: 470 (as subgenus of Thelairosoma Villeneuve, 1916). Type species: Thelairosoma gracilis Mesnil, 1952, by original designation [Oriental].
anaphe (Curran, 1927).—Afrotropical: Cameroon, D.R. Congo, Kenya, Malawi, Nigeria, Tanzania, Zimbabwe.
Sturmia anaphe Curran, 1927e: 447. Holotype male (BMNH). Type locality: Tanzania, Morogoro.
Note: Sturmia anaphe, described by
cardinalis (Mesnil, 1949).—Afrotropical: D.R. Congo.
Drino cardinalis Mesnil, 1949a: 91. Holotype, unspecified sex [male, examined by PC] (
potina (Curran, 1927).—Afrotropical: South Africa.
Sturmia potina Curran, 1927f: 118. Holotype male (
Genus THELAIROSOMA Villeneuve, 1916
THELAIROSOMA Villeneuve, 1916c: 499. Type species: Thelairosoma fumosum Villeneuve, 1916, by monotypy.
SEYRIGOMYIA Mesnil, 1944: 11. Type species: Seyrigomyia fulvella Mesnil, 1944, by original designation.
LESPESIOPSIS Mesnil, 1954c: 471 (as subgenus of Thelairosoma Villeneuve, 1916). Type species: Thelairosoma (Lespesiopsis) coerulescens Mesnil, 1954, by monotypy.
THELAIROXENIS Mesnil, 1954c: 472 (as subgenus of Thelairosoma Villeneuve, 1916). Type species: Thelairosoma (Thelairoxenis) pallidum Mesnil, 1954, by original designation.
angustifrons (Villeneuve, 1916).—Afrotropical: D.R. Congo, Malawi, Mozambique, Nigeria, South Africa, Tanzania, Uganda.
Sturmia (Blepharipoda) angustifrons Villeneuve, 1916c: 478. Syntypes, 3 males and 1 female (1 male in
atrum Mesnil, 1970.—Afrotropical: Madagascar.
Thelairosoma (Thelairosoma) atrum Mesnil, 1970b: 101. Holotype male (
brunnescens (Villeneuve, 1934).—Afrotropical: Rwanda, Uganda.
Erycia brunnescens Villeneuve, 1934d: 69. Lectotype female (
carbonatum (Mesnil, 1944).—Afrotropical: Madagascar.
Seyrigomyia carbonata Mesnil, 1944: 13. Holotype male (
coerulescens Mesnil, 1954.—Afrotropical: Burundi, D.R. Congo, Rwanda, Tanzania.
Thelairosoma (Lespesiopsis) coerulescens Mesnil, 1954c: 471. Holotype male (
comatum Villeneuve, 1938.—Afrotropical: Uganda.
Thelairosoma comatum Villeneuve, 1938b: 2. Holotype male (
diaphanum Mesnil, 1954.—Afrotropical: D.R. Congo.
Thelairosoma (Thelairoxenis) diaphanum Mesnil, 1954c: 472. Holotype male (
flavipalpe Villeneuve, 1938.—Afrotropical: D.R. Congo.
Thelairosoma flavipalpe Villeneuve, 1938b: 3. Holotype male (
fulvellum (Mesnil, 1944).—Afrotropical: Madagascar.
Seyrigomyia fulvella Mesnil, 1944: 12. Holotype, unspecified sex (
fumosum Villeneuve, 1916c: 500.—Afrotropical: D.R. Congo, Ghana, Malawi, Mozambique (new record,
Thelairosoma fumosum Villeneuve, 1916c: 500. Lectotype male (
hybridum Mesnil, 1970.—Afrotropical: Madagascar.
Thelairosoma (Seyrigomyia) hybrida Mesnil, 1970b: 103. Holotype male (
ingrami Mesnil, 1970.—Afrotropical: Uganda.
Thelairosoma (Seyrigomyia) ingrami Mesnil, 1970b: 103. Holotype male (
longicorne Mesnil, 1954.—Afrotropical: Zimbabwe.
Thelairosoma (Thelairoxenis) longicorne Mesnil, 1954c: 473. Holotype male (BMNH). Type locality: Zimbabwe, Harare [as “Salisbury”].
lutescens Mesnil, 1954.—Afrotropical: Malawi, South Africa, Zimbabwe.
Thelairosoma (Seyrigomyia) lutescens Mesnil, 1954c: 474. Holotype, unspecified sex (BMNH, not located by D. Whitmore, pers. comm.). Type locality: South Africa.
major Mesnil, 1970.—Afrotropical: Madagascar.
