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Catalogue
Catalogue of the type material of Scarabaeoidea (Coleoptera) deposited in the Research Institute of Evolutionary Biology, Tokyo, Japan
expand article infoNaoki Kaneko, Kaoru Wada§
‡ Tokyo University of Agriculture, Tokyo, Japan
§ Meisei University, Tokyo, Japan
Open Access

Abstract

A detailed catalogue of the type material of the Scarabaeoidea housed in the Research Institute of Evolutionary Biology, Tokyo, Japan is listed. The catalogue includes the data of the type material of four families and 111 species.

Keywords

Allotype, Geotrupidae, holotype, Lucanidae, neotype, paratype, Scarabaeidae, Trogidae

Introduction

In this report we present a catalogue of the type material of Scarabaeoidea housed in the Research Institute of Evolutionary Biology, Tokyo, Japan (RIEB). The beginning of the RIEB collection dates back to 1950, when the first natural objects were deposited at the Research Institute of Thremmatology of the Tokyo University of Agriculture (RIT) established that year. RIT was transformed into an incorporated foundation in 1974 and renamed Research Institute of Evolutionary Biology (Yamamoto 2000). The Lepidoptera collection in the RIEB was listed in detail by Aoki et al. (2005, 2008) and Yamaguchi et al. (2006, 2008, 2009, 2011, 2017), but the Coleoptera collection has never been catalogued. This creates a problem because well-organized museum collections and published type specimen catalogues are very important to facilitate taxonomic and systematic investigations of animals.

The majority of the types of Scarabaeidae deposited in the RIEB originate from the collection of Mr Yoshikazu Miyake (1926–2003), a well-known Japanese amateur specialist on Scarabaeidae, who was also the author of those taxa. A part of types of Miyake’s species is also housed in the National Museum of Nature and Science, Tokyo (NMNS) and the Nagaoka Municipal Science Museum, Niigata (NMSM). Unfortunately, some of his types are missing.

In addition, other researchers that contributed to the Scarabaeoidea type collection in RIEB are: Dirk Ahrens, Aleš Bezděk, Toshiyuki Ichikawa, Takeshi Itoh (Takeshi Matsumoto), Kimio Masumoto, Sizumu Nomura, Teruo Ochi, and Kaoru Wada.

Materials and methods

The scientific names of higher taxa in this catalogue follow Scholtz and Grebennikov (2016). The structure of each entry is as follows:

– original combination of the taxon name;

– original combination and spelling of the taxon name, followed by the author, year of description, and pagination;

– type material, number of specimens (including sex, if known), and exact label data. A double slash ‘//’ indicates separate labels and single slash ‘/’ indicates lines within each label. The words in Japanese or Chinese were transcribed into Roman alphabet. Paratypes with discrepancies between collection data on the label and data quoted in the original description are indicated by ‘ [[]]’;

– remarks on types condition (given only for holotypes and neotypes);

– current taxonomic status;

– remarks, if any.

The majority of holotypes and neotypes were photographed with a Nikon D7200 camera using a Nikon AF-S DX Micro NIKKOR 40 mm f/2.8 G lens, and some types were photographed with a KEYENCE VHX-1000 Digital Microscope.

Catalogue

Family Geotrupidae

Subfamily Geotrupinae

Genus Sinogeotrupes

Sinogeotrupes taiwanus Miyake & Yamaya

Sinogeotrupes taiwanus Miyake & Yamaya, 1995: 32−34.

Note

The following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Paratypes

3 exs.: 1 ex. ‘Wusha / –VI–1941 / R. Boloudo // Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’. 1 ex. ‘Wu sha / Formosa / 21. Viii. 1941 / Col. Bumua Tin // Geotrupes / subshiatelluo FAIRMAIR / DET. Y. MIYAKE // Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’. 1 ex. ‘台湾= taiwan 台中= taichû / 関刀山= guandaoshan / 1993. V. 31 // Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’.

Current status

Phelotrupes (Sinogeotrupes) taiwanus (Miyake and Yamaya, 1995), see Nikolajev et al. (2016).

Remark

In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 3 exs. ‘Tattaka / Formosa / 10. Vi. 1965 / T. Shirozu // Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’. 1 ex. ‘(FORMOSA) / Mt Rata / Vi–Vii–1972 / L. F, Hang // Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’. 1 ex. ‘(FORMOSA) / Mt Rata / –Vii–1972 / L. F, Hang // Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’. 2 exs. ‘ [Taiwan] / 台中= taichû 関刀山= guandaoshan / 3–VI–1993 // Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’. 1 ♂ ‘ [Taiwan] / 台中= taichû 関刀山= guandaoshan / 3–VI–1993 // Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’. 3 exs. ‘Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’.

Family Lucanidae

Subfamily Lucaninae

Genus Aegus

Aegus laevicollis nakanei Ichikawa & Imanishi

Aegus laevicollis nakanei Ichikawa & Imanishi, 1976: 9.

Note

The following paratype is deposited in RIEB (ex coll. Y. Kusui):

Paratype

1 ex.: [[1 ♀ ‘Mt. Yonahadake, / Kunigami-son, / Is. Okinawa-Hontō, / 28. VI. 1972 (the date given in the original description is 16–17. VI. 1970), / HEIKICHI IRIE leg., // KUSUI // Paratype’]].

Current status

Aegus nakanei nakanei Ichikawa and Imanishi, 1976, see Huang and Chen (2017).

Family Scarabaeidae

Subfamily Aphodiinae

Genus Aphodius

Aphodius (Agoliinus) tanakai Masumoto

Aphodius (Agoliinus) tanakai Masumoto, 1981: 73−74.

Note

The following paratype is deposited in RIEB (ex coll. K. Masumoto):

Paratype

1 ex.: ‘Nikko / Date: 23 V 1981 / K. MASUMOTO leg. // Paratype / Aphodius (Agr) / tanakai MASUMOTO’.

Current status

Agoliinus tanakai (Masumoto, 1981), see Dellacasa et al. (2016).

Aphodius (Phaeaphodius) himalorectus Ahrens & Stebnicka

Aphodius (Phaeaphodius) himalorectus Ahrens & Stebnicka, 1997: 12−13.

Note

The following paratypes are deposited in RIEB (ex coll. D. Ahrens):

Paratypes

2 exs.: ‘Phalante 20. 3. / unt. Nebelwaldstufe / 2100–2300 m // PARATYPUS / Aphodius (Phaeaphodius) himalorectus sp. n. / det. D. AHRENS 1994’.

Current status

Phaeaphodius himalorectus (Ahrens and Stebnicka, 1997), see Dellacasa et al. (2016).

Aphodius (Sinodiapterna) zeni Ochi

Aphodius (Sinodiapterna) zeni Ochi, 1991: 53−55.

Note

The following paratype is deposited in RIEB (ex coll. T. Ochi):

Paratype

1 ex.: ‘SUNGKAN / C. TAIWAN / 29–V. 1987 / K. LAH // PARATYPE / Aphodius (Sinodiapterna) / zeni / OCHI 1991’.

Current status

Sinodiapterna zeni (Ochi, 1991), see Dellacasa et al. (2016).

Genus Rhyparus

Rhyparus kitanoi Miyake

Figure 1A

Rhyparus kitanoi Miyake, 1982: 65−67.

Note

The holotype and following specimen are deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘C, Sata /〔KYUSHU〕/ Japan / 25, VI 1971 / T. Kitano leg. // PARATYPE // Holotype: / Rhyparus / kitanoi / Y. MIYAKE, 1992’. (Fig. 1A)

Figure 1. 

Habitus of holotype specimens. A Rhyparus kitanoi (Miyake) B Adoretus (Lepadoretus) ubonensis (Miyake, Yamaguchi et Aoki) C Anomala flavoguttata Miyake D Anomala ohmomoi Miyake, Yamaguchi et Aoki E Anomala thailandiana Miyake, Yamaguchi et Akiyama F Callistethus isidai Miyake G Malaia castanoptera Miyake H Malaia macassara Miyake.

Type condition

The right and left protarsus and the right antenna of the holotype are missing.

Current status

Rhyparus kitanoi kitanoi Miyake, 1982, see Ochi (2001).

Remark

In addition to the holotype mentioned above, the following specimen labeled as paratype is not designated in the original description: 1 ex.: ‘Sesonoura / Is. Shimokoshiki / Kagoshima-pref. / 18. VI. 1982 / S. Imasaka leg. // Paratype: Rhyparus / kitanoi / Y. MIYAKE, 1992’.

Subfamily Scarabaeinae

Genus Caccobius

Caccobius bawangensis Ochi, Kon & Kikuta

Caccobius bawangensis Ochi, Kon & Kikuta, 1997: 262−264.

Note

The following paratype is deposited in RIEB (ex coll. T. Ochi):

Paratype

1 ex.: ‘Mt. Bawang / Sarawak S. / VIII. 1991 / M. Kon Coll. // PARATYPE / Caccobius / bawangensis OCHI, KON and KIKUTA’.

Current status

Valid species.

Genus Copris

Copris (Microcopris) mindorensis Ochi

Copris (Microcopris) mindorensis Ochi, 1992: 13.

Note

The following paratype is deposited in RIEB (ex coll. T. Ochi):

Paratype

1 ex.: ‘MT. HALKON / IS. MINDRO / 28–XII. 1988 / D. MOHAGAN // PARATYPE / Copris / (Microcopris) / mindorensis / OCHI, 1992.

Current status

Valid species.

Genus Onthophagus

Onthophagus setchan Masumoto

Onthophagus setchan Masumoto, 1984: 80−81.

Note

The following paratype is deposited in RIEB (ex coll. K. Masumoto):

Paratype

1 ex.: ‘FENGCHIFU / FORMOSA / 7–VII. 74. / T. OCHI // PARATYPE / Onthophagus / setchan / MASUMOTO (1984)’.

Current status

Valid species.

Onthophagus (Parascatonomus) acuticollis sakishimanus Nomura

Onthophagus (Parascatonomus) acuticollis sakishimanus Nomura, 1976a: 29.

Note

The following paratypes are deposited in RIEB (ex coll. S. Nomura):

Paratypes

6 exs.: 3 ♂ ‘Kanpira / Iriomote Is. / Yaeyama Iss. / 12. VII. 1975 / R. YANO leg. // ♂ // Onthophagus (P.) / acuticollis / sakishimanus / NOMURA (1976)’. 3 ♀ ‘Kanpira / Iriomote Is. / Yaeyama Iss. / 12. VII. 1975 / R. YANO leg. // ♀ // Onthophagus (P.) / acuticollis / sakishimanus / NOMURA (1976)’.

Current status

Parascatonomus (Necramator) acuticollis sakishimanus (Nomura, 1976), see Ochi and Kon (2017).

Onthophagus (Parascatonomus) murasakianus Nomura

Onthophagus (Parascatonomus) murasakianus Nomura, 1976a: 30.

Note

The following paratype is deposited in RIEB (ex coll. S. Nomura):

Paratype

1 ex.: ‘Kanpira, / Iriomote Is. / Yaeyama Iss. / 12. VII 1975 / R. YANO leg. // PARATYPE / Onthophagus (P.) / murasakianus / NOMURA (1976)’.

Current status

Parascatonomus (Necramator) murasakianus (Nomura, 1976), see Ochi and Kon (2017).

Onthophagus (Parascatonomus) takedai Ochi & Araya

Onthophagus (Parascatonomus) takedai Ochi & Araya, 1992: 99−100.

Note

The following paratype is deposited in RIEB (ex coll. T. Ochi):

Paratype

1 ex.: ‘QUESON. N. P / IS. LUZON / 10–VIII. 1981 / K. SUGINO // PARATYPE / Onthophagus / (Parascatonomus) / takedai / OCHI et ARAYA, 1992’.

Current status

Parascatonomus (Granulidorsum) takedai (Ochi et Araya, 1992), see Ochi and Kon (2017).

Onthophagus (Proagoderus) watanabei Ochi & Kon

Onthophagus (Proagoderus) watanabei Ochi & Kon, 2002: 306−311.

Note

The following paratype is deposited in RIEB (ex coll. T. Ochi):

Paratype

1 ex.: ‘SEPIROK / N. BORNEO / 4–VII. 1985 / M. KON // PARATYPE / Onthophagus / (Proagoderus) / watanabei / OCHI and KON. 2002’.

Current status

Proagoderus watanabei (Ochi et Kon, 2002), see Philips (2016).

Onthophagus (Strandius) hiurai Ochi

Onthophagus (Strandius) hiurai Ochi, 1984: 63−65.

Note

The following paratype is deposited in RIEB (ex coll. T. Ochi):

Paratype

1 ex.: ‘PILU / Taiwan / 21, VIII, 1979 / K. Sugino leg. // PARATYPE / Onthophagus (St) / hiurai / OCHI (1984)’.

Current status

Onthophagus (Phanaeomorphus) potanini hiurai Ochi, 1984, see Ziani and Bezděk (2016).

Subfamily Rutelinae

Genus Adoretus

Adoretus (Lepadoretus) ubonensis Miyake, Yamaguchi & Aoki

Figure 1B

Adoretus (Lepadoretus) ubonensis Miyake, Yamaguchi, Aoki & Akiyama, 2002: 62−63.

Note

The holotype is deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘Shirinthorn (light) / Ubon Ratchatani / E–THAILAND / 27. Feb. 1999 / S. Yamaguchi and / T. Aoki leg. // No. 1 / ♂ // Holotype: / Adoretus / ubonensis // Holotype / Adoretus (Lepadoretus) ubonensis / Y. MIYAKE, YAMAGUCHI / et AOKI 2002’. (Fig. 1B)

Type condition

The left mesotarsus and the left metaleg of the holotype are missing.

Current status

Junior subjective synonym of Adoretus compressus (Weber, 1801), see Kobayashi (2018).

