Ecuador, Napo: Reserva Étnica Waorani, 1 Km S. Onkone Gare Camp, 220m, 0°30'10"S, 76°26'0"W, fogging terre firma forest, 12.II.1995, T. L. Erwin et al. leg.

Holotype male, mounted on one slide; thorax in a separate microvial. Original label: Ecuador. Napo. Reserva Étnica Waorani, 1 Km S. Onkone Gare Camp. 220m. 12.II.1995. 0°30'10"S: 76°26'0"W. Fogging terre firma forest. T. L. Erwin et al. (EPN, slide 160, vial 160).

This species is dedicated to the Ecuadorian entomologist Freddy Rubén Bravo Quijano, of the Universidade Estadual de Feira de Santana, Bahia, Brazil, in recognition to his important contributions in the taxonomy of Neotropical Psychodidae (Diptera), also for the support to AMSN, to pursue a career studying Psocodea, ‘Psocoptera’.

Differing from the known species of Triplocania, in having the hypandrium with side sclerites fused proximally to the central piece, and having two forked posterior projections, horn shaped; also by having a U–shaped phallosome, a phallobase with lateral extensions covering partly the anterior pairs of endophallic sclerites, and in having ornamented areas on the endophallus.

Color. Compound eyes black, ocelli hyaline, with ochre centripetal crescents; head pattern (Fig. 1). Scape and pedicel pale yellow, f₁–f₃ pale yellow, with apex white. Mx4 pale brown. Femora pale yellow; tibiae yellow, distally pale brown; tarsomere 1 pale yellow, tarsomeres 2–3 pale brown. Forewing with a brown marginal band from R₄₊₅ to almost CuA₂; a brown, almost rectangular band, well pigmented proximally and distally, weakly pigmented in the middle, from the apex of the areola postica to posterior end of the pterostigma; pterostigma with proximal and distal brown bands. Proximal third of forewing dark brown, limited posteriorly by level of crossvein Rs–M, as illustrated (Fig. 2). Veins brown. Hindwing with area below CuP dark brown, and a marginal pale brown band from R₄₊₅ to almost CuA (Fig. 3).

Figures 1–7. 

Triplocania bravoi sp. n. (Holotype male). 1 Front view of head 2 Forewing 3 Hindwing 4 Lacinial tip 5 Hypandrium 6 Phallosome in dorsal view 7 Clunium, paraprocts and epiproct. Scales in mm.

As in diagnosis, plus the following: compound eyes without interommatidial setae. Outer cusp of lacinial tip broad, with five denticles (Fig. 4). Forewing pterostigma long, widest in the middle. Areola postica very wide basally, slanted posteriorly, tall proximally with apex round and narrow, distally sinuous and low; R₂₊₃ and R₄₊₅ sinuous, M stem slightly concave proximally, M_1, M₂ and M₃ sinuous. Hindwing Rs almost straight, R₂₊₃ straight, R₄₊₅ almost straight, M sinuous. Hypandrium of three sclerites, central piece anteriorly convex, with side projections almost parallel to side sclerites, forked posterior projections densely setose, other setae as illustrated (Fig. 5). Phallosome U-shaped, side struts independent, V shaped, fused posteriorly to external parameres, these stout, bearing a field of pores distally, each with an small projection heavily sclerotized distally; three pairs of endophallic sclerites, a posterior pair, curved outwards distally, close to the ends of the external parameres, a mesal pair, originating from behind the side struts, crossing behind the external parameres, distally dilated, with a small, acuminate projection apically; anterior pair stout, curved inwards, apically pointed and proximally rounded; phallobase with anterior border concave; lateral extensions covering partly the anterior pair of endophallic sclerites; endophallus membranous, with areas thickened and ornamented as illustrated (Fig. 6). Paraprocts broad, a field of setae along inner margin, other setae on apex; sensory fields with 27–28 trichobothria on basal rosettes (Fig. 7). Epiproct wide based, posteriorly rounded, with three large mesal setae, next to anterior margin, other setae as illustrated (Fig. 7).

(in microns). FW: 3289, HW: 2108, F: 774, T: 1377, t1: 285, t2: 59, t3: 117, f1: 466, f2: 397, f3: 270, Mx4: 201, IO: 440, D: 332, d: 186, PO: 0.56.

