Erwinanillus baehri sp. nov.

Included species strongly characterised by the presence of longitudinal elytral grooves; male protarsi with two dilated tarsomeres and profemora unarmed; metafemora dentate and metatrochanters short and stout in male and female. Labial tooth absent. Head characterised by temples with a series of long excess setae. One elytral discal seta present. Aedeagus with right paramere bearing two apical setae; left bearing only one seta. This genus can be distinguished from Externanilus Baehr & Main, 2016 by the presence of grooves on elytral disc.

Included species of small-medium size (TL mm 1.40–1.42), anophthalmous. Depigmented and poorly sclerified integument covered with a sparse pubescence.

Head robust, almost hypertrophic, slightly narrower than the pronotum, with a series of excess long setae; labium toothless, mentum articulated, not fused with the submentum. Antennae moniliform, without particular features.

Pronotum subquadrate, with sides distinctly sinuate in the basal third. Basal angles subsquare and acuminate; basal border as wide as the anterior border; presence of two marginal setae, the posterior one placed before the basal angles.

Elytra oval and slightly elongated, separately rounded, not truncate and not emarginated apically; poorly convex and with a longitudinal groove. Elytral striae missing (except for the sutural stria). Lateral margin, distinctly crenelated from the humeral area to the apical third. Scutellar pore present, large and umbilicate; umbilicate series of type B (sensu Jeannel 1963; Giachino and Vailati 2011); disc bearing one discal seta.

Legs relatively short and stumpy. Profemora unarmed, metafemora dentate, metatrochanters normal, two dilated protarsomeres in the male.

Aedeagus small, median lobe moderately elongated, apex acuminate with apical blade very evident. Parameres stocky, bearing one apical seta (left) or two apical setae (right). Endophallus without sclerified phanerae.

Dedicated to the memory of Terry Lee Erwin, renowned carabid beetle specialist and explorer of world biodiversity. Name composed of Erwin and the genus name Anillus. The gender of the name is masculine.

For the time being, the genus is monotypic and only E. baehri sp. nov. belongs to this genus.

Figures 1–3. 

Erwinanillus baehri gen. et sp. nov., HT ♂. 1 habitus 2 right metafemur and metatrochanter in ventral view 3 aedeagus in left lateral view. Scale bars: 0.1 mm.

WA, Southern Goldfields region, Forrestania, 80 km E of Hyden, Cosmic Boy Mine, 32°29'19"S, 119°40'49"E.

HT ♂, WA, Southern Goldfields region, Forrestania, 80 km E of Hyden, Cosmic Boy Mine, 32°29'19"S, 119°40'49"E, 06 April 2009, Rockwater, bore WWMB4, hauling. Western Australian Museum Entomology Reg. no. 72025 (WAM). PTT: 1 ♀ (remains), WA, Southern Goldfields region, Forrestania, 80 km E of Hyden, Cosmic Boy Mine, 32°29'18.84"S, 119°40'48.69"E (WGS84), Rockwater, 6 April 2009, Stygo net haul. (RW-WWMB4-LN5105), Western Australian Museum Entomology Reg. no. 82637 (WAM); 1 spec. (remains), WA, Southern Goldfields region, Forrestania, 80 km E of Hyden, Cosmic Boy Mine, 32°29'18.84"S, 119°40'48.69"E (WGS84), Rockwater, 6 April 2009, Stygo net haul. (RW-WWMB4-LN5105), Western Australian Museum Entomology Reg. no. 82638 (CGi).

Small species, with longitudinal elytral grooves; male protarsi with two dilated tarsomeres and profemora unarmed; metafemora dentate and metatrochanters short and stout in male and female. Head characterised by temples with a series of excess long setae. One elytral discal seta present. Aedeagus with right paramere bearing two apical setae; left bearing only one seta.

TL 1.40–1.42 mm. Body elongated, depigmented, fulvo-testaceous with elytra and abdomen lighter, yellow-testaceous; integument shiny with evident microsculpture and spread pubescence.

Head robust, almost hypertrophic, slightly narrower than the pronotum. Labium toothless, mentum articulate. Antennae robust, moniliform, short, not exceeding the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; anterior margin of the epistome subrectilinear.

Pronotum subquadrate (max. width / max. length ratio = 1.05), with the maximum width on the anterior fourth, narrowed at the base, with the sides poorly arcuate anteriorly, sinuate before the base. Anterior angles rounded, weakly prominent; posterior angles subsquare and acuminate at tips. Disc convex, with a long and sparse pubescence; median groove very shallow, hardly evident. Marginal groove wide and flat, slightly enlarged near the base; anterior marginal setae placed inside the marginal groove, almost on the anterior fifth; basal setae before the posterior angles.

Legs short and stout, two protarsomeres dilated and without adhesive phanerae in male. Metatrochanters short and stout, metafemora dentate (Fig. 2).

Elytra oval, slightly elongated (max. length / max. width ratio = 1.72), with maximum width in the middle, not emarginated before apex. Disc poorly convex, provided with an evident longitudinal groove running more or less between the scutellar pore and the 9^th pore of the umbilicate series; integument shiny, with evident microsculpture and short, dense, upright pubescence. Humeri poorly marked, rounded. Post-humeral margin denticulate, with a very thin but distinct crenulation to the apical third; elytral apices separately rounded. Marginal groove wide and evident almost to the 8^th pore of the umbilicate series.

Chaetotaxy : scutellar pore large, foveate. Umbilicate series with the first three pores of the humeral group not equidistant, with 1^st and 2^nd closer together than 2^nd and 3^rd; 4^th pore clearly farther and placed at the end of the basal third of the elytron; 5^th pore placed before the apical third of the elytra; 5^th and 6^th ones spaced out ca. half distance between the 6^th and 7^th; 7^th slightly and 8^th nearly moved onto the disc; 7^th and 8^th slightly closer to each other than the 8^th and 9^th. One discal seta, placed before the 7^th pore of the umbilicate series.

Aedeagus (Fig. 3) small, abruptly arcuate in the basal part; median lobe moderately elongated with ventral margin gently arcuate up to the acuminate apex, with apical blade very evident. Endophallus without an evident lamella copulatrix. Parameres stocky in the basal part and relatively poorly elongated, not reaching the distal third. Right paramere bearing two apical setae; left bearing only one seta.

Dedicated to the memory of Martin Baehr, renowned beetle taxonomist, in honour of his contributions to the knowledge of Australian ground beetles.

Erwinanillus baehri sp. nov. is known only from bore WWMB4, 80 km E of Hyden, Forrestania, Southern Goldfields region, WA.

Gracilanillus longulus Baehr & Main, 2016.

Baehr and Main’s description (2016) is correct and detailed, although we noted a few inaccuracies and their description omits some important characters fundamental to the systematics of Anillini which are described as follows. Baehr and Main (2016) describe three setae on the elytral disc of Gracilanillus genus, and later in the paper they specify that G. longulus Baehr & Main, 2016, G. cockingi Baehr & Main, 2016 and G. cordatus Baehr & Main, 2016 bear only two setae, G. vixsulcatus Baehr & Main, 2016 bear only one seta, and that in G. minutus Baehr & Main, 2016 and G. currani Baehr & Main, 2016 have none. The two additional Gracilanillus species that we describe herein bear two setae, and we suggest that considering the difficulty of observing all setae if specimens are not examined using the methods as described in this paper, possibly all species in this genus share the character “two setae on the elytral disc”.

Baehr and Main (2016) refer to a “faint” labial tooth, while the two species described here do not possess this character. The labial tooth is not an important character at genus level (Giachino 2005; Giachino and Vailati 2011), and also it is very difficult to see if the specimen is not properly prepared. Baehr and Main (2016) do not mention the position of the elytral umbilicate setae, but only talk about “very long setae”. The position of these setae is fundamental (Jeannel 1963; Giachino 2005; Giachino and Vailati 2011), hence we specify the position, type “B” (sensu Jeannel 1963; Giachino and Vailati 2011). For additional details see descriptions.

Seven species currently belong to this genus:

Gracilanillus hirsutus sp. nov.

Gracilanillus pannawonicanus sp. nov.

Gracilanillus longulus Baehr & Main, 2016

Gracilanillus cockingi Baehr & Main, 2016

Gracilanillus cordatus Baehr & Main, 2016

Gracilanillus vixsulcatus Baehr & Main, 2016

Gracilanillus currani Baehr & Main, 2016

Figures 4–7. 

Gracilanillus spp.: habitus HT ♀ (4, 6) left metafemur and metatrochanter in ventral view HT ♀ (5, 7) 4, 5 G. hirsutus sp. nov. 6, 7 G. pannawonicanus sp. nov. Scale bars: 0.1 mm.

WA, Pilbara, 44 km W of Pannawonica, Robe Valley, Mesa A Mine, 21°39'52"S, 115°53'48"E.

HT ♀, WA, Pilbara, 44 km W of Pannawonica, Robe Valley, Mesa A Mine (bore MEARC4038), 21°39'52"S, 115°53'48"E (GPS), 10 June–3 Aug. 2006; litter trap L. Mould, M. Greenham; Western Australian Museum Entomology Reg. no. 64216 (WAM).

Gracilanillus hirsutus sp. nov. can be distinguished from G. pannawonicanus sp. nov. in the presence of long and sparse pubescence longitudinally aligned on the elytral disc, head with many excess setae, and for the standard position of the ninth pore of the umbilicate series.

TL 1.39 mm. Body elongated, depigmented, yellow-testaceous; shiny integument with evident microsculpture and pubescence.

Head robust, hypertrophic, slightly narrower than pronotum; bearing a number of excess setae. Labium toothless, mentum articulate. Antennae robust, moniliform, and short, not exceeding the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; anterior margin of the epistome subrectilinear.

Pronotum subquadrate (max. width / max. length ratio = 1.05), with maximum width in the middle, and with basal border as wide as anterior border; sides slightly but regularly arcuate from anterior to posterior, not sinuate, but strictly emarginated before the basal angles. Anterior angles rounded, poorly prominent; posterior angles subsquare. Disc convex, with long and sparse pubescence; median groove very shallow, hardly evident. Marginal groove wide and flat, slightly enlarged near the base; anterior marginal setae placed inside marginal groove, almost on anterior fifth; basal setae almost before the posterior angles.

Legs short and stout, with metatrochanters short and stout and metafemora not dentate (Fig. 5).

Elytra subrectangular, elongated (max. length / max. width ratio = 1.84), with maximum width at the base of the posterior third, not emarginated before apex. Disc slightly convex, with evident longitudinal groove running more or less between the scutellar pore and the 9^th pore of the umbilicate series; integument shiny with evident microsculpture and long, sparse, upright pubescence, longitudinally aligned. Humeri well marked; post-humeral margin denticulate, with distinct crenulations all the way to the apical third; elytral apices separately rounded. Marginal groove wide and evident almost up to the 9^th pore of the umbilicate series.

Chaetotaxy : scutellar pore large, foveate. Umbilicate series with first three pores of humeral group equidistant; 4^th pore farther and placed at the end of the basal fourth of elytron; 5^th pore placed before the apical third of elytra; 5^th and 6^th ones spaced out ca. half distance of 6^th and 7^th; 8^th slightly displaced on the disc; 7^th and 8^th closer to each other than 8^th and 9^th. Two discal setae, first placed before the 5^th pore of the umbilicate series, second one placed just before 8^th pore.

Male. Unknown.

The name comes from the Latin hirsutus (= hairy, hirsute) to recognise the presence of many excess setae on the head.

Gracilanillus hirsutus sp. nov. occurs only at the type locality in Mesa A Mine, Robe River Valley, 44 km W of Pannawonica, Pilbara, WA.

WA, Pilbara, 11 km SSE of Pannawonica, Robe Valley, Mesa K Mine, 21°43'11"S, 116°15'43"E.

HT ♀, WA, Pilbara, 11 km SSE of Pannawonica, Robe Valley, Mesa K Mine, (bore MEK1701), 21°43'11"S, 116°15'43"E (GPS), 10 June-3 Aug. 2006; litter trap L. Mould, M. Greenham; Western Australian Museum Entomology Reg. no. 64215 (WAM).

Gracilanillus pannawonicanus sp. nov. can be distinguished from G. hirsutus for the presence of a short and dense pubescence on the elytral disc, not directly longitudinally aligned, for the absence of excess setae on the head, and for the position of the ninth pore of the umbilicate series, which is placed quite forward, before the end of the elytral groove.

TL 1.43 mm. Body elongated, depigmented, yellow-testaceous; shiny integument with evident microsculpture and pubescence.

Head robust, almost hypertrophic, slightly narrower than pronotum, excess setae absent. Labium toothless, articulated mentum. Antennae robust, moniliform, short, not exceeding the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; anterior margin of the epistome subrectilinear.

Pronotum subquadrate (max. width / max. length ratio = 1.07), maximum width at the basal anterior third, and basal border as wide as anterior border; sides slightly but regularly arcuate from anterior to posterior angles, strictly sinuate just before basal angles. Anterior angles rounded, slightly prominent; posterior angles subsquare and acuminate at vertex. Disc convex, with short and very sparse pubescence; median groove very shallow, hardly evident. Marginal groove wide and flat, slightly enlarged near the base; anterior marginal setae placed inside the marginal groove, almost on the anterior fifth; basal setae nearly before posterior angles.

Legs short and stout, with metatrochanters short and stout and metafemora non dentate (Fig. 7).

Elytra subrectangular, elongated (max. length / max. width ratio = 1.95), with maximum width in the middle, not emarginated in preapical zone. Disc slightly convex, with evident longitudinal groove running more or less between the scutellar pore and the 2^nd discal pore; integument shiny, with evident microsculpture, and short, very dense, and upright pubescence, not longitudinally aligned. Humeri well marked but rounded; post-humeral margin denticulate, with distinct crenulation up to the apical fourth; elytral apices separately and acutely rounded. Marginal groove wide and evident almost all the way to the 8^th pore of the umbilicate series.

Chaetotaxy : scutellar pore large, foveate. Umbilicate series with the first three pores of the humeral group equidistant; 4^th pore farther and placed at the end of the basal third of the elytron; 5^th pore placed before the apical third of the elytra; 5^th and 6^th ones spaced out ca. half of the distance between 6^th and 7^th; 8^th slightly displaced onto the disc; 7^th and 8^th slightly closer to each other than the 8^th and 9^th; 9^th pore of the umbilicate series placed well forward (compare to the standard position). Two discal setae, first placed just after the 5^th pore of the umbilicate series, second one placed just after the 8^th pore.