Genus Anomala

Anomala flavoguttata Miyake

Figure 1C

Anomala flavoguttata Miyake, 2000: 108−109.

Note

The holotype is deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘Sapa=Chapa / N. Vietnam / 20. V. 1994 / N. KATSURA // Holotype: Anomala / flavoguttata / Y. MIYAKE, 2000’. (Fig. 1C)

Type condition

The aedeagus of the holotype is pinned separately.

Current status

Valid species.

Anomala ohmomoi Miyake, Yamaguchi & Aoki

Figure 1D

Anomala ohmomoi Miyake, Yamaguchi, Aoki & Akiyama, 2002: 58−60.

Note

The holotype and the following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘Ban Lao Kram / Mukdahan (light) / E–THAILAND沢= sawa / 8. Mar. 1999 / S. Yamaguchi and / T. Aoki leg. // Holo Type: / Anomala / ohmomoi // Holotype / Anomala ohmomoi / Y. MIYAKE, YAMAGUCHI / et AOKI 2002’. (Fig. 1D)

Paratypes

4 exs.: ‘Ban Lao Kram / Mukdahan (light) / E–THAILAND沢= sawa / 8. Mar. 1999 / S. Yamaguchi and / T. Aoki leg. // Paratype / Anomala ohmomoi / Y. MIYAKE, YAMAGUCHI / et AOKI 2002’.

Current status

Valid species.

Anomala thailandiana Miyake, Yamaguchi & Akiyama

Figure 1E

Anomala thailandiana Miyake, Yamaguchi, Aoki & Akiyama, 2002: 60−61.

Note

The holotype is deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘Don Far Huan (light) / Ubon Ratchathani / E–THAILAND / 17. Sep. 1999 / S. Yamaguchi and / M. Akiyama leg. [white label, front] // Apigoni / 5 [white label, back] // Holotype / Anomala thailandiana / Y. MIYAKE, YAMAGUCHI / et AKIYAMA 2002’. (Fig. 1E)

Type condition

The aedeagus of the holotype is pinned separately.

Current status

Valid species.

Genus Callistethus

Callistethus isidai Miyake

Figure 1F

Callistethus isidai Miyake, 1987: 5−6.

Note

The holotype and the following paratype are deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘ [Taiwan] / Lishan to Pilu / 31–VII–1972 / Y. Miyake leg. // Holotype: / Callistethus isidai / Y. MYAKE, 1987’. (Fig. 1F)

Paratype

1 ex.: ‘ [Taiwan] / Lishan to Pilu / 31–VII–1972 / Y. Miyake leg. // Paratype / Callistethus isidia / Y. MIYAKE, 1987’.

Type condition

The right and left metatarsi of the holotype are missing.

Current status

Callistethus plagiicollis isidai Miyake, 1987, see Zorn and Bezděk (2016).

Remark

The habitus photograph in the original description does not agree with the holotype. This is also the case with some other species described by Miyake. Apparently, he did not intend to provide the photographs of the holotypes. Also, as noted below, we were unable to trace some specimens illustrated in the original descriptions.

Genus Kibakoganea

Kibakoganea ohtanii Miyake & Muramoto

Kibakoganea ohtanii Miyake & Muramoto, 1992: 22−24.

Note

The following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Paratypes

2 exs.: [[1 ♂, 1 ♀ ‘Tam Dao / N. VIETNAM / 28–30, IV, 1991 (the date given in the original description is 28–30. III. 1991) / M. Fujioka and / R. Muramoto – leg. // PARATYPE / Kibakoganea / ohtanii / Y. MIYAKE et / MURAMOTO, 1992.’]].

Current status

Junior subjective synonym of Kibakoganea sexmaculata (Kraatz, 1900), see Miyake (2003).

Genus Malaia

Malaia castanoptera Miyake

Figure 1G

Malaia castanoptera Miyake, 2000: 112.

Note

The holotype is deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘Malino, Celebes / Indonesia / 21 Nov 1944 // malino / 2603.11.21 // Holotype: Malaia / castanoptera / Y. MIYAKE, 2000’. (Fig. 1G)

Paratypes

7 exs.: 1 ♂, 3 ♀ ‘Malino, Celebes / Indonesia / 21 Nov 1944 // malino / 2603.11.21 // Paratype: Malaia / castanoptera / Y. MIYAKE, 2000’. 2 exs. ‘Malino, Celebes / Indonesia / 21 Nov 1944 // malino / 2603.11.21 // Paratype: Malaia / castanoptera / Y. MIYAKE, 2000’. 1 ex. ‘malino / 2603.11.21 // Paratype: Malaia / castanoptera / Y. MIYAKE, 2000’.

Current status

Valid species.

Remark

The habitus photo in the original description belongs to the female paratype with the label ‘Malino, Celebes / Indonesia / 21 Nov 1944 // malino / 2603.11.21’.

Malaia cyanea Miyake

Malaia cyanea Miyake, 1996: 38−39.

Note

The following paratypes are deposited is RIEB (ex coll. Y. Miyake):

Paratypes

2 exs.: 1 ♂ ‘32 km from / Palopo / Surawesi / 10. VI. 1982 // Paratype: / Malaia cyanea / Y. MIYAKE, 1996’. 1 ♀ ‘Punchak–Lombanan / Mamasa, / S. SULAWESI / 1991. AUG. 10. / N. Kashiwai–leg. // Paratype: / Malaia cyanea / Y. MIYAKE, 1996’.

Current status

Valid species.

Malaia macassara Miyake

Figure 1H

Malaia macassara Miyake, 2000: 109−110.

Note

The holotype and the following paratype are deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘Lampbatter / Sulawesi / 23–25. VIII. 1999 / M. Takai // Holotype: Malaia / macassara / Y. MIYAKE, 2000’. (Fig. 1H)

Paratype

1ex.: 1 ♂ ‘Lampbatter / Sulawesi / 23–25. XII. 1999 / M. Takai leg. // Paratype: Malaia / macassara / Y. MIYAKE, 2000’.

Type condition

The left protarsus and metatarsus of the holotype are missing.

Current status

Valid species.

Malaia rufofemorata Miyake

Malaia rufofemorata Miyake, 1996: 40−41.

Note

The following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Paratypes

4 exs.: 1 ♂, 1 ♀ ‘Puncak, Palopo / VI–VII, 1989 / Central SULAWESI // Paratype: Malaia / rufofemorata / Y. MIYAKE, 1996’. 2 ♂ ‘Palu Palolo / VIII. 1990 / Central SULAWESI // Paratype: Malaia / rufofemorata / Y. MIYAKE, 1996’.

Current status

Valid species.

Remark

In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 2 ♂, 1 ♀ ‘Palolo, Palu / 01–10/XI. 1990 / native coll. / CELEBES // Paratype: Malaia / rufofemorata / Y. MIYAKE, 1996’.

Malaia sulawesi Miyake

Malaia sulawesi Miyake, 1996: 41−42.

Note

The following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Paratypes

58 exs.: 1 ♂ ‘Mt. Pedamaran / Tana Toraja / 1985. III–17 / South SULAWESI / Soma, K. leg. // Paratype: / Malaia / sulawesi / Y. MIYAKE, 1996’. 1 ex. ‘Puncak Palopo / 1989. III. 31 / Centr. CELEBES // Paratype: / Malaia / sulawesi / Y. MIYAKE, 1996’. 3 exs. ‘Palu Pulu / C. Sulawesi / II. 1989 // Paratype: / Malaia / sulawesi / Y. MIYAKE, 1996’. [[44 exs. ‘Mt. Pedamaran / S. Sulawesi (real data is C. Sulawesi) / 20. X. 1983 (real date is 20. V. 1983) / K. Soma leg. // Paratype: / Malaia / sulawesi / Y. MIYAKE, 1996’]]. [[9 exs. ‘Mt. Pedamaran / Tana Toraja / 1983. X–20 (real date is 1983. V–20) / South SULAWESI (real data is Central SULAWESI) / Soma, K. leg. // Paratype: / Malaia / sulawesi / Y. MIYAKE, 1996’]].

Current status

Valid species.

Malaia taoi Miyake

Malaia taoi Miyake, 1996: 39−40.

Note

The following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Paratypes

4 exs.: 1 ♂ ‘37 km from / Palopo C.- / Sulawesi / 9. IV. 1982 // Paratype: / Malaia / taoi / Y. MIYAKE, 1996’. 1 ♀ ‘32 km from / Palopo / Surawesi / 10. VI. 1982 // 37 km from / Palopo C.- / Sulawesi / 10. VI. 1982 // Paratype: / Malaia / taoi / Y. MIYAKE, 1996’. 1 ♂ ‘37 km from / Palopo C.- / Sulawesi / 11. IV. 1982 // Paratype: / Malaia / taoi / Y. MIYAKE, 1996’. 1 ♀ ‘32 km from / Palopo / Surawesi / 13. VI. 1982 // 37 km from / Palopo C.- / Sulawesi / 13. VI. 1982 // Paratype: / Malaia / taoi / Y. MIYAKE, 1996’.

Current status

Valid species.

Malaia tondanoensis Miyake

Malaia tondanoensis Miyake, 1996: 42−43.

Note

The following paratype is deposited in RIEB (ex coll. Y. Miyake):

Paratype

4exs.: ‘Tondano / N. Sulawesi / 10–IV–1989 / Y. Miyake leg. // Paratype: / Malaia / tondanoensis / Y. MIYAKE, 1996’.

[[1 ex. ‘Tangkoko N. / N. Sulawesi / 10–III–1989 (the date given in the original description is 10–IV–1989) / Y. Miyake leg. // Paratype: / Malaia / tondanoensis / Y. MIYAKE, 1996’]]. [[2 exs. ‘Tondano / N. Sulawesi / 9–IV–1989 (the date given in the original description is 10–IV–1989) / Y. Miyake leg. // Paratype: / Malaia / tondanoensis / Y. MIYAKE, 1996’]].

Current status

Valid species.

Malaia toraja Miyake

Figure 2A

Malaia toraja Miyake, 2000: 110−111.

Note

The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘Mt. Pedamaran / Tana Traja / 20–X. 1983 / K. SOHMA lgd. / South SULAWESI // Holotype: Malaia / toraja / Y. MIYAKE, 2000’. (Fig. 2A)

Figure 2. 

Habitus of holotype and neotype specimens. A Malaia toraja Miyake B Mimela kitanoi Miyake C Mimela marginipennis Miyake D Spinanomala moritai Miyake E Peltonotus morio Burmeister F Peltonotus morio sawaii Miyake G Apogonia ohmomoi Miyake H Apogonia ovata Miyake et Yamaguchi.

Paratypes

27 exs.: 1 ♂ ‘Mt. Pedamaran / Tana Traja / 20–X. 1983 / K. SOHMA lgd. // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 9 exs. ‘Mt. Pedamaran / Tana Traja / 20–X. 1983 / K. SOHMA lgd. // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 2 exs. ‘Mt. Pedamaran / S. Sulawesi / 20. X. 1983 / K. Soma leg. // Paratype: Malaia / toraja / Y. MIYAKE, 2000’.8 exs. ‘Puncak, Palopo / VI–VII, 1989 / Central SULAWESI // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 2 exs. ‘Central SULAWESI / Puncak, Palopo / VI–VII, 1989 // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 1 ♂ ‘Puncak Palopo / 1989. / Centr. CELEBES // SI [yellow label] // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 4 exs. ‘Puncak Palopo / 1989. / Centr. CELEBES // SI [yellow label] // Paratype: Malaia / toraja / Y. MIYAKE, 2000’.

Current status

Valid species.

Remark

The habitus photo in the original description belongs to the female paratype with the label ‘Mt. Pedamaran / Tana Toraja / S. Sulawesi / 7, VIII, 1988 / K. Soma leg.’.

In addition to the paratypes listed above, the following specimens labeled as the paratypes are not designated in the original description: 1 ♀ ‘Mt. Pedamaran / Tana Toraja / S. Sulawesi / 7, VIII, 1988 / K. Soma leg. // SI [yellow label] // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 1 ex. ‘(Pulu Pulu) / C. Sulawesi / INDONESIA / Dec. 1988 // [yellow label] // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 2 exs. ‘Puncak, Palopo / Centr. CELEBES / 1989. I / INDONESIA // [yellow label] // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 1 ex. ‘Puncak Palopo / 1989. III. 31 / Centr. CELEBES // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 3 exs. ‘Palu, Palolo / VII 1989 / Central SULAWESI // Paratype: Malaia / toraja / Y. MIYAKE, 2000’.

Genus Mimela

Mimela kitanoi Miyake

Figure 2B

Mimela kitanoi Miyake, 1987: 7−8.

Note

The holotype is deposited in RIEB (ex coll. K. Wada):

Holotype

(♀). ‘Larashan / Hsinchu Hs. / VI. 1975, / Fang leg // Holotype: / Mimela kitanoi / Y. MIYAKE, 1987 // Kaoru WADA / Collection 2001’. (Fig. 2B)

Current status

Junior subjective synonym of Mimela flavocincta Lin, 1966, see Zorn and Bezděk (2016).

Mimela marginipennis Miyake

Figure 2C

Mimela marginipennis Miyake, 1994: 151.

Note

The holotype is deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘[Borneo] / Crocker Range / 1800−1400m alt. / nr. Keningau C. / X. 1988. M. ITOI leg. // Holotype: / Mimela / marginipennis / Y. MIYAKE, 1994’. (Fig. 2C)

Type condition

The aedeagus of the holotype is pinned separately, and the left mesotarsus is missing.

Current status

Junior subjective synonym of Mimela margarita Arrow, 1910, see Wada (2001).

Mimela sawaii Miyake

Mimela sawaii Miyake, 1994: 149−150.

