Catalogue |
Corresponding author: Naoki Kaneko ( naoki.1993062z@gmail.com ) Academic editor: Andrey Frolov
© 2020 Naoki Kaneko, Kaoru Wada.
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation:
Kaneko N, Wada K (2020) Catalogue of the type material of Scarabaeoidea (Coleoptera) deposited in the Research Institute of Evolutionary Biology, Tokyo, Japan. ZooKeys 958: 35-89. https://doi.org/10.3897/zookeys.958.52799
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A detailed catalogue of the type material of the Scarabaeoidea housed in the Research Institute of Evolutionary Biology, Tokyo, Japan is listed. The catalogue includes the data of the type material of four families and 111 species.
Allotype, Geotrupidae, holotype, Lucanidae, neotype, paratype, Scarabaeidae, Trogidae
In this report we present a catalogue of the type material of Scarabaeoidea housed in the Research Institute of Evolutionary Biology, Tokyo, Japan (RIEB). The beginning of the RIEB collection dates back to 1950, when the first natural objects were deposited at the Research Institute of Thremmatology of the Tokyo University of Agriculture (RIT) established that year. RIT was transformed into an incorporated foundation in 1974 and renamed Research Institute of Evolutionary Biology (
The majority of the types of Scarabaeidae deposited in the RIEB originate from the collection of Mr Yoshikazu Miyake (1926–2003), a well-known Japanese amateur specialist on Scarabaeidae, who was also the author of those taxa. A part of types of Miyake’s species is also housed in the National Museum of Nature and Science, Tokyo (NMNS) and the Nagaoka Municipal Science Museum, Niigata (NMSM). Unfortunately, some of his types are missing.
In addition, other researchers that contributed to the Scarabaeoidea type collection in RIEB are: Dirk Ahrens, Aleš Bezděk, Toshiyuki Ichikawa, Takeshi Itoh (Takeshi Matsumoto), Kimio Masumoto, Sizumu Nomura, Teruo Ochi, and Kaoru Wada.
The scientific names of higher taxa in this catalogue follow
– original combination of the taxon name;
– original combination and spelling of the taxon name, followed by the author, year of description, and pagination;
– type material, number of specimens (including sex, if known), and exact label data. A double slash ‘//’ indicates separate labels and single slash ‘/’ indicates lines within each label. The words in Japanese or Chinese were transcribed into Roman alphabet. Paratypes with discrepancies between collection data on the label and data quoted in the original description are indicated by ‘ [[]]’;
– remarks on types condition (given only for holotypes and neotypes);
– current taxonomic status;
– remarks, if any.
The majority of holotypes and neotypes were photographed with a Nikon D7200 camera using a Nikon AF-S DX Micro NIKKOR 40 mm f/2.8 G lens, and some types were photographed with a KEYENCE VHX-1000 Digital Microscope.
Genus Sinogeotrupes
Sinogeotrupes taiwanus Miyake & Yamaya, 1995: 32−34.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
3 exs.: 1 ex. ‘Wusha / –VI–1941 / R. Boloudo // Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’. 1 ex. ‘Wu sha / Formosa / 21. Viii. 1941 / Col. Bumua Tin // Geotrupes / subshiatelluo FAIRMAIR / DET. Y. MIYAKE // Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’. 1 ex. ‘台湾= taiwan 台中= taichû / 関刀山= guandaoshan / 1993. V. 31 // Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’.
Phelotrupes (Sinogeotrupes) taiwanus (Miyake and Yamaya, 1995), see
In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 3 exs. ‘Tattaka / Formosa / 10. Vi. 1965 / T. Shirozu // Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’. 1 ex. ‘(FORMOSA) / Mt Rata / Vi–Vii–1972 / L. F, Hang // Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’. 1 ex. ‘(FORMOSA) / Mt Rata / –Vii–1972 / L. F, Hang // Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’. 2 exs. ‘ [Taiwan] / 台中= taichû 関刀山= guandaoshan / 3–VI–1993 // Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’. 1 ♂ ‘ [Taiwan] / 台中= taichû 関刀山= guandaoshan / 3–VI–1993 // Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’. 3 exs. ‘Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’.
Genus Aegus
Aegus laevicollis nakanei Ichikawa & Imanishi, 1976: 9.
The following paratype is deposited in RIEB (ex coll. Y. Kusui):
1 ex.: [[1 ♀ ‘Mt. Yonahadake, / Kunigami-son, / Is. Okinawa-Hontō, / 28. VI. 1972 (the date given in the original description is 16–17. VI. 1970), / HEIKICHI IRIE leg., // KUSUI // Paratype’]].
Aegus nakanei nakanei Ichikawa and Imanishi, 1976, see
Genus Aphodius
Aphodius (Agoliinus) tanakai Masumoto, 1981: 73−74.
The following paratype is deposited in RIEB (ex coll. K. Masumoto):
1 ex.: ‘Nikko / Date: 23 V 1981 / K. MASUMOTO leg. // Paratype / Aphodius (Agr) / tanakai MASUMOTO’.
Agoliinus tanakai (Masumoto, 1981), see
Aphodius (Phaeaphodius) himalorectus Ahrens & Stebnicka, 1997: 12−13.
The following paratypes are deposited in RIEB (ex coll. D. Ahrens):
2 exs.: ‘Phalante 20. 3. / unt. Nebelwaldstufe / 2100–2300 m // PARATYPUS / Aphodius (Phaeaphodius) himalorectus sp. n. / det. D. AHRENS 1994’.
Phaeaphodius himalorectus (Ahrens and Stebnicka, 1997), see
Aphodius (Sinodiapterna) zeni Ochi, 1991: 53−55.
The following paratype is deposited in RIEB (ex coll. T. Ochi):
1 ex.: ‘SUNGKAN / C. TAIWAN / 29–V. 1987 / K. LAH // PARATYPE / Aphodius (Sinodiapterna) / zeni /
Sinodiapterna zeni (Ochi, 1991), see
Rhyparus kitanoi Miyake, 1982: 65−67.
The holotype and following specimen are deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘C, Sata /〔KYUSHU〕/ Japan / 25, VI 1971 / T. Kitano leg. // PARATYPE // Holotype: / Rhyparus / kitanoi / Y. MIYAKE, 1992’. (Fig.
Habitus of holotype specimens. A Rhyparus kitanoi (Miyake) B Adoretus (Lepadoretus) ubonensis (Miyake, Yamaguchi et Aoki) C Anomala flavoguttata Miyake D Anomala ohmomoi Miyake, Yamaguchi et Aoki E Anomala thailandiana Miyake, Yamaguchi et Akiyama F Callistethus isidai Miyake G Malaia castanoptera Miyake H Malaia macassara Miyake.
The right and left protarsus and the right antenna of the holotype are missing.
Rhyparus kitanoi kitanoi Miyake, 1982, see
In addition to the holotype mentioned above, the following specimen labeled as paratype is not designated in the original description: 1 ex.: ‘Sesonoura / Is. Shimokoshiki / Kagoshima-pref. / 18. VI. 1982 / S. Imasaka leg. // Paratype: Rhyparus / kitanoi / Y. MIYAKE, 1992’.
Genus Caccobius
Caccobius bawangensis Ochi, Kon & Kikuta, 1997: 262−264.
The following paratype is deposited in RIEB (ex coll. T. Ochi):
1 ex.: ‘Mt. Bawang / Sarawak S. / VIII. 1991 / M. Kon Coll. // PARATYPE / Caccobius / bawangensis OCHI, KON and KIKUTA’.
Valid species.
Copris (Microcopris) mindorensis Ochi, 1992: 13.
The following paratype is deposited in RIEB (ex coll. T. Ochi):
1 ex.: ‘MT. HALKON / IS. MINDRO / 28–XII. 1988 / D. MOHAGAN // PARATYPE / Copris / (Microcopris) / mindorensis / OCHI, 1992.
Valid species.
Onthophagus setchan Masumoto, 1984: 80−81.
The following paratype is deposited in RIEB (ex coll. K. Masumoto):
1 ex.: ‘FENGCHIFU / FORMOSA / 7–VII. 74. / T. OCHI // PARATYPE / Onthophagus / setchan /
Valid species.
Onthophagus (Parascatonomus) acuticollis sakishimanus Nomura, 1976a: 29.
The following paratypes are deposited in RIEB (ex coll. S. Nomura):
6 exs.: 3 ♂ ‘Kanpira / Iriomote Is. / Yaeyama Iss. / 12. VII. 1975 / R. YANO leg. // ♂ // Onthophagus (P.) / acuticollis / sakishimanus / NOMURA (1976)’. 3 ♀ ‘Kanpira / Iriomote Is. / Yaeyama Iss. / 12. VII. 1975 / R. YANO leg. // ♀ // Onthophagus (P.) / acuticollis / sakishimanus / NOMURA (1976)’.
Parascatonomus (Necramator) acuticollis sakishimanus (Nomura, 1976), see
Onthophagus (Parascatonomus) murasakianus Nomura, 1976a: 30.
The following paratype is deposited in RIEB (ex coll. S. Nomura):
1 ex.: ‘Kanpira, / Iriomote Is. / Yaeyama Iss. / 12. VII 1975 / R. YANO leg. // PARATYPE / Onthophagus (P.) / murasakianus / NOMURA (1976)’.
Parascatonomus (Necramator) murasakianus (Nomura, 1976), see
Onthophagus (Parascatonomus) takedai Ochi & Araya, 1992: 99−100.
The following paratype is deposited in RIEB (ex coll. T. Ochi):
1 ex.: ‘QUESON. N. P / IS. LUZON / 10–VIII. 1981 / K. SUGINO // PARATYPE / Onthophagus / (Parascatonomus) / takedai / OCHI et ARAYA, 1992’.
Parascatonomus (Granulidorsum) takedai (Ochi et Araya, 1992), see
Onthophagus (Proagoderus) watanabei Ochi & Kon, 2002: 306−311.
The following paratype is deposited in RIEB (ex coll. T. Ochi):
1 ex.: ‘SEPIROK / N. BORNEO / 4–VII. 1985 / M. KON // PARATYPE / Onthophagus / (Proagoderus) / watanabei /
Proagoderus watanabei (Ochi et Kon, 2002), see
Onthophagus (Strandius) hiurai Ochi, 1984: 63−65.
The following paratype is deposited in RIEB (ex coll. T. Ochi):
1 ex.: ‘PILU / Taiwan / 21, VIII, 1979 / K. Sugino leg. // PARATYPE / Onthophagus (St) / hiurai /
Onthophagus (Phanaeomorphus) potanini hiurai Ochi, 1984, see
Genus Adoretus
Adoretus (Lepadoretus) ubonensis Miyake, Yamaguchi, Aoki & Akiyama, 2002: 62−63.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘Shirinthorn (light) / Ubon Ratchatani / E–THAILAND / 27. Feb. 1999 / S. Yamaguchi and / T. Aoki leg. // No. 1 / ♂ // Holotype: / Adoretus / ubonensis // Holotype / Adoretus (Lepadoretus) ubonensis / Y. MIYAKE, YAMAGUCHI / et AOKI 2002’. (Fig.
The left mesotarsus and the left metaleg of the holotype are missing.