Male. Unknown.

The name of the species derives from the town Pannawonica near the type locality Mesa K Mine, in the Pilbara region.

Gracilanillus pannawonicanus sp. nov. is known so far only from the type locality Mesa K Mine, in the Robe River Valley, 11 km SSE of Pannawonica, Pilbara, WA.

Gregorydytes ophthalmianus sp. nov.

Genus characterised by normal metatrochanters, elytra reduced at tip and lacking longitudinal grooves, 9^th pore of the umbilicate series in normal position (placed after the 8^th one) and two discal setae; basal border of pronotum remarkably narrower than anterior border and sides distinctly sinuate before the basal angles; aedeagus with median lobe subrectilinear, basal bulb extremely reduced and parameres bearing one apical seta. Labial tooth lacking.

Species small (TL mm 1.02–1.05) and anophthalmous. Integument depigmented, poorly sclerified, and covered with sparse pubescence.

Head of normal size, narrower than pronotum; mandibles short and simple, without hyperplasias. Maxillary palpi ovoidal, swollen. Labium transverse, articulated; mentum not fused with submentum. Labial tooth absent. Antennae moniliform, without particular features.

Pronotum slightly transverse, with sides distinctly sinuate at the basal third. Basal angles obtuse and rounded; basal border remarkably narrower than anterior border; presence of two marginal setae, the posterior one placed at the basal angles.

Elytra subrectangular and elongated, separately rounded, depressed and without longitudinal groove, and slightly truncated but not emarginated at apex. Elytral striae absent (except sutural stria). Lateral margin starting from the humeral area, distinctly crenulate to at least half-length. Scutellar pore present, large and umbilicate; umbilicate series of type B (sensu Jeannel 1963; Giachino and Vailati 2011); disc bearing two discal setae.

Legs relatively short and stumpy. Unarmed pro- and metafemora, normal metatrochanters, two dilated protarsomeres in the male.

Aedeagus small, median lobe short and subrectilinear with basal bulb extremely reduced. Parameres long, bearing one apical seta. Endophallus without sclerified phanerae.

Gregorydytes is a compound noun: in honour of Francis Thomas Gregory, the first European explorer who discovered Ophthalmia Range (type locality) in 1876, and dytes meaning diver in Greek. Gender name masculine.

Only G. ophthalmianus sp. nov. belongs to this new genus.

Figures 8, 9. 

Gregorydytes ophthalmianus gen. et sp. nov., HT ♂ 8 habitus 9 aedeagus in lateral view. Scale bars: 0.1 mm.

WA, Pilbara, 25 km W of Newman, Ophthalmia Range, 23°16'41.39"S, 119°24'13.28"E.

HT ♂, WA, Pilbara, 25 km W of Newman, Ophthalmia Range, 23°16'41.39"S, 119°24'13.28"E (WGS84), P. Bell, N. Coen, 25 June 2010, Trog. net scrape; (BHP020_EXR1691-LN:9124), Western Australian Museum Entomology Reg. no. 82611 (WAM). PTT: 1 ♀, WA, Pilbara, 25 km W of Newman, Ophthalmia Range, 23°16'41.39"S, 119°24'13.28"E (WGS84), P. Bell, N. Coen, 14 August 2010, Trog. net scrape (BHP020_EXR1691-LN:9362), Western Australian Museum Entomology Reg. no. 82612 (CGi).

Gregorydytes ophthalmianus sp. nov. differs from other Australian Anillini by the characters highlighted in the genus diagnosis.

TL mm 1.02 ♂–1.05 ♀. Body moderately elongated, depigmented, yellow-testaceous; integument shiny, with evident microsculpture and pubescence.

Head robust, not hypertrophic, narrower than pronotum; excess setae absent. Labium toothless, mentum articulated. Antennae robust, moniliform, short, hardly exceeding the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; subrectilinear anterior margin of epistome.

Pronotum transverse (max. width / max. length ratio = 1.25), with maximum width at the base of the anterior fourth, and basal border remarkably narrower than anterior border; sides distinctly and regularly arcuate anteriorly, sinuate before basal angles. Anterior angles rounded, not prominent; posterior angles obtuse and rounded. Disc convex, with very sparse pubescence of medium length; median groove very shallow, hardly evident. Marginal groove wide and flat, slightly enlarged near the base; anterior marginal setae placed inside the marginal groove, almost on the anterior fourth; basal setae at posterior angles.

Legs short and stout, with metatrochanters short and stout and metafemora non dentate. Two protarsomeres dilated and without adhesive phanerae in male.

Elytra subrectangular (max. length / max. width ratio = 1.55), with maximum width at the base of the posterior third, subtruncate but not emarginated before apex. Disc slightly convex, without longitudinal groove; integument shiny, with evident microsculpture and very sparse and upright pubescence of medium length, longitudinally aligned. Humeri well marked but rounded; post-humeral margin denticulate, with distinct crenulation up to the apical third; elytral apices separately rounded. Marginal groove wide and evident almost up to the 9^th pore of the umbilicate series.

Chaetotaxy : large and foveate basal umbilicate pore. Umbilicate series with the first three pores of the humeral group very closed to each other and equidistant; 4^th pore farther and placed at the end of the basal third of the elytron; 5^th pore placed at the base the apical third of the elytron; 5^th and 6^th ones spaced from each other equidistant with 6^th and 7^th; 5^th, 7^th and 8^th displaced onto the disc; 7^th and 8^th spaced from each other as 8^th and 9^th. Two discal setae, the first placed at the level of the 3^rd pore of the umbilicate series, the second one placed just before the 8^th pore.

Aedeagus (Fig. 9) small, median lobe short, stout, subrectilinear, with basal bulb extremely reduced; ventral margin gently bisinuated; apical blade evident. Endophallus without an evident lamella copulatrix. Left paramere elongated, reaching the aedeagal distal fourth and bearing only one seta; right paramere lost during the preparation of the specimen.

The name arises from the type locality Ophthalmia Range in the Pilbara region.

Gregorydytes ophthalmianus sp. nov. is known so far only from the type locality Ophthalmia Range, 25 km W of Newman, Pilbara, WA.

Pilbaraphanus chichesterianus sp. nov.

Genus characterised by very long and sharp metatrochanters; dentate metafemora; elytra not reduced at tip; elytral disc bearing three setae and longitudinal grooves absent; 9^th pore of the umbilicate series in the normal position (placed after the 8^th); aedeagus with median lobe not very curved, size of basal bulb normal and parameres bearing two apical setae. Labial tooth present and smooth. Differs from Magnanillus Baehr, 2017 by the absence of longitudinal elytral grooves.

Species of medium size (TL mm 1.37–2.18) and anophthalmous. Integument depigmented but well sclerified and covered with sparse pubescence.

Head from normal to large size, slightly narrower than pronotum; mandibles short and simple, without hyperplasias. Maxillary palpi ovoidal and swollen. Labium transverse and articulated; mentum not fused with submentum. Labial tooth present and smooth. Antennae moniliform, without particular features.

Pronotum subquadrate, with sides not or only slightly sinuate at the basal third, smooth or denticulated before basal angles. Basal angles right, sharp, not rounded; basal border as wide as, or only slightly narrower, than the anterior border; presence of two marginal setae, the posterior one placed just before basal angles.

Elytra subrectangular and elongated, separately rounded and depressed, not truncate and not emarginated apically, with longitudinal groove absent. Elytral striae absent (except sutural stria). Lateral margin starting from humeral area, distinctly crenulate up to at least half-length. Scutellar pore present, large and umbilicate; umbilicate series of type B (sensu Jeannel 1963; Giachino and Vailati 2011); disc bearing three discal setae.

Legs relatively short and stumpy. Unarmed profemora; posterior edge of metafemora dentate, metatrochanters very long and sharp, two dilated protarsomeres in the male.

Aedeagus relatively large, median lobe long and not very curved, with basal bulb of normal size. Parameres long and bearing two apical setae. Endophallus with poorly sclerified phanerae.

The genus name combines the Pilbara name with the suffix -phanus to recognise the genus Illaphanus. Gender masculine.

Only two species currently belong to this genus:

Pilbaraphanus chichesterianus sp. nov.

Pilbaraphanus bilybarianus sp. nov.

Figures 10–14. 

Pilbaraphanus gen. et spp. nov., habitus (10, 13) right metafemur and metatrochanter in ventral view (11, 14) aedeagus in lateral view (12) 10–12 P. chichesterianus sp. nov., HT ♂ 13, 14 P. bilybarianus sp. nov. HT ♀. Scale bars: 0.1 mm.

WA, Pilbara, 100 km E of Port Hedland, Chichester Ranges, Quarry 8, 21°59'43.58"S, 119°0'38"E.

HT ♂, WA, Pilbara, 100 km E of Port Hedland, Chichester Ranges, Quarry 8, 21°59'43.58"S, 119°0'38"E, (WGS84), P. Bell, 3 Jun. 2008, Stygo Net Haul. (Q8-EXR1622R-LN1735), Western Australian Museum Entomology Reg. no. 82635 (WAM).

Pilbaraphanus chichesterianus sp. nov. and P. bilybarianus sp. nov. are closely related and share the characters indicated in the genus diagnosis. P. chichesterianus sp. nov. differs from P. bilybarianus sp. nov. by its bigger body size, shorter metatrochanters, and more transverse pronotum.

TL mm 2.18. Body elongated and depigmented, yellow-testaceous; integument shiny, with evident microsculpture and short pubescence.

Head robust, hypertrophic, slightly narrower than pronotum; without excess setae. Labium with a smooth tooth, mentum articulated. Antennae robust, moniliform, short, not reaching the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; anterior margin of the epistome subrectilinear.

Pronotum subsquare (max. width / max. length ratio = 1.12), with maximum width at the base of the anterior fifth, and with basal border remarkably wider than anterior border; sides slightly and irregularly arcuate in the anterior part, subrectilinear before basal angles. Anterior angles acute, prominent; posterior angles squared and not rounded. Disc convex, with very sparse pubescence of medium length; median groove very shallow, hardly evident. Marginal groove wide and flat, slightly enlarged near the base; anterior marginal setae placed inside the marginal groove, almost on the anterior fifth; basal setae placed inside the disk and before posterior angles.

Legs long and slender, with metatrochanters long and acuminate, but not curved, and metafemora dentate; metatrochanters (Fig. 11) shorter than femoral tooth. Two protarsomeres dilated and without adhesive phanerae in males. Left pro-, left and right meso-, and left metalegs missing in the HT ♂.

Elytra perfectly subrectangular (max. length / max. width ratio = 1.80), not truncate and not emarginated before apex. Disc convex, longitudinal grooves absent; integument shiny, with evident microsculpture, and very short, sparse and upright, pubescence longitudinally aligned. Humeri well marked, gently rounded; post-humeral margin denticulate, with distinct crenulation up to the apical third; elytral apices separately rounded. Marginal groove wide and evident almost up to the 8^th pore of the umbilicate series.

Chaetotaxy : scutellar pore large and foveate. Umbilicate series with the first three pores of the humeral group very closed to each other and equidistant; 4^th pore farther and placed at the end of the basal third of the elytron; 5^th pore placed before the base of the apical third of the elytron; 5^th and 6^th ones spaced from each other half the distance from 6^th and 7^th; 7^th and 8^th displaced onto the disc; 7^th and 8^th spaced to each other as the 8^th and 9^th. Three discal setae, the first placed before the 4^th pore of the umbilicate series, the second and third ones placed respectively just before the 5^th and 8^th umbilicate pores.

Aedeagus (Fig. 12) large, median lobe long, stout, subrectilinear, with basal bulb extremely reduced; ventral margin gently curved; apical blade evident but short. Endophallus without an evident lamella copulatrix, but with two crossing apical slightly sclerified stripes. Left paramere elongated, reaching the aedeagal distal third and bearing two setae; right paramere lost during the preparation of the specimen.

The species name comes from the Chichester Range, where the type locality (Quarry 8) is located.

Pilbaraphanus chichesterianus sp. nov. is known so far only from the type locality Quarry 8 in the Chichester Range, 100 km E of Port Hedland, Pilbara, WA.

WA, Pilbara, 60 km N of Tom Price, Solomon Mining Area, Kings deposit, 22°09'31.44"S, 117°51'50.9E.

HT ♀, WA, Pilbara, 60 km N of Tom Price, Solomon Mining Area, Kings deposit, 22°09'31.44"S, 117°51'50.9E (WGS84), P. Bell, E.S. Volschenk, 24.Jan. 2010; Trog. net scrape (FMG005_SM0347_10:7877 Western Australian Museum Entomology Reg. no. 82607 (WAM).

Pilbaraphanus bilybarianus sp. nov. and P. chichesterianus sp. nov. are closely related and share the characters described in the genus diagnosis. P. bilybarianus sp. nov. differs from P. chichesterianus sp. nov. by its smaller body size, longer metatrochanters, and less transverse pronotum.

TL mm 1.37. Body elongated and depigmented, yellow; integument shiny, with evident microsculpture and short pubescence.

Head robust, hypertrophic, narrower than pronotum; excess setae absent. Labium with smooth tooth, mentum articulated. Antennae robust, moniliform, short, reaching the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; anterior margin of the epistome subrectilinear.

Pronotum subsquare (max. width / max. length ratio = 1.03), with the maximum width at the base of the anterior fourth, and with basal border remarkably wider than anterior border; sides poorly and not regularly arcuate in the anterior part, gently sinuate in the basal half and slightly dentate before basal angles. Anterior angles obtuse, prominent; posterior angles squared, gently rounded. Disc convex, with very sparse pubescence of medium length; median groove very shallow, hardly evident. Marginal groove wide and flat, enlarged near the base; anterior marginal setae placed inside the marginal groove, almost on the anterior fourth; basal setae not placed inside on the disk, but before the posterior angles.

Legs long and slender, with metatrochanters long and acuminate, gently curved and metafemora dentate; metatrochanters (Fig. 14) as long as femoral tooth. All left legs missing in the HT ♀.

Elytra perfectly subrectangular (max. length / max. width ratio = 1.91), not truncated, only slightly emarginated before the apex. Disc convex, longitudinal grooves absent; integument shiny with evident microsculpture, and very short, sparse, upright pubescence not longitudinally aligned. Humeri well marked, gently rounded; post-humeral margin denticulate, with distinct crenulation up to the apical third; elytral apices separately rounded. Marginal groove wide and evident almost up to the 9^th pore of the umbilicate series.