Thelairosoma (Seyrigomyia) major Mesnil, 1970b: 102. Holotype male (
melancholicum Mesnil, 1970.—Afrotropical: Madagascar.
Thelairosoma (Seyrigomyia) melancholica Mesnil, 1970b: 102. Holotype male (
obversum Villeneuve, 1943.—Afrotropical: Zimbabwe.
Thelairosoma obversum Villeneuve, 1943a: 40. Syntypes, 3 males (not located). Type locality: Zimbabwe, Harare [as “Salisbury”].
pallidum Mesnil, 1954.—Afrotropical: D.R. Congo, Malawi, Nigeria.
Thelairosoma (Thelairoxenis) pallidum Mesnil, 1954c: 472. Holotype male (
palposum Villeneuve, 1938.—Afrotropical: “W. Afr. to E. Afr. & sthn Afr.” (
Thelairosoma palposum Villeneuve, 1938b: 2. Syntypes, 1 male and 1 female (1 male in
pulchellum (Mesnil, 1944).—Afrotropical: Madagascar.
Seyrigomyia pulchella Mesnil, 1944: 13. Holotype, unspecified sex (
quadriguttatum (Mesnil, 1944).—Afrotropical: Madagascar.
Seyrigomyia quadriguttata Mesnil, 1944: 12. Holotype, unspecified sex [male, see
rosatum Villeneuve, 1943.—Afrotropical: Malawi.
Thelairosoma rosatum Villeneuve, 1943a: 39. Holotype female (not located). Type locality: Malawi, Mt. Mulanje [as “Mt. Mlanje”].
triste Mesnil, 1970.—Afrotropical: Madagascar.
Thelairosoma (Seyrigomyia) tristis Mesnil, 1970b: 102. Holotype male (
varipes Villeneuve, 1943.—Afrotropical: Malawi.
Thelairosoma varipes Villeneuve, 1943a: 39. Syntypes, 3 males and 4 females (not located). Type locality: Malawi.
Genus THELYCONYCHIA Brauer & Bergenstamm, 1889
THELYCONYCHIA Brauer & Bergenstamm, 1889: 89 [also 1890: 21]. Type species: Masicera (Ceromasia) solivaga Rondani, 1861, by monotypy.
TORINAMYIA Mesnil, 1959: 2. Type species: Torinamyia delicatula Mesnil, 1959, by monotypy.
delicatula (Mesnil, 1959).—Afrotropical: Tanzania, Uganda.
Torinamyia delicatula Mesnil, 1959: 2. Holotype male (
solivaga (Rondani, 1861).—Afrotropical: Botswana, U.A. Emirates, Yemen. Palaearctic: C. Asia, Europe (all except British Is., Scand.), Japan, M. East (Israel), Pal. China, Russia (E. Siberia, S. Far East), Transcaucasia. Oriental: Pakistan.
Masicera (Ceromasia) solivaga Rondani, 1861b: 24. Type(s), male (MZF,
Note: Thelyconychia solivaga (Rondani, 1861) of current authors is likely a species complex but is treated here as a single species pending further study.
Genus THELYMYIOPS Mesnil, 1950
THELYMYIOPS Mesnil, 1950b: 10 (as subgenus of Carcelia Robineau-Desvoidy, 1830). Type species: Carcelia coniformis Villeneuve, 1941, by monotypy. Status n.
Note: Thelymyiops Mesnil, 1950 was treated as a subgenus of Carcelia Robineau-Desvoidy, 1830 by
coniformis (Villeneuve, 1941).—Afrotropical: D.R. Congo, Ghana, Tanzania, Uganda.
Carcelia coniformis Villeneuve, 1941b: 124. Holotype female (
Unplaced species of Eryciini
varicornis Curran, 1940.—Afrotropical: Zambia, Zimbabwe.
Phorocera varicornis Curran, 1940: 7. Holotype female (
Note: Phorocera varicornis Curran, 1940 was treated as an “Unplaced species of Goniinae” [= Exoristinae] by
Tribe ETHILLINI
Genus AMNONIA Kugler, 1971
AMNONIA Kugler, 1971: 71. Type species: Amnonia carmelitana Kugler, 1971, by original designation.
carmelitana Kugler, 1971.—Afrotropical: Ethiopia (new record,
Amnonia carmelitana Kugler, 1971: 71. Holotype male (
Note: Amnonia carmelitana Kugler, 1971 is newly recorded from the Afrotropical Region.
deemingi Zeegers, 2010.—Afrotropical: U.A. Emirates.