Junior subjective synonym of Adoretus compressus (Weber, 1801), see
Anomala flavoguttata Miyake, 2000: 108−109.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘Sapa=Chapa / N. Vietnam / 20. V. 1994 / N. KATSURA // Holotype: Anomala / flavoguttata / Y. MIYAKE, 2000’. (Fig.
The aedeagus of the holotype is pinned separately.
Valid species.
Anomala ohmomoi Miyake, Yamaguchi, Aoki & Akiyama, 2002: 58−60.
The holotype and the following paratypes are deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘Ban Lao Kram / Mukdahan (light) / E–THAILAND沢= sawa / 8. Mar. 1999 / S. Yamaguchi and / T. Aoki leg. // Holo Type: / Anomala / ohmomoi // Holotype / Anomala ohmomoi / Y. MIYAKE, YAMAGUCHI / et AOKI 2002’. (Fig.
4 exs.: ‘Ban Lao Kram / Mukdahan (light) / E–THAILAND沢= sawa / 8. Mar. 1999 / S. Yamaguchi and / T. Aoki leg. // Paratype / Anomala ohmomoi / Y. MIYAKE, YAMAGUCHI / et AOKI 2002’.
Valid species.
Anomala thailandiana Miyake, Yamaguchi, Aoki & Akiyama, 2002: 60−61.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘Don Far Huan (light) / Ubon Ratchathani / E–THAILAND / 17. Sep. 1999 / S. Yamaguchi and / M. Akiyama leg. [white label, front] // Apigoni / 5 [white label, back] // Holotype / Anomala thailandiana / Y. MIYAKE, YAMAGUCHI / et AKIYAMA 2002’. (Fig.
The aedeagus of the holotype is pinned separately.
Valid species.
Callistethus isidai Miyake, 1987: 5−6.
The holotype and the following paratype are deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘ [Taiwan] / Lishan to Pilu / 31–VII–1972 / Y. Miyake leg. // Holotype: / Callistethus isidai / Y. MYAKE, 1987’. (Fig.
1 ex.: ‘ [Taiwan] / Lishan to Pilu / 31–VII–1972 / Y. Miyake leg. // Paratype / Callistethus isidia / Y. MIYAKE, 1987’.
The right and left metatarsi of the holotype are missing.
Callistethus plagiicollis isidai Miyake, 1987, see
The habitus photograph in the original description does not agree with the holotype. This is also the case with some other species described by Miyake. Apparently, he did not intend to provide the photographs of the holotypes. Also, as noted below, we were unable to trace some specimens illustrated in the original descriptions.
Kibakoganea ohtanii Miyake & Muramoto, 1992: 22−24.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
2 exs.: [[1 ♂, 1 ♀ ‘Tam Dao / N. VIETNAM / 28–30, IV, 1991 (the date given in the original description is 28–30. III. 1991) / M. Fujioka and / R. Muramoto – leg. // PARATYPE / Kibakoganea / ohtanii / Y. MIYAKE et / MURAMOTO, 1992.’]].
Junior subjective synonym of Kibakoganea sexmaculata (Kraatz, 1900), see Miyake (2003).
Malaia castanoptera Miyake, 2000: 112.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘Malino, Celebes / Indonesia / 21 Nov 1944 // malino / 2603.11.21 // Holotype: Malaia / castanoptera / Y. MIYAKE, 2000’. (Fig.
7 exs.: 1 ♂, 3 ♀ ‘Malino, Celebes / Indonesia / 21 Nov 1944 // malino / 2603.11.21 // Paratype: Malaia / castanoptera / Y. MIYAKE, 2000’. 2 exs. ‘Malino, Celebes / Indonesia / 21 Nov 1944 // malino / 2603.11.21 // Paratype: Malaia / castanoptera / Y. MIYAKE, 2000’. 1 ex. ‘malino / 2603.11.21 // Paratype: Malaia / castanoptera / Y. MIYAKE, 2000’.
Valid species.
The habitus photo in the original description belongs to the female paratype with the label ‘Malino, Celebes / Indonesia / 21 Nov 1944 // malino / 2603.11.21’.
Malaia cyanea Miyake, 1996: 38−39.
The following paratypes are deposited is RIEB (ex coll. Y. Miyake):
2 exs.: 1 ♂ ‘32 km from / Palopo / Surawesi / 10. VI. 1982 // Paratype: / Malaia cyanea / Y. MIYAKE, 1996’. 1 ♀ ‘Punchak–Lombanan / Mamasa, / S. SULAWESI / 1991. AUG. 10. / N. Kashiwai–leg. // Paratype: / Malaia cyanea / Y. MIYAKE, 1996’.
Valid species.
Malaia macassara Miyake, 2000: 109−110.
The holotype and the following paratype are deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘Lampbatter / Sulawesi / 23–25. VIII. 1999 / M. Takai // Holotype: Malaia / macassara / Y. MIYAKE, 2000’. (Fig.
1ex.: 1 ♂ ‘Lampbatter / Sulawesi / 23–25. XII. 1999 / M. Takai leg. // Paratype: Malaia / macassara / Y. MIYAKE, 2000’.
The left protarsus and metatarsus of the holotype are missing.
Valid species.
Malaia rufofemorata Miyake, 1996: 40−41.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
4 exs.: 1 ♂, 1 ♀ ‘Puncak, Palopo / VI–VII, 1989 / Central SULAWESI // Paratype: Malaia / rufofemorata / Y. MIYAKE, 1996’. 2 ♂ ‘Palu Palolo / VIII. 1990 / Central SULAWESI // Paratype: Malaia / rufofemorata / Y. MIYAKE, 1996’.
Valid species.
In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 2 ♂, 1 ♀ ‘Palolo, Palu / 01–10/XI. 1990 / native coll. / CELEBES // Paratype: Malaia / rufofemorata / Y. MIYAKE, 1996’.
Malaia sulawesi Miyake, 1996: 41−42.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
58 exs.: 1 ♂ ‘Mt. Pedamaran / Tana Toraja / 1985. III–17 / South SULAWESI / Soma, K. leg. // Paratype: / Malaia / sulawesi / Y. MIYAKE, 1996’. 1 ex. ‘Puncak Palopo / 1989. III. 31 / Centr. CELEBES // Paratype: / Malaia / sulawesi / Y. MIYAKE, 1996’. 3 exs. ‘Palu Pulu / C. Sulawesi / II. 1989 // Paratype: / Malaia / sulawesi / Y. MIYAKE, 1996’. [[44 exs. ‘Mt. Pedamaran / S. Sulawesi (real data is C. Sulawesi) / 20. X. 1983 (real date is 20. V. 1983) / K. Soma leg. // Paratype: / Malaia / sulawesi / Y. MIYAKE, 1996’]]. [[9 exs. ‘Mt. Pedamaran / Tana Toraja / 1983. X–20 (real date is 1983. V–20) / South SULAWESI (real data is Central SULAWESI) / Soma, K. leg. // Paratype: / Malaia / sulawesi / Y. MIYAKE, 1996’]].
Valid species.
Malaia taoi Miyake, 1996: 39−40.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
4 exs.: 1 ♂ ‘37 km from / Palopo C.- / Sulawesi / 9. IV. 1982 // Paratype: / Malaia / taoi / Y. MIYAKE, 1996’. 1 ♀ ‘32 km from / Palopo / Surawesi / 10. VI. 1982 // 37 km from / Palopo C.- / Sulawesi / 10. VI. 1982 // Paratype: / Malaia / taoi / Y. MIYAKE, 1996’. 1 ♂ ‘37 km from / Palopo C.- / Sulawesi / 11. IV. 1982 // Paratype: / Malaia / taoi / Y. MIYAKE, 1996’. 1 ♀ ‘32 km from / Palopo / Surawesi / 13. VI. 1982 // 37 km from / Palopo C.- / Sulawesi / 13. VI. 1982 // Paratype: / Malaia / taoi / Y. MIYAKE, 1996’.
Valid species.
Malaia tondanoensis Miyake, 1996: 42−43.
The following paratype is deposited in RIEB (ex coll. Y. Miyake):
4exs.: ‘Tondano / N. Sulawesi / 10–IV–1989 / Y. Miyake leg. // Paratype: / Malaia / tondanoensis / Y. MIYAKE, 1996’.
[[1 ex. ‘Tangkoko N. / N. Sulawesi / 10–III–1989 (the date given in the original description is 10–IV–1989) / Y. Miyake leg. // Paratype: / Malaia / tondanoensis / Y. MIYAKE, 1996’]]. [[2 exs. ‘Tondano / N. Sulawesi / 9–IV–1989 (the date given in the original description is 10–IV–1989) / Y. Miyake leg. // Paratype: / Malaia / tondanoensis / Y. MIYAKE, 1996’]].
Valid species.
Malaia toraja Miyake, 2000: 110−111.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘Mt. Pedamaran / Tana Traja / 20–X. 1983 / K. SOHMA lgd. / South SULAWESI // Holotype: Malaia / toraja / Y. MIYAKE, 2000’. (Fig.
27 exs.: 1 ♂ ‘Mt. Pedamaran / Tana Traja / 20–X. 1983 / K. SOHMA lgd. // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 9 exs. ‘Mt. Pedamaran / Tana Traja / 20–X. 1983 / K. SOHMA lgd. // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 2 exs. ‘Mt. Pedamaran / S. Sulawesi / 20. X. 1983 / K. Soma leg. // Paratype: Malaia / toraja / Y. MIYAKE, 2000’.8 exs. ‘Puncak, Palopo / VI–VII, 1989 / Central SULAWESI // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 2 exs. ‘Central SULAWESI / Puncak, Palopo / VI–VII, 1989 // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 1 ♂ ‘Puncak Palopo / 1989. / Centr. CELEBES // SI [yellow label] // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 4 exs. ‘Puncak Palopo / 1989. / Centr. CELEBES // SI [yellow label] // Paratype: Malaia / toraja / Y. MIYAKE, 2000’.
Valid species.
The habitus photo in the original description belongs to the female paratype with the label ‘Mt. Pedamaran / Tana Toraja / S. Sulawesi / 7, VIII, 1988 / K. Soma leg.’.
In addition to the paratypes listed above, the following specimens labeled as the paratypes are not designated in the original description: 1 ♀ ‘Mt. Pedamaran / Tana Toraja / S. Sulawesi / 7, VIII, 1988 / K. Soma leg. // SI [yellow label] // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 1 ex. ‘(Pulu Pulu) / C. Sulawesi / INDONESIA / Dec. 1988 // [yellow label] // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 2 exs. ‘Puncak, Palopo / Centr. CELEBES / 1989. I / INDONESIA // [yellow label] // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 1 ex. ‘Puncak Palopo / 1989. III. 31 / Centr. CELEBES // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 3 exs. ‘Palu, Palolo / VII 1989 / Central SULAWESI // Paratype: Malaia / toraja / Y. MIYAKE, 2000’.
Mimela kitanoi Miyake, 1987: 7−8.
The holotype is deposited in RIEB (ex coll. K. Wada):
(♀). ‘Larashan / Hsinchu Hs. / VI. 1975, / Fang leg // Holotype: / Mimela kitanoi / Y. MIYAKE, 1987 // Kaoru WADA / Collection 2001’. (Fig.