Chaetotaxy : scutellar pore large and foveate. Umbilicate series with the first three pores of the humeral group very closed to each other and equidistant; 4^th pore farther and placed at the end of the basal third of the elytron; 5^th pore placed before the base of the apical third of the elytron; 5^th and 6^th ones spaced from each other as half distance from 6^th and 7^th; 7^th and 8^th displaced onto the disc; 7^th and 8^th spaced from each other as the 8^th and 9^th. Three discal setae, the first placed before the 4^th pore of the umbilicate series, the second one placed just before the 5^th, and the third one placed at the level of the 7^th pore of the umbilicate series.

Male. Unknown.

The species name derives from Bilybara, aboriginal name that refers to the Pilbara region.

Pilbaraphanus bilybarianus sp. nov. is known only from the type locality (Kings deposit, which is part of the Solomon Mining Area), 60 km N of Tom Price, Pilbara, WA.

Macranillus bennetti Baehr & Main, 2016: 61.

This genus is characterised by: metatrochanters very long and sharp; metafemora dentate; elytra not reduced at tip; elytral disc with longitudinal grooves and bearing three setae; 9^th pore of the umbilicate series in normal position (placed after the 8^th one); aedeagus with median lobe gently curved, size of basal bulb normal and parameres bearing two apical setae. Labial tooth lacking. Differs from Pilbaraphanus by the presence of a longitudinal elytral grooves and the lack of labial tooth.

The description of this genus by Baehr and Main (2016) (sub Macranillus), although very accurate, does not consider some peculiar characters present in species belonging to Macranillus, in particular: the elytral chaetotaxy, the absence of labial tooth, the presence of teeth on the posterior margin of metafemora, and the elongated and sharpened shape of the metatrochanters. The presence of the last two characters in Magnanillus was confirmed (M. Baehr pers. comm. 2018). The genus is redescribed below.

Species of medium-large size (TL mm 1.44–2.51) and anophthalmous. Integument depigmented but well sclerified and covered with sparse pubescence.

Head size normal to large, well narrower, or just narrower, than pronotum; mandibles short and simple, without hyperplasias. Maxillary palps ovoidal, swollen. Labium transverse, articulated; mentum not fused with submentum. Labial tooth absent. Antennae moniliform, without particular features.

Pronotum subquadrate, with sides not, or only slightly, sinuate at the basal third, smooth or denticulated before basal angles. Basal angles right or obtuse, sharp, not rounded; base as wide as, or only slightly narrower, than anterior margin; presence of two marginal setae, the posterior one placed just before the basal angles.

Elytra subrectangular and elongated, separately rounded, not truncated and not emarginated apically; depressed, with a longitudinal groove. Elytral striae absent (except sutural stria). Lateral margin, starting from the humeral area and distinctly crenulate up to at least half-length. Scutellar pore present, large and umbilicate; umbilicate series of type B (sensu Jeannel 1963; Giachino and Vailati 2011); disc bearing three discal setae.

Legs relatively short and stumpy. Profemora unarmed; metafemora dentate on the posterior edge, metatrochanters very long and sharp, two dilated protarsomeres in the male.

Aedeagus relatively large, median lobe long and slightly curved, with basal bulb of normal size. Parameres long, each bearing two apical setae. Endophallus with poorly sclerified phanerae.

Currently ten species belong to this genus:

Magnanillus firetailianus sp. nov.

Magnanillus quartermaini (Baehr & Main, 2016)

Magnanillus sabae sp. nov.

Magnanillus salomonis sp. nov.

Magnanillus regalis sp. nov.

Magnanillus serenitatis sp. nov.

Magnanillus bennetti (Baehr & Main, 2016)

Magnanillus pearsoni (Baehr & Main, 2016)

Magnanillus maini (Baehr & Main, 2016)

Magnanillus magnus (Baehr & Main, 2016)

The key to species of Magnanillus Baehr, 2017 lack clear information on important diagnostic characters such as metafemora and metatrochanter morphology (see genus redescription). For the identification of the described species of Magnanillus refer to Baehr and Main (2016).

WA, Pilbara, 50 km N of Tom Price, Solomon Mining Area, Firetail deposit, 22°07'25.3"S, 117°53'59.2"E.

HT ♂, WA, Pilbara, 50 km N of Tom Price, Solomon Mining Area, Firetail Mine (drill hole FT0430), 22°07'25.3"S, 117°53'59.2"E (WGS84); G. Pearson, D. Main, 20 May 2010; Trog. Trap., Western Australian Museum Entomology Reg. no. 82657 (WAM). PTT: 6 ♂♂ 3 ♀♀ (+ remains of 1 ♂ 2 ♀♀), WA, Pilbara, 50 km N of Tom Price, Solomon Mining Area, Firetail Mine (drill hole FT0430), 22°07'25.3"S, 117°53'59.2"E (WGS84), G. Pearson, D. Main, 20 May 2010, Trog. Trap., Western Australian Museum Entomology Reg. no. 82658–82662 (WAM, CGi); 1 ♀, WA, Pilbara, 50 km N of Tom Price, Solomon Mining Area, Firetail Mine (drill hole FT1455), 22°08'53.1"S, 117°55'37.8"E (WGS84) G. Pearson, D. Main, 19 May 2010, Trog. Trap., Western Australian Museum Entomology Reg. no. 82663 (WAM).

Magnanillus firetailianus sp. nov. is easily distinguishable from the other species of this genus (except M. quartermaini (Baehr & Main, 2016)) by its pronotum with basal border narrower than anterior border. It also differs from M. sabae sp. nov., M. salomonis sp. nov. and M. regalis n. sp by much longer metatrochaters, extended over the position of the femoral tooth. M. firetailianus sp. nov. differs from M. serenitatis sp. nov. by its bigger size, while it differs from M. quartermaini by a less transverse pronotum.

TL mm 1.44–1.46 ♂♂, 1.48–1.50 ♀♀. Body elongate, depigmented, yellow; integument shiny, with evident microsculpture, and short pubescence.

Head robust, hypertrophic, slightly narrower than pronotum; without excess setae. Labium without tooth, mentum articulated. Antennae robust, moniliform, short, reaching the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; subrectilinear anterior margin of epistome.

Pronotum sub-rectangular (max. width / max. length ratio = 1.14), with maximum width at the base of the anterior fifth, and with basal border remarkably narrower than anterior border; sides slightly and irregularly arcuate in the anterior part, subrectilinear at the basal half, gently sinuate and slightly dentate before basal angles. Anterior angles obtuse, slightly prominent; posterior angles squared, acute. Disc convex, with very sparse pubescence of medium length; median groove very shallow, hardly evident. Marginal groove wide and flat, enlarged near the base; anterior marginal setae placed inside the marginal groove, almost on the anterior fifth; basal setae not inside the disk, but placed before posterior angles.

Legs long and slender, with metatrochanters long and acuminate, gently curved and metafemora dentate; metatrochanters (Fig. 16) slightly longer than femoral tooth. Two dilated protarsomeres in males.

Elytra perfectly subrectangular (max. length / max. width ratio = 1.80), not truncated and not emarginated before the apex. Disc convex, with longitudinal grooves; integument shiny, with evident microsculpture, very short, sparse, and upright, pubescence, not longitudinally aligned. Humeri well marked, gently rounded; post-humeral margin denticulate, with distinct crenulation up to the apical third; elytral apices separately rounded. Marginal groove wide and evident almost up to the 8^th pore of the umbilicate series.

Chaetotaxy : scutellar pore large and foveate. Umbilicate series with the first three pores of the humeral group very closed to each other and equidistant; 4^th pore farther and placed at the end of the basal third of the elytron; 5^th pore placed before the base of the apical third of the elytra; 5^th and 6^th ones spaced out ca. half of the distance between 6^th and 7^th; 7^th and 8^th displaced onto the disc; 7^th and 8^th spaced from each other as the 8^th and 9^th. Three discal setae, the first placed before the 4^th pore of the umbilicate series, second one placed just before the 5^th, the third one placed at the 7^th pore.

Aedeagus (Fig. 17) large, median lobe long, stout, gently curved, with basal bulb small, but tight and evident; ventral margin gently curved from basal bulb to apex; apical blade evident but short. Endophallus without an evident lamella copulatrix, but with two small, apical, Y-shaped, crossing and slightly sclerified stripes. Left paramere elongated, not reaching the distal third and bearing two setae; right paramere shorter and bearing two apical setae.

The name comes from the type locality in the Solomon Mining Area, the Firetail deposit, in the Pilbara region.

Magnanillus firetailianus sp. nov. is known only from the type locality (Firetail deposit) 50 km N of Tom Price, Pilbara, WA.

Figures 15–17. 

Magnanillus firetalianus sp. nov., HT ♂ 15 habitus 16 right metafemur and metatrochanter in ventral view 17 aedeagus in lateral view. Scale bars: 0.1 mm.

WA, Pilbara, 60 km NW of Tom Price, Solomon Mining Area, Serenity deposit, 22°8'4.02"S, 117°32'3.444E.

HT ♂, WA, Pilbara, 60 km NW of Tom Price, Solomon Mining Area, Serenity mine, (drill hole SMD0057), 22°8'4.02"S, 117°32'3.444E (WGS84), N. Coen, S. Catomore, 19 Apr. 2011, Trog. net scrape, Western Australian Museum Entomology Reg. no. 82651 (WAM). PTT: 1 ♀, WA, Pilbara, 60 km NW of Tom Price, Solomon Mining Area, Serenity mine, (drill hole SR0126), 22°11'22.70"S, 117°32'28.68E (WGS84), N. Coen, S. Catomore, 30 May 2011, Trog. net scrape, Western Australian Museum Entomology Reg. no. 82652 (WAM); 1 ♀, WA, Pilbara, 60 km NW of Tom Price, Solomon Mining Area, Sheila Valley, (drill hole SV0577), 22°13'1.236"S, 117°40'55.2E (WGS84), N. Coen, S. Catomore, 18 Apr. 2011, Trog. Net scrape, Western Australian Museum Entomology Reg. no. 82640 (WAM);1 ♂, WA, Pilbara, 60 km NW of Tom Price, Solomon Mining Area, Sheila Valley, (drill hole SV0443), 22°14'45.59"S, 117°38'46.71E (WGS84), S. Eberhard, S. Catomore, 05 Oct. 2010, Stygo net haul., Western Australian Museum Entomology Reg. no. 82615 (CGi); 1 ♀, WA, Pilbara, 50 km N of Tom Price, Solomon Mining Area, Zion mine, (drill hole SM0282) 22°12'00.8"S, 117°57'16.71E (WGS84), E.S. Volschenk, N. Krawczyk, 01.Mar. 2010, Trog. Trap, Western Australian Museum Entomology Reg. no. 82610 (WAM).

Magnanillus. sabae sp. nov. can be easily distinguished from M. firetailianus sp. nov. by its base of pronotum ca. as large as the pronotal apex. It can be distinguished from M. salomonis sp. nov., M. regalis sp. nov., and M. serenitatis sp. nov. by its short metatrochanters not reaching the femoral tooth. It differs from M. pearsoni (Baehr & Main, 2016) by longer elytra and more transverse pronotum.

TL mm 2.42–2.46 ♂♂, 2.48–2.51 ♀♀. Body elongated, depigmented, testaceous; integument shiny, with evident microsculpture and very short pubescence.

Head robust, narrower than pronotum, excess setae absent. Labium without tooth, mentum articulated. Antennae robust, moniliform, short, reaching the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; anterior margin of the epistome subrectilinear.

Pronotum sub-squared (max. width / max. length ratio = 1.14), with maximum width at the base of the anterior fourth, and basal border slightly wider than anterior border; sides slightly and irregularly arcuate in the anterior part, subrectilinear in the basal half, not sinuate and slightly denticulate before basal angles. Anterior angles obtuse, slightly prominent; posterior angles obtuse, blunted. Disc convex, with very short and sparse pubescence; median groove very shallow, hardly evident. Marginal groove wide and flat, enlarged near the base; anterior marginal setae placed inside the marginal groove, almost on the anterior fifth; basal setae not inside on the disk, but placed before the posterior angles.

Legs long and slender, with metatrochanters long, acuminate and subrectilinear and metafemora dentate; metatrochanters (Fig. 19) shorter than femoral tooth. Two dilated protarsomeres in males.

Elytra subrectangular, relatively short (max. length / max. width ratio = 1.63), not truncated and very slightly emarginated before apex. Disc convex, with longitudinal grooves; very short, sparse, and upright pubescence not longitudinally aligned. Humeri well marked, obtuse; post-humeral margin denticulate, with a distinct crenulation down to the apical third; elytral apices separately rounded. Marginal groove wide and evident almost up to the 7^th pore of the umbilicate series.

Chaetotaxy : scutellar pore large and foveate. Umbilicate series with the first three pores of the humeral group very closed to each other and equidistant; 4^th pore farther and placed at the end of the basal third of the elytron; 5^th pore placed before the base of the apical third of the elytron; 5^th and 6^th ones spaced out ca. 1/3 of the distance from 6^th and 7^th; 7^th and 8^th displaced onto the disc; 7^th and 8^th spaced from each other as the 8^th and 9^th. Three discal setae, first placed before the 4^th pore of the umbilicate series, second one placed just before the 5^th, third one placed before the 7^th pore.

Aedeagus (Fig. 20) large, median lobe long, stout, gently curved, with basal bulb small but evident; ventral margin gently curved from basal bulb to apex; apical blade poorly evident, very short. Endophallus without an evident lamella copulatrix, but with two small, apical, subparallel, and slightly sclerified stripes. Left paramere elongate, not reaching the aedeagal distal third and bearing two setae; right paramere shorter and bearing two apical setae.

The name is to remind one of the Solomon mining area and the different deposits where this species occurs, and it originates from the mythological Queen of Sheba (in Latin Sheba = Saba). According to tradition, the Queen of Sheba visited the Kingdom of Solomon with valuable gifts for its king.

Magnanillus sabae sp. nov. is known from different deposits of the Solomon Mining Area, 50–60 km N/NW of Tom Price, Pilbara, WA.

Figures 18–20. 

Magnanillus sabae sp. nov., HT ♂ 18 habitus 19 right metafemur and metatrochanter in ventral view 20 aedeagus in lateral view. Scale bars: 0.1 mm.

WA, Pilbara, 50 km N of Tom Price, Solomon Mining Area, Kings deposit, 22°12'00.8"S, 117°57'16.71E.