Note

The following paratype is deposited in RIEB (ex coll. Y. Miyake):

Paratype

1 ex.: 1 ♂ ‘Crocker Range / (1000–1400 m) / Nr. Keningau C. / N. Borneo, – / 15–V–1989 // [blue label] // Paratype: / Mimela / sawaii / Y. MIYAKE, 1994’.

Current status

Junior subjective synonym of Mimela pallidicauda Arrow, 1910, see Wada (2001).

Mimela suspecta Miyake

Mimela suspecta Miyake, 1994: 150.

Note

The following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Paratype

2ex.: 1 ♂ ‘Mt. Trus Madi / (SW Slopa 1200m alt.) / Sabah, MALAYSIA / 14–IV–1993 / Minoru SAWAI leg. // Paratype: / Mimela / suspecta / Y. MIYAKE, 1994’. [[1 ex. ‘Mt. Trus Madi / (SW Slopa 1200m alt.) / Sabah, MALAYSIA / 18–IV–1993 (the date given in the original description is 4–IV–1993) / Minoru SAWAI leg. // [yellow label] // Paratype: / Mimela / suspecta / Y. MIYAKE, 1994’]].

Current status

Valid species.

Remark

In addition to the paratypes mentioned above, the following specimen labeled as paratype is not designated in the original description: 1 ♂ ‘Trus Madi Mts. / near Keningau / Saba MALAYSIA / 1993. iv. 17–30. // F //Paratype: / Mimela / suspecta / Y. MIYAKE, 1994’.

Genus Popillia

Popillia iwasei Miyake

Popillia iwasei Miyake, 1996: 43−44.

Note

The following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Paratypes

6 exs.: 2 ♀ ‘Puncak, Palopo / Centr. CELEBES / 1989. I / INDONESIA // Paratype: / Popillia / iwasei / Y. MIYAKE, 1996’. 2 ♀ ‘Puncak Palopo / C. Sulawesi / 1. 1989 // Paratype: / Popillia / iwasei / Y. MIYAKE, 1996’. 1 ♀ ‘Puncak Palopo / C. Sulawesi / 1989. V / INDONESIA // SI [yellow label] // Paratype: / Popillia / iwasei / Y. MIYAKE, 1996’. [[1 ♂ ‘Palolo (the locality given in the original description is Palopo) / II. 1991 (the date given in the original description and Telnov and Zorn (2019) are I. 1989) // Paratype: / Popillia / iwasei / Y. MIYAKE, 1996’]].

Current status

Valid species.

Genus Spilopopillia

Spilopopillia rugosa Miyake

Spilopopillia rugosa Miyake, 1989a: 180−181.

Note

The following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Paratypes

18 exs.: [[1 ex. ‘ [Malaya] / Fraser’s Hill / 29–III–1977 (the date given in the original description is 28. III. 1977) / Y. Miyake leg. // Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’]]. [[1 ex. ‘ [Malaya] / Freaser’s Hill (the locality given in the original description is Fraser’s Hill) / 30–III–1977 (the date given in the original description is 28. III. 1977) / Y. Miyake leg. // Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’]]. 6 exs. ‘ [Malaya] / Tanah Rata / 30–III–1974 / Y, Miyake leg // Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’. 9 exs. ‘TANARATA / MALAYA / 30 III 1974 / Y. Miyake leg. // Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’. [[1 ex. ‘Gentig / Hiland (the locality given in the original description is Genting Highland) MALAYA / 5. IV. 1974 (the date given in the original description is 5. IV. 1975) / Y Miyake leg. // Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’]].

Current status

Valid species.

Remark

In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 8 exs. ‘Tanah Rata / MALAYA / 1. III. 1977 / Y. Miyake // Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’. 5 exs. ‘TANARATA / MALAYA / 28, 29, 30 III 1974 / Y. Miyake leg. // Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’. 1 ex. ‘84. IV // Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’. 2 exs. ‘Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’.

Genus Spinanomala

Spinanomala moritai Miyake

Figure 2D

Spinanomala moritai Miyake, 1996: 38.

Note

The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘Tanah Rata / MALAYA / IV. 1984 / M. Hata // Holotype: / Spinanomala / moritai / Y. MIYAKE, 1996’. (Fig. 2D)

Paratypes

11 exs.: 1 ex. ‘Tanah Rata / MALAYA / I–II. 1973 / MORITA, Kozo // Paratype: Spinanomala / moritai / Y. MIYAKE, 1996’. 2 ♂, 3 ♀ ‘Tanah Rata / MALAYA / 1. III. 1977 / Y. Miyake // Paratype: Spinanomala / moritai / Y. MIYAKE, 1996’. 3 ♂, 2 ♀ ‘Tanah Rata / MALAYA / IV. 1984 / M. Hata // Paratype: Spinanomala / moritai / Y. MIYAKE, 1996’.

Type condition

The aedeagus of the holotype is pinned separately and the right metaleg is missing.

Current status

Valid species.

Remark

The collecting data of the specimen labelled as the holotype do not match the original description. There are two male paratypes (‘Tanah Rata / MALAYA / 1. III. 1977 / Y. Miyake // Paratype: Spinanomala / moritai / Y. MIYAKE, 1996’) that match the holotype data.

In addition to the paratypes mentioned above, the following specimen labeled as the paratype is not designated in the original description: 1 ex. ‘フレーザーヒル= furêzâhiru / MALAYA / 4 – 6. IV. 1977 / M. Hata // Paratype: Spinanomala / moritai / Y. MIYAKE, 1996’.

Subfamily Dynastinae

Genus Papuana

Papuana timorensis Miyake & Yamaya

Papuana timorensis Miyake & Yamaya, 1999: 103−104.

Note

The following paratype is deposited in RIEB (ex coll. Y. Miyake):

Paratype

1 ex.: 1 ♂ ‘Timor Is. / Indonesia / V. 1993 / Native Coll. // Timor / Is. [aedeagus mount] // Paratype: / Papuana / timorensis YAMAYA / Y. MIYAKE, 1999’.

Current status

Valid species.

Genus Peltonotus

Peltonotus morio Burmeister

Figure 2E

Peltonotus morio Burmeister, 1847: 75.

Note

The neotype is deposited in RIEB (ex coll. K. Wada):

Neotype

(♂). ‘NEPAL・HIMALAYA / 10. 5. 1993 Chhachok Madi– / Khola/ Gandaki (Kaski– / Destr.) 1300m lg: D. Ahrens //Peltonotus / morio Burn. / det: D. Ahrens 1997 // Neotype / Peltonotus / morio Burmeister, 1847 / Jameson and Wada, 2004’. (Fig. 2E)

Type condition

The aedeagus of the neotype is pinned separately.

Current status

Valid species.

Remark

The neotype was designated by Jameson and Wada (2004).

Peltonotus morio sawaii Miyake

Figure 2F

Peltonotus morio sawaii Miyake, 2000: 112−113.

Note

The holotype and the following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘Doi Suthep / Chieng Mai / N–THAILAND/ 22. V. 1983 / leg. M. Sawai // 12 // P. morio / Burmi. [white label, front] // Cameron / Highland / Pahang [white label, back] // Holotype: Peltonotus / morio sawaii / Y. MIYAKE, 2000’. (Fig. 2F)

Paratypes

4 exs.: 2 ♂, 1 ♀ ‘Doi Suthep / Chieng Mai / N–THAILAND / 22. V. 1983 / leg. M. Sawai // Paratype: Peltonotus / morio sawaii / Y. MIYAKE, 2000 // P. morio /Burmeister’. 1 ♀ ‘Doi Suthep / Chieng Mai / N–THAILAND / 22. V. 1983 / leg. M. Sawai // Paratype: Peltonotus / morio sawaii / Y. MIYAKE, 2000’.

Type condition

The aedeagus of the holotype is pinned separately.

Current status

Junior subjective synonym of Peltonotus morio Burmeister, 1847, see Krell and Bezděk (2016).

Peltonotus podocrassus Jameson & Wada

Peltonotus podocrassus Jameson & Wada, 2004: 34−37.

Note

The following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Paratypes

7 exs.: 2 exs. ‘ [Malaya] / Tanah Rata / 30–III–1976 / Y. Miyake leg. // Paratype: Peltonotus peninsularis / Y. MIYAKE, 2000 // Paratype / Peltonotus podocrassus / Jameson and Wada, 2004’. 1 ♂ ‘39 miles from / Tapah, Malaya / 30–III–1976 / Y. Miyake leg. // Paratype: Peltonotus peninsularis / Y. MIYAKE, 2000 // Paratype / Peltonotus podocrassus / Jameson and Wada, 2004’. 1 ♂ ‘TANARATA / MALAYA / 30. III. 1976 / Y. Miyake leg. // Paratype: Peltonotus peninsularis / Y. MIYAKE, 2000 // Paratype / Peltonotus podocrassus / Jameson and Wada, 2004’. 1 ♂ ‘V. R〔C. H.〕/ Malaya / 21. III. 1978 // 46 // Paratype: Peltonotus peninsularis / Y. MIYAKE, 2000 // Paratype / Peltonotus podocrassus / Jameson and Wada, 2004’. 1 ex. [Malaya] / Tanah Rata / 1–IV–1984 / Y. Miyake leg. // Paratype: Peltonotus peninsularis / Y. MIYAKE, 2000 // Paratype / Peltonotus podocrassus / Jameson and Wada, 2004’. 1 ex. ‘TANARATA / MALAYA / 1. IV. 1984 / Y. Miyake // Paratype: Peltonotus peninsularis / Y. MIYAKE, 2000 // Paratype / Peltonotus podocrassus / Jameson and Wada, 2004’.

Current status

Valid species.

Peltonotus sakaii Miyake & Yamaya

Peltonotus sakaii Miyake & Yamaya, 1994: 39−40, 42.

Note

The following paratype is deposited in RIEB (ex coll. Y. Miyake):

Paratype

1 ex.: ‘Mt. Serapi / SARAWAKU / 1991/III. // 20 // Paratype: / Peltonotus sakaii / Y. MIYAKE et YAMAYA, 1994 // Peltonotus / similis Arrow, 1931 / det. Kaoru Wada, 2004’.

Current status

Junior subjective synonym of Peltonotus similis Arrow, 1931, see Krajcik (2012).

Remark

In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 1 ♂ ‘Mt. Serapi / Kuching, Sarawak / IX. 1990 / BORNEO // Paratype: / Peltonotus sakaii / Y. MIYAKE et YAMAYA, 1994 // Peltonotus / similis Arrow, 1931 / det. Kaoru Wada, 2004’. 1 ex. ‘Lembah Anai / W. Sumatra / VI. 1990 / INDONESIA // [yellow label] // Paratype: / Peltonotus sakaii / Y. MIYAKE et YAMAYA, 1994 // Peltonotus / similis Arrow, 1931 / det. Kaoru Wada, 2004’. 1 ♂ ‘Kimanis Road / Near Keningau / N. Borneo / 3–V–1994 // Paratype: / Peltonotus sakaii / Y. MIYAKE et YAMAYA, 1994 // Peltonotus / similis Arrow, 1931 / det. Kaoru Wada, 2004’. 2 exs. ‘Kimanis Road / Near Keningau / N. Borneo / 3–V–1994 // Paratype: / Peltonotus sakaii / Y. MIYAKE et YAMAYA, 1994 // Peltonotus / similis Arrow, 1931 / det. Kaoru Wada, 2004’.

Peltonotus sulawesiensis Wada

Peltonotus sulawesiensis Wada, 1990: 1−2.

Note

The following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Paratypes

4 exs.: ‘Tondano N. Celebes / INDONESIA / V 1988 / leg. N. Nisikawa // PARATYPE / Peltonotus sulawesiensis / Kaoru WADA, 1990’.

Current status

Junior subjective synonym of Neohyphus celebesus Heller, 1896, see Kuijten (1994).

Subfamily Melolonthinae

Genus Amiserica

Amiserica rufidula Nomura

Amiserica rufidula Nomura, 1974: 84.

Note

The following paratypes are deposited in RIEB (ex coll. S. Nomura):

Paratypes

2 exs.: 1 ♂ ‘Li-shan / Taiwan / 29. VII. 1974 / Y. Miyake // ♂ // PARATYPE / Amiserica / rufidula / NOMURA (1974)’. 1 ♂ ‘Li-shan / Taiwan / 1. VIII. 1974 / Y. Miyake // ♂ // PARATYPE / Amiserica / rufidula / NOMURA (1974)’.

Current status

Valid species.

Genus Apogonia

Apogonia fujiokai Miyake & Yamaya

Apogonia fujiokai Miyake & Yamaya, 1997: 6−7.

Note

The following paratype is deposited in RIEB (ex coll. Y. Miyake):

Paratype

1 ex.: ‘Flores Is. / Indonesia / May. 1995 // Y // Paratype: / Apogonia fujiokai / Y. MIYAKE et YAMAYA, 1997’.

Current status

Valid species.

Apogonia fusciventris Miyake & Yamaya

Apogonia fusciventris Miyake & Yamaya, 1997: 5−6.

Note

The following paratype is deposited in RIEB (ex coll. Y. Miyake):

Paratype

1 ex.: ‘Vangvieng / P. Vietiane / LAO. P. D. R. / 25–26. MAY. 1994. / K. Miura–leg. // Y // Paratype: Apogonia / fuscoventris / Y. MIYAKE et YAMAYA, 1997’.

Current status

Junior subjective synonym of of Apogonia striatipennis Frey, 1971, see Kobayashi (2010).

Remark

The species name on the paratype label is ‘fuscoventris’, which is invalid. Probably, the name was confused with fusciventris by Y. Miyake.

Apogonia hongkongica Miyake

Apogonia hongkongica Miyake, 1989b: 38−39.