Junior subjective synonym of Mimela flavocincta Lin, 1966, see
Mimela marginipennis Miyake, 1994: 151.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘[Borneo] / Crocker Range / 1800−1400m alt. / nr. Keningau C. / X. 1988. M. ITOI leg. // Holotype: / Mimela / marginipennis / Y. MIYAKE, 1994’. (Fig.
The aedeagus of the holotype is pinned separately, and the left mesotarsus is missing.
Junior subjective synonym of Mimela margarita Arrow, 1910, see
Mimela sawaii Miyake, 1994: 149−150.
The following paratype is deposited in RIEB (ex coll. Y. Miyake):
1 ex.: 1 ♂ ‘Crocker Range / (1000–1400 m) / Nr. Keningau C. / N. Borneo, – / 15–V–1989 // [blue label] // Paratype: / Mimela / sawaii / Y. MIYAKE, 1994’.
Junior subjective synonym of Mimela pallidicauda Arrow, 1910, see
Mimela suspecta Miyake, 1994: 150.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
2ex.: 1 ♂ ‘Mt. Trus Madi / (SW Slopa 1200m alt.) / Sabah, MALAYSIA / 14–IV–1993 / Minoru SAWAI leg. // Paratype: / Mimela / suspecta / Y. MIYAKE, 1994’. [[1 ex. ‘Mt. Trus Madi / (SW Slopa 1200m alt.) / Sabah, MALAYSIA / 18–IV–1993 (the date given in the original description is 4–IV–1993) / Minoru SAWAI leg. // [yellow label] // Paratype: / Mimela / suspecta / Y. MIYAKE, 1994’]].
Valid species.
In addition to the paratypes mentioned above, the following specimen labeled as paratype is not designated in the original description: 1 ♂ ‘Trus Madi Mts. / near Keningau / Saba MALAYSIA / 1993. iv. 17–30. // F //Paratype: / Mimela / suspecta / Y. MIYAKE, 1994’.
Popillia iwasei Miyake, 1996: 43−44.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
6 exs.: 2 ♀ ‘Puncak, Palopo / Centr. CELEBES / 1989. I / INDONESIA // Paratype: / Popillia / iwasei / Y. MIYAKE, 1996’. 2 ♀ ‘Puncak Palopo / C. Sulawesi / 1. 1989 // Paratype: / Popillia / iwasei / Y. MIYAKE, 1996’. 1 ♀ ‘Puncak Palopo / C. Sulawesi / 1989. V / INDONESIA // SI [yellow label] // Paratype: / Popillia / iwasei / Y. MIYAKE, 1996’. [[1 ♂ ‘Palolo (the locality given in the original description is Palopo) / II. 1991 (the date given in the original description and
Valid species.
Spilopopillia rugosa Miyake, 1989a: 180−181.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
18 exs.: [[1 ex. ‘ [Malaya] / Fraser’s Hill / 29–III–1977 (the date given in the original description is 28. III. 1977) / Y. Miyake leg. // Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’]]. [[1 ex. ‘ [Malaya] / Freaser’s Hill (the locality given in the original description is Fraser’s Hill) / 30–III–1977 (the date given in the original description is 28. III. 1977) / Y. Miyake leg. // Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’]]. 6 exs. ‘ [Malaya] / Tanah Rata / 30–III–1974 / Y, Miyake leg // Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’. 9 exs. ‘TANARATA / MALAYA / 30 III 1974 / Y. Miyake leg. // Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’. [[1 ex. ‘Gentig / Hiland (the locality given in the original description is Genting Highland) MALAYA / 5. IV. 1974 (the date given in the original description is 5. IV. 1975) / Y Miyake leg. // Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’]].
Valid species.
In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 8 exs. ‘Tanah Rata / MALAYA / 1. III. 1977 / Y. Miyake // Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’. 5 exs. ‘TANARATA / MALAYA / 28, 29, 30 III 1974 / Y. Miyake leg. // Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’. 1 ex. ‘84. IV // Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’. 2 exs. ‘Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’.
Spinanomala moritai Miyake, 1996: 38.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘Tanah Rata / MALAYA / IV. 1984 / M. Hata // Holotype: / Spinanomala / moritai / Y. MIYAKE, 1996’. (Fig.
11 exs.: 1 ex. ‘Tanah Rata / MALAYA / I–II. 1973 / MORITA, Kozo // Paratype: Spinanomala / moritai / Y. MIYAKE, 1996’. 2 ♂, 3 ♀ ‘Tanah Rata / MALAYA / 1. III. 1977 / Y. Miyake // Paratype: Spinanomala / moritai / Y. MIYAKE, 1996’. 3 ♂, 2 ♀ ‘Tanah Rata / MALAYA / IV. 1984 / M. Hata // Paratype: Spinanomala / moritai / Y. MIYAKE, 1996’.
The aedeagus of the holotype is pinned separately and the right metaleg is missing.
Valid species.
The collecting data of the specimen labelled as the holotype do not match the original description. There are two male paratypes (‘Tanah Rata / MALAYA / 1. III. 1977 / Y. Miyake // Paratype: Spinanomala / moritai / Y. MIYAKE, 1996’) that match the holotype data.
In addition to the paratypes mentioned above, the following specimen labeled as the paratype is not designated in the original description: 1 ex. ‘フレーザーヒル= furêzâhiru / MALAYA / 4 – 6. IV. 1977 / M. Hata // Paratype: Spinanomala / moritai / Y. MIYAKE, 1996’.
Genus Papuana
Papuana timorensis Miyake & Yamaya, 1999: 103−104.
The following paratype is deposited in RIEB (ex coll. Y. Miyake):
1 ex.: 1 ♂ ‘Timor Is. / Indonesia / V. 1993 / Native Coll. // Timor / Is. [aedeagus mount] // Paratype: / Papuana / timorensis YAMAYA / Y. MIYAKE, 1999’.
Valid species.
Peltonotus morio Burmeister, 1847: 75.
The neotype is deposited in RIEB (ex coll. K. Wada):
(♂). ‘NEPAL・HIMALAYA / 10. 5. 1993 Chhachok Madi– / Khola/ Gandaki (Kaski– / Destr.) 1300m lg: D. Ahrens //Peltonotus / morio Burn. / det: D. Ahrens 1997 // Neotype / Peltonotus / morio Burmeister, 1847 / Jameson and Wada, 2004’. (Fig.
The aedeagus of the neotype is pinned separately.
Valid species.
The neotype was designated by
Peltonotus morio sawaii Miyake, 2000: 112−113.
The holotype and the following paratypes are deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘Doi Suthep / Chieng Mai / N–THAILAND/ 22. V. 1983 / leg. M. Sawai // 12 // P. morio / Burmi. [white label, front] // Cameron / Highland / Pahang [white label, back] // Holotype: Peltonotus / morio sawaii / Y. MIYAKE, 2000’. (Fig.
4 exs.: 2 ♂, 1 ♀ ‘Doi Suthep / Chieng Mai / N–THAILAND / 22. V. 1983 / leg. M. Sawai // Paratype: Peltonotus / morio sawaii / Y. MIYAKE, 2000 // P. morio /Burmeister’. 1 ♀ ‘Doi Suthep / Chieng Mai / N–THAILAND / 22. V. 1983 / leg. M. Sawai // Paratype: Peltonotus / morio sawaii / Y. MIYAKE, 2000’.
The aedeagus of the holotype is pinned separately.
Junior subjective synonym of Peltonotus morio Burmeister, 1847, see
Peltonotus podocrassus Jameson & Wada, 2004: 34−37.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
7 exs.: 2 exs. ‘ [Malaya] / Tanah Rata / 30–III–1976 / Y. Miyake leg. // Paratype: Peltonotus peninsularis / Y. MIYAKE, 2000 // Paratype / Peltonotus podocrassus / Jameson and Wada, 2004’. 1 ♂ ‘39 miles from / Tapah, Malaya / 30–III–1976 / Y. Miyake leg. // Paratype: Peltonotus peninsularis / Y. MIYAKE, 2000 // Paratype / Peltonotus podocrassus / Jameson and Wada, 2004’. 1 ♂ ‘TANARATA / MALAYA / 30. III. 1976 / Y. Miyake leg. // Paratype: Peltonotus peninsularis / Y. MIYAKE, 2000 // Paratype / Peltonotus podocrassus / Jameson and Wada, 2004’. 1 ♂ ‘V. R〔C. H.〕/ Malaya / 21. III. 1978 // 46 // Paratype: Peltonotus peninsularis / Y. MIYAKE, 2000 // Paratype / Peltonotus podocrassus / Jameson and Wada, 2004’. 1 ex. [Malaya] / Tanah Rata / 1–IV–1984 / Y. Miyake leg. // Paratype: Peltonotus peninsularis / Y. MIYAKE, 2000 // Paratype / Peltonotus podocrassus / Jameson and Wada, 2004’. 1 ex. ‘TANARATA / MALAYA / 1. IV. 1984 / Y. Miyake // Paratype: Peltonotus peninsularis / Y. MIYAKE, 2000 // Paratype / Peltonotus podocrassus / Jameson and Wada, 2004’.
Valid species.
Peltonotus sakaii Miyake & Yamaya, 1994: 39−40, 42.
The following paratype is deposited in RIEB (ex coll. Y. Miyake):
1 ex.: ‘Mt. Serapi / SARAWAKU / 1991/III. // 20 // Paratype: / Peltonotus sakaii / Y. MIYAKE et YAMAYA, 1994 // Peltonotus / similis Arrow, 1931 / det. Kaoru Wada, 2004’.
Junior subjective synonym of Peltonotus similis Arrow, 1931, see
In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 1 ♂ ‘Mt. Serapi / Kuching, Sarawak / IX. 1990 / BORNEO // Paratype: / Peltonotus sakaii / Y. MIYAKE et YAMAYA, 1994 // Peltonotus / similis Arrow, 1931 / det. Kaoru Wada, 2004’. 1 ex. ‘Lembah Anai / W. Sumatra / VI. 1990 / INDONESIA // [yellow label] // Paratype: / Peltonotus sakaii / Y. MIYAKE et YAMAYA, 1994 // Peltonotus / similis Arrow, 1931 / det. Kaoru Wada, 2004’. 1 ♂ ‘Kimanis Road / Near Keningau / N. Borneo / 3–V–1994 // Paratype: / Peltonotus sakaii / Y. MIYAKE et YAMAYA, 1994 // Peltonotus / similis Arrow, 1931 / det. Kaoru Wada, 2004’. 2 exs. ‘Kimanis Road / Near Keningau / N. Borneo / 3–V–1994 // Paratype: / Peltonotus sakaii / Y. MIYAKE et YAMAYA, 1994 // Peltonotus / similis Arrow, 1931 / det. Kaoru Wada, 2004’.
Peltonotus sulawesiensis Wada, 1990: 1−2.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
4 exs.: ‘Tondano N. Celebes / INDONESIA / V 1988 / leg. N. Nisikawa // PARATYPE / Peltonotus sulawesiensis / Kaoru WADA, 1990’.
Junior subjective synonym of Neohyphus celebesus Heller, 1896, see
Genus Amiserica
Amiserica rufidula Nomura, 1974: 84.