HT ♂, WA, Pilbara, 50 km N of Tom Price, Solomon Mining Area, Kings mine, 22°12'00.8"S, 117°57'16.71E (WGS84), E.S. Volschenk, N. Krawczyk, 01.March 2010, Trog. trap (FMG005_SM0282_10:8313), WA Museum Entomology Reg. no. 82609 (WAM). PTT: 1 ♂ (remains) 6 ♀♀ and 3 spec. (♂♀?),WA, Pilbara, 60 km NW of Tom Price, Solomon Mining Area, Serenity mine, 22°11'9.24"S, 117°32'49.524E (WGS84), N. Coen, S. Catomore, 20.04.2011, Stygo net haul (FMG008_SOM0039_11:0879 Western Australian Museum Entomology Reg. no. 82641–82650 (WAM, CGi); 1 ♀, WA, Pilbara, 50 km N of Tom Price, Solomon Mining Area, Firetail Mine, 22°07'32.2"S, 117°29'34.6"E (WGS84), G. Pearson and D. Main, 13"Sept 2010, Stygo. net haul (HPRC0211) Western Australian Museum Entomology Reg. no. 82664 (WAM); 1 ♂ 1 ♀ (remains) 1 spec. (remains), WA, Pilbara, 50 km N of Tom Price, Solomon Mining Area, Firetail Mine, 22°06'44.2"S, 117°53'28.8"E (WGS84), G. Pearson and D. Main, 3 Mar. 2010, Trog. trap (FT0541) Western Australian Museum Entomology Reg. no. 82654–82656 (WAM); 1 ♀ (remains), WA, Pilbara, 60 km NW of Tom Price, Solomon Mining Area, Sheila Valley, 22°12'00.39"S, 117°42'14.52E (WGS84), M. Weerheim, S. Catomore, 9 Dec. 2010, Trog. net scrape, (FMG006_SV0267_10:0955), Western Australian Museum Entomology Reg. no. 82613 (WAM); 1 spec. (remains), WA, Pilbara, 60 km NW of Tom Price, Solomon Mining Area, Sheila Valley, 22°14'45.59"S, 117°38'46.71E (WGS84), S. Eberhard, S. Catomore, 05 Oct. 2010, Stygo net haul. (FMG006_SV0443_10:0490), Western Australian Museum Entomology Reg. no. 82614 (WAM).

Magnanillus salomonis sp. nov. is easily distinguishable from M. firetailianus sp. nov. by its protonum with basal border ca. as wide as the anterior border. It can be distinguished from M. sabae sp. nov. by its longer metatrochanters, reaching the femoral tooth. It can be distinguished from M. serenitatis sp. nov. by its shorter metatrochanters, not overreaching the femoral tooth. It can be distinguished from M. regalis sp. nov. by its more transverse pronotum, and the clearly curved apex of the metatrochanters. It can be distinguished from M. benneti (Baehr & Main, 2016), M. pearsoni (Baehr & Main, 2016), M. maini (Baehr & Main, 2016), M. magnus (Baehr & Main, 2016) and M. quartermaini (Baehr & Main, 2016) by its more transverse pronotum.

TL mm 2.20–2.22 ♂♂, 2.25–2.28 ♀♀. Body elongated, depigmented, testaceous; integument shiny with evident microsculpture and very short pubescence.

Head relatively small, narrower than pronotum; excess setae absent. Labium without tooth, mentum articulated. Antennae robust, submoniliform, short, reaching the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; anterior margin of the epistome subrectilinear.

Pronotum transverse (max. width / max. length ratio = 1.16), with maximum width at the base of the anterior fourth, and basal border wider than anterior border; sides slightly and irregularly arcuate in anterior part, subrectilinear in the basal half, not sinuate and denticulate before basal angles. Anterior angles obtuse, slightly prominent; posterior angles right, acute. Disc convex, with short and very sparse pubescence; median groove very shallow, hardly evident. Marginal groove wide and flat, enlarged near the base; anterior marginal setae placed inside the marginal groove, almost on the anterior fourth; basal setae slightly located internally on the disk and placed before the posterior angles.

Legs long and slender, with metatrochanters long, acuminate, and gently curved and metafemora dentate; metatrochanters (Fig. 22) as long as the femoral tooth. Anterior legs missing in all male specimens.

Elytra subrectangular, relatively short (max. length / max. width ratio = 1.69), not truncate and not emarginated before the apex. Disc convex, with longitudinal grooves; integument shiny with evident microsculpture, very short, sparse, and upright, pubescence, not longitudinally aligned. Humeri well marked, obtuse; post-humeral margin denticulate, with distinct crenulations up to the base of the apical third; elytral apices separately rounded. Marginal groove wide and evident almost up to the 8^th pore of the umbilicate series.

Chaetotaxy : scutellar pore large and foveate. Umbilicate series with the first three pores of the humeral group very closed to each other and equidistant; 4^th pore farther and placed at the end of the basal third of the elytron; 5^th pore placed at the base of the apical third of the elytron; 5^th and 6^th ones spaced out ca. half of the distance between 6^th and 7^th; 7^th and 8^th displaced onto the disc; 7^th and 8^th spaced from each other as the 8^th and the 9^th. Three discal setae, first placed before the 4^th pore of the umbilicate series, second one placed in the middle of the elytron, the third one placed before the 7^th pore.

Aedeagus (Fig. 23) large, median lobe long, stout, gently curved, with basal bulb small but tight and evident; ventral margin gently curved from basal bulb to apex; apical blade poorly evident, very short. Endophallus without an evident lamella copulatrix, but with two very small, apical, and slightly sclerified, ovoidal areas. Left paramere elongate, reaching the distal third and bearing two setae; right paramere shorter and bearing two apical setae.

The name comes from type locality “Solomon mining area” and the mythological King of Solomon (in Latin Salomon).

Magnanillus salomonis sp. nov. is known from different deposits of the Solomon Mining Area, 50–60 km N/NW of Tom Price, Pilbara, WA.

Figures 21–23. 

Magnanillus salomonis sp. nov., HT ♂ 21 habitus 22 left metafemur and metatrochanter in dorsal view 23 aedeagus in lateral view. Scale bars: 0.1 mm.

WA, Pilbara, 50 km N of Tom Price, Solomon Mining Area, Kings deposit, 22°07'45.5"S, 117°52'24.3"E.

HT ♂, WA, Pilbara, 50 km N of Tom Price, Solomon Mining Area, Kings Mine, 22°07'45.5"S, 117°52'24.3"E (WGS84), G. Pearson and D. Main, 12 Jan 2010. Trog. net scrape (SM3175), Western Australian Museum Entomology Reg. no. 82653 (WAM). PTT: 1 ♂ 2 ♀♀ (remains), WA, Pilbara, 50 km N of Tom Price, Solomon Mining Area, Kings Mine, 22°07'45.5"S, 117°52'24.3"E (WGS84), G. Pearson and D. Main, 12 Jan 2010. Trog. net scrape (SM3175), Western Australian Museum Entomology Reg. no. 82653 (WAM, CGi).

Magnanillus regalis sp. nov. is easily distinguishable from M. firetailianus sp. nov. and M. quartermaini (Baehr & Main, 2016) by its pronotum with basal border ca. as wide as the anterior border. It can be distinguished from M. sabae sp. nov. by its longer metatrochanters, reaching the femoral tooth. It differs from M. serenitatis sp. nov. by its shorter metatrochanters, not overreaching the femoral tooth. It can be distinguished from M. salomonis sp. nov. by its less transverse pronotum and the straight apex of its metatrochaters.

TL mm 2.25. Body elongate, depigmented, yellow-testaceous; integument shiny with evident microsculpture and short pubescence.

Head relatively large, narrower than pronotum; with two couples of excess setae on the vertex as in Fig. 24. Labium without tooth, mentum articulated. Antennae robust, submoniliform, very short, not reaching the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; anterior margin of the epistome subrectilinear.

Pronotum sub-squared (max. width / max. length ratio = 1.14 maximum width at the base of the anterior fourth, and basal border slightly wider than anterior border; sides slightly and irregularly arcuate in anterior part, subrectilinear at the basal half, not sinuate but with an evident tooth before basal angles. Anterior angles obtuse, slightly prominent; posterior angles right, acute. Disc convex, with very sparse pubescence of medium length; median groove very shallow, hardly evident. Marginal groove wide and flat, enlarged near the base; anterior marginal setae placed inside the marginal groove, almost on the anterior fourth; basal setae slightly placed internally on the disk and before the posterior angles.

Legs long and slender, with metatrochanters long, acuminate, subrectilinear and metafemora dentate; metatrochanters (Fig. 25) reaching the femoral tooth. Two dilated protarsomeres, without adhesive phanerae in males.

Elytra subrectangular, elongate (max. length / max. width ratio = 1.83), not truncated and only slightly emarginated before apex. Disc convex, with longitudinal grooves; integument shiny, with evident microsculpture and very short, longitudinally aligned, upright pubescence. Humeri well marked, obtuse; post-humeral margin denticulate, with distinct crenulations up to the base of the apical third; elytral apices separately rounded. Marginal groove wide and evident almost up to the 8^th pore of the umbilicate series.

Chaetotaxy : scutellar pore large and foveate. Umbilicate series with the first three pores of the humeral group very closed to each other and equidistant; 4^th pore farther and placed at the end of the basal third of the elytron; 5^th pore placed at the base the apical third of the elytron; 7^th pore very forward, placed near the 6^th pore; 6^th and 7^th pores closer than 5^th and 6^th; 8^th displaced onto the disc; 7^th and 8^th spaced out ca. the 5^th and 6^th. Three discal setae, first placed before the 4^th pore of the umbilicate series, second one placed just before the 5^th pore, third one placed after the 7^th pore.

Aedeagus (Fig. 26) large, median lobe long, slender, gently curved, with basal bulb small but tight and evident; ventral margin gently curved from basal bulb to apex; apical blade poorly evident, short. Endophallus without an evident lamella copulatrix, but with small, Y-shaped, apical, slightly sclerified stripe. Left paramere elongate, reaching the distal third and bearing two setae; right paramere shorter and bearing two apical setae.

The name comes from the Latin word regalis = royal, and it reminds the type locality “King” deposit in the Solomon Mining Area.

Magnanillus regalis sp. nov. is known only from the type locality (Kings deposit) in the Solomon Mining Area, 50 km N of Tom Price, Pilbara, WA.

Figures 24–26. 

Magnanillus regalis sp. nov., HT ♂ 24 habitus 25 right metafemur and metatrochanter in ventral view 26 aedeagus in lateral view. Scale bars: 0.1 mm.

WA, Pilbara, 65 km NW of Tom Price, Serenity mining area, Champion deposit, 22°07'5.1"S, 117°26'12.5"E.

HT ♂, WA, Pilbara, 65 km NW of Tom Price, Serenity mining area, Champion deposit, 22°07'5.1"S, 117°26'12.5"E (WGA84), J. Quartermaine and D. Main, 13 Jun 2011, Trog. trap (HPRC0712), Western Australian Museum Entomology Reg. no. 82667 (WAM). PTT: 2 ♂♂, WA, Pilbara, 50 km N of Tom Price, Solomon Mining Area, Firetail Mine, 22°09'0.0"S, 117°28'55.3"E (WGS84), J. Cocking, D. Main, 23 June 2010, Trog. net scrape (HPRC0243), Western Australian Museum Entomology Reg. no. 82665–82666 (WAM, CGi); 1 ♀, WA, Pilbara, 65 km NW of Tom Price, Serenity mining area, Delta deposit, 22°08'31.0"S, 117°28'1.5"E (WGS84), J. Cocking, D. Main, 19 August 2010, Trog. trap (HPRC2004), Western Australian Museum Entomology Reg. no. 82668 (WAM).

Magnanillus serenitatis sp. nov. is easily distinguishable from M. firetailianus sp. nov. by its pronotum with basal border as wide as the anterior border. It can be distinguished from M. sabae sp. nov., M. salomonis sp. nov. and M. regalis sp. nov. by its longer metatrochanters, overreaching the femoral tooth.

TL mm 2.27–2.30 ♂♂, 2.32 ♀. Body elongate, depigmented, testaceous; integument shiny, with evident microsculpture and very short pubescence.

Head relatively small, narrower than pronotum; without excess setae. Labium without tooth, mentum articulated. Antennae robust, submoniliform, short, reaching the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; anterior margin of the epistome subrectilinear.

Pronotum transverse (max. width / max. length ratio = 1.23), maximum width at the base of the anterior third, and basal border slightly wider than anterior border; sides slightly and irregularly arcuate in anterior part, very poorly arcuate in the basal half, not sinuate and denticulate before the basal angles. Anterior angles obtuse, slightly prominent; posterior angles right, acute. Disc convex, with very short and sparse pubescence; median groove very shallow, hardly evident. Marginal groove wide and flat, enlarged near the base; anterior marginal setae placed inside the marginal groove, almost on the anterior fourth; basal setae slightly placed inside the disk and before the posterior angles.

Legs long and slender, with metatrochanters long, acuminate, and curved and metafemora dentate; metatrochanters (Fig. 28) longer than femoral tooth. Two dilated protarsomeres, without adhesive phanerae in males.

Elytra subrectangular, relatively short (max. length / max. width ratio = 1.67), not truncated and only very slightly emarginated before apex. Disc convex, with longitudinal grooves; shiny integument with evident microsculpture and very short, sparse, upright pubescence, not longitudinally aligned. Humeri well marked but rounded; post-humeral margin denticulate, with distinct crenulations up to the base of the apical fourth; elytral apices separately rounded. Marginal groove wide and evident almost down to the 7^th pore of the umbilicate series.

Chaetotaxy : scutellar pore large and foveate. Umbilicate series with the first three pores of the humeral group very closed to each other and nearly equidistant; 4^th pore farther and placed at the end of the basal third of the elytron; 5^th pore placed at the base of the apical third of the elytron; 5^th and 6^th ones spaced out ca. half of the distance between the 6^th and the 7^th; 7^th and 8^th displaced onto the disc; 7^th and 8^th spaced from each other as the 8^th and 9^th. Three discal setae, first placed before the 4^th pore of the umbilicate series, second one placed after the midpoint of the elytron, third one placed before the 7^th pore.

Aedeagus (Fig. 29) large, median lobe long, slender, gently curved, with basal bulb small but tight and evident; ventral margin gently curved from basal bulb to apex; apical blade poorly evident, very short. Endophallus without an evident lamella copulatrix, but with very small, V-shaped, apical, slightly sclerified stripe. Left paramere elongate, reaching the distal third and bearing two setae; right paramere shorter and bearing two apical setae.