Note

The following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Paratypes

11 exs.: 1 ♂ ‘Hong Kong / China / C. K. Yu / 1974. V. [white label, front] // Native Collector / legt. [white label, back] // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’. 1 ♂ ‘Hong Kong / China / V. 1974 / C. K. Yu [white label, front] // Legt. / Native / Collector [white label, back] // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’. 1 ♂ ‘Hong Kong / China / V. 1974 / C. K. Yu [white label, front] // Native Collector [white label, back] // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’. 1 ♀ ‘♀ // Hong Kong / China / V. 1974 / C. K. Yu [white label, front] // Legt. / Native / Collector // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’. 1 ♀ ‘♀ // Hong Kong / China / V. 1974 / C. K. Yu [white label, front] // Native / Collector / legt. // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’. 1 ♀ ‘Hong Kong / 1974 / From C. K. Yu ♀ // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’. 1 ex. ‘Hong Kong / China / V. 1974 / C. K. Yu [white label, front] // Legt. / native / Collector [white label, back] // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’. 2 exs. ‘Hang Kong / China / V. 1974 / C. K. Yu Col [white label, front] // Native / Collector / legt. // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’. 2 exs. ‘Hong Kong / V. 1974 / From C. K. Yu // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’.

Current status

Valid species.

Remark

In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 1ex. ‘Hong Kong / 1975 / From. C. K-Yu // Paratype: Apogonia / hongkongica / Y. MIYAKE, 1989’. 1 ♀ ‘Hong Kong / ♀ 1975 [white label, front] // from / C. K. Yu [white label, back] // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’.

Apogonia inconstans Kobayashi & Bezděk

Apogonia inconstans Kobayashi & Bezděk, 2011: 64−65.

Note

The following paratypes are deposited in RIEB (ex coll. A. Bezděk):

Paratypes

4 exs.: 1 ♂ ‘1985. 5. 18−19. / Wieng Ko Sai Nat. / Park, Phrae Prov., / N-Thailand / Col. S. Ohmomo [white label, front] // 5/18 夜= yoru [white label, back] // PARATYPE / Apogonia / inconstans / Kobayashi et Bezděk, 2011’. 1 ex. ‘1985. 5. 18−19. / Wieng Ko Sai Nat. / Park, Phrae Prov., / N-Thailand / Col. S. Ohmomo [white label, front] // 5/18 夜= yoru [white label, back] // PARATYPE / Apogonia / inconstans / Kobayashi et Bezděk, 2011’. 1 ex. ‘1985. 5. 18−19. / Wieng Ko Sai Nat. / Park, Phrae Prov., / N-Thailand / Col. S. Ohmomo [white label, front] // 夜= yoru 5/18 [white label, back] // PARATYPE / Apogonia / inconstans / Kobayashi et Bezděk, 2011’. 1 ex. ‘1985. 5. 18−19. / Wieng Ko Sai Nat. / Park, Phrae Prov., / N-Thailand / Col. S. Ohmomo [white label, front] // night [white label, back] // PARATYPE / Apogonia / inconstans / Kobayashi et Bezděk, 2011’.

Current status

Valid species.

Apogonia ohmomoi Miyake

Figure 2G

Apogonia ohmomoi Miyake, 1989: 177−178.

Note

The holotype is deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘1985. 5. 18−19. / Wieng Ko Sai Nat. / Park, Phrae Prov., / N−Thailand / Col. S. Ohmomo [white label, front] // 夜= yoru 5/18 [white label, back] // 14 // Holotype: / Apogonia / scabra / Y. MIYAKE // Apogonia ohmomoi / Miyake, 1989 / HOLOTYPUS, ♂ / A. Bezděk det. 2011’. (Fig. 2G)

Type condition

The aedeagus of the holotype is pinned separately, and the right antenna, right and left protarsus, right mesoclaw, left mesotarsus, right metatarsus and mesonotum are missing.

Current status

Valid species.

Remark

The label ‘Apogonia scabra’ was apparently accidentally put by Miyake.

Apogonia ovata Miyake & Yamaguchi

Figure 2H

Apogonia ovata Miyake & Yamaguchi, 1998: 25−27.

Note

The holotype is deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘Kaeng Tana / Khong Chiam / Ubonratchathani / [NE-THAILAND] / 29. Aug. 1997 / S. Yamaguchi leg. // Holotype / Apogonia ovata / Y. MIYAKE et YAMAGUCHI 1998’. (Fig. 2H)

Type condition

The aedeagus of the holotype is pinned separately.

Current status

Senior synonym of Apogonia miyakei Bezděk, 2004, see Bezděk (2004).

Apogonia terminalis Miyake, Yamaguchi & Akiyama

Figure 3A

Apogonia terminalis Miyake, Yamaguchi & Akiyama, 2002: 56−58.

Note

The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘Don Far Huan (light) / Ubon Ratchathani / E-THAILAND / 17. Sep. 1999 / S. Yamaguchi & / M. Akiyama leg. // Holotype / Apogonia terminalis / Y. MIYAKE, YAMAGUCHI / et AKIYAMA 2002’. (Fig. 3A)

Figure 3. 

Habitus of holotype specimens. A Apogonia terminalis Miyake, Yamagushi et Akiyama B Apogonia unidentata Miyake, Yamaguchi et Akiyama C Dichelomorpha sublineata Miyake D Holomelia gigantea Miyake E Holotrichia loochooana umebayashii Miyake F Holotrichia yamayai Miyake et Yamaguchi G Hoplia choui Miyake H Hoplia nakanei Miyake.

Paratypes

4exs.: 1 ♂ ‘Ban Lao Kram / Mukdahan (light) / NE-THAILAND / 5. Sep. 1999 / S. Yamaguchi & / M. Akiyama leg. // Paratype / Apogonia terminalis / Y. MIYAKE, YAMAGUCHI / et AKIYAMA 2002’. 3 ♀ ‘Don Far Huan (light) / Ubon Ratchathani / E-TAILAND / 17. Sep. 1999 / S. Yamaguchi & / M. Akiyama leg. // ♀ // Paratype / Apogonia terminalis / Y. MIYAKE, YAMAGUCHI / et AKIYAMA 2002’.

Type condition

The aedeagus of the holotype is pinned separately, and the left proclaw, right and left mesotarsus, right metaclaw and left metatarsus are missing.

Current status

Valid species.

Apogonia unidentata Miyake, Yamaguchi & Akiyama

Figure 3B

Apogonia unidentata Miyake, Yamaguchi & Akiyama, 2002: 55−56.

Note

The holotype and following paratypes are deposited in RIEB (ex coll. Miyake):

Holotype

(♂). ‘Ban Lao Kram / Mukdahan (light) / NE-THAILAND / 5. Sep. 1999 / S. Yamaguchi & / M. Akiyama leg. // Holotype / Apogonia / unidentata // Holotype / Apogonia unidentata / Y. MIYAKE, YAMAGUCHI / et AKIYAMA 2002’. (Fig. 3B)

Paratypes

6 exs.: 4exs. ‘Ban Lao Kram / Mukudahan (light) / NE-THAILAND / 5. Sep. 1999 / S. Yamaguchi & / M. Akiyama leg. // Paratype / Apogonia unidentata / Y. MIYAKE, YAMAGUCHI / et AKIYAMA 2002’. 1 ex. ‘Ban Lao Kram / Mukudahan (light) / NE-THAILAND / 5. Sep. 1999 / S. Yamaguchi & / M. Akiyama leg. // 63 // Paratype / Apogonia unidentata / Y. MIYAKE, YAMAGUCHI / et AKIYAMA 2002’. 1 ex. ‘Ban Lao Kram / Mukudahan (light) / NE-THAILAND / 14. Sep. 1999 / S. Yamaguchi & / M. Akiyama leg. / シミレア= shimirea // Paratype / Apogonia unidentata / Y. MIYAKE, YAMAGUCHI / et AKIYAMA 2002’.

Type condition

The aedeagus of the holotype is pinned separately, and the right protarsus and right and left metatarsi are missing.

Current status

Valid species.

Genus Dichelomorpha

Dichelomorpha sublineata Miyake

Figure 3C

Dichelomorpha sublineata Miyake, 1994: 140−141.

Note

The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘Is. Tioman, east of / Malay Pennisular March. 1993, nat, col // 1 [aedeagus mount] // Holotype: / Dichelomorpha / sublineata / Y. MIYAKE, 1994’. (Fig. 3C)

Paratypes

34 exs.: 10 ♂ ‘Is. Tioman, east of / Malay Pennisular / March. 1993. nat, col // Paratype: / Dichelomorpha / sublineata / Y. MIYAKE, 1994’. 14 exs. ‘Is. Tioman, east of / Malay Pennisular / March. 1993. nat, col // Paratype: / Dichelomorpha / sublineata / Y. MIYAKE, 1994’. 1 ♂ ‘Is. Tioman, east of / Malay Pennisular / March. 1993. nat, col // 11 // Paratype: / Dichelomorpha / sublineata / Y. MIYAKE, 1994’. 1 ex. ‘Is. Tioman, east of / Malay Pennisular / March. 1993. nat, col // 4 // Paratype: / Dichelomorpha / sublineata / Y. MIYAKE, 1994’. [[8 exs. ‘Tioman Is. / E. of Malaya / III–1933 (the date given in the original description is III–1993), W. // Paratype: / Dichelomorpha / sublineata / Y. MIYAKE, 1994’]].

Type condition

The aedeagus of the holotype is pinned separately. The right mesotarsus is missing.

Current status

Valid species.

Remark

Miyake (1994) did not provide a photograph of the holotype. Apparently he provided a photograph of a paratype, but we could not trace that specimen.

Genus Holomelia

Holomelia gigantea Miyake

Figure 3D

Holomelia gigantea Miyake, 1996: 35−36.

Note

The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake).

Holotype

(♂). ‘19 miles from / Tapah, Malaya / 1–II–1974 / Y. Miyake leg [white label, front] // Wu TET KONG / legt. [white label, back] // 29・5 // 家= ie [aedeagus mount] // Holotype: / Holomelia / gigantea / Y. MIYAKE, 1996’. (Fig. 3D)

Paratypes

2 exs.: 1 ♂ ‘Cameron H. / MALAYA / Jan–Feb 1974 / Y. Miyake leg. // 30.5 // Paratype: / Holomelia gigantea / Y. MIYAKE, 1996 // 19 miles from / Tapah, Malaya / 1–II–1974 [white label, front] // Wu Tet Kong legt. [white label, back]’. 1 ex. ‘19 miles from / Tapah, Malaya / 1–II–1974 [white label, front] // Wu Tet Kong legt. [white label, back] // Paratype: / Holomelia gigantea / Y. MIYAKE, 1996’.

Type condition

The aedeagus of the holotype is pinned separately.

Current status

Valid species.

Remark

Miyake (1996) did not provide a photograph of the holotype. Apparently he provided a photograph of a paratype, but we could not trace that specimen.

Genus Holotrichia

Holotrichia loochooana umebayashii Miyake

Figure 3E

Holotrichia loochooana umebayashii Miyake, 1986a: 1−2.

Note

The holotype is deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘〔Loo-Choo〕/ Okinoerabuzima / 21. IV. 1957. / K. Umebayashi // 7 [white label, mounted on a part of tarsus] // HOLOTYPE / Holotrichia / loochooana / umebayashii / Y. MIYAKE, 1986’. (Fig. 3E)

Type condition

The aedeagus of the holotype is pinned separately. The left protarsus and the right and left metatarsus are missing.

Current status

Junior subjective synonym of Nigrotrichia loochooana okinawana (Nomura, 1964), see Matsumoto (2016).

Remark

The collecting date in the original description is 1.V.1957, which disagrees with the label data.

Holotrichia yamayai Miyake & Yamaguchi

Figure 3F

Holotrichia yamayai Miyake & Yamaguchi, 1998: 27−28.

Note

The holotype and following paratype are deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘North Thailand / Chiangmai-Saraphee / 15th, June, 1996 / Native collector // Holotype / Holotricchia yamayai / Y. MIYAKE et YAMAGUCHI 1998’. (Fig. 3F)

Paratype

1 ex.: 1 ♀ ‘Kaeng Tana / Khong Chiam / Ubonratohathani / [NE-THAILAND] / 27. Aug. 1997 / S. Yamaguchi leg. // Paratype / Holotrichia yamayai / Y. MIYAKE et YAMAGUCHI 1998’.

Type condition

The aedeagus of the holotype is pinned separately.

Current status

Valid species.

Genus Hoplia

Hoplia choui Miyake

Figure 3G

Hoplia choui Miyake, 1986b: 203−204.

Note

The holotype is deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘Sozan / Formosa / 26. V. 1940 / Coll. M. Uno // HOLOTYPE / Hoplia / Choui Y. MIYAKE / 1986’. (Fig. 3G)

Type condition

Abdomen only.

Current status

Valid species.

Remark

Most part of the holotype specimen was destroyed by Anthrenus verbasci.

Hoplia nakanei Miyake

Figure 3H

Hoplia nakanei Miyake, 1986b: 207.