The following paratypes are deposited in RIEB (ex coll. S. Nomura):
2 exs.: 1 ♂ ‘Li-shan / Taiwan / 29. VII. 1974 / Y. Miyake // ♂ // PARATYPE / Amiserica / rufidula /
Valid species.
Apogonia fujiokai Miyake & Yamaya, 1997: 6−7.
The following paratype is deposited in RIEB (ex coll. Y. Miyake):
1 ex.: ‘Flores Is. / Indonesia / May. 1995 // Y // Paratype: / Apogonia fujiokai / Y. MIYAKE et YAMAYA, 1997’.
Valid species.
Apogonia fusciventris Miyake & Yamaya, 1997: 5−6.
The following paratype is deposited in RIEB (ex coll. Y. Miyake):
1 ex.: ‘Vangvieng / P. Vietiane / LAO. P. D. R. / 25–26. MAY. 1994. / K. Miura–leg. // Y // Paratype: Apogonia / fuscoventris / Y. MIYAKE et YAMAYA, 1997’.
Junior subjective synonym of of Apogonia striatipennis Frey, 1971, see
The species name on the paratype label is ‘fuscoventris’, which is invalid. Probably, the name was confused with fusciventris by Y. Miyake.
Apogonia hongkongica Miyake, 1989b: 38−39.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
11 exs.: 1 ♂ ‘Hong Kong / China / C. K. Yu / 1974. V. [white label, front] // Native Collector / legt. [white label, back] // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’. 1 ♂ ‘Hong Kong / China / V. 1974 / C. K. Yu [white label, front] // Legt. / Native / Collector [white label, back] // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’. 1 ♂ ‘Hong Kong / China / V. 1974 / C. K. Yu [white label, front] // Native Collector [white label, back] // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’. 1 ♀ ‘♀ // Hong Kong / China / V. 1974 / C. K. Yu [white label, front] // Legt. / Native / Collector // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’. 1 ♀ ‘♀ // Hong Kong / China / V. 1974 / C. K. Yu [white label, front] // Native / Collector / legt. // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’. 1 ♀ ‘Hong Kong / 1974 / From C. K. Yu ♀ // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’. 1 ex. ‘Hong Kong / China / V. 1974 / C. K. Yu [white label, front] // Legt. / native / Collector [white label, back] // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’. 2 exs. ‘Hang Kong / China / V. 1974 / C. K. Yu Col [white label, front] // Native / Collector / legt. // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’. 2 exs. ‘Hong Kong / V. 1974 / From C. K. Yu // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’.
Valid species.
In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 1ex. ‘Hong Kong / 1975 / From. C. K-Yu // Paratype: Apogonia / hongkongica / Y. MIYAKE, 1989’. 1 ♀ ‘Hong Kong / ♀ 1975 [white label, front] // from / C. K. Yu [white label, back] // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’.
Apogonia inconstans Kobayashi & Bezděk, 2011: 64−65.
The following paratypes are deposited in RIEB (ex coll. A. Bezděk):
4 exs.: 1 ♂ ‘1985. 5. 18−19. / Wieng Ko Sai Nat. / Park, Phrae Prov., / N-Thailand / Col. S. Ohmomo [white label, front] // 5/18 夜= yoru [white label, back] // PARATYPE / Apogonia / inconstans / Kobayashi et Bezděk, 2011’. 1 ex. ‘1985. 5. 18−19. / Wieng Ko Sai Nat. / Park, Phrae Prov., / N-Thailand / Col. S. Ohmomo [white label, front] // 5/18 夜= yoru [white label, back] // PARATYPE / Apogonia / inconstans / Kobayashi et Bezděk, 2011’. 1 ex. ‘1985. 5. 18−19. / Wieng Ko Sai Nat. / Park, Phrae Prov., / N-Thailand / Col. S. Ohmomo [white label, front] // 夜= yoru 5/18 [white label, back] // PARATYPE / Apogonia / inconstans / Kobayashi et Bezděk, 2011’. 1 ex. ‘1985. 5. 18−19. / Wieng Ko Sai Nat. / Park, Phrae Prov., / N-Thailand / Col. S. Ohmomo [white label, front] // night [white label, back] // PARATYPE / Apogonia / inconstans / Kobayashi et Bezděk, 2011’.
Valid species.
Apogonia ohmomoi Miyake, 1989: 177−178.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘1985. 5. 18−19. / Wieng Ko Sai Nat. / Park, Phrae Prov., / N−Thailand / Col. S. Ohmomo [white label, front] // 夜= yoru 5/18 [white label, back] // 14 // Holotype: / Apogonia / scabra / Y. MIYAKE // Apogonia ohmomoi / Miyake, 1989 / HOLOTYPUS, ♂ / A. Bezděk det. 2011’. (Fig.
The aedeagus of the holotype is pinned separately, and the right antenna, right and left protarsus, right mesoclaw, left mesotarsus, right metatarsus and mesonotum are missing.
Valid species.
The label ‘Apogonia scabra’ was apparently accidentally put by Miyake.
Apogonia ovata Miyake & Yamaguchi, 1998: 25−27.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘Kaeng Tana / Khong Chiam / Ubonratchathani / [NE-THAILAND] / 29. Aug. 1997 / S. Yamaguchi leg. // Holotype / Apogonia ovata / Y. MIYAKE et YAMAGUCHI 1998’. (Fig.
The aedeagus of the holotype is pinned separately.
Senior synonym of Apogonia miyakei Bezděk, 2004, see
Apogonia terminalis Miyake, Yamaguchi & Akiyama, 2002: 56−58.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘Don Far Huan (light) / Ubon Ratchathani / E-THAILAND / 17. Sep. 1999 / S. Yamaguchi & / M. Akiyama leg. // Holotype / Apogonia terminalis / Y. MIYAKE, YAMAGUCHI / et AKIYAMA 2002’. (Fig.
Habitus of holotype specimens. A Apogonia terminalis Miyake, Yamagushi et Akiyama B Apogonia unidentata Miyake, Yamaguchi et Akiyama C Dichelomorpha sublineata Miyake D Holomelia gigantea Miyake E Holotrichia loochooana umebayashii Miyake F Holotrichia yamayai Miyake et Yamaguchi G Hoplia choui Miyake H Hoplia nakanei Miyake.
4exs.: 1 ♂ ‘Ban Lao Kram / Mukdahan (light) / NE-THAILAND / 5. Sep. 1999 / S. Yamaguchi & / M. Akiyama leg. // Paratype / Apogonia terminalis / Y. MIYAKE, YAMAGUCHI / et AKIYAMA 2002’. 3 ♀ ‘Don Far Huan (light) / Ubon Ratchathani / E-TAILAND / 17. Sep. 1999 / S. Yamaguchi & / M. Akiyama leg. // ♀ // Paratype / Apogonia terminalis / Y. MIYAKE, YAMAGUCHI / et AKIYAMA 2002’.
The aedeagus of the holotype is pinned separately, and the left proclaw, right and left mesotarsus, right metaclaw and left metatarsus are missing.
Valid species.
Apogonia unidentata Miyake, Yamaguchi & Akiyama, 2002: 55−56.
The holotype and following paratypes are deposited in RIEB (ex coll. Miyake):
(♂). ‘Ban Lao Kram / Mukdahan (light) / NE-THAILAND / 5. Sep. 1999 / S. Yamaguchi & / M. Akiyama leg. // Holotype / Apogonia / unidentata // Holotype / Apogonia unidentata / Y. MIYAKE, YAMAGUCHI / et AKIYAMA 2002’. (Fig.
6 exs.: 4exs. ‘Ban Lao Kram / Mukudahan (light) / NE-THAILAND / 5. Sep. 1999 / S. Yamaguchi & / M. Akiyama leg. // Paratype / Apogonia unidentata / Y. MIYAKE, YAMAGUCHI / et AKIYAMA 2002’. 1 ex. ‘Ban Lao Kram / Mukudahan (light) / NE-THAILAND / 5. Sep. 1999 / S. Yamaguchi & / M. Akiyama leg. // 63 // Paratype / Apogonia unidentata / Y. MIYAKE, YAMAGUCHI / et AKIYAMA 2002’. 1 ex. ‘Ban Lao Kram / Mukudahan (light) / NE-THAILAND / 14. Sep. 1999 / S. Yamaguchi & / M. Akiyama leg. / シミレア= shimirea // Paratype / Apogonia unidentata / Y. MIYAKE, YAMAGUCHI / et AKIYAMA 2002’.
The aedeagus of the holotype is pinned separately, and the right protarsus and right and left metatarsi are missing.
Valid species.
Dichelomorpha sublineata Miyake, 1994: 140−141.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘Is. Tioman, east of / Malay Pennisular March. 1993, nat, col // 1 [aedeagus mount] // Holotype: / Dichelomorpha / sublineata / Y. MIYAKE, 1994’. (Fig.
34 exs.: 10 ♂ ‘Is. Tioman, east of / Malay Pennisular / March. 1993. nat, col // Paratype: / Dichelomorpha / sublineata / Y. MIYAKE, 1994’. 14 exs. ‘Is. Tioman, east of / Malay Pennisular / March. 1993. nat, col // Paratype: / Dichelomorpha / sublineata / Y. MIYAKE, 1994’. 1 ♂ ‘Is. Tioman, east of / Malay Pennisular / March. 1993. nat, col // 11 // Paratype: / Dichelomorpha / sublineata / Y. MIYAKE, 1994’. 1 ex. ‘Is. Tioman, east of / Malay Pennisular / March. 1993. nat, col // 4 // Paratype: / Dichelomorpha / sublineata / Y. MIYAKE, 1994’. [[8 exs. ‘Tioman Is. / E. of Malaya / III–1933 (the date given in the original description is III–1993), W. // Paratype: / Dichelomorpha / sublineata / Y. MIYAKE, 1994’]].
The aedeagus of the holotype is pinned separately. The right mesotarsus is missing.
Valid species.
Holomelia gigantea Miyake, 1996: 35−36.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake).
(♂). ‘19 miles from / Tapah, Malaya / 1–II–1974 / Y. Miyake leg [white label, front] // Wu TET KONG / legt. [white label, back] // 29・5 // 家= ie [aedeagus mount] // Holotype: / Holomelia / gigantea / Y. MIYAKE, 1996’. (Fig.
2 exs.: 1 ♂ ‘Cameron H. / MALAYA / Jan–Feb 1974 / Y. Miyake leg. // 30.5 // Paratype: / Holomelia gigantea / Y. MIYAKE, 1996 // 19 miles from / Tapah, Malaya / 1–II–1974 [white label, front] // Wu Tet Kong legt. [white label, back]’. 1 ex. ‘19 miles from / Tapah, Malaya / 1–II–1974 [white label, front] // Wu Tet Kong legt. [white label, back] // Paratype: / Holomelia gigantea / Y. MIYAKE, 1996’.
The aedeagus of the holotype is pinned separately.
Valid species.
Holotrichia loochooana umebayashii Miyake, 1986a: 1−2.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘〔Loo-Choo〕/ Okinoerabuzima / 21. IV. 1957. / K. Umebayashi // 7 [white label, mounted on a part of tarsus] // HOLOTYPE / Holotrichia / loochooana / umebayashii / Y. MIYAKE, 1986’. (Fig.