Ecuador, Napo: Reserva Étnica Waorani, 1 Km S. Onkone Gare Camp, 220m, 0°30'10"S: 76°26'0"W, fogging terre firma forest, 12.II.1995, T. L. Erwin et al. leg.

Holotype male, mounted on slides; thorax in a separate microvial. Original label: Ecuador. Napo. Reserva Étnica Waorani, 1 Km S. Onkone Gare Camp. 220m. 12.II.1995. 0°30'10"S: 76°26'0"W. Fogging terre firma forest. T. L. Erwin et al. Paratype: 1 male, same data as the holotype (EPN, slides 163–164, vials 163–164).

This species is dedicated to Dr. Terry L. Erwin, of the Smithsonian Institution, Washington, D. C., USA., in recognition to his seminal studies in biodiversity, in estimating the number of arthropods on this planet, in systematics and biology of the Carabidae, and for making available for study to ANGA, the psocid specimens collected by his team in Napo, Ecuador, by canopy fogging.

Differing from the known species of Triplocania, in having the hypandrium with side sclerites and central piece similar in size; central piece with two short, lateral posterior projections, and two short, blunt ended, median posterior projections, leaving between them a small concavity, in having the external parameres with a distinct lobe apically on the inner side and, in having two pairs of endophallic sclerites.

Color. Body pale brown, with ochre spots as indicated below. Compound eyes black, ocelli hyaline, with ochre centripetal crescents; head pattern (Fig. 8). Scape brown and pedicel yellow, f₁–f₃ yellow, with apices white. Mx4 brown. Tergal lobes of meso and methathorax brown, pleura with ochre spots above the level of the coxae; dark brown bands on proximal and distal ends of coxae, femora yellow with three brown equidistant bands, a middle one, and one on each end of the femur; tibiae pale brown, tarsomeres 1–3 yellow. Forewing with an irregular, submarginal brown band from R₂₊₃ to areola postica, this with a small brown spot proximally, and a dark brown spot between its apex and M, a dark brown spot below the proximal end of CuA, and a brown spot at confluence of CuP–1A; a pale brown spot between proximal ends of R₄₊₅–M₁; pterostigma with brown bands anteriorly and posteriorly; veins brown, with brown spots at wing margin (Fig. 9). Hindwing almost hyaline, veins brown, with a pale brown spot at confluence of CuP and wing margin (Fig. 10).

Figures 8–14. 

Triplocania erwini sp. n. (Holotype male). 8 Front view of head 9 Forewing 10 Hindwing 11 Lacinial tip. 12 Hypandrium 13 Phallosome in dorsal view 14 Clunium, paraprocts and epiproct. Scales in mm.

As in diagnosis plus the following: compound eyes without interommatidial setae. Outer cusp of lacinial tip broad, with five denticles (Fig. 11). Forewing pterostigma basally narrow, wider in the middle; areola postica wide basally, slightly slanted posteriorly; R₂₊₃ and R₄₊₅ sinuous, M stem slightly concave proximally, M_1, M₂ and M₃ sinuous. Hindwing Rs almost straight, R₂₊₃ and R₄₊₅ straight, M sinuous. Hypandrium with two large setae posteriorly between each lateral-median posterior projections (Fig. 12). Phallosome with side struts independent, V shaped, fused posteriorly to external parameres, these stout, with pores posteriorly; the distal lobe of each heavily sclerotized; anterior pair of endophallic sclerites elongate, almost touching anteriorly, inserted on a membranous, V shaped, thickened area, lying distally behind the external parameres, ending in a rounded apex; posterior pair of endophallic sclerites triangular, each anteriorly concave, close to the inner border of the external parameres, with apices sinuous.

(in microns). FW: 3302, HW: 2270, F: 645, T: 882, t1: 330, t2: 63, t3: 115, f1: 519, f2: 384, f3: 330, Mx4: 239, IO: 398, D: 326, d: 194, PO: 0.59.

Brazil, Rondônia: Ariquemes, Rio ji Paraná, 90°44'S: 61°52'W, Malaise trap. 28.I.1986, J. A. Rafael leg.