The name comes from the Serenity Valley, type locality of the species, in the Pilbara region.

Magnanillus serenitatis sp. nov. is known only from two drill holes in the Serenity Valley, 65 km N/NW of Tom Price, Pilbara, WA.

Figures 27–29. 

Magnanillus serenitatis sp. nov., HT ♂ 27 habitus 28 right metafemur and metatrochanter in ventral view 29 aedeagus in lateral view. Scale bars: 0.1 mm.

Neoillaphanus callawanus sp. nov.

Genus characterised by: metatrochanters long and sharp; metafemora non dentate; elytra not reduced at tip; elytral disc with longitudinal grooves and bearing one seta; 9^th pore of the umbilicate series in normal position (placed after the 8^th one); aedeagus with median lobe curved, size of the basal bulb normal and parameres bearing two apical setae. Labial tooth absent.

Species of medium size (TL mm 1.93–2.04) and anophthalmous. Integument depigmented but well sclerified and covered with sparse pubescence.

Head large but narrower than pronotum; mandibles short and simple, without hyperplasias. Maxillary palpi ovoidal, swollen. Labium transverse, articulated; mentum not fused with submentum. Labial tooth absent. Antennae moniliform, without particular features.

Pronotum cordiform, sides sinuate at the basal third, denticulated before basal angles. Basal angles right, sharp at tips, not rounded; basal border narrower than anterior border; presence of two marginal setae, the posterior one placed before basal angles.

Elytra ovoidal short, separately rounded, not truncated and apically not emarginated; convex, with longitudinal grooves. Elytral striae absent (except sutural stria). Lateral margin starting from the humeral area, distinctly crenulate up to its half-length.

Scutellar pore present, large and umbilicate; umbilicate series of type B (sensu Jeannel 1963; Giachino and Vailati 2011); disc bearing one seta.

Legs relatively long and slender. Pro- and metafemora unarmed; metatrochanters long and sharp, two dilated protarsomeres in the male.

Aedeagus relatively large, median lobe long and curved with basal bulb of normal size. Parameres long, each of them bearing two apical setae. Endophallus with very small and poorly sclerified phanerae.

The name is derived by the combination of the Greek prefix Neo- (means new) and the name Illaphanus. Gender masculine.

Currently only one species belongs to this genus: N. callawanus sp. nov.

Figures 30–33. 

Neoillaphanus callawanus gen. et sp. nov. 30 habitus HT ♂ 31 right metafemor and metatrochanter in ventral view, HT ♂ 32 aedeagus in lateral view, HT ♂ 33 aedeagus in lateral view, PT ♂. Scale bars: 0.1 mm.

WA, Pilbara, 200 km E of Port Hedland, Goldsworthy Mining Area, Callawa deposit, 20°38'16.74"S, 120°18'14.33"E.

HT ♂, WA, Pilbara, 200 km E of Port Hedland, Goldsworthy Mining Area, Callawa Mine (drill hole CA0013R), 20°38'16.74"S, 120°18'14.33"E (WGS84), 29 Jul. 2008, P. Bell, Trog trap, Western Australian Museum Entomology Reg. no. 82631 (WAM). PTT: 1 ♂, WA, Pilbara, 200 km E of Port Hedland, Goldsworthy Mining Area, Callawa Mine (drill hole CA0011R) 20°38'16.74"S, 120°18'14.33"E (WGS84), 9 July 2009, P. Bell, Trog net scrape, Western Australian Museum Entomology Reg. no. 82629 (WAM); 1 ♀, WA, Pilbara, 200 km E of Port Hedland, Goldsworthy Mining Area, Callawa Mine (drill hole CA0021R), 20°38'16.74"S, 120°18'14.33"E (WGS84), 29 July 2008, P. Bell, Stygo net haul, Western Australian Museum Entomology Reg. no. 82628 (WAM); 1 ♂, WA, Pilbara, 200 km E of Port Hedland, Goldsworthy Mining Area, Callawa Mine (drill hole CA0011R), 20°38'16.74"S, 120°18'14.33"E (WGS84), 12 June 2009, P. Bell, Stygo net haul, Western Australian Museum Entomology Reg. no. 82630 (CGi); 1 ♀, WA, Pilbara, 200 km E of Port Hedland, Goldsworthy Mining Area, Callawa Mine (drill hole CA0102R), 20°38'16.74"S, 120°18'14.33"E (WGS84), 29 April 2008, P. Bell, Stygo net haul, Western Australian Museum Entomology Reg. no. 82627 (WAM); 1 ♀, WA, Pilbara, 200 km E of Port Hedland, Goldsworthy Mining Area, Callawa Mine (drill hole CA0124R), 20°38'53"S, 114°17'57"E, 28 July 2008, Subterranean Ecology, hauling, Western Australian Museum Entomology Reg. no. 72011 (CGi); 1 ♂, WA, Pilbara, 200 km E of Port Hedland, Goldsworthy Mining Area, Callawa Mine (drill hole CA0008R), 20°38'34"S, 120°18'02"E, 31 May 2009, Subterranean Ecology, hauling, Western Australian Museum Entomology Reg. no. 72012 (WAM); 1 ♀, WA, Pilbara, 200 km E of Port Hedland, Goldsworthy Mining Area, Callawa Mine, 20°38'46"S, 120°17'50"E, 12 June 2009, Subterranean Ecology, bore CA0019R, scraping, Western Australian Museum Entomology Reg. no. 72013 (WAM).

Species identified by the characters listed in the genus diagnosis.

TL mm 1.93–1.97 ♂♂, 2.00–2.04 ♀♀. Body elongated, depigmented, testaceous; integument shiny, with evident microsculpture and medium length pubescence.

Head large, narrower than pronotum; without excess setae. Labium without tooth, mentum articulated. Antennae robust, moniliform, short, reaching the base of the pronotum when stretched backwards. Fronto-clypeal furrow distinct; anterior margin of the epistome subrectilinear.

Pronotum cordiform (max. width / max. length ratio = 1.13), maximum width at the base of the anterior third, and basal border narrower than anterior border; sides slightly and regularly arcuate in anterior part, subrectilinear at the basal half, sinuate and denticulate before basal angles. Anterior angles obtuse, prominent; posterior angles right, sharp at tips. Disc convex, with sparse pubescence of medium length; median groove very shallow, hardly evident. Marginal groove wide and flat, poorly enlarged near the base; anterior marginal setae placed inside the marginal groove, almost on the anterior fifth; basal setae not on the disk and placed before posterior angles.

Legs long and slender, with metatrochanters long and acuminate but not curved and metafemora unarmed; metatrochanters (Fig. 31) as long as 2/3 of the femoral length. Two poorly dilated protarsomeres, without adhesive phanerae in males.

Elytra ovoidal, relatively short (max. length / max. width ratio = 1.8), maximum width at the base of the posterior third, not truncated and not emarginated in preapical zone. Disc convex, with longitudinal grooves; integument shiny with evident microsculpture and pubescence of medium length, sparse and upright, not longitudinally aligned. Humeri rounded; post-humeral margin denticulate, with distinct crenulations up to half-length; elytral apices separately rounded. Marginal groove wide and evident almost up to the 7^th pore of the umbilicate series.

Chaetotaxy : scutellar pore large, foveate. Umbilicate series with the first three pores of the humeral group very closed to each other and nearly equidistant; 4^th pore farther and placed at the end of the basal third of the elytron; 5^th pore placed just before the base the apical third of the elytron; 5^th and 6^th ones spaced out ca. half of the distance between 6^th and 7^th; 7^th and 8^th displaced onto the disc; 7^th and 8^th spaced out ca. as the 8^th and 9^th. One discal seta placed just before the 7^th pore of the umbilicate series.

Aedeagus (Figs 32, 33) large, median lobe long, slender, curved, with basal bulb tight and evident; ventral margin regularly curved from basal bulb to apex; apical blade evident, but short. Endophallus without an evident lamella copulatrix, but with very small, preapical, slightly sclerified stripe. Left paramere elongate, reaching the distal third of median lobe and bearing two setae; right paramere shorter and bearing two apical setae.

The name comes from the Callawa Ridge (type locality) in the NE of the Pilbara region.

Neoillaphanus callawanus sp. nov. is known only from a few drill holes on the Callawa Ridge, 200 km E of Port Hedland, Pilbara, WA.

Kimberleytyphlus carrboydianus sp. nov.

Genus of the “Illaphanus phyletic series” (sensu Giachino 2005), with species strongly characterised by normal metatrochanters; metafemora non dentate; elytra not reduced at tip; elytral disc without longitudinal grooves and bearing three setae, scaly microsculpture and 9^th pore of the umbilicate series in normal position (placed after the 8^th one); aedeagus with median lobe subrectilinear, size of basal bulb normal and parameres each bearing one apical seta. Labial tooth absent.

Species of a medium size (TL mm 1.57–1.59) and anophthalmous. Integument depigmented but well sclerified, with strong microsculpture and covered with short and sparse pubescence.

Head large but slightly narrower than pronotum; mandibles short and simple, without hyperplasias. Maxillary palpi ovoidal, swollen. Labium transverse, articulated; mentum not fused with the submentum. Labial tooth absent. Antennae moniliform, without particular features.

Pronotum trapezoidal, with sides not sinuate in the basal third, denticulated at the basal third. Basal angles obtuse, sharp, not rounded; basal border slightly narrower than anterior border; presence of two marginal setae, the posterior one placed near basal angles.

Elytra subrectangular elongate, separately rounded, not truncated and apically only slightly emarginated, convex, without longitudinal grooves. Elytral striae absent (except sutural stria). Lateral margin starting from the humeral area and distinctly crenulate up to the level of 8^th–9^th pores of the umbilicate series. Scutellar pore present, large and umbilicate; umbilicate series of type B (sensu Jeannel 1963; Giachino and Vailati 2011); disc bearing three setae.

Legs relatively long and slender. Pro- and metafemora unarmed; metatrochanters normal, two slightly dilated protarsomeres, without adhesive phanerae in males.

Aedeagus relatively small, median lobe long, subrectilinear with basal bulb of normal size. Parameres long, bearing one apical seta. Endophallus without any sclerified phanerae.

The name combines the name Kimberley (region where the type locality is located) and the Greek suffix -typhlos (meaning blind).

Currently only K. carrboydianus sp. nov. belongs to this genus.

Figures 34, 35. 

Kimberleytyphlus carrboydianus gen. et sp. nov., HT ♂ 34 habitus 35 aedeagus in lateral view. Scale bars: 0.1 mm.

WA, Kimberley, Carr-Boyd Ranges, 530 km SE Darwin, 16°37'26.02"S, 128°15'32.36"E.

HT ♂, WA, Carr-Boyd Ranges, 530 km SE Darwin, Kimberley, WA, 16°37'26.02"S, 128°15'32.36"E (WGS84), Animal Plant Mineral (APM), 12 August 2009, Trog. net scrape (APM-KMGTROG 7-LN7274), Western Australian Museum Entomology Reg. no. 82632 (WAM). PTT: 1 ♂, WA, Kimberley, Carr-Boyd Ranges, 530 km SE Darwin, 16°37'52.59"S, 128°15'32.83"E (WGS84), Animal Plant Mineral (APM), 12 Aug. 2009, Trog. net scrape (APM-KMGTROG 8-LN7276), Western Australian Museum Entomology Reg. no. 82633 (CGi).

Species identified by the characters listed in the genus diagnosis.

TL mm 1.57–1.59 ♂♂. Body elongate, depigmented, testaceous; integument with medium length pubescence. Microsculpture evident and very strong: composed by isodiametric meshes on pronotal disc; scaly on basal part of elytral disc, and longitudinally oriented hollow points on apical elytral disc (Fig. 34).

Head large, slightly narrower than pronotum; without excess setae. Labium without tooth, mentum articulated. Antennae robust, moniliform, short, reaching the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; anterior margin of the epistome subrectilinear.

Pronotum trapezoidal (max. width / max. length ratio = 1.00), maximum width at the base of the anterior fourth, with basal border narrower than anterior border; sides subrectilinear, anteriorly slightly but regularly arcuate, not arcuate and not sinuate at the basal half, denticulated at the basal third. Anterior angles obtuse, rounded and not prominent; posterior angles obtuse, but evident. Disc convex, with sparse pubescence of medium length; median groove very shallow, hardly evident. Marginal groove wide and flat, enlarged near the base; anterior marginal setae placed inside the marginal groove, almost on the anterior fourth; basal setae not on the disk and placed at posterior angles.

Legs long and slender, with metatrochanters normal and metafemora unarmed. Two poorly dilated protarsomeres, without adhesive phanerae in males.

Elytra subrectangular, very elongated (max. length / max. width ratio = 2.20), maximum width at the base of the posterior third, not truncated and slightly emarginated before apex. Disc convex, without longitudinal grooves; integument with sparse and upright pubescence, longitudinally aligned. Humeri well marked; post-humeral margin denticulate, with distinct crenulations up to the 8^th and 9^th pores of the umbilicate series; elytral apices separately rounded. Marginal groove wide and evident almost up to the 9^th pore of the umbilicate series.

Chaetotaxy : scutellar pore small and foveate. Umbilicate series with the first three pores of the humeral group very closed to each other and nearly equidistant; 4^th pore farther and placed at the end of the basal third of the elytron; 5^th pore placed just after the middle of the elytron; 5^th and 6^th ones spaced out ca. 1/4 of the distance between 6^th and 7^th; 7^th and 9^th placed onto the disc; 7^th and 8^th spaced out ca. double of the distance from 8^th and 9^th; 9^th placed at the level of the 8^th pore. Three discal setae, first placed at the level of the 2^nd pore of the umbilicate series, second one placed at the level of the base of the posterior fourth of elytron, third one placed just before the level of the 9^th pore.

Aedeagus (Fig. 35) relatively small, median lobe long, slender, subrectilinear, with basal bulb tight and evident; ventral margin weakly curved from basal bulb to apex; apical blade evident, but short. Endophallus without any sclerified phanerae. Left paramere very elongated, reaching the distal fifth of the median lobe, and bearing only one seta; right paramere shorter than left one, bearing one apical seta.

The name derives from the name of one of the early European explorers to visit the region named as the Carr-Boyd Ranges (type locality). In 1883 William Henry James Carr-Boyd became second-in-command of an expedition led by W. J. O’Donnell on behalf of the Cambridge Downs Pastoral Association; their purpose was to explore the country around the Cambridge Gulf, and to establish a sheep station. The party of six men, including a cook and an Aboriginal boy, twenty-six horses and provisions for six months, left Katherine on 26 March 1883. O’Donnell named the impressive Carr Boyd Range after his second-in-command on 26 May.