The aedeagus of the holotype is pinned separately. The left protarsus and the right and left metatarsus are missing.
Junior subjective synonym of Nigrotrichia loochooana okinawana (Nomura, 1964), see
The collecting date in the original description is 1.V.1957, which disagrees with the label data.
Holotrichia yamayai Miyake & Yamaguchi, 1998: 27−28.
The holotype and following paratype are deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘North Thailand / Chiangmai-Saraphee / 15th, June, 1996 / Native collector // Holotype / Holotricchia yamayai / Y. MIYAKE et YAMAGUCHI 1998’. (Fig.
1 ex.: 1 ♀ ‘Kaeng Tana / Khong Chiam / Ubonratohathani / [NE-THAILAND] / 27. Aug. 1997 / S. Yamaguchi leg. // Paratype / Holotrichia yamayai / Y. MIYAKE et YAMAGUCHI 1998’.
The aedeagus of the holotype is pinned separately.
Valid species.
Hoplia choui Miyake, 1986b: 203−204.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘Sozan / Formosa / 26. V. 1940 / Coll. M. Uno // HOLOTYPE / Hoplia / Choui Y. MIYAKE / 1986’. (Fig.
Abdomen only.
Valid species.
Most part of the holotype specimen was destroyed by Anthrenus verbasci.
Hoplia nakanei Miyake, 1986b: 207.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘ (Taiwan) / Jibbon / 2. iv. 1970 / Y. Miyake // 1 // HOLOTYPE / Hoplia / nakanei Y. MIYAKE’. (Fig.
29 exs.: 4 exs. ‘ [Taiwan] / 六亀= liugui.石山= shi shan / 9–VI–1976 / Y. Miyake leg. // Paratype / Hoplia / nakanei / Y. Miyake, 1986’. 5 exs. ‘ [Taiwan] / Liukuei 石山= shi shan / 9–VI–1976 / Y. Miyake leg. // Paratype / Hoplia / nakanei / Y. Miyake, 1986’. 1 ex. ‘ [Taiwan] / 六亀= liugui.石山= shi shan / 9–VI–1976 / 陳文竜= chen wenzhao leg. // Paratype / Hoplia nakanei / Y. Miyake, 1986’. 4 exs. ‘ [Taiwan] / Baibara / –V–1963 / T. Shirozu leg. // Paratype / Hoplia / nakanei / Y. Miyake, 1986’. 1 ex. ‘ [Taiwan] / Baibara / –V–1963 / Shirozu leg. // Paratype / Hoplia / nakanei / Y. Miyake, 1986’. 1 ex. ‘ [Taiwan] / Baibara / –V–1963 / T. Shirozu // Paratype: Hoplia / nakanei Y. Miyake, 1986’. 1 ♂ ‘六亀= liugui.石山= shi shan / alt. 2000 m / 9. VI. 1976 // Paratype: / Hoplia / nakanei / Y. MIYAKE, 1986’. 1 ex. ‘FENCHIFO / FORMOSA / 24. V. 1975 / K. MATSUDA // Paratype: / Hoplia nakanei / Y. MIYAKE, 1986’. 1 ex. ‘Swakang / Taiwan / 15. VI. 1970 / Fukuda // Paratype: / Hoplia nakanei / 1986. Y. MIYAKE’. 1 ex. ‘NANSHANCHI / TAIWAN / 1. IV. 1970 / H. NOMURA // PARATYPE / Hoplia / nakanei / Y. MIYAKE, 1986’. [[4 exs. ‘ [Taiwan] / Baibara / V. 1965 (the date given in the original description is V. 1963) / T. Shirozu // Paratype / Hoplia nakanei / Y. Miyake’]]. [[2 exs. ‘ [Taiwan] / Baibara / V. 1965 (the date given in the original description is V. 1963) / T. Shirozu // Paratype / Hoplia nakanei / Y. Miyake, 1986’]]. [[1 ♂ ‘ [Taiwan] / Baibara / V. 1965 (the date given in the original description is V. 1963) / Col. T. Shirozu // Paratype / Hoplia nakanei / Y. Miyake, 1986’]]. [[1 ex. ‘ [Taiwan] / Baibara / V. 1965 (the date given in the original description is V. 1963) / Col. T. Shirozu // Paratype / Hoplia nakanei / Y. Miyake, 1986’]]. [[1 ex. ‘ [Taiwan] / Baibara / V. 1965 (the date given in the original description is V. 1963) / Col. T. Shirozu // Paratype / Hoplia nakanei / 1986. Y. Miyake’]].
The aedeagus of the holotype is pinned separately.
Valid species.
The label data the holotype do not match the original description. There is one male paratype (‘六亀= liugui.石山= shi shan / alt. 2000 m / 9. VI. 1976 // Paratype: / Hoplia / nakanei / Y. MIYAKE, 1986’) that agrees with the data of the holotype. Probably the type labels were mixed up and this specimen should be considered the holotype.
In addition to the paratypes listed above, the following specimens labeled as paratypes are not designated in the original description: 1 ex. ‘ [Taiwan] / Liukuei / –V–1963 / T. Shirozu col. // Paratype / Hoplia / nakanei / Y. Miyake, 1986’. 1 ex. ‘APR II. 1971 / 榮華= ronghua / B-S. CHANG // PARATYPE / Hoplia / nakanei / Y. MIYAKE, 1986’. 1 ex. ‘Paratype / Hoplia nakanei / Y. MIYAKE, 1986’.
Hoplia nengkaoshana Miyake, 1986b: 211.
The allotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Allotype (♀). ‘Allotype // mt. Noko (2650 m) / V. 20–VI. 2, 1966 // 1 // PARATYPE / Hoplia / nengkaoshana / Y. MIYAKE, 1986’.
3ex.: 1 ♂ ‘mt. Nōhō / C-Formosa / V–20 ~ Vi–21 1966 // PARATYPE / Hoplia / nengkaoshana / Y. MIYAKE, 1986’. 1 ♂ ‘MT. HÔFAN / FORMOSA / 5–V. 73 / T. OCHI // 1 // PARATYPE / Hoplia / nengkaoshana / Y. MIYAKE, 1986 // Hoplia / inornata / H. Kobayashi, 1990 / Det. H. Kobayashi, 2015’. 1 ex. ‘JUL. 16. 1966 / 合欢山= hehuan shan / B. S. Chang // PARATYPE / nengkaoshana / Y. MIYAKE, 1986 // Hoplia / inornata / H. Kobayashi, 1990 / Det. H. Kobayashi, 2015’.
Valid species.
In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 1 ex. ‘台中= tai chung 和平郷= heping xiang 早等村= zaodeng cun / 横貫公路= hengguan gonglu. 梨山站= lishan zhan / 海抜= kaibatsu 1679–2196 m. / 13–16. V. 1958. 陶家駒= tao jiayu // PARATYPE / Hoplia / nengkaoshana / Y. MIYAKE, 1986’. 1 ex. ‘AUG 13. 1970 / 新坡= xinpo / B-S. CHANG // PARATYPE / Hoplia / nengkaoshana / Y. MIYAKE, 1986’.
Hoplia simillima Miyake, 1986b: 206−207.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘ [Taiwan] / Liukuei 石山= shi shan / 9–VI–1979 / Y. Miyake leg. // HOLOTYPE / Hoplia / simillima / Y. MIYAKE, 1986’. (Fig.
5 exs.: 3 exs. ‘(FORMOSA) / Baibara / –V–1969 // Paratype / Hoplia / simillima / 1989, Y. Miyake’. 2 exs. ‘(FORMOSA) / Baibara / –V–1969 / T. Shirozu // Paratype / Hoplia / simillima / 1989, Y. Miyake’.
Valid species.
In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 1 ex. ‘ [Taiwan] / Liukuei 石山= shi shan / 9–VI–1979 / Y. Miyake leg. // Paratype / Hoplia / simillima / 1989, Y. Miyake’. 1 ♂ ‘石= ishi // [Taiwan] / 六亀= liugui 石山= shi shan / 9–VI–1976 / Y. Miyake leg. // Paratype / Hoplia / simillima / 1989, Y. Miyake’. 1 ex. ‘(FORMOSA) / Mt. Ali / 14 V 1968 / Y. HAYASHI // Paratype’.
Hoplia taiwana Miyake, 1986b: 204.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘HOLOTYPE / Hoplia / Taiwana Y. MIYAKE / 1986’. (Fig.
9 exs.: 1 ex. ‘NANSHANCHI / TAIWAN / 3. IV. 1970 / H. NOMURA // PARATYPE / Hoplia taiwana / Y. MIYAKE, 1986’. 1 ex. ‘NANSHANCHI / TAIWAN / 4. IV. 1970 / H. NOMURA // PARATYPE / Hoplia taiwana / Y. MIYAKE, 1986’. 1 ex. ‘NANSHANCHI / TAIWAN / 5. IV. 1970 / Y. KIYOYAMA // PARATYPE / Hoplia taiwana / Y. MIYAKE, 1986’. 1 ex. ‘SUNGKANG / FORMOSA / 29. VI. 1971 / Y. MAEDA // PARATYPE / Hoplia taiwana / Y. MIYAKE, 1986’. 1 ex. ‘NANSHANCHI / TAIWAN / 31. III. 1981 / Y. YAMAMOTO // Paratype / Hoplia / taiwana / Y. MIYAKE, 1986’. 1 ex. ‘NANSHANCHI / TAIWAN / 1. IV. 1981 / F. KIMURA // Paratype / Hoplia / taiwana Y. MIYAKE / (1968)’. 1 ex. ‘NANSHANCHI / TAIWAN / 2. IV. 1981 / F. KIMURA // Paratype / Hoplia / taiwana Y. MIYAKE’. 1 ex. ‘NANSHANCHI / TAIWAN / 2. IV. 1981 / F. KIMURA // Paratype / Hoplia taiwana / 1986 Y. MIYAKE’. 1 ex. ‘NANSHANCHI / TAIWAN / 3. IV. 1981 / Y. YAMAMOTO // PARATYPE / Hoplia taiwana / Y. MIYAKE, 1986’.
Valid species.
The locality label data of the holotype are missing but the specimen agrees with the original description.
In addition to the paratypes mentioned above, the following specimens labeled as paratypes were not designated in the original description: 3 exs. ‘ [Taiwan] / Liukuei 石山= shi shan / 9–VI–1976 / Y. Miyake leg. // PARATYPE / Hoplia / taiwana / Y. MIYAKE, 1986’. 1 ex. ‘ [Taiwan] / 六亀= liugui 石山= shi shan / 9–VI–1976 / Y. Miyake leg. // PARATYPE / Hoplia / taiwana / Y. MIYAKE, 1986’. 2 exs. ‘Mt. YUSHAN / TAIWAN / 8. VI. 1980 / M. YAMAMOTO // Paratype / Hoplia / taiwana Y. MIYAKE’. 4 exs. ‘TAIWAN / 陽明山= mt. yangmingshan / 20. IV. 1956 / COL. K. S. LIN // Paratype / Hoplia / taiwana / Y. MIYAKE, 1986’. 1 ex. ‘LUSHAN / FORMOSA / 13–IV. 74 / S. TAKEDA // PARATYPE / Hoplia / taiwana / Y. MIYAKE, 1986’. 1 ex. ‘榮華= ronghua / APR. 7. 1971 / BS, CHANG // PARATYPE / Hoplia taiwana / Y. MIYAKE, 1986’. 1 ex. ‘ [Taiwana] / Baibara / –V–1965 / T. Shirozu leg. // Paratype: Hoplia / taiwana Y. MIYAKE. 1986’.