Note

The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘ (Taiwan) / Jibbon / 2. iv. 1970 / Y. Miyake // 1 // HOLOTYPE / Hoplia / nakanei Y. MIYAKE’. (Fig. 3H)

Paratypes

29 exs.: 4 exs. ‘ [Taiwan] / 六亀= liugui.石山= shi shan / 9–VI–1976 / Y. Miyake leg. // Paratype / Hoplia / nakanei / Y. Miyake, 1986’. 5 exs. ‘ [Taiwan] / Liukuei 石山= shi shan / 9–VI–1976 / Y. Miyake leg. // Paratype / Hoplia / nakanei / Y. Miyake, 1986’. 1 ex. ‘ [Taiwan] / 六亀= liugui.石山= shi shan / 9–VI–1976 / 陳文竜= chen wenzhao leg. // Paratype / Hoplia nakanei / Y. Miyake, 1986’. 4 exs. ‘ [Taiwan] / Baibara / –V–1963 / T. Shirozu leg. // Paratype / Hoplia / nakanei / Y. Miyake, 1986’. 1 ex. ‘ [Taiwan] / Baibara / –V–1963 / Shirozu leg. // Paratype / Hoplia / nakanei / Y. Miyake, 1986’. 1 ex. ‘ [Taiwan] / Baibara / –V–1963 / T. Shirozu // Paratype: Hoplia / nakanei Y. Miyake, 1986’. 1 ♂ ‘六亀= liugui.石山= shi shan / alt. 2000 m / 9. VI. 1976 // Paratype: / Hoplia / nakanei / Y. MIYAKE, 1986’. 1 ex. ‘FENCHIFO / FORMOSA / 24. V. 1975 / K. MATSUDA // Paratype: / Hoplia nakanei / Y. MIYAKE, 1986’. 1 ex. ‘Swakang / Taiwan / 15. VI. 1970 / Fukuda // Paratype: / Hoplia nakanei / 1986. Y. MIYAKE’. 1 ex. ‘NANSHANCHI / TAIWAN / 1. IV. 1970 / H. NOMURA // PARATYPE / Hoplia / nakanei / Y. MIYAKE, 1986’. [[4 exs. ‘ [Taiwan] / Baibara / V. 1965 (the date given in the original description is V. 1963) / T. Shirozu // Paratype / Hoplia nakanei / Y. Miyake’]]. [[2 exs. ‘ [Taiwan] / Baibara / V. 1965 (the date given in the original description is V. 1963) / T. Shirozu // Paratype / Hoplia nakanei / Y. Miyake, 1986’]]. [[1 ♂ ‘ [Taiwan] / Baibara / V. 1965 (the date given in the original description is V. 1963) / Col. T. Shirozu // Paratype / Hoplia nakanei / Y. Miyake, 1986’]]. [[1 ex. ‘ [Taiwan] / Baibara / V. 1965 (the date given in the original description is V. 1963) / Col. T. Shirozu // Paratype / Hoplia nakanei / Y. Miyake, 1986’]]. [[1 ex. ‘ [Taiwan] / Baibara / V. 1965 (the date given in the original description is V. 1963) / Col. T. Shirozu // Paratype / Hoplia nakanei / 1986. Y. Miyake’]].

Type condition

The aedeagus of the holotype is pinned separately.

Current status

Valid species.

Remark

The label data the holotype do not match the original description. There is one male paratype (‘六亀= liugui.石山= shi shan / alt. 2000 m / 9. VI. 1976 // Paratype: / Hoplia / nakanei / Y. MIYAKE, 1986’) that agrees with the data of the holotype. Probably the type labels were mixed up and this specimen should be considered the holotype.

In addition to the paratypes listed above, the following specimens labeled as paratypes are not designated in the original description: 1 ex. ‘ [Taiwan] / Liukuei / –V–1963 / T. Shirozu col. // Paratype / Hoplia / nakanei / Y. Miyake, 1986’. 1 ex. ‘APR II. 1971 / 榮華= ronghua / B-S. CHANG // PARATYPE / Hoplia / nakanei / Y. MIYAKE, 1986’. 1 ex. ‘Paratype / Hoplia nakanei / Y. MIYAKE, 1986’.

Hoplia nengkaoshana Miyake

Hoplia nengkaoshana Miyake, 1986b: 211.

Note

The allotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Allotype (♀). ‘Allotype // mt. Noko (2650 m) / V. 20–VI. 2, 1966 // 1 // PARATYPE / Hoplia / nengkaoshana / Y. MIYAKE, 1986’.

Paratypes

3ex.: 1 ♂ ‘mt. Nōhō / C-Formosa / V–20 ~ Vi–21 1966 // PARATYPE / Hoplia / nengkaoshana / Y. MIYAKE, 1986’. 1 ♂ ‘MT. HÔFAN / FORMOSA / 5–V. 73 / T. OCHI // 1 // PARATYPE / Hoplia / nengkaoshana / Y. MIYAKE, 1986 // Hoplia / inornata / H. Kobayashi, 1990 / Det. H. Kobayashi, 2015’. 1 ex. ‘JUL. 16. 1966 / 合欢山= hehuan shan / B. S. Chang // PARATYPE / nengkaoshana / Y. MIYAKE, 1986 // Hoplia / inornata / H. Kobayashi, 1990 / Det. H. Kobayashi, 2015’.

Current status

Valid species.

Remark

In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 1 ex. ‘台中= tai chung 和平郷= heping xiang 早等村= zaodeng cun / 横貫公路= hengguan gonglu. 梨山站= lishan zhan / 海抜= kaibatsu 1679–2196 m. / 13–16. V. 1958. 陶家駒= tao jiayu // PARATYPE / Hoplia / nengkaoshana / Y. MIYAKE, 1986’. 1 ex. ‘AUG 13. 1970 / 新坡= xinpo / B-S. CHANG // PARATYPE / Hoplia / nengkaoshana / Y. MIYAKE, 1986’.

Hoplia simillima Miyake

Figure 4A

Hoplia simillima Miyake, 1986b: 206−207.

Note

The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘ [Taiwan] / Liukuei 石山= shi shan / 9–VI–1979 / Y. Miyake leg. // HOLOTYPE / Hoplia / simillima / Y. MIYAKE, 1986’. (Fig. 4A)

Figure 4. 

Habitus of holotype specimens. A Hoplia simillima Miyake B Hoplia taiwana Miyake C Maladera hiranoi Miyake D Maladera kusuii Miyake E Nematophylla sugiharai Miyake F Pseudohoplia shibatai makiharai Miyake G Pseudohoplia shibatai matsudai Miyake H Paratrichius kyushuensis Miyake.

Paratypes

5 exs.: 3 exs. ‘(FORMOSA) / Baibara / –V–1969 // Paratype / Hoplia / simillima / 1989, Y. Miyake’. 2 exs. ‘(FORMOSA) / Baibara / –V–1969 / T. Shirozu // Paratype / Hoplia / simillima / 1989, Y. Miyake’.

Current status

Valid species.

Remark

In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 1 ex. ‘ [Taiwan] / Liukuei 石山= shi shan / 9–VI–1979 / Y. Miyake leg. // Paratype / Hoplia / simillima / 1989, Y. Miyake’. 1 ♂ ‘石= ishi // [Taiwan] / 六亀= liugui 石山= shi shan / 9–VI–1976 / Y. Miyake leg. // Paratype / Hoplia / simillima / 1989, Y. Miyake’. 1 ex. ‘(FORMOSA) / Mt. Ali / 14 V 1968 / Y. HAYASHI // Paratype’.

Hoplia taiwana Miyake

Figure 4B

Hoplia taiwana Miyake, 1986b: 204.

Note

The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘HOLOTYPE / Hoplia / Taiwana Y. MIYAKE / 1986’. (Fig. 4B)

Paratypes

9 exs.: 1 ex. ‘NANSHANCHI / TAIWAN / 3. IV. 1970 / H. NOMURA // PARATYPE / Hoplia taiwana / Y. MIYAKE, 1986’. 1 ex. ‘NANSHANCHI / TAIWAN / 4. IV. 1970 / H. NOMURA // PARATYPE / Hoplia taiwana / Y. MIYAKE, 1986’. 1 ex. ‘NANSHANCHI / TAIWAN / 5. IV. 1970 / Y. KIYOYAMA // PARATYPE / Hoplia taiwana / Y. MIYAKE, 1986’. 1 ex. ‘SUNGKANG / FORMOSA / 29. VI. 1971 / Y. MAEDA // PARATYPE / Hoplia taiwana / Y. MIYAKE, 1986’. 1 ex. ‘NANSHANCHI / TAIWAN / 31. III. 1981 / Y. YAMAMOTO // Paratype / Hoplia / taiwana / Y. MIYAKE, 1986’. 1 ex. ‘NANSHANCHI / TAIWAN / 1. IV. 1981 / F. KIMURA // Paratype / Hoplia / taiwana Y. MIYAKE / (1968)’. 1 ex. ‘NANSHANCHI / TAIWAN / 2. IV. 1981 / F. KIMURA // Paratype / Hoplia / taiwana Y. MIYAKE’. 1 ex. ‘NANSHANCHI / TAIWAN / 2. IV. 1981 / F. KIMURA // Paratype / Hoplia taiwana / 1986 Y. MIYAKE’. 1 ex. ‘NANSHANCHI / TAIWAN / 3. IV. 1981 / Y. YAMAMOTO // PARATYPE / Hoplia taiwana / Y. MIYAKE, 1986’.

Current status

Valid species.

Remark

The locality label data of the holotype are missing but the specimen agrees with the original description.

In addition to the paratypes mentioned above, the following specimens labeled as paratypes were not designated in the original description: 3 exs. ‘ [Taiwan] / Liukuei 石山= shi shan / 9–VI–1976 / Y. Miyake leg. // PARATYPE / Hoplia / taiwana / Y. MIYAKE, 1986’. 1 ex. ‘ [Taiwan] / 六亀= liugui 石山= shi shan / 9–VI–1976 / Y. Miyake leg. // PARATYPE / Hoplia / taiwana / Y. MIYAKE, 1986’. 2 exs. ‘Mt. YUSHAN / TAIWAN / 8. VI. 1980 / M. YAMAMOTO // Paratype / Hoplia / taiwana Y. MIYAKE’. 4 exs. ‘TAIWAN / 陽明山= mt. yangmingshan / 20. IV. 1956 / COL. K. S. LIN // Paratype / Hoplia / taiwana / Y. MIYAKE, 1986’. 1 ex. ‘LUSHAN / FORMOSA / 13–IV. 74 / S. TAKEDA // PARATYPE / Hoplia / taiwana / Y. MIYAKE, 1986’. 1 ex. ‘榮華= ronghua / APR. 7. 1971 / BS, CHANG // PARATYPE / Hoplia taiwana / Y. MIYAKE, 1986’. 1 ex. ‘ [Taiwana] / Baibara / –V–1965 / T. Shirozu leg. // Paratype: Hoplia / taiwana Y. MIYAKE. 1986’.

Genus Maladera

Maladera hiranoi Miyake

Figure 4C

Maladera hiranoi Miyake, 1986b: 3−5.

Note

The holotype is deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘三京= mikyô / 21/VII–82 // HOLOTYPE / Maladera / hiranoi MIYAKE / (1986) // Maladera / impressthorax / hiranoi Y. MIYAKE / DET. Y. MIYAKE, 199 // ムナクボビロウドコガネ= munakubobirôdokogane / Maladera (japonica–G) / impressithorax / Nomura, 1973 / Det. H. Hirasawa 2012’. (Fig. 4C)

Current status

Junior subjective synonym of Maladera okinoerabuana Kobayashi, 1978, see Ahrens and Bezděk (2016).

Remark

The date on the collecting data label of the holotype is 21.VII.1982, but the collecting date quoted in the original description is 22.VII.1982. Probably, Miyake made a typographical error when writing the description.

Maladera kobayashii Nomura

Maladera kobayashii Nomura, 1974: 106−107.

Note

The following paratypes are deposited in RIEB (ex coll. S. Nomura):

Paratypes

2 exs.: 1 ♂ ‘Li-shan / Taiwan / 29. VII. 1974 / Y. Miyake // ♂ // PARATYPE / Maladera (s. str) / kobayashii / NOMURA (1974)’. 1 ♀ ‘Li-shan / Taiwan / 29. VII. 1974 / Y. Miyake // ♀ // PARATYPE / Maladera / kobayashii / NOMURA (1974)’.

Current status

Valid species.

Maladera kusuii Miyake

Figure 4D

Maladera kusuii Miyake, 1986b: 2−3.

Note

The holotype is deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘三宅= miyake / コガネムシ上科標準図鑑= koganemushijyôka-hyôjyunzukan / ビロウドコガネ族= birôdokoganezoku / 図鑑使用標本= zukanshiyôhyôhon / 執筆担当= shippitsutantô:平沢 伴明= hirasawa tomoaki / 学研教育出版= gakkenkyôikushuppan (2012) // GAKIYA / Iheya-jima / Aug. 11 1977 / leg. Y Kusui // イヘヤビロウドコガネ= iheyabirôdokogane / Maladera (japonica–G) / kusuii / Y. Miyake, 1986 / Det. H. Hirasawa 2012 // Holotype: / Maladera kusuii / Y. MIYAKE, 1986’.

Type condition

The aedeagus of the holotype is pinned separately. The right and left protarsi, the left mesotarsus, and the right and left metatarsi are missing, but some of the missing parts are pinned separately.

Current status

Valid species.

Maladera lishana Miyake

Maladera lishana Miyake, 1989b: 37−38.

Note

The following paratype is deposited in RIEB (ex coll. Y. Miyake):

Paratype

1 ex.: 1 ♀ ‘Lishan / Taiwan / 28. VII. 1973 / Y. Miyake // Paratype: / Maladera / lishana / Y. MIYAKE, 1989’.

Current status

Valid species.

Maladera nanshanchiana Nomura

Maladera nanshanchiana Nomura, 1974: 111−112.

Note

The following paratypes are deposited in RIEB (ex coll. S. Nomura):

Paratypes

2 exs.: 1 ex. ‘Puli 24 // PARATYPE / Maladera / nanshanchiana / NOMURA (1974)’. 1 ♀ ‘Puli 24 // ♀ // PARATYPE / Maladera / nanshanchiana / NOMURA (1974) // ナンザンケイビロウドコガネ= nanzenkeibirodokogane / Maladera (Cephaloserica) / nanshanchiana / Nomura, 1974 / Det. H. Hirasawa 2012’.