Holotype male, mounted on slides, with thorax in a separate microvial. Original label: Brasil. Rondonte [Rondônia]. Ariquemes, Rio ji Paraná. 28.I.1986. 90°44'S: 61°52'W. Malaise trap. J. A. Rafael. Paratypes: 1 female and 3 males, same data as the holotype (INPA, slides 57–61, vials 57–61).

The specific name refers to the proximity of this species to T. lamasi Silva–Neto, Rafael & García Aldrete.

Differing from T. lamasi in having the posterior sclerite of the hypandrium thicker in the middle, with the posterior projection more than twice as long; sickle-shaped lateral projections distal to the anterior sclerite barely reaching the inner margins of the lateral sclerites.

Color. Body yellowish brown, with dark brown spots as indicated below. Compound eyes black, ocelli hyaline, with ochre centripetal crescents; head pattern (Fig. 15). Scape and pedicel pale brown; flagellomeres pale yellow. Mx4 pale yellow. Tergal lobes of meso- and metathorax reddish brown; episternum of mesothorax ochre. Coxae, trochanters and femora creamy white, tibiae and tarsomeres pale yellow. Forewings hyaline, as illustrated (Fig. 16); veins brown. Hindwing (Fig. 17), hyaline throughout, veins brown.

Figures 15–21. 

Triplocania lamasoides sp. n. (Holotype male). 15 Front view of head 16 Forewing 17 Hindwing 18 Lacinial tip 19 Clunium, paraprocts and epiproct 20 Hypandrium 21 Phallosome in dorsal view. Scales in mm.

As in diagnosis, plus the following: compound eyes with interommatidial setae. Outer cusp of lacinial tip broad, with six denticles (Fig. 18). Forewing pterostigma long, widest in the middle. Areola postica wide basally, slightly slanted posteriorly, apex round, narrow. R₂₊₃ and R₄₊₅ sinuous, M stem concave, M₁ almost straight, M₂ sinuous, M₃ branched, the branching point closer to M than to the wing margin. Hindwing Rs almost straight. Hypandrium (Fig. 19) of four sclerites, anterior piece broad, setose, bearing distally two sickle-shaped lateral projections, heavily sclerotized at both ends, and having also a well defined, setose sclerotized area in the middle; posterior sclerite concave anteriorly, with a long, slender posterior projection in the middle, flanked by two large, broadly triangular lateral sclerites. Phallosome (Fig. 20) with side struts independent, V shaped, fused posteriorly to external parameres, these stout, each with an elongate projection on inner margin, with field of pores; three pairs of endophallic sclerites; anterior pair long, slender and curved, mesal pair wide proximally, narrowing distally, pointed, and posterior pair parallel to the inner margin of the external parameres, with three acuminate projections distally. Paraprocts broad, wide proximally, narrowing to round apex; with a field of short setae along inner margin, other setae as illustrated; sensory fields with 30–31 trichobothria on basal rosettes (Fig. 21). Epiproct mesally with an almost elliptic protuberance, with a field of setae posteriorly, and three large mesal setae next to anterior margin (Fig. 21).

(in microns). FW: 3710, HW: 2465, F: 910, T: 1493, t1: 622, t2: 77, t3: 132, f1: 556, f2: 455, f3: 390, Mx4: 170, IO: 470, D: 395, d: 210, PO: 0.53.

Color. Essentially as in the male.

Fore- and hind- wings (Figs 22, 23) same as in the male. Subgenital plate broad, V shaped, pigmented area wide, setae as illustrated (Fig. 24); Gonapophyses: V₁ long, slender, heavily sclerotized; V₂₊₃ stout, heeled, narrow anteriorly and wider in the middle, with three large setae on outer lobe as illustrated, distal process stout, sinuous, distally blunt, with a field of microsetae (Fig. 25). Ninth sternum broad, with two distinct areas, the anterior one unpigmented, with a concavity anteriorly and posteriorly in the middle; posterior area pigmented, thicker than the anterior one, with a strongly sclerotized band latero-posteriorly, and a small, strongly pigmented area mesally on each side. Paraprocts broad, almost triangular, wide proximally, narrowing to round apex, setose posteriorly as illustrated, sensory fields with 26–27 trichobothria on basal rosettes (Fig. 26). Epiproct missing.

Figures 22–26. 

Triplocania lamasoides sp. n. (Paratype female). 22 Forewing 23 Hindwing 24 Subgenital plate 25 Gonapophyses and Ninth sternum 26 Left paraproct Scales in mm.