Maladera hiranoi Miyake, 1986b: 3−5.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘三京= mikyô / 21/VII–82 // HOLOTYPE / Maladera / hiranoi MIYAKE / (1986) // Maladera / impressthorax / hiranoi Y. MIYAKE / DET. Y. MIYAKE, 199 // ムナクボビロウドコガネ= munakubobirôdokogane / Maladera (japonica–G) / impressithorax / Nomura, 1973 / Det. H. Hirasawa 2012’. (Fig.
Junior subjective synonym of Maladera okinoerabuana Kobayashi, 1978, see
The date on the collecting data label of the holotype is 21.VII.1982, but the collecting date quoted in the original description is 22.VII.1982. Probably, Miyake made a typographical error when writing the description.
Maladera kobayashii Nomura, 1974: 106−107.
The following paratypes are deposited in RIEB (ex coll. S. Nomura):
2 exs.: 1 ♂ ‘Li-shan / Taiwan / 29. VII. 1974 / Y. Miyake // ♂ // PARATYPE / Maladera (s. str) / kobayashii /
Valid species.
Maladera kusuii Miyake, 1986b: 2−3.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘三宅= miyake / コガネムシ上科標準図鑑= koganemushijyôka-hyôjyunzukan / ビロウドコガネ族= birôdokoganezoku / 図鑑使用標本= zukanshiyôhyôhon / 執筆担当= shippitsutantô:平沢 伴明= hirasawa tomoaki / 学研教育出版= gakkenkyôikushuppan (2012) // GAKIYA / Iheya-jima / Aug. 11 1977 / leg. Y Kusui // イヘヤビロウドコガネ= iheyabirôdokogane / Maladera (japonica–G) / kusuii / Y. Miyake, 1986 / Det. H. Hirasawa 2012 // Holotype: / Maladera kusuii / Y. MIYAKE, 1986’.
The aedeagus of the holotype is pinned separately. The right and left protarsi, the left mesotarsus, and the right and left metatarsi are missing, but some of the missing parts are pinned separately.
Valid species.
Maladera lishana Miyake, 1989b: 37−38.
The following paratype is deposited in RIEB (ex coll. Y. Miyake):
1 ex.: 1 ♀ ‘Lishan / Taiwan / 28. VII. 1973 / Y. Miyake // Paratype: / Maladera / lishana / Y. MIYAKE, 1989’.
Valid species.
Maladera nanshanchiana Nomura, 1974: 111−112.
The following paratypes are deposited in RIEB (ex coll. S. Nomura):
2 exs.: 1 ex. ‘Puli 24 // PARATYPE / Maladera / nanshanchiana /
Valid species.
Maladera secreta horaiana Nomura, 1974: 112.
The following paratype is deposited in RIEB (ex coll. S. Nomura):
1 ex.: 1 ♂ ‘Lishan / C. Taiwan / 29. vii. 1973 / Y. Miyake // 三宅= miyake // PARATYPE / Maladera / secreta / horaiana /
Valid species.
Genus Melanomaladera
Melanomaladera yunnana Miyake & Yamaya, 2001: 38−39.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
2 exs.: 1 ♂ ‘China, N W Yunnan / Degen City env. / 3300 m alt. / 29. Jun. 1998 / A. Gorodinsky leg. // 13 // Paratype: / Melanoserica / yunnana’. 1 ♀ ‘China, N W Yunnan / Degen City env. / 3300 m alt. / 29. Jun. 1998 / A. Gorodinsky leg. // 13 // Paratype: / Melanoserica / yunnana’.
Archeohomaloplia yunnana (Miyake et Yamaya, 2001), see
The genus name on the paratype labels is ‘Melanoserica’, which is invalid. Probably, it was confused with ‘Melanomaladera’ by Y. Miyake.
Microserica ( Parvulomaladera) annapurnae Ahrens, 1995: 46−48.
The following paratype is deposited in RIEB (ex coll. D. Ahrens):
1 ex.: ‘NEPAL–HIMALAYA / Annapurna–Mts. / leg. Ahrens 1993 // Pisang bis / Manang, 30. 5. / 3000–3300 m // PARATYPEUS / Parvulomaladera / annapumae sp. n. / det. D. AHRENS 1994’.
Oxyserica pygidialis annapurnae (Ahrens, 1995), see
Microserica nitidipyga Nomura, 1974: 99.
The following paratype is deposited in RIEB (ex coll. S. Nomura):
1 ex.: ‘Lu-SHAN / FORMOSA / 27. IV. 1973 / K. MATSUDA // 三宅= miyake // Paratype / Microserica / nitidipyga /
Valid species.
Nematophylla sugiharai Miyake, 2000: 106−108.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘Kiau Gap / Mt. Kinabalu / Sabah, Borneo / 1. IV. 2000 / H. SUGIHARAI leg. // Holotype: Nematophylla / sugiharai / Y.
Junior subjective synonym of Nematophylla carinicollis Arrow, 1938, see
Nipponoserica pubiventris Nomura, 1976b: 189−190.
The following paratype is deposited in RIEB (ex coll. S. Nomura):
1 ex.: ‘Takashinohama, / Osaka Pref. / 3. VI. 1974 / Coll. Mitsuo Goto // at light // PARATYPE / Nipponoserica / pubiventris / NOMURA (1976)’.
Valid species.
Paramaladera makiharai Nomura, 1974: 101−102.
The following paratypes are deposited in RIEB (ex coll. S. Nomura):
4 exs.: 4 ♂ ‘Li-shan / Taiwan / 29. VII. 1974 / Y. Miyake // ♂ // Paratype / Paramaladera / makiharai /
Valid species.
Pseudohoplia shibatai Miyake, 1986b: 201.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
2 exs.: 1 ♂ ‘(FORMOSA)/ Mt. Ali / 15 V 1968 / Y. HAYASHI // Paratype // Hoplia / shibatai shibatai / Y. Miyake, 1986 / Det. H. Kobayashi, 2015’. 1 ex. ‘(FORMOSA)/ Mt. Ali / 15 V 1968 / Y. HAYASHI // Paratype // Hoplia / shibatai shibatai / Y. Miyake, 1986 / Det. H. Kobayashi, 2015’.
Hoplia shibatai shibatai (Miyake, 1986), see
In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 1 ex. ‘Mt. Arishan / Formosa / 1. V. 1973 / K. MATSUDA // Paratype // Paratype / Pseudohoplia / shibatai / Y. Miyake, 1986 // 贈台湾農試= zou-taiwan-noushi’. 1 ♂ ‘Houanchi- / Sundchuankang / Nantou Hsien / Taiwan / 25. VI. 1976 / H. Makihara leg. // Paratype // Paratype / Pseudohoplia / shibatai / Y. Miyake, 1986’. 1 ex. ‘Mt. Arishan / Formosa / 1. V. 1973 / K. MATSUDA // Paratype / Pseudohoplia / shibatai / Y. Miyake, 1986’. 1 ex. ‘MT. ARI / FORMOSA / 1. V. 1973 / K. MATSUDA // Paratype // Paratype / Pseudohoplia / shibatai / Y. Miyake, 1986’.
Pseudohoplia shibatai makiharai Miyake, 1986b: 202.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘Mt. Lalashan / Taoyuan Hsien / Taiwan / 21–24. V. 1980 / H. Makihara // HOLOTYPE / Pseudohoplia / shibatai makiharai / Y. MIYAKE, 1886’. (Fig.
The aedeagus of the holotype is pinned separately. The right antenna is missing.
Hoplia shibatai makiharai (Miyake, 1986), see
Pseudohoplia shibatai matsudai Miyake, 1986b: 202.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘Mt. Nōko / C-Formosa / v–20~vi–21’ 66 // HOLOTYPE / Pseudohoplia / shibatai matsudai / Y. MIYAKE, 1986’. (Fig.
The aedeagus of the holotype is pinned separately.
Junior subjective synonym of Hoplia shibatai shibatai (Miyake, 1986), see
Pseudomaladera nitidifrons Nomura, 1974: 96.
The following paratypes are deposited in RIEB (ex coll. S. Nomura):
2 exs.: 1 ♂ ‘FUNCHIHO / FORMOSA / 1. VIII. 1973 / H. OHASHI // ♂ // 三宅= miyake // Paratype / Pseudomaladera / nitidifrons / NOMURA, 1974’. 1 ♂ ‘MEIFENG / FORMOSA / 1. VIII. 1973 / T. OCHI // ♂ // Paratype / Pseudomaladera / nitidifrons / NOMURA, 1974’.
Valid species.
Sinoserica maculipennis Miyake & Yamaya, 2001: 40−41.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
2 exs.: ‘China, Yunnan / Boshan C. env. / 2100 m alt. / 10. VII. 1998 / A. Gorodinski // 12 // Paratype / Sinoserica / maculipennis’.
Trioserica maculipennis (Miyake et Yamaya, 2001), see
Serica fusifemorata Nomura, 1974: 91.
The following paratypes are deposited in RIEB (ex coll. S. Nomura):
11 exs.: 1 ♀ ‘Lishan / C. Taiwan / 29. vii. 1973 / Y. Miyake // ♀ // 三宅= miyake // Paratype / Serica / fusifemorata /
Valid species.
Serica gansuensis Miyake & Yamaya, 2001: 35−36.
The following paratype is deposited in RIEB (ex coll. Y. Miyake):
1 ex.: ‘China, Gangu / Mts. Minshan / nr. Wudn. 2100 / 1. VI. 1997 / A. Gorodinski // 4 // Paratype: / Serica / gansuensis’.
Valid species.
Sericania miyakei Nomura, 1960: 60.
The following paratype is deposited in RIEB (ex coll. S. Nomura):
1 ex.: ‘Paratype // TASHIRO / CHIKUGO / 4. V. 1952 / Y. MIYAKE // Sericania / miyakei /
Valid species.
Sophrops takatoshii Itoh, 1990: 9−10.
The following paratypes are deposited in RIEB (ex coll. T. Itoh):
3 exs.: ‘1989–4–17 / 平良植物園= hirara-syokubutsuen /(宮古島)= miyako-jima / leg. T. Ueno // Paratype / Sophrops / takatoshii / T. Itoh’.
Valid species.
Taiwanoserica elongata Nomura, 1974: 86.
The following paratype is deposited in RIEB (ex coll. S. Nomura):
1 ex.: 1 ♂ ‘MT. ARI / FORMOSA / 30. VII. 1973 / H. OHASHI // ♂ // 三宅= miyake // Paratype / Taiwanoserica / elongata /
Serica (Taiwanoserica) elongata (Nomura, 1974), see
Taiwanoserica lishana Nomura, 1974: 87.