Current status

Valid species.

Maladera secreta horaiana Nomura

Maladera secreta horaiana Nomura, 1974: 112.

Note

The following paratype is deposited in RIEB (ex coll. S. Nomura):

Paratype

1 ex.: 1 ♂ ‘Lishan / C. Taiwan / 29. vii. 1973 / Y. Miyake // 三宅= miyake // PARATYPE / Maladera / secreta / horaiana / NOMURA (1974)’.

Current status

Valid species.

Genus Melanomaladera

Melanomaladera yunnana Miyake & Yamaya

Melanomaladera yunnana Miyake & Yamaya, 2001: 38−39.

Note

The following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Paratypes

2 exs.: 1 ♂ ‘China, N W Yunnan / Degen City env. / 3300 m alt. / 29. Jun. 1998 / A. Gorodinsky leg. // 13 // Paratype: / Melanoserica / yunnana’. 1 ♀ ‘China, N W Yunnan / Degen City env. / 3300 m alt. / 29. Jun. 1998 / A. Gorodinsky leg. // 13 // Paratype: / Melanoserica / yunnana’.

Current status

Archeohomaloplia yunnana (Miyake et Yamaya, 2001), see Ahrens and Bezděk (2016).

Remark

The genus name on the paratype labels is ‘Melanoserica’, which is invalid. Probably, it was confused with ‘Melanomaladera’ by Y. Miyake.

Genus Microserica

Microserica (Parvulomaladera) annapurnae Ahrens

Microserica ( Parvulomaladera) annapurnae Ahrens, 1995: 46−48.

Note

The following paratype is deposited in RIEB (ex coll. D. Ahrens):

Paratype

1 ex.: ‘NEPAL–HIMALAYA / Annapurna–Mts. / leg. Ahrens 1993 // Pisang bis / Manang, 30. 5. / 3000–3300 m // PARATYPEUS / Parvulomaladera / annapumae sp. n. / det. D. AHRENS 1994’.

Current status

Oxyserica pygidialis annapurnae (Ahrens, 1995), see Ahrens and Bezděk (2016).

Microserica nitidipyga Nomura

Microserica nitidipyga Nomura, 1974: 99.

Note

The following paratype is deposited in RIEB (ex coll. S. Nomura):

Paratype

1 ex.: ‘Lu-SHAN / FORMOSA / 27. IV. 1973 / K. MATSUDA // 三宅= miyake // Paratype / Microserica / nitidipyga / NOMURA (1974) // Microserica / nitidipyga NOM. / DET. Y. MIYAKE, 1994’.

Current status

Valid species.

Genus Nematophylla

Nematophylla sugiharai Miyake

Figure 4E

Nematophylla sugiharai Miyake, 2000: 106−108.

Note

The holotype is deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘Kiau Gap / Mt. Kinabalu / Sabah, Borneo / 1. IV. 2000 / H. SUGIHARAI leg. // Holotype: Nematophylla / sugiharai / Y. MIYAKE 2000’. (Fig. 4E)

Current status

Junior subjective synonym of Nematophylla carinicollis Arrow, 1938, see Krajcik (2012).

Genus Nipponoserica

Nipponoserica pubiventris Nomura

Nipponoserica pubiventris Nomura, 1976b: 189−190.

Note

The following paratype is deposited in RIEB (ex coll. S. Nomura):

Paratype

1 ex.: ‘Takashinohama, / Osaka Pref. / 3. VI. 1974 / Coll. Mitsuo Goto // at light // PARATYPE / Nipponoserica / pubiventris / NOMURA (1976)’.

Current status

Valid species.

Genus Paramaladera

Paramaladera makiharai Nomura

Paramaladera makiharai Nomura, 1974: 101−102.

Note

The following paratypes are deposited in RIEB (ex coll. S. Nomura):

Paratypes

4 exs.: 4 ♂ ‘Li-shan / Taiwan / 29. VII. 1974 / Y. Miyake // ♂ // Paratype / Paramaladera / makiharai / NOMURA (1974)’.

Current status

Valid species.

Genus Pseudohoplia

Pseudohoplia shibatai Miyake

Pseudohoplia shibatai Miyake, 1986b: 201.

Note

The following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Paratypes

2 exs.: 1 ♂ ‘(FORMOSA)/ Mt. Ali / 15 V 1968 / Y. HAYASHI // Paratype // Hoplia / shibatai shibatai / Y. Miyake, 1986 / Det. H. Kobayashi, 2015’. 1 ex. ‘(FORMOSA)/ Mt. Ali / 15 V 1968 / Y. HAYASHI // Paratype // Hoplia / shibatai shibatai / Y. Miyake, 1986 / Det. H. Kobayashi, 2015’.

Current status

Hoplia shibatai shibatai (Miyake, 1986), see Kobayashi (2017).

Remark

In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 1 ex. ‘Mt. Arishan / Formosa / 1. V. 1973 / K. MATSUDA // Paratype // Paratype / Pseudohoplia / shibatai / Y. Miyake, 1986 // 贈台湾農試= zou-taiwan-noushi’. 1 ♂ ‘Houanchi- / Sundchuankang / Nantou Hsien / Taiwan / 25. VI. 1976 / H. Makihara leg. // Paratype // Paratype / Pseudohoplia / shibatai / Y. Miyake, 1986’. 1 ex. ‘Mt. Arishan / Formosa / 1. V. 1973 / K. MATSUDA // Paratype / Pseudohoplia / shibatai / Y. Miyake, 1986’. 1 ex. ‘MT. ARI / FORMOSA / 1. V. 1973 / K. MATSUDA // Paratype // Paratype / Pseudohoplia / shibatai / Y. Miyake, 1986’.

Pseudohoplia shibatai makiharai Miyake

Figure 4F

Pseudohoplia shibatai makiharai Miyake, 1986b: 202.

Note

The holotype is deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘Mt. Lalashan / Taoyuan Hsien / Taiwan / 21–24. V. 1980 / H. Makihara // HOLOTYPE / Pseudohoplia / shibatai makiharai / Y. MIYAKE, 1886’. (Fig. 4F)

Type condition

The aedeagus of the holotype is pinned separately. The right antenna is missing.

Current status

Hoplia shibatai makiharai (Miyake, 1986), see Kobayashi (2017).

Pseudohoplia shibatai matsudai Miyake

Figure 4G

Pseudohoplia shibatai matsudai Miyake, 1986b: 202.

Note

The holotype is deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘Mt. Nōko / C-Formosa / v–20~vi–21’ 66 // HOLOTYPE / Pseudohoplia / shibatai matsudai / Y. MIYAKE, 1986’. (Fig. 4G)

Type condition

The aedeagus of the holotype is pinned separately.

Current status

Junior subjective synonym of Hoplia shibatai shibatai (Miyake, 1986), see Kobayashi (2017).

Genus Pseudomaladera

Pseudomaladera nitidifrons Nomura

Pseudomaladera nitidifrons Nomura, 1974: 96.

Note

The following paratypes are deposited in RIEB (ex coll. S. Nomura):

Paratypes

2 exs.: 1 ♂ ‘FUNCHIHO / FORMOSA / 1. VIII. 1973 / H. OHASHI // ♂ // 三宅= miyake // Paratype / Pseudomaladera / nitidifrons / NOMURA, 1974’. 1 ♂ ‘MEIFENG / FORMOSA / 1. VIII. 1973 / T. OCHI // ♂ // Paratype / Pseudomaladera / nitidifrons / NOMURA, 1974’.

Current status

Valid species.

Genus Sinoserica

Sinoserica maculipennis Miyake & Yamaya

Sinoserica maculipennis Miyake & Yamaya, 2001: 40−41.

Note

The following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Paratypes

2 exs.: ‘China, Yunnan / Boshan C. env. / 2100 m alt. / 10. VII. 1998 / A. Gorodinski // 12 // Paratype / Sinoserica / maculipennis’.

Current status

Trioserica maculipennis (Miyake et Yamaya, 2001), see Ahrens and Bezděk (2016).

Genus Serica

Serica fusifemorata Nomura

Serica fusifemorata Nomura, 1974: 91.

Note

The following paratypes are deposited in RIEB (ex coll. S. Nomura):

Paratypes

11 exs.: 1 ♀ ‘Lishan / C. Taiwan / 29. vii. 1973 / Y. Miyake // ♀ // 三宅= miyake // Paratype / Serica / fusifemorata / NOMURA (1974)’. 2 ♀ ‘Li-shan / C. Taiwan / 26. VII. 1974 / Y. Miyake // ♀ // Paratype / Serica / fusifemorata / S. NOMURA (1974)’. 2 ♂ ‘Li-shan / C. Taiwan / 29. VII. 1974 / Y. Miyake // ♂ // Paratype / Serica / fusifemorata / S. NOMURA (1974)’. 2 ♀ ‘Li-shan / C. Taiwan / 29. VII. 1974 / Y. Miyake // ♀ // Paratype / Serica / fusifemorata / S. NOMURA (1974)’. 1 ♂ ‘Li-shan / C. Taiwan / 1. VIII. 1974 / Y. Miyake // ♂ // Paratype / Serica / fusifemorata / S. NOMURA (1974)’. 2 ♀ ‘Li-shan / C. Taiwan / 1. VIII. 1974 / Y. Miyake // ♀ // Paratype / Serica / fusifemorata / S. NOMURA (1974)’. 1 ♀ ‘27 // ♀ // Paratype / Serica / fusifemorata / S. NOMURA (1974)’.

Current status

Valid species.

Serica gansuensis Miyake & Yamaya

Serica gansuensis Miyake & Yamaya, 2001: 35−36.

Note

The following paratype is deposited in RIEB (ex coll. Y. Miyake):

Paratype

1 ex.: ‘China, Gangu / Mts. Minshan / nr. Wudn. 2100 / 1. VI. 1997 / A. Gorodinski // 4 // Paratype: / Serica / gansuensis’.

Current status

Valid species.

Genus Sericania

Sericania miyakei Nomura

Sericania miyakei Nomura, 1960: 60.

Note

The following paratype is deposited in RIEB (ex coll. S. Nomura):

Paratype

1 ex.: ‘Paratype // TASHIRO / CHIKUGO / 4. V. 1952 / Y. MIYAKE // Sericania / miyakei / NOMURA (1960)’.

Current status

Valid species.

Genus Sophrops

Sophrops takatoshii Itoh

Sophrops takatoshii Itoh, 1990: 9−10.

Note

The following paratypes are deposited in RIEB (ex coll. T. Itoh):

Paratypes

3 exs.: ‘1989–4–17 / 平良植物園= hirara-syokubutsuen /(宮古島)= miyako-jima / leg. T. Ueno // Paratype / Sophrops / takatoshii / T. Itoh’.

Current status

Valid species.

Genus Taiwanoserica

Taiwanoserica elongata Nomura

Taiwanoserica elongata Nomura, 1974: 86.

Note

The following paratype is deposited in RIEB (ex coll. S. Nomura):

Paratypes

1 ex.: 1 ♂ ‘MT. ARI / FORMOSA / 30. VII. 1973 / H. OHASHI // ♂ // 三宅= miyake // Paratype / Taiwanoserica / elongata / NOMURA (1974)’.

Current status

Serica (Taiwanoserica) elongata (Nomura, 1974), see Ahrens and Bezděk (2016).

Taiwanoserica lishana Nomura

Taiwanoserica lishana Nomura, 1974: 87.

Note

The following paratypes are deposited in RIEB (ex coll. S. Nomura):

Paratypes

22 exs.: 2 ♂ ‘Lishan / C. Taiwan / 30. vii. 1970 / Y. Miyake // ♂ // PARATYPE / Taiwanoserica / lishana / S. Nomura (1974)’. 6 ♂ ‘Lishan / C-Taiwan / 29. vii. 1973 / Y. Miyake // ♂ // PARATYPE / Taiwanoserica / lishana / S. Nomura (1974)’. 4 ♀ ‘Lishan / C-Taiwan / 29. vii. 1973 / Y. Miyake // ♀ // PARATYPE / Taiwanoserica / lishana / S. Nomura (1974)’. 4 ♂ ‘Li-shan / Taiwan / 29. VII. 1974 / Y. Miyake // ♂ // PARATYPE / Taiwanoserica / lishana / S. Nomura (1974)’. 1 ♀ ‘Li-shan / Taiwan / 29. VII. 1974 / Y. Miyake // ♀ // PARATYPE / Taiwanoserica / lishana / S. Nomura (1974)’. 4 ♂ ‘Li-shan / Taiwan / 1. VII. 1974 // Y. Miyake // ♂ // PARATYPE / Taiwanoserica / lishana / S. Nomura (1974)’. 1 ♀ ‘Li-shan / Taiwan / 1. VII. 1974 // Y. Miyake // ♀ // PARATYPE / Taiwanoserica / lishana / S. Nomura (1974)’.

Current status

Serica (Taiwanoserica) lishana (Nomura, 1974), see Ahrens and Bezděk (2016).

Genus Trichomaladera

Trichomaladera elongata Nomura

Trichomaladera elongata Nomura, 1974: 93−94.

Note

The following paratype is deposited in RIEB (ex coll. S. Nomura):

Paratype

1 ex.: 1 ♂ ‘Paratype / Trichomaladera / elongata / NOMURA (1974)’.

Current status

Valid species.

Remark

The collecting data label of the paratype is missing.

Genus Trichoserica

Trichoserica elongata nitididorsis Nomura

Trichoserica elongata nitididorsis Nomura, 1971: 70−71.

Note

The following paratype is deposited in RIEB (ex coll. S. Nomura):

Paratype

1 ex.: ‘Mt. Hiko / [KYUSHU] / 27. 7. 1960 / TAKEISHI // Paratype / Trichoserica / elongata / nitididorsis / Nomura’.