(in microns). FW: 3723, HW: 2560, F: 890, T: 1385, t1: 607, t2: 58, t3: 121.

Brazil, Mato Grosso: Chapada dos Guimarães, 23-30.XI.1983. A. Yamamoto leg.

Holotype male, mounted on slides; thorax in a separate microvial. Original label: Brasil. Mato Grosso. Chapada dos Guimarães. 23-30.XI.1983. A. Yamamoto. Paratypes: 1 male, Original label: Brasil. Rondonte [Rondônia]. Ariquemes. Rio ji Paraná. 28.I.1986. 09°44'S: 61°52´W. Malaise trap. J. A. Rafael. (INPA, slides 112–113, vials 112–113).

The specific name refers to the characteristic of the hypandrium, having three posterior projections.

Differing from the known species of Triplocania, in having the central piece of the hypandrium with three posterior projections, a middle one, pointed, setose, flanked by lateral, strongly sclerotized, glabrous acuminate projections. Phallosome with a transverse, strongly sclerotized mesal bridge, biconcave anteriorly, convex posteriorly, widest in the middle, narrowing to the sides; four pairs of endophallic sclerites; external parameres distally with an elliptic papillose field.

Color. Body yellow, with ochre spots as indicated below. Compound eyes black, ocelli hyaline, with ochre centripetal crescents. Head pattern (Fig. 27). Scape brown, pedicel pale brown, f₁ anteriorly pale brown, posteriorly yellow, apex white, f₂ yellow. Mx4 brown. Tergal lobes of meso and methathorax pale brown, pleura yellow; femora pale yellow, tibiae pale brown, tarsomeres 1–3 yellow. Forewing with an irregular, submarginal pale brown band from R₂₊₃ to posterior end of areola postica, this with a small brown spot proximally, and a dark brown spot between its apex and M; a triangular brown area next to CuA, and a brown spot at confluence of CuP–1A; pterostigma with brown bands anteriorly and posteriorly; veins brown, with brown spots at wing margin (Fig. 28). Hindwing almost hyaline, with small brown spots distally on veins M, R₂₊₃ and R_4+5; with a pale brown spot at confluence of CuP and wing margin; veins brown (Fig. 29).

Figures 27–33. 

Triplocania trifida sp. n. (Holotype male). 27 Front view of head 28 Forewing 29 Hindwing 30 Lacinial tip 31 Hypandrium 32 Phallosome in dorsal view 33 Clunium, paraprocts and epiproct. Scales in mm.

As in diagnosis, plus the following: compound eyes without interommatidial setae. Outer cusp of lacinial tip broad, with five denticles (Fig. 30). Forewing pterostigma wider in the middle, narrow anteriorly; Rs convex, R₂₊₃ almost straight proximally and concave distally, R₄₊₅ sinuous, M stem slightly concave proximally, then almost straight, M₂–M₃ sinuos, areola postica wide basally, slightly slanted posteriorly; hindwing Rs almost straight, R₂₊₃ and R₄₊₅ straight, M sinuous. Hypandrium of three sclerites; side sclerites, large, irregular, with setae as illustrated (Fig. 31). Phallosome with side struts independent, V shaped, fused posteriorly to external parameres, these stout, with pores posteriorly. Anterior pair of endophallic sclerites elongate, sinuous, distally acuminate; a central pair, narrow, elongate, sinuous, heavily sclerotized, with anterior end blunt, posteriorly dilated, lance-shaped; a lateral pair biramous, with inner arms long, curved out, wide based, extended posteriorly and distally acuminate, and outer arms wider in the middle, narrowing at the ends, posteriorly shaped like a bird’s head; posterior pair, small, curved outwards, distally acuminate (Fig. 32). Paraprocts broad, almost triangular, sensory fields with 20–31 trichobothria on basal rosettes, setae as illustrated (Fig. 33). Epiproct trapeziform, wide anteriorly, with sides converging towards a straight posterior border, three mesal setae near anterior border, other setae as illustrated (Fig. 33).

(in microns). FW: 3804, HW: 2569, F: 947, T: 1511, t1: 643, t2: 65, t3: 125; f1: 478, f2: 308, IO: 453, D: 328, d: 219, PO: 0.67.