The following paratypes are deposited in RIEB (ex coll. S. Nomura):
22 exs.: 2 ♂ ‘Lishan / C. Taiwan / 30. vii. 1970 / Y. Miyake // ♂ // PARATYPE / Taiwanoserica / lishana / S.
Serica (Taiwanoserica) lishana (Nomura, 1974), see
Trichomaladera elongata Nomura, 1974: 93−94.
The following paratype is deposited in RIEB (ex coll. S. Nomura):
1 ex.: 1 ♂ ‘Paratype / Trichomaladera / elongata /
Valid species.
The collecting data label of the paratype is missing.
Trichoserica elongata nitididorsis Nomura, 1971: 70−71.
The following paratype is deposited in RIEB (ex coll. S. Nomura):
1 ex.: ‘Mt. Hiko / [KYUSHU] / 27. 7. 1960 / TAKEISHI // Paratype / Trichoserica / elongata / nitididorsis / Nomura’.
Serica nitididorsis (Nomura, 1971), see
Trichoserica incurvata Nomura, 1971: 67−68.
The following paratypes are deposited in RIEB (ex coll. S. Nomura):
8 exs.: 1 ex. ‘Aomori-pref / 17–vii–1956 / S. Kimotog // Paratype / Trichoserica / incurvata /
Serica incurvata (Nomura, 1971), see
Trichoserica nitidiceps Nomura, 1971: 69−70.
The following paratype is deposited in RIEB (ex coll. S. Nomura):
1 ex.: 1 ♂ ‘(Kyushu) / Mt. Sobo / 23. V. 1952 / Y. Miyake //Paratype / Trichoserica / nitidiceps / NOMURA’.
Junior subjective synonym of Serica inexspectata (Kontkanen, 1956), see
In addition to the paratype mentioned above, the following specimen labeled as paratype is not designated in the original description: 1 ex. ‘Mt. Kuju / Bungo / 28. vii. 1961 / Y. Miyake // Paratype / Trichoserica / nitidiceps /
Trichoserica opacifrons Nomura, 1971: 68−69.
The following paratypes are deposited in RIEB (ex coll. S. Nomura):
3 exs.: 1 ♂ ‘Mt. Hiko / Buzen / 1. VIII. 1953 / Y. Miyake // Paratype / Trichoserica / opacifrons / NOMURA’. 1 ex. ‘mt. Kujū / Hokkeein / 27. VII. 1961 / Y. Miyake Coll // Paratype / Trichoserica / opacifrons / NOMURA’. 1 ♂ ‘ [Kyushu] / Yakushima / 21. Vi. 1954 / Col. K. Matsuda // Paratype / Trichoserica / opacifrons / NOMURA // Ophthalmoserica/ matsudai / sp.n. / DET. Y. MIYAKE’.
Senior objective synonym of Serica planifrons Nomura, 1972, see
Genus Anomalocera
Anomalocera paotao Masumoto & Sakai, 1987a: 51−54.
The following paratypes are deposited in RIEB (ex coll. Masumoto):
2 exs.: 1 ex. ‘Lishan / Taiwan / 28. VII. 1974 / Y. Miyake // Paratype / Anomalocera / paotao MAS. et SAK.’. 1 ex. ‘Lishan / C, -Taiwan / 29 vii. 1973 / Y. Miyake // Paratype / Anomalocera / paotao MAS. et Sak.’.
Diphyllomorpha paotao (Masumoto et Sakai, 1987), see
Rhomborhina kurosawai Masumoto & Sakai, 1987b: 104−105, 107.
The following paratypes are deposited in RIEB (ex coll. Masumoto):
3 exs.: 2 ♂ ‘FORMOSA / tattaka / 9, VIII 1971 / H. OHARA // Paratype / Rhomborrhina / kurosawai / MASUMOTO et K. SAKAI’. [[1 ♂ ‘Tsuifeng / C. Tsiwan / 26. vii. 1970 (the date given in the original description is 27.vii.1970) // Y. Miyake // Paratype / Rhomborrhina / kurosawai / MASUMOTO et K. SAKAI’]].
Valid species.
Thaumastopeus chicheryi tenuigenius Miyake & Yamaya, 1995: 38.
The following paratype is deposited in RIEB (ex coll. Y. Miyake):
1 ex.: 1 ♂ ‘N. Keningau / N. Borneo / 30. V. 1994 // Paratype: Thaumastopeus / chicheryi tenuigenius / Y. MIYAKE et YAMAYA, 1995’.
Valid species.
Paratrichius hatai Miyake, 1989: 41−42.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
4 exs.: [[‘Tanah Rata / MALAYA / 7. IV. 1972 (the date given in the original description is 7.IV.1978) / M. Hata // Paratype: / Paratrichius / hatai / Y. MIYAKE, 1989’]].
Valid species.
Paratrichius kyushuensis Miyake, 1990: 30−32.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘MT. SOBO / BUNGO / 28. vii. 1954 / Y. MIYAKE // Holotype: / Paratrichius / kyushuensis / Y. MIYAKE, 1990’. (Fig.
5 exs.: 1 ex. ‘(Kyushu) / Mt. Sobo / 24. VII. 1950 / Y. Miyake // Paratype: Paratrichius / kyushuensis / Y. MIYAKE, 1990’. 1 ♂ ‘(Kyushu) / Mt. Sobo / 24. VII. 1952 / Y. Miyake // Paratype: Paratrichius / kyushuensis / Y. MIYAKE, 1990’. 1 ex. ‘(Kyushu) / Mt. Ichi Fu Sa / 28 VII 1952 / Col. S. Kimoto // Paratype: Paratrichius / kyushuensis / Y. MIYAKE, 1990’. 1 ex. 1 ♂ ‘(Kyushu) / Mt. Sobo / 24. VII. 1953 / K. MATSUDA // Paratype: Paratrichius / kyushuensis / Y. MIYAKE, 1990’.
The left metatarsus of the holotype is missing.
Valid species.
The date on the label of the holotype is 28.VII.1954 while that in the original description is 28.VII.1952.
In addition to the paratypes mentioned above, the following specimen labeled as paratype is not designated in the original description.: 1 ex. ‘⑦ // Paratype: Paratrichius / kyushuensis / Y. MIYAKE, 1990’.
Tibiotrichius vietnamensis Miyake, 1996: 44.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘M. ITO leg / Sapa=Chapa / N. Vietnam / 23–V–1995 // サパ= sapa [aedeagus mount] // Holotype: Tibiotrichius / vietnamensis Y. MIYAKE, 1996’. (Fig.
The aedeagus of the holotype is pinned separately.
Valid species.
Genus Charitovalgus
Charitovalgus banzai Sawada, 1941: 8−9.
The neotype is deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘Hourng Shoei ken / near Liukuei, / Kao Hsung Hsien / VII. 1972 W. Chen leg // Holotype: Neotype / Charitovalgus / banzai SAWADA / Y. MIYAKE, 1993’. (Fig.
The right and left metatarsi of the neotype are missing.
Valid species.
The neotype was designated by Miyake (1993).
Dasyvalgus annamensis Miyake, 1993a: 21−22.
The holotype and following paratype are deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘4. 4 / 203 // Iluon Ma Thuot / Annam M. ITO // Holotype: Dasyvalgus / Annamensis / Y. MIYAKE, 1993’. (Fig.
1 ex.: 1 ♂ ‘Iluon Ma Thuot / Annam M. ITO // Paratype: Dasyvalgus / annamensis / Y. MIYAKE, 1993’.
Valid species.
The locality on the collecting data label of the holotype is ‘Iluon Ma Thuot’, but the locality given in the original description is ‘Ban Me Thuot’. However, the habitus of the holotype and the figure in the original description match. Probably, Miyake made a typographical error when writing the description.
Dasyvalgus castaneodorsalis Miyake, 1993a: 24−25.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘〔Thailand〕/ Pupin–Doi Phui / 19–IV–1985 / M. TAO leg. // Holotype: Dasyvalgus / casraneodorsalis / Y. MIYAKE, 1993’. (Fig.
6 exs.: 2 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 19–IV–1985 / M. TAO leg. // Paratype: Dasyvalgus / castaneodorsalis / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 21–IV–1985 / M. TAO leg. // Paratype: Dasyvalgus / castaneodorsalis / Y. MIYAKE, 1993’. 2 ♂ ‘〔Thailand〕/ Pupin / 21–IV–1985 / M. TAO // Paratype: Dasyvalgus / castaneodorsalis / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Doi Suthep / 10–V–1985 / M. TAO leg. // Paratype: Dasyvalgus / castaneodorsalis / Y. MIYAKE, 1993’.
The aedeagus of the holotype is pinned separately.
Valid species.
Dasyvalgus decamaculatus Miyake, 1993a: 22−23.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘Iluon Ma Thuot / Annam M. ITO // Holotype: / Dasyvalgus / decamaculatus / Y. MIYAKE, 1993’. (Fig.
2 exs.: 1 ex. ‘Near Chang Mai / N. Thailand / 7–V–1994 / K. KUME leg. // Paraype: Dasyvalgus / decamaculatus / Y. MIYAKE, 1993’. 1 ex. ‘Paratype: Dasyvalgus / decamaculatus / Y. MIYAKE, 1993’.
The right and left metatarsus are missing.
Valid species.
The locality on the collecting data label of the holotype is ‘Iluon Ma Thuot’, but the locality given in the original description is ‘Ban Me Thuot’. However, the habitus of the holotype and the figure in the original description match. Probably, Miyake caused a typographical error when writing the description.
Dasyvalgus flavicauda Miyake, Yamaguchi, Aoki et Akiyama, 2002: 65−66.
The holotype and following paratype are deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘Ban Lao Kram / Mukdahan / E-THAILAND / 17. Feb. 1999 / S. Yamaguchi and / T. Aoki leg. // Holotype: / Dasyvalgus / flavicauda // Holotype / Dasyvalgus flavicauda / Y. MIYAKE, YAMAGUCHI / et AOKI 2002’. (Fig.
1 ex.: 1 ♂ ‘Phu Huai Sing / Mukdahan / E–THAILAND / 19. Feb. 1999 / S. Yamaguchi and / T. Aoki leg. // Paratype / Dasyvalgus flavicauda / Y. MIYAKE, YAMAGUCHI / et AOKI 2002’.
Valid species.
Dasyvalgus macacus Miyake, 1993a: 24.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘〔Thailand〕Pupin–Doi Phui / 19–IV–1985 / M. TAO leg. // Holotype: Dasyvalgus / macacus / Y. MIYSKE, 1993’. (Fig.
4 exs.: 1 ♂ ‘〔Thailand〕/ Pupin-Doi Phui / 1–V–1985 / M. TAO leg. // Paratype: Dasyvalgus / macacus / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Pupin-Doi Phui / 5–V–1985 / M. TAO leg. // Paratype: Dasyvalgus / macacus / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Pupin-Doi Phui / 12–V–1985 / M. TAO leg. // Paratype: Dasyvalgus / macacus / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Pupin-Doi Phui / 21–V–1985 / M. TAO leg. // Paratype: Dasyvalgus / macacus / Y. MIYAKE, 1993’.
Valid species.
The habitus photograph in the original description does not refer to the holotype.