Current status

Serica nitididorsis (Nomura, 1971), see Ahrens and Bezděk (2016).

Trichoserica incurvata Nomura

Trichoserica incurvata Nomura, 1971: 67−68.

Note

The following paratypes are deposited in RIEB (ex coll. S. Nomura):

Paratypes

8 exs.: 1 ex. ‘Aomori-pref / 17–vii–1956 / S. Kimotog // Paratype / Trichoserica / incurvata / Nomura (1971) // Aomori / 17. VII. 1956 / S. Kimoto’. 1 ♂ ‘17. VII. 1956 / S. Kimoto // Paratype / Trichoserica / incurvata / Nomura (1971)’. 2 exs. ‘Aomori-pref / 19–vii–1956 / S. Kimotog // Paratype / Trichoserica / incurvata / Nomura (1971) // Kimoto S. / 19. VII. 1956’. 1 ex. ‘Aomori-pref / 19–vii–1956 / S. Kimotog // Paratype / Trichoserica / incurvata / Nomura (1971) // 19. VII. 1956 / S. Kimoto’. 2 exs. ‘Sensuidani 1200 m / Okutama Tokyo / 21–VII–1968 / Coll. H. Yuasa // Paratype / Trichoserica / incurvata / Nomura (1971)’. 1 ex. ‘Okutama / 21. VII. 1968 / YUASA // Paratype / Trichoserica / incurvata / Nomura (1971)’.

Current status

Serica incurvata (Nomura, 1971), see Ahrens and Bezděk (2016).

Trichoserica nitidiceps Nomura

Trichoserica nitidiceps Nomura, 1971: 69−70.

Note

The following paratype is deposited in RIEB (ex coll. S. Nomura):

Paratype

1 ex.: 1 ♂ ‘(Kyushu) / Mt. Sobo / 23. V. 1952 / Y. Miyake //Paratype / Trichoserica / nitidiceps / NOMURA’.

Current status

Junior subjective synonym of Serica inexspectata (Kontkanen, 1956), see Ahrens and Bezděk (2016).

Remark

In addition to the paratype mentioned above, the following specimen labeled as paratype is not designated in the original description: 1 ex. ‘Mt. Kuju / Bungo / 28. vii. 1961 / Y. Miyake // Paratype / Trichoserica / nitidiceps / NOMURA (1971)’.

Trichoserica opacifrons Nomura

Trichoserica opacifrons Nomura, 1971: 68−69.

Note

The following paratypes are deposited in RIEB (ex coll. S. Nomura):

Paratypes

3 exs.: 1 ♂ ‘Mt. Hiko / Buzen / 1. VIII. 1953 / Y. Miyake // Paratype / Trichoserica / opacifrons / NOMURA’. 1 ex. ‘mt. Kujū / Hokkeein / 27. VII. 1961 / Y. Miyake Coll // Paratype / Trichoserica / opacifrons / NOMURA’. 1 ♂ ‘ [Kyushu] / Yakushima / 21. Vi. 1954 / Col. K. Matsuda // Paratype / Trichoserica / opacifrons / NOMURA // Ophthalmoserica/ matsudai / sp.n. / DET. Y. MIYAKE’.

Current status

Senior objective synonym of Serica planifrons Nomura, 1972, see Ahrens and Bezděk (2016). Nomura (1972) proposed Serica planifrons as a replacement name for Serica opacifrons (Nomura, 1971), a junior secondary homonym of Serica opacifrons Fairmaire, 1891.

Subfamily Cetoniinae

Genus Anomalocera

Anomalocera paotao Masumoto & Sakai

Anomalocera paotao Masumoto & Sakai, 1987a: 51−54.

Note

The following paratypes are deposited in RIEB (ex coll. Masumoto):

Paratypes

2 exs.: 1 ex. ‘Lishan / Taiwan / 28. VII. 1974 / Y. Miyake // Paratype / Anomalocera / paotao MAS. et SAK.’. 1 ex. ‘Lishan / C, -Taiwan / 29 vii. 1973 / Y. Miyake // Paratype / Anomalocera / paotao MAS. et Sak.’.

Current status

Diphyllomorpha paotao (Masumoto et Sakai, 1987), see Bezděk (2016).

Genus Rhomborhina

Rhomborhina kurosawai Masumoto & Sakai

Rhomborhina kurosawai Masumoto & Sakai, 1987b: 104−105, 107.

Note

The following paratypes are deposited in RIEB (ex coll. Masumoto):

Paratypes

3 exs.: 2 ♂ ‘FORMOSA / tattaka / 9, VIII 1971 / H. OHARA // Paratype / Rhomborrhina / kurosawai / MASUMOTO et K. SAKAI’. [[1 ♂ ‘Tsuifeng / C. Tsiwan / 26. vii. 1970 (the date given in the original description is 27.vii.1970) // Y. Miyake // Paratype / Rhomborrhina / kurosawai / MASUMOTO et K. SAKAI’]].

Current status

Valid species.

Genus Thaumastopeus

Thaumastopeus chicheryi tenuigenius Miyake & Yamaya

Thaumastopeus chicheryi tenuigenius Miyake & Yamaya, 1995: 38.

Note

The following paratype is deposited in RIEB (ex coll. Y. Miyake):

Paratype

1 ex.: 1 ♂ ‘N. Keningau / N. Borneo / 30. V. 1994 // Paratype: Thaumastopeus / chicheryi tenuigenius / Y. MIYAKE et YAMAYA, 1995’.

Current status

Valid species.

Genus Paratrichius

Paratrichius hatai Miyake

Paratrichius hatai Miyake, 1989: 41−42.

Note

The following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Paratypes

4 exs.: [[‘Tanah Rata / MALAYA / 7. IV. 1972 (the date given in the original description is 7.IV.1978) / M. Hata // Paratype: / Paratrichius / hatai / Y. MIYAKE, 1989’]].

Current status

Valid species.

Paratrichius kyushuensis Miyake

Figure 4H

Paratrichius kyushuensis Miyake, 1990: 30−32.

Note

The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘MT. SOBO / BUNGO / 28. vii. 1954 / Y. MIYAKE // Holotype: / Paratrichius / kyushuensis / Y. MIYAKE, 1990’. (Fig. 4H)

Paratypes

5 exs.: 1 ex. ‘(Kyushu) / Mt. Sobo / 24. VII. 1950 / Y. Miyake // Paratype: Paratrichius / kyushuensis / Y. MIYAKE, 1990’. 1 ♂ ‘(Kyushu) / Mt. Sobo / 24. VII. 1952 / Y. Miyake // Paratype: Paratrichius / kyushuensis / Y. MIYAKE, 1990’. 1 ex. ‘(Kyushu) / Mt. Ichi Fu Sa / 28 VII 1952 / Col. S. Kimoto // Paratype: Paratrichius / kyushuensis / Y. MIYAKE, 1990’. 1 ex. 1 ♂ ‘(Kyushu) / Mt. Sobo / 24. VII. 1953 / K. MATSUDA // Paratype: Paratrichius / kyushuensis / Y. MIYAKE, 1990’.

Type condition

The left metatarsus of the holotype is missing.

Current status

Valid species.

Remark

The date on the label of the holotype is 28.VII.1954 while that in the original description is 28.VII.1952.

In addition to the paratypes mentioned above, the following specimen labeled as paratype is not designated in the original description.: 1 ex. ‘⑦ // Paratype: Paratrichius / kyushuensis / Y. MIYAKE, 1990’.

Genus Tibiotrichius

Tibiotrichius vietnamensis Miyake

Figure 5A

Tibiotrichius vietnamensis Miyake, 1996: 44.

Note

The holotype is deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘M. ITO leg / Sapa=Chapa / N. Vietnam / 23–V–1995 // サパ= sapa [aedeagus mount] // Holotype: Tibiotrichius / vietnamensis Y. MIYAKE, 1996’. (Fig. 5A)

Figure 5. 

Habitus of holotype specimens. A Tibiotrichius vietnamensis Miyake B Charitovalgus banzai Sawada C Dasyvalgus annamensis Miyake D Dasyvalgus castaneodorsalis Miyake E Dasyvalgus decamaculatus Miyake F Dasyvalgus flavicauda Miyake G Dasyvalgus macacus Miyake H Dasyvalgus multicus Miyake.

Type condition

The aedeagus of the holotype is pinned separately.

Current status

Valid species.

Subfamily Valginae

Genus Charitovalgus

Charitovalgus banzai Sawada

Figure 5B

Charitovalgus banzai Sawada, 1941: 8−9.

Note

The neotype is deposited in RIEB (ex coll. Y. Miyake):

Neotype

(♂). ‘Hourng Shoei ken / near Liukuei, / Kao Hsung Hsien / VII. 1972 W. Chen leg // Holotype: Neotype / Charitovalgus / banzai SAWADA / Y. MIYAKE, 1993’. (Fig. 5B)

Type condition

The right and left metatarsi of the neotype are missing.

Current status

Valid species.

Remark

The neotype was designated by Miyake (1993).

Genus Dasyvalgus

Dasyvalgus annamensis Miyake

Figure 5C

Dasyvalgus annamensis Miyake, 1993a: 21−22.

Note

The holotype and following paratype are deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘4. 4 / 203 // Iluon Ma Thuot / Annam M. ITO // Holotype: Dasyvalgus / Annamensis / Y. MIYAKE, 1993’. (Fig. 5C)

Paratype

1 ex.: 1 ♂ ‘Iluon Ma Thuot / Annam M. ITO // Paratype: Dasyvalgus / annamensis / Y. MIYAKE, 1993’.

Current status

Valid species.

Remark

The locality on the collecting data label of the holotype is ‘Iluon Ma Thuot’, but the locality given in the original description is ‘Ban Me Thuot’. However, the habitus of the holotype and the figure in the original description match. Probably, Miyake made a typographical error when writing the description.

Dasyvalgus castaneodorsalis Miyake

Figure 5D

Dasyvalgus castaneodorsalis Miyake, 1993a: 24−25.

Note

The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘〔Thailand〕/ Pupin–Doi Phui / 19–IV–1985 / M. TAO leg. // Holotype: Dasyvalgus / casraneodorsalis / Y. MIYAKE, 1993’. (Fig. 5D)

Paratypes

6 exs.: 2 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 19–IV–1985 / M. TAO leg. // Paratype: Dasyvalgus / castaneodorsalis / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 21–IV–1985 / M. TAO leg. // Paratype: Dasyvalgus / castaneodorsalis / Y. MIYAKE, 1993’. 2 ♂ ‘〔Thailand〕/ Pupin / 21–IV–1985 / M. TAO // Paratype: Dasyvalgus / castaneodorsalis / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Doi Suthep / 10–V–1985 / M. TAO leg. // Paratype: Dasyvalgus / castaneodorsalis / Y. MIYAKE, 1993’.

Type condition

The aedeagus of the holotype is pinned separately.

Current status

Valid species.

Dasyvalgus decamaculatus Miyake

Figure 5E

Dasyvalgus decamaculatus Miyake, 1993a: 22−23.

Note

The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘Iluon Ma Thuot / Annam M. ITO // Holotype: / Dasyvalgus / decamaculatus / Y. MIYAKE, 1993’. (Fig. 5E)

Paratypes

2 exs.: 1 ex. ‘Near Chang Mai / N. Thailand / 7–V–1994 / K. KUME leg. // Paraype: Dasyvalgus / decamaculatus / Y. MIYAKE, 1993’. 1 ex. ‘Paratype: Dasyvalgus / decamaculatus / Y. MIYAKE, 1993’.

Type condition

The right and left metatarsus are missing.

Current status

Valid species.

Remark

The locality on the collecting data label of the holotype is ‘Iluon Ma Thuot’, but the locality given in the original description is ‘Ban Me Thuot’. However, the habitus of the holotype and the figure in the original description match. Probably, Miyake caused a typographical error when writing the description.

Dasyvalgus flavicauda Miyake, Yamaguchi et Aoki

Figure 5F

Dasyvalgus flavicauda Miyake, Yamaguchi, Aoki et Akiyama, 2002: 65−66.

Note

The holotype and following paratype are deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘Ban Lao Kram / Mukdahan / E-THAILAND / 17. Feb. 1999 / S. Yamaguchi and / T. Aoki leg. // Holotype: / Dasyvalgus / flavicauda // Holotype / Dasyvalgus flavicauda / Y. MIYAKE, YAMAGUCHI / et AOKI 2002’. (Fig. 5F)

Paratype

1 ex.: 1 ♂ ‘Phu Huai Sing / Mukdahan / E–THAILAND / 19. Feb. 1999 / S. Yamaguchi and / T. Aoki leg. // Paratype / Dasyvalgus flavicauda / Y. MIYAKE, YAMAGUCHI / et AOKI 2002’.

Current status

Valid species.

Dasyvalgus macacus Miyake

Figure 5G

Dasyvalgus macacus Miyake, 1993a: 24.

Note

The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘〔Thailand〕Pupin–Doi Phui / 19–IV–1985 / M. TAO leg. // Holotype: Dasyvalgus / macacus / Y. MIYSKE, 1993’. (Fig. 5G)

Paratypes

4 exs.: 1 ♂ ‘〔Thailand〕/ Pupin-Doi Phui / 1–V–1985 / M. TAO leg. // Paratype: Dasyvalgus / macacus / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Pupin-Doi Phui / 5–V–1985 / M. TAO leg. // Paratype: Dasyvalgus / macacus / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Pupin-Doi Phui / 12–V–1985 / M. TAO leg. // Paratype: Dasyvalgus / macacus / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Pupin-Doi Phui / 21–V–1985 / M. TAO leg. // Paratype: Dasyvalgus / macacus / Y. MIYAKE, 1993’.