Kimberleytyphlus carrboydianus sp. nov. occurs only at the type locality Carr-Boyd Ranges, 530 km SE Darwin, Kimberley, WA.

Austranillus macleayi (Lea, 1906)

Genus of the “Illaphanus phyletic series” (sensu Giachino 2005), with species characterised by: absence of longitudinal elytral grooves, pronotum with basal border wider than anterior border and denticulate sides before basal angles, elytral disc with two setae, 8^th pore of the umbilicate series after the 9^th (sensu Giachino and Vailati 2011), labium toothless, median lobe of the aedeagus long and weakly curved, and parameres very long and slender.

In the original description of the genus Giachino (2005) does not mention the forward position of the 9^th pore of the umbilicate series. The omission was due to the lost seta corresponding to the 9^th pore, and to the extended length of secondary setae (1^st, 3^rd, 4^th, 5^th, 7^th, and 8^th) in A. macleayi. The study of the umbilicate series of A. jinayrianus sp. nov. indicates, according to Giachino and Vailati (2011) that the 8^th pore is actually placed backwards in respect to the 9^th.

Currently, two species belong to this genus:

Austranillus macleayi (Lea, 1906)

Austranillus jinayrianus sp. nov.

Figures 36, 37. 

Austranillus jinayrianus sp. nov., HT ♂ 36 habitus 37 aedeagus in lateral view. Scale bars: 0.1 mm.

WA, Pilbara, 110 km NW of Newman, Area C Mining Area, Jinayri deposit, 22°58'22.4"S, 119°15'37.9"E.

HT ♂, WA, Pilbara, 110 km NW of Newman, Area C Mining Area, Jinayri Mine, 22°58'22.4"S, 119°15'37.9"E (WGS84), J. Cocking and M. Scanlon, 25 Nov. 2008. Trog. trap (JIN0744), Western Australian Museum Entomology Reg. no. 82674 (WAM).

Very large species (TL mm 2.98), easily distinguishable from A. macleayi by its larger size (mm 2.98 vs. mm 1.76 in A. macleayi), by the position of the 4^th pore of the umbilicate series placed further, and by the position of the anterior discal pore which is placed at the base of the posterior third of elytra.

TL mm 2.98. Body elongated, depigmented, fulvo-testaceous; integument shiny, with evident microsculpture and sparse and short pubescence.

Head very small, much narrower than pronotum. Labium toothless. Antennae long and robust, not moniliform (with relatively elongated antennomeres), considerably exceeding the base of the pronotum when stretched backwards. Fronto-clypeal furrow indistinct; anterior margin of the epistome subrectilinear.

Pronotum slightly transverse (max. width / max. length ratio = 1.13) with maximum width at the anterior third, and a very wide basal border, wider than the anterior border. Pronotum sides anteriorly clearly arcuate, weakly sinuate and denticulate before the base, base laterally distinctly emarginated. Anterior angles rounded, not prominent. Posterior angles protruding, acute at the vertex. Disc slightly convex, with a short and relatively dense pubescence; median groove very shallow, slightly marked. Marginal groove relatively wide and flat, very enlarged near the base; anterior marginal setae inserted inside the marginal groove, on approximately the anterior third; basal setae inserted almost on the posterior angles.

Legs long and slender, with metatrochanters normal and metafemora unarmed. Two asymmetrically dilated protarsomeres, with one row of adhesive phanerae in male.

Elytra subrectangular, very elongated (max. length/max. width ratio = 1.77), with maximum width in the middle, very slightly emarginated before apex; sides slightly sinuate at the base of the anterior fourth. Disc poorly convex; integument shiny, with evident microsculpture and short, dense, upright pubescence. Humeri extremely marked, almost right angle; post-humeral margin denticulate, with a distinct crenulation up to the base of the apical third of the elytron; elytral apices separately rounded. Marginal groove wide and evident up to over the 9^th pore of the umbilicate series.

Chaetotaxy : scutellar pore large, foveate. Umbilicate series with pores of the humeral group not equidistant; 4^th pore slightly displaced towards the disc, not so far from the third pore and inserted just on the basal fourth of the elytron; 5^th pore placed just before the apical third of the elytron; 5^th and 6^th pores spaced out, ca. 2/3 of the 6^th and 7^th; 7^th, 8^th and 9^th pores almost equidistant and slightly displaced onto the disc, 8^th pore placed after the 9^th one. Two discal setae, first placed at the base of the apical third, second one placed at the level of the 9^th pore.

Aedeagus (Fig. 37) large, median lobe long, slender, subrectilinear, with basal bulb poorly evident; ventral margin poorly and gently curved from the basal bulb to the apex; apical blade evident, but short and emarginated in the lower edge. Endophallus with a median, C-shaped, sclerified phanera. Parameres slender and very elongated, reaching the distal seventh of the median lobe, and bearing three setae only; right paramere slightly shorter than the left.

The name of the species comes from the type locality of the Jinayri deposit.

Austranillus jinayrianus sp. nov. is known only from the type locality, Jinayri deposit, in the Pilbara region, 110 km NW of Newman, WA.

Gracilanillus vixsulcatus Baehr & Main, 2016

Species of this genus are strongly characterised by: lacking longitudinal elytral grooves, pronotum with basal border as wide as the anterior border, sides posteriorly not sinuate, and denticulate before, or near, the basal angles, elytral disc bearing one (posterior) seta placed at the level, or after, the 7^th umbilicate pore, 8^th pore of the umbilicate series located after the 9^th (sensu Giachino and Vailati 2011), toothed labium, median lobe of the aedeagus long and weakly curved with parameres very long and slender.

Differs from Gracilanillus Baehr & Main, 2016 by bearing a single discal seta on the elytral disc (2 in Gracilanillus), and for the absence of longitudinal elytral grooves.

Included medium size species (TL mm 1.29–2.11), and anophthalmous. Integument depigmented but well sclerified, with strong microsculpture and covered by short and moderately dense pubescence.

Head large, slightly narrower than pronotum; mandibles short and simple, without hyperplasias. Maxillary palps ovoidal, swollen. Labium transverse, articulated; mentum not fused with submentum. Labial tooth present. Antennae moniliform, without particular features.

Pronotum squared, sides not sinuate on the basal third, denticulated on the basal third or just before the posterior angles. Basal angles right, sharp, not rounded; basal border as wide as the anterior border; two marginal setae, posterior seta placed near the basal angles.

Elytra elongated and subrectangular, separately rounded, not truncate and not apically emarginated; convex, without longitudinal grooves. Elytral striae absent (except for sutural stria). Lateral margin starting from the humeral area, distinctly serrulate up to the level of 8^th-9^th pores of the umbilicate series.

Scutellar pore present, large and umbilicate; umbilicate series of type B (sensu Jeannel 1963; Giachino and Vailati 2011) with the 8^th pore placed after the 9^th; disc bearing one seta.

Legs relatively long and slender. Pro- and metafemora unarmed; metatrochanters normal, two slightly dilated protarsomeres, without adhesive phanerae, in the male.

Aedeagus relatively small, median lobe long, subrectilinear with basal bulb of normal size. Parameres long, bearing two apical setae. Endophallus with a sinuate, slightly sclerified, apical phanera.

Gilesdytes: name composed by two sections, the first one (Giles) dedicated to William Ernest Powell Giles (20 July 1835 – 13 November 1897), Australian explorer who in 1876 named the Ophthalmia Range (type locality of the genus); and the second one (dytes) for diver.

Currently three species belong to this genus:

Gilesdytes vixsulcatus (Baehr & Main, 2016)

Gilesdytes ethelianus sp. nov.

Gilesdytes pardooanus sp. nov.

Figures 38–42. 

Gilesdytes gen. et spp. nov., habitus (38, 40, 41) aedeagus in lateral view (39, 42) 38, 39 G. vixsulcatus (Baehr and Main), ♂ 40 G. pardooanus sp. nov., HT ♀ 41, 42 G. ethelianus sp. nov., HT ♂. Scale bars: 0.1 mm.

WA, Pilbara, Marillana Creek, c. 85 km NNW Newman, 22°41'53.30"S, 119°20'28.80"E.

1 ♂, WA, Pilbara, 90 km NW of Newman, near Yandi mining area, Ministers North deposit, 22°49'26.54"S, 119°05'23.89"E (WGS84), 20 Feb. 2009, P. Bell, G. Perina, Trap, (BHP010-EXR1154-MNL:7056) Western Australian Museum Entomology Reg. no. 82619 (WAM).

Baehr and Main (2016) description of this species is based on only one female specimen. They assigned the genus Gracilanillus based on the presence of inconspicuous longitudinal grooves at the base of elytra. The study of a male morphologically comparable to the habitus drawing provided by Baehr and Main (2016) and collected from a locality close to the Gracilanillus vixsulcatus type locality, allowed the description of the male of this species, and also the clarification of the taxon’s systematics. Based on some characters considered fundamental in the Anillini taxonomy (Giachino 2005; Giachino and Vailati 2011), the species is here assigned to Gilesdytes gen. nov.

Small species (TL mm 1.29), easily distinguishable from G. ethelianus sp. nov. by the discal pore located towards the elytral edge, and from G. pardooanus sp. nov. by the lateral edge of the pronotum smooth, non-denticulate, near the basal angle.

TL 1.29 mm. Body elongated, depigmented, yellow-testaceous; integument shiny, with evident microsculpture, and covered with sparse and relatively long pubescence.

Head relatively large, narrower than the pronotum. Labium bearing a median tooth. Antennae short, robust, moniliform, just exceeding the base of the pronotum when stretched backwards. Fronto-clypeal furrow slightly distinct; subrectilinear anterior margin of epistome.

Pronotum subrectangular, slightly transverse (max. width / max. length ratio = 1.04), maximum width at the anterior third, very wide basal border, slightly wider than anterior border. Pronotum sides anteriorly poorly arcuate, posteriorly subrectilinear, gently convergent, and laterally smooth, non-denticulate, but distinctly emarginated before the base. Anterior angles rounded, not prominent; posterior angles not protruding externally, rounded at the vertex. Disc slightly convex, with sparse and relatively long pubescence; median groove very shallow, slightly marked. Marginal groove relatively wide and flat, much enlarged near the base; anterior marginal setae inserted inside the marginal groove, approximately on the anterior third; basal setae inserted approximately on the posterior angles.

Legs long and slender, with metatrochanters normal and metafemora unarmed. Males with two dilated protarsomeres without adhesive phanerae.

Elytra subrectangular, very elongated (max. length/max. width ratio = 1.94), maximum width at the anterior 7^th, not emarginated before apex. Disc poorly convex; integument shiny, with evident microsculpture and relatively long, sparse, upright pubescence. Humeri marked, almost right; post-humeral margin denticulate, with distinct crenulations up to the base of the apical 5^th of the elytron; elytral apices not separately rounded. Marginal groove wide and evident up to the 8^th pore of the umbilicate series.

Chaetotaxy : scutellar pore large, foveate. Umbilicate series with 1^st, 2^nd, and 3^rd pores of the humeral group not equidistant, 2^nd and 3^rd pores closest; 4^th pore clearly farther from the 3^rd one and placed at the end of the basal third of the elytron; 5^th pore placed just after the middle length of the elytron; 5^th and 6^th pores spaced out ca. 1/3 of the distance from 6^th and 7^th; 7^th, 8^th, and 9^th pores almost equidistant and slightly displaced onto the disc, 8^th pore placed after the 9^th one. A single discal seta placed laterally towards the edge of the disc and located between the 7^th and 9^th pores.

Aedeagus (Fig. 39) relatively small, median lobe long, slender, subrectilinear, with evident basal bulb; ventral margin poorly and gently curved from the basal bulb to the apex; apical blade evident, but short. Endophallus with a median, sinuate, sclerified phanera. Left paramere slender and very elongated, reaching the distal fifth of the median lobe, and bearing two setae; right paramere shorter than the left one.

Female. See Baehr and Main (2016).

Gilesdytes vixsulcatus is known from two localities approximately 85–90 km NW of Newman, Pilbara, WA. The distance between the two collection points is ca. 30 km.

WA, Pilbara, 100 km E of Port Hedland, Pardoo Mine, 20°17'19.72"S, 119°10'38.88"E.

HT ♀, WA, Pilbara, 100 km E of Port Hedland, Pardoo Mine, (WGS84) 20°17'19.72"S, 119°10'38.88"E, N. Stevens, 02 Oct. 2007, Trog. Trap (PDRC779-LN743), Western Australian Museum Entomology Reg. no. 82634 (WAM).

Medium sized species (TL mm 2.11), easily distinguishable from G. ethelianus sp. nov. by the discal pore placed towards the elytral edge, and from G. vixsulcatus by the denticulate lateral edge of the pronotum before the basal angle.

TL 2.11 mm. Body elongated, depigmented, testaceous; integument shiny, with evident microsculpture, covered with very sparse and short pubescence.

Head relatively large, narrower than pronotum. Labium bearing a median tooth. Antennae short and delicate, moniliform, just exceeding the base of the pronotum when stretched backwards. Fronto-clypeal furrow slightly distinct; anterior margin of the epistome subrectilinear.

Pronotum subrectangular, slightly transverse (max. width / max. length ratio = 1.05), maximum width at the anterior third, with very wide basal border, slightly wider than anterior border, pronotum sides anteriorly poorly arcuate, and laterally-posteriorly denticulate before the basal angles. Anterior angles obtuse, slightly prominent; posterior angles sub squared not protruding, gently rounded at the vertex. Disc slightly convex, with very sparse and short pubescence; median groove very shallow, faintly marked. Marginal groove relatively wide and flat, very enlarged near the base; anterior marginal setae inserted inside marginal groove, approximately on the anterior fifth; basal setae inserted approximately on the posterior angles.

Legs long and slender, with metatrochanters normal and metafemora unarmed.

Elytra subrectangular, very elongated (max. length/max. width ratio = 1.92), maximum width at the anterior 3^rd, slightly emarginated before apex. Disc poorly convex; integument shiny, with evident microsculpture and very short, sparse and upright pubescence. Humeri hardly marked, almost right; post-humeral margin denticulate, with distinct crenulations up to the 9^th pore of the umbilicate series; elytral apices not separately rounded. Marginal groove wide and evident up to the 8^th pore of the umbilicate series.