Dasyvalgus makiharai Miyake, 1985: 11−12.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
1 ex.: ‘MEIFENG / FORMOSA / 3–VII. 74 / T, OCHI // Paratype: / Dasyvalgus / makiharai Y.
Valid species.
In addition to the paratype mentioned above, the following specimens labeled as paratype are not designated in the original description: 1 ex. ‘松崗= songgang / 27~29. VII 1968 / K. YAMAMOTO // Paratype: Dasyvalgus / makiharai / Y. MIYAKE, 1985’. 1 ♂ ‘SUNG KANG / FORMOSA / 30–VI, 74 / T. OCHI // Paratype: / Dasyvalgus / makiharai Y.
Dasyvalgus minahasanus Miyake, 1989b: 42−43.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
78 exs.: 75 exs. ‘Tondano / N. Sulawesi / 10–IV–1989 / Y. Miyake leg. // Paratype: / Dasyvalgus / minahasanus / Y. MIYAKE, 1989’. 3 exs. ‘Tondano / N. Sulawesi / 9–IV–1989 / Y. Miyake leg. // Paratype: / Dasyvalgus / minahasanus / Y. MIYAKE, 1989’.
Valid species.
In addition to the paratypes mentioned above, the following specimen labeled as paratypes is not designated in the original description: 1 ex. ‘Tondano / N. Sulawesi / 11–IV–1989 / Y. Miyake leg. // Paratype: / Dasyvalgus / minahasanus / Y. MIYAKE, 1989’.
Dasyvalgus multicus Miyake, 1993a: 25−26.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘ [Thailand] / プピン-プイ= pupin-pui / 5–V–1985 / M. Tao leg. // Holotype: Dasyvalgus / multicus / Y. MIYAKE, 1993’. (Fig.
The aedeagus of the holotype is pinned separately. The right and left protarsus are missing.
Valid species.
Dasyvalgus nudis Miyake, 1994: 153.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘Cao Bang / N. Vietnam / 9–11. VI, 1993 / M. ITO leg // Holotype: Dasyvalgus / nudis / Y. MIYAKE, 1994’. (Fig.
9 exs.: 3 ♂ ‘Cao Bang / N. Vietnam / 8–10 VI, 1993 / M. ITO leg. // Paratype: / Dasyvalgus / nudis / Y. MIYAKE, 1994’. 3 ♂ ‘Cao Bang / N. Vietnam / 10. VI, 1993 / M. ITO leg. // Paratype: / Dasyvalgus / nudis / Y. MIYAKE, 1994’. [[2 ♂ ‘Cao Bang / N. Vietnam / 9–11. VI, 1993 (real date is 8–10. VI. 1993) / M. ITO leg. // Paratype: / Dasyvalgus / nudis / Y. MIYAKE, 1994’]]. [[1 ex. ‘Cao Bang / N. Vietnam / 11. VI, 1993 (real date is 8–10. VI. 1993) / M. ITO leg. // Paratype: / Dasyvalgus / nudis / Y. MIYAKE, 1994’]].
The aedeagus of the holotype is pinned separately. The right antenna is missing.
Valid species.
The date on the collecting data label of the holotype is 9–11,VI,1993, but the collecting date quoted in the original description is 8–10,VI,1993. However, the habitus of the holotype and the figure in the original description match. Probably, Miyake caused a typographical error when writing the description.
Dasyvalgus rubrothoracicus Miyake, 1993a: 23−24.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘〔Thailand〕/ Pupin–Doi Phui / 21–IV–1985 / M. TAO leg. // Holotype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’. (Fig.
11 exs.: 1 ex. ‘Doi Sutehp / Thailand / 28–IV–1985 / M Tao leg. // TDvA // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 1–V–1985 / M. TAO leg. // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 1–V–1985 / M. TAO leg. // TDvA // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 5–V–1985 / M. TAO leg. // TDvA // TDvA // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 11–V–1985 / M. TAO leg. // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’. [[1 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 17–V–1985 (the date given in the original description is 17–IV–1985) / M. TAO leg. // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’]]. [[1 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 21–V–1985 (the date given in the original description is 21–IV–1985) / M. TAO leg. // TDvA // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’]]. [[1 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 21–V–1985 (the date given in the original description is 21–IV–1985) / M. TAO leg. // TDvA // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’]]. [[1 ♂‘〔Thailand〕/ Pupin-Doi-Phui / 23–V–1985 (the date given in the original description is 23–IV–1985) / M. TAO leg. // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’]]. [[2 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 28–V–1985 (the date given in the original description is 28–IV–1985) / M. TAO leg. // TDvA // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’]].
The aedeagus of the holotype is pinned separately.
Valid species.
The date on the collecting data label of the holotype is 21–IV–1985, but the collecting date quoted in the original description is 5–V–1985. However, the habitus of the holotype and the figure in the original description match. Probably, Miyake caused a typographical error when writing the description.
Dasyvalgus similis Miyake, 1985: 8−9.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
3 exs.: 1 ♂ ‘蓬来= hôrai / 20, 21 VIII 1968 / H. MAKIHARA // Paratype’. 1 ♂ ‘FUNCHIIFO / FORMOSA / 1. VI. 1970 / Y. KIYOYAMA // Paratype // Paratype / Dasyvalgus / similis, Y.
Senior objective synonym of Dasyvalgus taiwanus Miyake, 1991. Miyake in
In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 1 ex. ‘LIUKUEI / FORMOSA / 3. X. 1970 / Y. KIYOYAMA // 16 // Paratype // Paratype / Dasyvalgus / similis Y. MIYAKE, 1985’. 1 ex. ‘NANSHANCHI / FORMOSA / 27. VI. 1971 / Y. MAEDA // Paratype // Paratype / Dasyvalgus / similis Y. MIYAKE, 1985’. 1 ex. ‘紅水坑= hongshuikeng / Liukuei / 11. V. 1978 / Chen. W. // Paratype’. 1 ♂ ‘紅水坑= hongshuikeng / Liukuei / 11. V. 1978 / Chen. W. // Paratype’. 1 ex. ‘紅水坑= hongshuikeng / Liukuei / 11. V. 1978 / W. Chen // Paratype // Paratype / Dasyvalgus / similis Y. MIYAKE, 1985’. 1 ex. ‘Paratype’.
Dasyvalgus wadai Miyake, 1985: 9−10.
The following paratype is deposited in RIEB (ex coll. Y. Miyake):
1 ex.: 1 ♂ ‘Paratype / Dasyvalgus / wadai Y. MIYAKE, 1985’.
Valid species.
This paratype has no collecting data label.
Hybovalgus matsudai Miyake, 1985: 6.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘Paratype // Holotype / Hybovalgus / matsudai Y.
The right protarsus is missing.
Valid species.
There is no data label attached to the holotype, but the habitus of the holotype matches the figure in the original description. If confidence is placed in the pinned holotype label and the habitus, the above holotype specimen is the true holotype.
Neovalgus formosanus Miyake, 1985: 3.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘六亀= liugui.石山= shi shan / 9. VI. 1976 / 2000m // Holotype: / Neovalgus / formosanus / Y, MIYAKE, 1985’. (Fig.
2 exs.: 1 ex. ‘ [Taiwan] / Baibara, mt. Puli / –V–1965 // Paratype: Neovalgus / formosanus Y. MIYAKE, 1985’. 1 ex. ‘ [Taiwan] / Liukuei / 16–V–1978 // PARATYPE / Neovalgus / formosanus / Y. MIYAKE, 1985’.
Valid species.
The date on the collecting label of the holotype is 9.VI.1976, but the collecting date quoted in the original description is 16.V.1978. However, the habitus of the holotype matches the figure in the original description. Probably, Miyake caused a typographical error when writing the description.
In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 3 exs. ‘ [Taiwan] / Baibara, mt. Puli / –V–1965 // PARATYPE / Neovalgus / formosanus / Y. MIYAKE, 1985’. 1 ex. ‘Formosa / Wushe / V. 1965 / T. Shiroz / Paratype: Neovalgus / formosanus / Y. MIYAKE, 1985’. 1 ex. ‘六 亀= liugui, 石山= shi shan / 2000 m / 9–VI–1976 / Paratype / Neovalgus / formosanus / Y. MIYAKE, 1985’. 1 ex. ‘Paratype: / Neovalgus / formosanus / Y. MIYAKE, 1985’.
Oreoderus quadrimaculatus Miyake, Yamaguchi, Aoki et Akiyama, 2002: 64−65.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘Phu Huai Sing / Mukdahan / E-THAILAND / 19. Feb. 1999 / S. Yamahuchi and T. Aoki leg. // Holotype: / Oreoderus / quadrumaculatus // Or. 4ml / Coll. // Holotype / Oreoderus quadrimaculatus / Y. MIYAKE, YAMAGUCHI / et AOKI 2002’. (Fig.
The aedeagus of the holotype is pinned separately. The right protarsus is missing.
Valid species.
Tarsovalgus hatai Miyake, 1993b: 32−33.
The holotype and following paratype are deposited in RIEB (ex coll. Y. Miyake):
(♂). ‘ [Malaya] / – –1992 // Holotype: Tarsovalgus / hatai Y. MIYAKE, 1993’. (Fig.
1 ex.: ‘ [Malaya] / – –1992 // Paratype: / Tarsovalgus / hatai / Y. MIYAKE, 1993’.
Valid species.
Trox kyotensis Ochi & Kawahara, 2000: 53−56.
The following paratype is deposited in RIEB (ex coll. T. Ochi):
1 ex.: ‘YAMAZAKI- / CHO, KYOTO. / 5–IV, 1999 / M. KAWAHARA // PARATYPE / Trox / kyotensis /
Trox (Niditrox) kyotensis Ochi et Kawahara, 2000, see
Trox sabulosus fujiokai Ochi, 2000: 43−44.
The following paratype is deposited in RIEB (ex coll. T. Ochi):
1 ex.: ‘Yukyuzan-Park, Nagaoka / Niigata Pref., Japan / 28. V. 1997 / Y and M. Kawahara leg // PARATYPE / Trox / Sabulosus / Subsp. fujiokai / OCHI, 2000’.
Valid species.
Trox setifer horiguchii Ochi & Kawahara, 2002: 54−55.
The following paratype is deposited in RIEB (ex coll. T. Ochi):
1 ex.: 1♂ ‘MEBORO-DAM / KAMIAGATA. T. / IS. TSUSHIMA / NAGASAKI. J. P / 25–V. 2002 / Y and M. KAWAHARA // 対県= tsu-ken上県町= kamiagata-chô / 目保呂ダム= meborodamu / 25. V. 2002 / Y and M. Kawahara // ♂ // PARATYPE / Trox / setifer / subsp. horiguchii /
Trox horiguchii Ochi and Kawahara, 2002, see
We are grateful to Toshiaki Aoki (RIEB), Shûhei Yamagushi (RIEB), Reiichiro Katsuyama (RIEB), Shûhei Nomura (NMNS), and Kônosuke Hoshino (NMSM) for their help in the type material survey. Our thanks go to Hao Xu (Mianyang Normal University) for providing the information about Miyake’s type material. We express our sincere gratitude to Carsten Zorn (Gnoien) for reviewing the manuscript.