Current status

Valid species.

Remark

The habitus photograph in the original description does not refer to the holotype.

Dasyvalgus makiharai Miyake

Dasyvalgus makiharai Miyake, 1985: 11−12.

Note

The following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Paratype

1 ex.: ‘MEIFENG / FORMOSA / 3–VII. 74 / T, OCHI // Paratype: / Dasyvalgus / makiharai Y. MIYAKE / (1985)’.

Current status

Valid species.

Remark

In addition to the paratype mentioned above, the following specimens labeled as paratype are not designated in the original description: 1 ex. ‘松崗= songgang / 27~29. VII 1968 / K. YAMAMOTO // Paratype: Dasyvalgus / makiharai / Y. MIYAKE, 1985’. 1 ♂ ‘SUNG KANG / FORMOSA / 30–VI, 74 / T. OCHI // Paratype: / Dasyvalgus / makiharai Y. MIYAKE / (1985)’.

Dasyvalgus minahasanus Miyake

Dasyvalgus minahasanus Miyake, 1989b: 42−43.

Note

The following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Paratypes

78 exs.: 75 exs. ‘Tondano / N. Sulawesi / 10–IV–1989 / Y. Miyake leg. // Paratype: / Dasyvalgus / minahasanus / Y. MIYAKE, 1989’. 3 exs. ‘Tondano / N. Sulawesi / 9–IV–1989 / Y. Miyake leg. // Paratype: / Dasyvalgus / minahasanus / Y. MIYAKE, 1989’.

Current status

Valid species.

Remark

In addition to the paratypes mentioned above, the following specimen labeled as paratypes is not designated in the original description: 1 ex. ‘Tondano / N. Sulawesi / 11–IV–1989 / Y. Miyake leg. // Paratype: / Dasyvalgus / minahasanus / Y. MIYAKE, 1989’.

Dasyvalgus multicus Miyake

Figure 5H

Dasyvalgus multicus Miyake, 1993a: 25−26.

Note

The holotype is deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘ [Thailand] / プピン-プイ= pupin-pui / 5–V–1985 / M. Tao leg. // Holotype: Dasyvalgus / multicus / Y. MIYAKE, 1993’. (Fig. 5H)

Type condition

The aedeagus of the holotype is pinned separately. The right and left protarsus are missing.

Current status

Valid species.

Dasyvalgus nudis Miyake

Figure 6A

Dasyvalgus nudis Miyake, 1994: 153.

Note

The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘Cao Bang / N. Vietnam / 9–11. VI, 1993 / M. ITO leg // Holotype: Dasyvalgus / nudis / Y. MIYAKE, 1994’. (Fig. 6A)

Figure 6. 

Habitus of holotype specimens. A Dasyvalgus nudis Miyake B Dasyvalgus rubrothoracicus Miyake C Hybovalgus matsudai Miyake D Neovalgus formosanus Miyake E Oreoderus quadrimaculatus Miyake F Tarsovalgus hatai Miyake.

Paratypes

9 exs.: 3 ♂ ‘Cao Bang / N. Vietnam / 8–10 VI, 1993 / M. ITO leg. // Paratype: / Dasyvalgus / nudis / Y. MIYAKE, 1994’. 3 ♂ ‘Cao Bang / N. Vietnam / 10. VI, 1993 / M. ITO leg. // Paratype: / Dasyvalgus / nudis / Y. MIYAKE, 1994’. [[2 ♂ ‘Cao Bang / N. Vietnam / 9–11. VI, 1993 (real date is 8–10. VI. 1993) / M. ITO leg. // Paratype: / Dasyvalgus / nudis / Y. MIYAKE, 1994’]]. [[1 ex. ‘Cao Bang / N. Vietnam / 11. VI, 1993 (real date is 8–10. VI. 1993) / M. ITO leg. // Paratype: / Dasyvalgus / nudis / Y. MIYAKE, 1994’]].

Type condition

The aedeagus of the holotype is pinned separately. The right antenna is missing.

Current status

Valid species.

Remark

The date on the collecting data label of the holotype is 9–11,VI,1993, but the collecting date quoted in the original description is 8–10,VI,1993. However, the habitus of the holotype and the figure in the original description match. Probably, Miyake caused a typographical error when writing the description.

Dasyvalgus rubrothoracicus Miyake

Figure 6B

Dasyvalgus rubrothoracicus Miyake, 1993a: 23−24.

Note

The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘〔Thailand〕/ Pupin–Doi Phui / 21–IV–1985 / M. TAO leg. // Holotype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’. (Fig. 6B)

Paratypes

11 exs.: 1 ex. ‘Doi Sutehp / Thailand / 28–IV–1985 / M Tao leg. // TDvA // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 1–V–1985 / M. TAO leg. // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 1–V–1985 / M. TAO leg. // TDvA // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 5–V–1985 / M. TAO leg. // TDvA // TDvA // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 11–V–1985 / M. TAO leg. // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’. [[1 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 17–V–1985 (the date given in the original description is 17–IV–1985) / M. TAO leg. // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’]]. [[1 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 21–V–1985 (the date given in the original description is 21–IV–1985) / M. TAO leg. // TDvA // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’]]. [[1 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 21–V–1985 (the date given in the original description is 21–IV–1985) / M. TAO leg. // TDvA // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’]]. [[1 ♂‘〔Thailand〕/ Pupin-Doi-Phui / 23–V–1985 (the date given in the original description is 23–IV–1985) / M. TAO leg. // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’]]. [[2 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 28–V–1985 (the date given in the original description is 28–IV–1985) / M. TAO leg. // TDvA // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’]].

Type condition

The aedeagus of the holotype is pinned separately.

Current status

Valid species.

Remark

The date on the collecting data label of the holotype is 21–IV–1985, but the collecting date quoted in the original description is 5–V–1985. However, the habitus of the holotype and the figure in the original description match. Probably, Miyake caused a typographical error when writing the description.

Dasyvalgus similis Miyake

Dasyvalgus similis Miyake, 1985: 8−9.

Note

The following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Paratypes

3 exs.: 1 ♂ ‘蓬来= hôrai / 20, 21 VIII 1968 / H. MAKIHARA // Paratype’. 1 ♂ ‘FUNCHIIFO / FORMOSA / 1. VI. 1970 / Y. KIYOYAMA // Paratype // Paratype / Dasyvalgus / similis, Y. MIYAKE / 1985’. 1 ♂ ‘FUNCHIIFO / FORMOSA / 1. VI. 1970 / Y. KIYOYAMA // Paratype’.

Current status

Senior objective synonym of Dasyvalgus taiwanus Miyake, 1991. Miyake in Miyake et al. (1991) proposed Dasyvalgus taiwanus as a replacement name for Dasyvalgus similis Miyake, 1985, a junior primary homonym of Dasyvalgus similis Moser, 1908.

Remark

In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 1 ex. ‘LIUKUEI / FORMOSA / 3. X. 1970 / Y. KIYOYAMA // 16 // Paratype // Paratype / Dasyvalgus / similis Y. MIYAKE, 1985’. 1 ex. ‘NANSHANCHI / FORMOSA / 27. VI. 1971 / Y. MAEDA // Paratype // Paratype / Dasyvalgus / similis Y. MIYAKE, 1985’. 1 ex. ‘紅水坑= hongshuikeng / Liukuei / 11. V. 1978 / Chen. W. // Paratype’. 1 ♂ ‘紅水坑= hongshuikeng / Liukuei / 11. V. 1978 / Chen. W. // Paratype’. 1 ex. ‘紅水坑= hongshuikeng / Liukuei / 11. V. 1978 / W. Chen // Paratype // Paratype / Dasyvalgus / similis Y. MIYAKE, 1985’. 1 ex. ‘Paratype’.

Dasyvalgus wadai Miyake

Dasyvalgus wadai Miyake, 1985: 9−10.

Note

The following paratype is deposited in RIEB (ex coll. Y. Miyake):

Paratype

1 ex.: 1 ♂ ‘Paratype / Dasyvalgus / wadai Y. MIYAKE, 1985’.

Current status

Valid species.

Remark

This paratype has no collecting data label.

Genus Hybovalgus

Hybovalgus matsudai Miyake

Figure 6C

Hybovalgus matsudai Miyake, 1985: 6.

Note

The holotype is deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘Paratype // Holotype / Hybovalgus / matsudai Y. MIYAKE / (1985)’.

Type condition

The right protarsus is missing.

Current status

Valid species.

Remark

There is no data label attached to the holotype, but the habitus of the holotype matches the figure in the original description. If confidence is placed in the pinned holotype label and the habitus, the above holotype specimen is the true holotype.

Genus Neovalgus

Neovalgus formosanus Miyake

Figure 6D

Neovalgus formosanus Miyake, 1985: 3.

Note

The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘六亀= liugui.石山= shi shan / 9. VI. 1976 / 2000m // Holotype: / Neovalgus / formosanus / Y, MIYAKE, 1985’. (Fig. 6D)

Paratypes

2 exs.: 1 ex. ‘ [Taiwan] / Baibara, mt. Puli / –V–1965 // Paratype: Neovalgus / formosanus Y. MIYAKE, 1985’. 1 ex. ‘ [Taiwan] / Liukuei / 16–V–1978 // PARATYPE / Neovalgus / formosanus / Y. MIYAKE, 1985’.

Current status

Valid species.

Remark

The date on the collecting label of the holotype is 9.VI.1976, but the collecting date quoted in the original description is 16.V.1978. However, the habitus of the holotype matches the figure in the original description. Probably, Miyake caused a typographical error when writing the description.

In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 3 exs. ‘ [Taiwan] / Baibara, mt. Puli / –V–1965 // PARATYPE / Neovalgus / formosanus / Y. MIYAKE, 1985’. 1 ex. ‘Formosa / Wushe / V. 1965 / T. Shiroz / Paratype: Neovalgus / formosanus / Y. MIYAKE, 1985’. 1 ex. ‘六 亀= liugui, 石山= shi shan / 2000 m / 9–VI–1976 / Paratype / Neovalgus / formosanus / Y. MIYAKE, 1985’. 1 ex. ‘Paratype: / Neovalgus / formosanus / Y. MIYAKE, 1985’.

Genus Oreoderus

Oreoderus quadrimaculatus Miyake, Yamaguchi et Aoki

Figure 6E

Oreoderus quadrimaculatus Miyake, Yamaguchi, Aoki et Akiyama, 2002: 64−65.

Note

The holotype is deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘Phu Huai Sing / Mukdahan / E-THAILAND / 19. Feb. 1999 / S. Yamahuchi and T. Aoki leg. // Holotype: / Oreoderus / quadrumaculatus // Or. 4ml / Coll. // Holotype / Oreoderus quadrimaculatus / Y. MIYAKE, YAMAGUCHI / et AOKI 2002’. (Fig. 6E)

Type condition

The aedeagus of the holotype is pinned separately. The right protarsus is missing.

Current status

Valid species.

Genus Tarsovalgus

Tarsovalgus hatai Miyake

Figure 6F

Tarsovalgus hatai Miyake, 1993b: 32−33.

Note

The holotype and following paratype are deposited in RIEB (ex coll. Y. Miyake):

Holotype

(♂). ‘ [Malaya] / – –1992 // Holotype: Tarsovalgus / hatai Y. MIYAKE, 1993’. (Fig. 6F)

Paratype

1 ex.: ‘ [Malaya] / – –1992 // Paratype: / Tarsovalgus / hatai / Y. MIYAKE, 1993’.

Current status

Valid species.

Family Trogidae

Genus Trox

Trox kyotensis Ochi & Kawahara

Trox kyotensis Ochi & Kawahara, 2000: 53−56.

Note

The following paratype is deposited in RIEB (ex coll. T. Ochi):

Paratype

1 ex.: ‘YAMAZAKI- / CHO, KYOTO. / 5–IV, 1999 / M. KAWAHARA // PARATYPE / Trox / kyotensis / OCHI and KAWAHARA / 2000’.

Current status

Trox (Niditrox) kyotensis Ochi et Kawahara, 2000, see Kalz (2019).

Trox sabulosus fujiokai Ochi

Trox sabulosus fujiokai Ochi, 2000: 43−44.

Note

The following paratype is deposited in RIEB (ex coll. T. Ochi):

Paratype

1 ex.: ‘Yukyuzan-Park, Nagaoka / Niigata Pref., Japan / 28. V. 1997 / Y and M. Kawahara leg // PARATYPE / Trox / Sabulosus / Subsp. fujiokai / OCHI, 2000’.

Current status

Valid species.

Trox setifer horiguchii Ochi & Kawahara

Trox setifer horiguchii Ochi & Kawahara, 2002: 54−55.

Note

The following paratype is deposited in RIEB (ex coll. T. Ochi):

Paratype

1 ex.: 1♂ ‘MEBORO-DAM / KAMIAGATA. T. / IS. TSUSHIMA / NAGASAKI. J. P / 25–V. 2002 / Y and M. KAWAHARA // 対県= tsu-ken上県町= kamiagata-chô / 目保呂ダム= meborodamu / 25. V. 2002 / Y and M. Kawahara // ♂ // PARATYPE / Trox / setifer / subsp. horiguchii / OCHI and KAWAHARA / 2002’.

Current status

Trox horiguchii Ochi and Kawahara, 2002, see Pittino (2006).

Acknowledgments

We are grateful to Toshiaki Aoki (RIEB), Shûhei Yamagushi (RIEB), Reiichiro Katsuyama (RIEB), Shûhei Nomura (NMNS), and Kônosuke Hoshino (NMSM) for their help in the type material survey. Our thanks go to Hao Xu (Mianyang Normal University) for providing the information about Miyake’s type material. We express our sincere gratitude to Carsten Zorn (Gnoien) for reviewing the manuscript.

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