Amnonia deemingi Zeegers, 2010: 674. Holotype male (
Genus CALLIETHILLA Shima, 1979
CALLIETHILLA Shima, 1979: 147. Type species: Calliethilla caerulea Shima, 1979, by original designation [Oriental].
hirta Cerretti, 2012.—Afrotropical: Uganda.
Calliethilla hirta Cerretti, 2012: 322. Holotype male (
Genus ETHILLA Robineau-Desvoidy, 1863
ETHILLA Robineau-Desvoidy, 1863a: 202. Type species: Tachina aemula Meigen, 1824, by original designation [Palaearctic].
ETHYLLA Mesnil, 1939d: 32. Unjustified emendation of Ethilla Robineau-Desvoidy, 1863 (see
adiscalis Mesnil, 1977.—Afrotropical: Madagascar.
Ethilla adiscalis Mesnil, 1977b: 173. Holotype male (
tenor (Curran, 1927).—Afrotropical: ?Angola, D.R. Congo, ?Kenya, ?Malawi.
Zenillia tenor Curran, 1927c: 5. Holotype male (
Note:
Possibly undescribed sp.: South Africa (BMNH,
Genus ETHYLLOIDES Verbeke, 1970
ETHYLLOIDES Verbeke, 1970: 286. Type species: Ethylloides emdeni Verbeke, 1970, by original designation.
emdeni Verbeke, 1970.—Afrotropical: South Africa.
Ethylloides emdeni Verbeke, 1970: 288. Holotype male (
Genus GYNANDROMYIA Bezzi, 1923
GYNANDROMYIA Bezzi, 1923: 97. Type species: Gynandromyia seychellensis Bezzi, 1923, by original designation.
ZENILLIANA Curran, 1927c: 3 (as subgenus of Zenillia Robineau-Desvoidy, 1830). Type species: Zenillia (Zenilliana) devastator Curran, 1927 (= Myxexorista habilis Brauer & Bergenstamm, 1891), by monotypy.
ZELINDOMYIA Verbeke, 1962a: 166. Type species: Zelindomyia grossa Verbeke, 1962, by original designation.
TRYPHEROSOMA Verbeke, 1962a: 167. Type species: Trypherosoma gilva Verbeke, 1962, by original designation.
Note: Trypherosoma Verbeke, 1962 and Zelindomyia Verbeke, 1962 were synonymized with Gynandromyia Bezzi, 1923 by
bafwankei Verbeke, 1962.—Afrotropical: D.R. Congo.
Gynandromyia bafwankei Verbeke, 1962a: 172. Lectotype male (
Note: The name Gynandromyia bafwankei was made available by
basilewskyi (Verbeke, 1960).—Afrotropical: Tanzania.
Zenilliana basilewskyi Verbeke, 1960: 337. Holotype male (
crypta (Verbeke, 1962).—Afrotropical: D.R. Congo.
Trypherosoma crypta Verbeke, 1962a: 167, 168. Holotype male (
fumigata (Verbeke, 1962).—Afrotropical: D.R. Congo.
Trypherosoma fumigata Verbeke, 1962a: 167, 168. Holotype male (
gilva (Verbeke, 1962).—Afrotropical: D.R. Congo.
Trypherosoma gilva Verbeke, 1962a: 167, 168. Holotype male (
grossa (Verbeke, 1962).—Afrotropical: D.R. Congo.
Zelindomyia grossa Verbeke, 1962a: 167. Holotype male (
habilis (Brauer & Bergenstamm, 1891).—Afrotropical: “widespread W. Afr., E. Afr. & sthn Afr.” (
Myxexorista habilis Brauer & Bergenstamm, 1891: 332 [also 1891: 28] (as “habilis Wd. litt. Coll. Wiedm.”). Type(s), male (
Zenillia (Zenilliana) devastator Curran, 1927c: 3. Holotype female (
Zenillia fuscicosta Curran, 1927c: 4. Holotype male (
invaginata (Villeneuve, 1939).—Afrotropical: D.R. Congo.
Zenilliana devastator invaginata Villeneuve, 1939: 9. Syntypes, 2 females (not located). Type localities: D.R. Congo, Orientale, Bafwakei [as “Bafwankei”, ca. 1°41′N 27°02′E, near Bomili] and Équateur, Irebu.
kibatiana Verbeke, 1962.—Afrotropical: D.R. Congo.