Chaetotaxy : scutellar pore large, foveate. Umbilicate series with 1^st, 2^nd, and 3^rd pores of the humeral group almost equidistant; 4^th pore slightly displaced onto the disc and clearly farther from the 3^rd pore, placed at the end of the basal third of the elytron; 5^th pore placed just after the middle length of the elytron; 5^th and 6^th ones spaced out ca. the 1/3 of the distance from 6^th and 7^th; 7^th, 8^th, and 9^th not equidistant, 8^th and 9^th pores closest, 8^th pore located after the 9^th one; 8^th slightly displaced onto the disc. One single discal seta laterally placed towards the edge, at the level of the 9^th pore.

Male. Unknown.

The name of the species comes from the type locality Pardoo, in the Pilbara region (WA).

Gilesdytes pardooanus sp. nov. is known only from the type locality (Pardoo Mine), 100 km E of Port Hedland, Pilbara, WA.

WA, Pilbara, 11 km ENE Newman, Ethel Gorge, 23°18'22.176S, 119°51'41.652E.

HT ♂ (remains), WA, Pilbara, 11 km ENE Newman, Ethel Gorge, 23°18'22.176S, 119°51'41.652E (WGS84), P. Bell, S. Catomore, 05 Nov. 2010, Stygo. Net haul; (BHP021_W262-10:0697e) Western Australian Museum Entomology Reg. no. 82608 (WAM).

The only specimen collected had head, prothorax and legs missing (excluding one trochanter), however the remaining characters available were not comparable with other species described, and the presence of aedeagus allowed us to describe the new species.

The taxon ethelianus sp. nov., based on elytra and aedeagus morphology, is provisionally assigned to the genus Gilesdytes.

Small sized species (estimated TL mm 1.30), easily distinguishable from G. pardooanus sp. nov. and from G. vixsulcatus by the discal pore placed in the middle of elytron, not located towards its edge.

Estimated TL mm 1.30.

Elytra (Fig. 41) subrectangular (total length = mm 0.77), very elongated (max. length/max. width ratio = 1.94), maximum width near the middle, slightly emarginated before apex. Disc poorly convex; shiny, integument testaceous, with evident microsculpture and very short, sparse and upright pubescence. Humeri hardly marked, almost right; post-humeral margin denticulate, with distinct crenulation up to the 8^th, 9^th pores of the umbilicate series; elytral apices not separately rounded. Marginal groove wide and evident up to the 7^th pore of the umbilicate series.

Chaetotaxy : scutellar pore large, foveate. Umbilicate series with the 1^st, 2^nd and 3^rd pores of the humeral group almost equidistant; 4^th pore clearly farther from the 3^rd, placed after the end of the basal third of the elytron; 5^th pore placed just after the middle length of the elytron; 5^th and 6^th pores spaced out ca. the 1/3 of the distance from 6^th and 7^th ones; 7^th, 8^th and 9^th pores not equidistant, with 8^th and 9^th pores closest. Due to setae missing it is impossible to determine if the 8^th pore is placed after the 9^th one; 7^th pore displaced onto the disc. One single discal seta not laterally located towards the edge and placed at the level of the 7^th pore.

Metatrochanters unarmed in the male.

Aedeagus (Fig. 42) relatively small, median lobe long, slender, subrectilinear, with basal bulb well-shaped; ventral margin poorly and gently curved from basal bulb to apex; apical blade short and poorly evident. Endophallus with a subapical, bisinuate, sclerified phanera. Parameres missing.

The name comes from the Ethel Gorge type locality.

Gilesdytes ethelianus sp. nov. is known only from the type locality (Ethel Gorge), 11 km ENE of Newman, Pilbara, WA.

Pilbaradytes abydosianus sp. nov.

Included species strongly characterised by: lacking longitudinal elytral grooves, pronotum basal border as wide as anterior border, with sides posteriorly not or only slightly sinuate, basal seta of pronotum absent, one elytral discal seta placed near elytral edge, 8^th pore of the umbilicate series placed after the 9^th one (sensu Giachino and Vailati 2011), metafemora not dentate, labium tooth absent, median lobe of the aedeagus long and curved with parameres long and slender. Differs from Magnanillus Baher, 2017 by the absence of elytral longitudinal grooves, and by a single elytral discal seta (3 setae in Magnanillus).

A genus of Anillini with species of a medium size (TL mm 2.12–2.23), anophthalmous. Integument depigmented but well sclerified, with strong microsculpture and covered with short and sparse pubescence.

Head as wide as, or narrower, than the pronotum base; mandibles short and simple, without hyperplasias. Maxillary palps ovoidal, swollen. Labium transverse, articulated; mentum not fused with submentum. Labial tooth absent. Antennae not strictly moniliform (with relatively elongated antennomeres).

Pronotum squared, with sides smooth or serrulate, not or only slightly sinuate towards the basal third. Basal angles right or obtuse, sharp, not rounded; basal border as wide as or wider than anterior margin; presence of only one marginal seta, posterior seta absent.

Elytra subrectangular, elongated, not separately rounded, not truncate and slightly emarginated apically; convex, without longitudinal grooves. Elytral striae missing (except for the sutural stria). Lateral margin, starting from the humeral area, hardly serrulate up to the level of 6^th pore of the umbilicate series.

Scutellar pore present, large and umbilicate; umbilicate series of type B (sensu Jeannel 1963; Giachino and Vailati 2011) with the 8^th pore placed after the 9^th one; disc bearing one seta strictly located near the elytral edge.

Legs relatively long and slender. In female, unarmed pro- and metafemora; metatrochanters normal; male legs unknown.

Aedeagus relatively large; median lobe long, curved, with basal bulb of normal size. Parameres long, slender, bearing two apical setae. Endophallus without sclerified phanerae.

Composite name coming from the Pilbara region with the suffix -dytes (diver). Gender masculine.

Currently two species belong to this genus:

Pilbaradytes abydosianus sp. nov.

Pilbaradytes webberianus sp. nov.

Figures 43–45. 

Pilbaradytes gen. et spp. nov., habitus (43, 45) aedeagus in lateral view (44) 43, 44 P. abydosianus sp. nov., HT ♂ 45 P. webberianus sp. nov., HT ♀. Scale bars: 0.1 mm.

WA, Pilbara, 100 km SE of Port Hedland, Abydos deposit, 21°08'31.1"S, 119°06'53.99"E.

HT ♂, WA, Pilbara, 100 km SE of Port Hedland, Abydos Mine, 21°08'31.1"S, 119°06'53.99"E (WGS84) 10 Nov. 2008, P. Bell, Trog trap; (ABY01_ABRC029-10LN6261), Western Australian Museum Entomology Reg. no. 82624 (WAM).

Medium sized species (TL mm 2.12), easily distinguishable from P. webberianus sp. nov. by smaller head and by serrulated lateral edges of pronotum.

TL mm 2.12. Body elongated, depigmented, yellow-testaceous; shiny integument, with evident microsculpture, covered with very sparse and short pubescence.

Head small, narrower than the base of pronotum. Labium without tooth. Antennae missing. Fronto-clypeal furrow slightly distinct; subrectilinear anterior margin of epistome.

Pronotum subrectangular, slightly transverse (max. width / max. length ratio = 1.22), maximum width at the base of the anterior third, very wide basal border, as wide as the anterior border, pronotum sides poorly arcuate anteriorly, subrectilinear and gently convergent posteriorly, laterally completely serrulate from the anterior seta to the basal angles, not emarginated before the base. Anterior angles rounded, only slightly prominent; posterior angles sharp and obtuse, not protruding. Disc slightly convex, with very sparse and short pubescence; median groove very shallow, slightly marked. Marginal groove wide and flat, very enlarged near the base; anterior marginal setae inserted inside the marginal groove, approximately on the anterior fifth; basal seta lacking.

Legs missing.

Elytra subrectangular, very elongated (max. length/max. width ratio = 1.79), maximum width at the middle, slightly emarginated in the pre-apical zone. Disc convex; shiny integument, with evident microsculpture and short, very sparse, upright pubescence. Humeri very marked, but rounded; post-humeral margin denticulate, with distinct crenulations up to 6^th pore of the umbilicate series; elytral apices not separately rounded. Marginal groove wide and evident up to the 8^th pore of the umbilicate series.

Chaetotaxy : scutellar pore large, foveate. Umbilicate series with 1^st, 2^nd, and 3^rd pores of the humeral group not equidistant, 2^nd and 3^rd pores closest; 4^th pore clearly farther from the 3^rd one and placed at the end of the basal third of the elytron; 5^th pore placed well after the middle length of the elytron; 5^th and 6^th pores spaced out ca. the 1/3 of the distance from 6^th and 7^th ones; 7^th, 8^th, and 9^th pores almost equidistant, 8^th pore slightly displaced onto the disc and placed after the 9^th one. One single discal seta laterally placed near the edge, midway between the 7^th and the 9^th pores.

Aedeagus (Fig. 44) relatively large, median lobe long, tubular, regularly curved, with basal bulb evident; ventral margin gently curved from the basal bulb to the apex; apical blade evident, but short. Endophallus without any sclerified phanerae. Left parameres slender and very elongated, reaching the distal fourth of the median lobe, and bearing two setae; right paramere stout and shorter than the left one, and bearing two setae.

Female. Unknown.

The name comes from the type locality Abydos deposit.

Pilbaradytes abydosianus sp. nov. is known only from the type locality 100 km SE of Port Hedland, Pilbara, WA.

WA, Pilbara, 150 km SE of Port Hedland, Mount Webber Mine, 21°32'16.81"S, 119°17'17.88"E.

HT ♀, WA, Pilbara, 150 km SE of Port Hedland, Mount Webber Mine, 21°32'16.81"S, 119°17'17.88"E (WGS84), E.S. Volschenk, S. Catomore, 05"Sept. 2010; Trog. trap (COF01-MW025-10:9656) Western Australian Museum Entomology Reg. no. 82616 (WAM). PTT: 1 ♀, WA, Pilbara, 150 km SE of Port Hedland, Mount Webber Mine, 21°32'08.09"S, 119°17'18.4"E (WGS84), E.S. Volschenk, S. Catomore, 04"Sept. 2010, Trog. trap (COF01-MW090-10:9708) Western Australian Museum Entomology Reg. no. 82617 (CGi).

Medium size species (TL mm 2.23), easily distinguishable from P. abydosianus sp. nov. by the larger head and the smooth lateral edges of the pronotum.

TL mm 2.23. Body elongated, depigmented, yellow-testaceous; shiny integument, with evident microsculpture, covered with very sparse and short pubescence.

Head small, approximately as wide as the base of the pronotum. Labium without tooth. Antennae with relatively elongated antennomeres, short, just exceeding the base of the pronotum when stretched backwards. Fronto-clypeal furrow slightly distinct; subrectilinear anterior margin of epistome.

Pronotum subrectangular, slightly transverse (max. width / max. length ratio = 1.23), maximum width at the base of the anterior fifth, with very wide basal border, as wide as the anterior border, pronotum sides poorly arcuate anteriorly, subrectilinear and very slightly sinuated posteriorly, laterally completely smooth from anterior seta to basal angles, not emarginated before the base. Anterior angles rounded, only slightly prominent; posterior angles right, sharp, not protruding. Disc slightly convex, with very sparse and short pubescence; median groove very shallow, slightly marked. Marginal groove narrow and flat, not enlarged near the base; anterior marginal setae inserted inside marginal groove, approximately on the anterior seventh; basal seta absent.

Legs relatively long and slender. In females, unarmed pro- and metafemora; normal metatrochanters.

Elytra subrectangular, very elongated (max. length/max. width ratio = 1.85), maximum width at the middle, lateral sides slightly emarginated at the end of the basal third and in the pre-apical zone. Disc convex; shiny integument, with evident microsculpture and short, sparse and upright pubescence. Humeri very marked, but rounded; post-humeral margin denticulate, with distinct crenulations up to 6^th pore of the umbilicate series; elytral apices not separately rounded. Marginal groove wide and evident up to the 9^th pore of the umbilicate series.

Chaetotaxy : scutellar pore large, foveate. Umbilicate series with 1^st, 2^nd, and 3^rd pores of the humeral group not equidistant, 2^nd and 3^rd pores closest; 4^th pore clearly farther from the 3^rd one and placed at the end of the basal third of the elytron; 5^th pore placed well after the middle length of the elytron; 5^th and 6^th pores spaced out ca. half of the distance from 6^th and 7^th; 7^th, 8^th, and 9^th pores almost equidistant, 8^th and 9^th pores closer to each other than 7^th and 9^th; 8^th pore placed after the 9^th one. One single discal seta laterally placed near the edge, midway between the 7^th and the 9^th pores.

Male. Unknown.

The name comes from the type locality Mount Webber, in the Pilbara region.

Pilbaradytes webberianus sp. nov. is known only from Mount Webber, 150 km SE of Port Hedland, Pilbara, WA.

Gracilanillus currani Baehr & Main, 2016

Included species strongly characterised by: longitudinal elytral grooves absent, pronotum with basal border as wide as or narrower than anterior border, and sides not or only slightly posteriorly sinuate, basal seta of pronotum present, one elytral discal seta present, 8^th pore of the umbilicate series placed after the 9^th pore (sensu Giachino and Vailati 2011), metafemora not dentate, labial tooth absent, median lobe of aedeagus long and curved, parameres long and slender. Bylibaraphanus differs from Magnanillus Baher, 2017 by elytra with a single discal seta (3 in Magnanillus) and smaller body size. It differs from Hesperanillus Baher & Main, 2016 by head narrower than pronotum, and pronotum less cordiform. It differs from Gracilanillus Baher & Main, 2016 by the absence of longitudinal elytral grooves.

Genus with small size species (TL mm 1.43–1.50), anophthalmous. Depigmented integument, medium sclerified, with strong microsculpture and dense pubescence.

Head wider than pronotum base; mandibles short and simple, without hyperplasias. Maxillary palps ovoidal, swollen. Labium transverse, articulated; mentum not fused with submentum. Labial tooth absent. Antennae moniliform.

Pronotum squared, pronotum sides smooth, not or only slightly sinuate at the basal third. Basal angles acute or obtuse, sharp or rounded; basal border as wide as, or narrower, than anterior border; basal seta present.

Elytra convex, subrectangular, elongated, separately rounded, not truncate, apically slightly emarginated, and without longitudinal grooves. Elytral striae missing (except for the sutural stria). Lateral margin, starting from the humeral area, hardly serrulate up to the level of the 7^th or 9^th pore of the umbilicate series.