Gynandromyia kibatiana Verbeke, 1962a: 172. Lectotype male (
Note: The name Gynandromyia kibatiana was made available by
mesnili Verbeke, 1962.—Afrotropical: Burundi.
Gynandromyia mesnili Verbeke, 1962a: 172. Holotype male (
Note: The name Gynandromyia mesnili was made available by
prima Verbeke, 1962.—Afrotropical: Ghana, Kenya, Malawi, South Africa, Uganda, Zimbabwe.
Gynandromyia prima Verbeke, 1962a: 172. Lectotype male (
Note: The name Gynandromyia prima was made available by
saegeri Verbeke, 1962.—Afrotropical: D.R. Congo.
Gynandromyia saegeri Verbeke, 1962a: 171. Holotype male (
Note: The name Gynandromyia saegeri was made available by
seychellensis Bezzi, 1923.—Afrotropical: Seychelles.
Gynandromyia seychellensis Bezzi, 1923: 98. Holotype female [not male as published,
Genus MYCTEROMYIELLA Mesnil, 1966
MYCTEROMYIA Mesnil, 1949b: 102. Nomen nudum (proposed after 1930 without designation of type species; no included species) (see
MYCTEROMYIA Mesnil, 1950c: 107 (junior homonym of Mycteromyia Philippi, 1865). Type species: Mycteromyia laetifica Mesnil, 1950, by monotypy (see
MYCTEROMYIELLA Mesnil, 1966: 232 (nomen novum for Mycteromyia Mesnil, 1950).
Undescribed sp.: Angola (BMNH,
Genus NEMORILLOIDES Brauer & Bergenstamm, 1891
NEMORILLOIDES Brauer & Bergenstamm, 1891: 355 [also 1891: 51]. Type species: Nemorilloides flaviventris Brauer & Bergenstamm, 1891, by monotypy.
carbonata Mesnil, 1952.—Afrotropical: D.R. Congo, South Africa.
Nemorilloides carbonata Mesnil, 1952a: 10. Holotype male (
flaviventris Brauer & Bergenstamm, 1891.—Afrotropical: South Africa.
Nemorilloides flaviventris Brauer & Bergenstamm, 1891: 356 [also 1891: 52]. Lectotype female (
Genus PARATRYPHERA Brauer & Bergenstamm, 1891
PARATRYPHERA Brauer & Bergenstamm, 1891: 328 [also 1891: 24]. Type species: Paratryphera handlirschii Brauer & Bergenstamm, 1891 (= Chetina palpalis Rondani, 1859), by monotypy [Palaearctic].
sordida (Villeneuve, 1916).—Afrotropical: Botswana, Kenya, South Africa, Tanzania, Uganda, Yemen.
Zenillia sordida Villeneuve, 1916c: 485. Holotype male (
sordia. Incorrect subsequent spelling of sordida Villeneuve, 1916 (
Note: Paratryphera sordida (Villeneuve, 1916) of current authors is likely a species complex but is treated here as a single species pending further study.
Possibly undescribed spp.: Kenya, South Africa (both records based on specimens in BMNH,
Genus PHOROCEROSOMA Townsend, 1927
PHOROCEROSOMA Townsend, 1927c: 61. Type species: Phorocerosoma forte Townsend, 1927 (= Masicera vicaria Walker, 1856), by original designation [Oriental].
aberrans Verbeke, 1962.—Afrotropical: Rwanda.
Phorocerosoma aberrans Verbeke, 1962a: 170. Holotype female (
Note: The name Phorocerosoma aberrans was made available by
albifacies Verbeke, 1962.—Afrotropical: Cameroon, D.R. Congo.
Phorocerosoma albifacies Verbeke, 1962a: 170. Lectotype female (
Note: The name Phorocerosoma albifacies was made available by
caparti Verbeke, 1962.—Afrotropical: Burundi, D.R. Congo, Tanzania, Uganda.
Phorocerosoma caparti Verbeke, 1962a: 171. Lectotype male (
Phorocerosoma vicina Verbeke, 1962b: 22 (junior objective synonym of Phorocerosoma caparti Verbeke, 1962; both names based on same name-bearing type). Holotype male (
Note: The name Phorocerosoma caparti was made available by
echinum Verbeke, 1962.—Afrotropical: D.R. Congo.
Phorocerosoma echina Verbeke, 1962a: 170. Holotype male (
Note: The name Phorocerosoma echina was made available by