Scutellar pore present, large and umbilicate; umbilicate series of type B (sensu Jeannel 1963; Giachino and Vailati 2011) with the 8^th pore placed after the 9^th one; disc bearing one seta (posterior) in the central area of the disc.

Legs relatively long and slender. Pro- and metafemora unarmed; metatrochanters normal; two protarsomeres slightly dilated and without adhesive phanerae in males.

Aedeagus relatively large; median lobe relatively long, curved, not restricted before the basal bulb, basal bulb of normal size. Parameres long, slender, bearing two or three apical setae. Endophallus with sclerified phanera.

Combined name formed by Bylibara (aboriginal noun for the Pilbara region) and the suffix -phanus taken from Illaphanus genus. The gender of the name is masculine.

The following two species belong to this genus:

Bylibaraphanus currani (Baehr & Main, 2016)

Bylibaraphanus cundalinianus sp. nov.

Figures 46–48. 

Bylibaraphanus gen. et spp. nov., habitus (46, 48) aedeagus in lateral view (47) 46, 47 B. currani (Baehr and Main), ♂ 48 B. cundalinianus sp. nov., HT ♀. Scale bars: 0.1 mm.

2 ♂♂ 3 ♀♀, WA, Pilbara, 65 km NW of Tom Price, Cloudbreak Mine, 22°20'26.40"S, 119°25'50.70"E (WGS84), J. Cocking, M. Scanlon, 15 Mar. 2011, Trog. Net scrape (GNGC05589), Western Australian Museum Entomology Reg. no. 8269–82673 (WAM, CGi).

Baehr and Main’s description (2016) is correct and detailed, although we noted a few inaccuracies and their description omits some important characters fundamental to the systematics of Anillini which, including male genitalia, are described as follows.

Species of small size (TL mm 1.47–1.50), easily distinguishable from B. cundalinianus sp. nov. by the basal angles of pronotum acute and protruding.

TL mm 1.49–1.50 ♂♂ 1.47–1.49 ♀♀. Body elongated, depigmented, yellow-testaceous; integument shiny, with evident microsculpture, covered with relatively long pubescence.

Head large, slightly narrower than the base of pronotum. Labium without tooth. Antennae short, just exceeding the base of the pronotum when stretched backwards. Fronto-clypeal furrow slightly distinct; subrectilinear anterior margin of epistome.

Pronotum subrectangular, slightly transverse (max. width / max. length ratio = 1.22), maximum width at the base of the anterior fourth, with very wide basal border, as wide as the anterior border, pronotum sides, anteriorly poorly arcuate, and posteriorly subrectilinear shortly and gently sinuated, laterally completely smooth from anterior seta to basal angles, not emarginated before the base. Anterior angles rounded, not prominent; posterior angles acute, sharp, protruding. Disc slightly convex, with very sparse and relatively long pubescence; median groove very shallow, slightly marked. Marginal groove narrow and flat, not particularly enlarged near the base; anterior marginal setae inserted inside marginal groove, at ca. the anterior third; basal setae inserted approximately on posterior angles.

Legs long and slender, with normal metatrochanters and unarmed metafemora; two protarsomeres slightly dilated and without adhesive phanerae in males.

Elytra subrectangular, very elongated (max. length/max. width ratio = 2.09), with parallel sides, slightly emarginated before apex. Disc convex; integument shiny, with evident microsculpture and short, very dense and upright pubescence. Humeri very marked, but rounded; post-humeral margin denticulate, with distinct crenulations up to 7^th pore of the umbilicate series; elytral apices separately rounded. Marginal groove narrow and evident up to the 7^th pore of the umbilicate series.

Chaetotaxy : scutellar pore large, foveate. Umbilicate series with 1^st, 2^nd, and 3^rd pores of the humeral group not equidistant, 2^nd and 3^rd pores closest; 4^th pore clearly farther from the 3^rd one and placed at the end of the basal third of the elytron; 5^th pore placed well after the middle length of the elytron; 5^th and 6^th pores spaced out ca. the half distance between 6^th and 7^th pores; 7^th, 8^th and 9^th pores slightly displaced onto the disc and not equidistant, 8^th placed after the 9^th one. One single discal seta in the central area of the disc and placed approximately half way between the 6^th and 7^th pores.

Aedeagus (Fig. 47) relatively large, median lobe, in lateral view, relatively long and triangularly restricted at apex, regularly curved and not restricted before the basal bulb, basal bulb of normal size. Ventral margin gently curved from basal bulb to apex, emarginated just before the apex; apical blade evident, very long. Endophallus with a large, concave, well sclerified phanera in the middle area. Left parameres slender and very elongated, reaching the distal fourth of the median lobe, and bearing two setae; right paramere shorter than left one and bearing three setae.

B. currani is known only from Cloudbreak Mine, 65 km NW of Tom Price, Pilbara, WA.

WA, Pilbara, 200 km E of Port Hedland, Yarrie Mining Area, Cundaline Ridge, 20°32'36"S, 120°09'35"E.

HT ♀, WA, Pilbara, 200 km E of Port Hedland, Yarrie Mining Area, Cundaline Ridge, 20°32'36"S, 120°09'35"E, 29 May 2009, Subterranean Ecology, bore-hole CU0046R, scraping, Western Australian Museum Entomology Reg. no. 72022 (WAM).

Small sized species (TL mm 1.43), easily distinguishable from B. currani by obtuse and not protruding basal angles of the pronotum.

TL mm 1.43. Body elongated, depigmented, yellow-testaceous; integument shiny, with evident microsculpture, covered with short pubescence.

Head large, slightly narrower than the base of the pronotum. Labium without tooth. Antennae short, just exceeding the base of the pronotum when stretched backwards. Fronto-clypeal furrow slightly distinct; subrectilinear anterior margin of epistome.

Pronotum subrectangular (max. width / max. length ratio = 1.06) maximum width at the base of the anterior fourth, with basal border narrower than the anterior border, pronotum sides, anteriorly poorly arcuate, posteriorly subrectilinear and slightly sinuated, laterally completely smooth from anterior seta to basal angles, not emarginated before the base. Anterior angles rounded, slightly prominent; posterior angles obtuse, rounded, not protruding. Disc slightly convex, with very sparse and relatively short pubescence; median groove very shallow, slightly marked. Marginal groove relatively narrow and flat, not particularly enlarged near the base; anterior marginal setae inserted inside the marginal groove, approximately on the anterior fifth; basal setae inserted approximately on posterior angles.

Legs long and slender, with metatrochanters normal and metafemora unarmed.

Elytra subrectangular, very elongated (max. length/max. width ratio = 1.98), with parallel sides, slightly emarginated before apex. Disc convex; integument shiny, with evident microsculpture and very short, very dense and upright pubescence. Humeri very marked, squared; post-humeral margin denticulate, with distinct crenulations up to the discal pore; elytral apices separately rounded. Marginal groove narrow and evident up to the discal pore.

Chaetotaxy : scutellar pore large, foveate. Umbilicate series with 1^st, 2^nd,and 3^rd pores of the humeral group almost equidistant; 4^th pore clearly farther from the 3^rd one and placed at the end of the basal third of the elytron; 5^th pore placed well after the middle length of the elytron; 5^th and 6^th pores spaced out ca. 1/3 of the distance between 6^th and 7^th pores; 7^th, 8^th, and 9^th pores not equidistant, 8^th placed after the 9^th one; 7^th slightly displaced onto the disc. One single discal seta in the central area of the disc and placed at ca. 2/3 of the distance between the 6^th and 7^th pores.

Male. Unknown.

The name comes from the type locality: Cundaline Ridge.

Bylibaraphanus cundalinianus sp. nov. is known only from the type locality Cundaline Ridge, in the Yarrie Mining Area, 200 km E of Port Hedland, Pilbara, WA.

Angustanillus striatipennis Baehr & Main, 2016

Included species strongly characterised by: body very elongated, posterior supraorbital seta and longitudinal elytral grooves absent, pronotum with basal border narrower than anterior border, and sides posteriorly distinctly crenelated, basal seta of pronotum present, one or two elytral discal seta present, 8^th pore of the umbilicate series located after the 9^th pore (sensu Giachino and Vailati 2011), metafemora dentate in males (state unknown in females), labial tooth absent, median lobe of the aedeagus long, slender and curved, parameres long and slender.

Describing Angustanillus Baehr and Main (2016) mention the presence of two setae on the elytral disc, one located in the fifth basal section, and one in the fourth apical section. The species we describe here has only one seta, situated in the posterior one. Considering the taxonomic importance of the number of setae in establishing the systematics of a taxon in Anillini (Giachino 2005; Giachno and Vailati 2011), together with the difficulties in identifying these setae without a proper preparation and observation with a microscope at high magnification, we consider the presence of one seta as the most probable character in this genus. However, in the genus diagnosis we account for the possible presence of one or two discal setae.

Additionally, in the genus diagnosis we indicate the presence of a large metafemoral tooth, at least in the male. The female of A. armatus sp. nov. is unknown to date, and the female holotype is known only for A. striatipennis Baehr and Main (2016), therefore still it is not possible to confirm the presence of this character state in both sexes of the two species. We hence re-describe the genus Angustanillus integrating the new characters presented by A. armatus sp. nov.

Included large size species (TL > mm 2.40), anophthalmous. Depigmented and sclerified integument, with strong microsculpture (sometimes formed by distinct, deep, large punctures) and covered with dense pubescence.

Head wider than the pronotum base; mandibles short and simple, without hyperplasias. Maxillary palps ovoidal, swollen. Labium transverse, articulated; mentum not fused with submentum. Labial tooth absent. Antennae not strictly moniliform (with relatively elongated antennomeres).

Pronotum elongated, with sides distinctly crenelated in the basal fourth, not sinuate in the basal third. Pronotum basal border slightly narrower than the anterior border; basal angles sharp and squared; basal seta present, placed before basal angles.

Elytra subrectangular and elongated, separately rounded, not or distinctly emarginated apically; convex, without longitudinal grooves. Elytral striae present (4 striae) or missing (except for the sutural stria). Lateral margin, starting from the humeral area, hardly serrulate up to the level of 8^th pore of the umbilicate series.

Scutellar pore present, large and umbilicate; umbilicate series of type B (sensu Jeannel 1963; Giachino and Vailati 2011), 8^th pore placed after the 9^th pore; disc bearing one (posterior) or two setae in the central area of the disc.

Legs relatively long and slender. Male metafemora distally armed with a long, internal spur; metatrochanters normal; two protarsomeres dilated and without adhesive phanerae in males.

Aedeagus relatively large; median lobe long, curved, not restricted before the basal bulb that is of normal size. Parameres relatively long and robust, bearing two setae: one apical and one subapical. Endophallus without sclerified phanerae.

The following two species belong to this genus:

Angustanillus striatipennis Baehr & Main, 2016

Angustanillus armatus sp. nov.

Figures 49–51. 

Angustanillus armatus sp. nov., HT ♂ 49 habitus 50 left metafemur and metatrochanter in dorsal view 51 aedeagus in lateral view. Scale bars: 0.1 mm.

WA, Pilbara, 38 km W of Pannawonica, Mesa B, 21°39'36"S, 115°57'20"E.

HT ♂, WA, Pilbara, 38 km W of Pannawonica, Mesa B (bore hole MEBRC0021), 21°39'36"S, 115°57'20"E (GPS WGS84), March-May 2005; M. Greenham, D. Kamien, L. Mould, Western Australian Entomology Reg. no. 64217 (WAM).

Large sized species (TL mm 2.66), easily distinguishable from A. striatipennis by: elytra without distinct striae, but with a series of short and longitudinally aligned setae; only one discal seta; pronotum and elytra surface without distinct punctures; and more elongated pronotum.

TL mm 2.66. Body elongated, depigmented, yellow-testaceous; integument shiny, with evident microsculpture, covered with very short pubescence.

Head large, slightly wider than the base of the pronotum. Labium without tooth. Antennae short, just exceeding the base of the pronotum when stretched backwards. Fronto-clypeal furrow of frontal clypeo slightly distinct; anterior margin of the epistome subrectilinear.

Pronotum elongated (max. width / max. length ratio = 0.90), maximum width at the middle, basal border slightly narrower than the anterior border, pronotum sides poorly arcuate, distinctly crenelated at the posterior fourth, emarginated before the base. Anterior angles rounded, not prominent; posterior angles sharp, squared, not protruding. Disc slightly convex, with very sparse and short pubescence; median groove very shallow, slightly marked. Marginal groove wide and flat, enlarged near the base; anterior marginal setae inserted inside the marginal groove, approximately on the anterior fifth; basal setae inserted before the posterior angles.

Legs long and slender, with metatrochanters normal and metafemora (Fig. 50) armed with a long, internal spur; two protarsomeres dilated and without adhesive phanerae in male.

Elytra subrectangular, very elongated (max. length/max. width ratio = 2.28), with parallel sides, slightly emarginated before apex. Disc convex; integument shiny, with evident microsculpture and short, longitudinally aligned and upright pubescence. Humeri very marked, squared; post-humeral margin denticulate, with a distinct crenulation up to the 8^th pore of the umbilicate series; elytral apices separately rounded. Marginal groove narrow and evident up to the 8^th pore of the umbilicate series.

Chaetotaxy : scutellar pore large, foveate. Umbilicate series with 1^st, 2^nd, and 3^rd pores of the humeral group almost equidistant; 4^th pore clearly farther from the 3^rd one and placed at the end of the basal third of the elytron; 5^th pore placed at the end of median third of the elytron; 5^th and 6^th pores spaced out ca. 1/4 of the distance from 6^th and 7^th pores; 7^th, 8^th, and 9^th pores not displaced onto the disc and not equidistant, 8^th pore placed after the 9^th one. One single discal seta in the central area of the disc and placed approximately at the level of the 9^th pore.

Aedeagus (Fig. 51) relatively large, median lobe, in lateral view, long and restricted at apex, regularly curved and not restricted before the basal bulb, basal bulb of normal size. Ventral margin curved from basal bulb to apex, gently emarginated just before the apex; apical blade evident, but short. Endophallus without sclerified phanerae. Parameres relatively long and robust, bearing two setae: one apical and one subapical; right paramere shorter than the left one.

The name comes from the Latin word armato (which means armed) to highlight the presence of a large spur on metafemora.

Angustanillus armatus sp. nov. is known only from the type locality Mesa B, 38 km W of Pannawonica, Pilbara, WA.