Holotype : ♂ (CAUC), China, Guizhou: Dashahe qianfengcun, 1300–1550 m, 18.viii.2004, Yajun Zhu. Paratypes: 1♀(CAUC), China, Guizhou: as holotype; 1♂, 1♀(CAUC), China, Guizhou: Dashahe qianfengcun, 1300–1550 m, 19.viii.2004, Yajun Zhu; 1♀(CAUC), China, Guizhou: Dashahe qianfengcun, 1350–1500 m, 17.viii.2004, Yajun Zhu; 1♂, 1♀(CAUC), China, Guizhou: Dashahe hebacun, 1200–1500 m, 20.viii.2004, Yajun Zhu.

Figures 1–5. 

Minettia (Minettiella) dashahensis sp. nov. Male 1 head, anterior view 2 wing 3 habitus, lateral view 4 thorax, dorsal view 5 abdomen, dorsal view.

The specific epithet is named for holotype locality in Guizhou, Dashahe.

Arista blackish brown except paler basally. Mesonotum 1 + 3 dorsocentral setae, acrostichal setulae in four irregular rows, with two pairs of strong acrostichal setulae before prescutellar setae. Epandrium broad, ventral apical angle with setae. Surstylus separated from epandrium and extended from interior of epandrium, curved in lateral view, with connective sclerite under cercus in posterior view. Aedeagal apodeme long, nearly as long as aedeagus.

Male. Body length 3.4–3.5 mm, wing length 3.7–3.8 mm. Female. Body length 3.0–3.4 mm, wing length 3.5–3.8 mm.

Head (Fig. 1) black. Face and parafacial with dense grayish white pruinescence. Frons blackish brown, anterior margin yellow; ocellar triangle black, ocellar setae developed, almost as long as posterior fronto-orbital setae and extending to anterior margin of frons; anterior fronto-orbital setae reclinate, shorter than posterior fronto-orbital setae. Gena ca. 1/6 height of eye. Antenna brownish yellow, first flagellomere blackish brown on apical 2/3 of dorsal margin, 1.7 times longer than high; arista blackish brown except paler basally, plumose with longest setula slightly longer than height of first flagellomere. Proboscis blackish brown, with pale yellow and black setulae; palpus black with black setulae.

Thorax (Fig. 4) black, with gray pruinescence. 1 + 3 dorsocentral setae, acrostichal setulae in four irregular rows, with two pairs of strong acrostichal setulae before prescutellar setae; a pair of prescutellar setae, shorter than anteriormost dorsocentral setae. One anepisternal seta and one katepisternal seta. Legs black, tibia brownish yellow at base, mid and hind tarsomeres dark yellow (sometimes fore tarsomeres also dark yellow). Fore femur with eight posterior dorsal setae and five posterior ventral setae; fore tibia with one dorsal preapical seta and one short apical ventral seta. Mid femur with three or four anterior setae and one short apical posterior seta; mid tibia with one strong dorsal preapical seta and one strong apical ventral seta. Hind femur with one preapical anterior dorsal seta; hind tibia with one weak dorsal preapical seta and one short apical ventral seta. Wing slight yellow and hyaline, pale yellow at base; costa with 2^nd (between R₁ and R₂₊₃), 3^rd (between R₂₊₃ and R₄₊₅), and 4^th (between R₄₊₅ and M₁) sections in proportion of 9.5 : 2.8 : 2.1; r-m on middle of discal cell; ultimate and penultimate sections of M₁ in proportion of 6.8 : 3.9; ultimate section of CuA₁ ca. 1/5 of penultimate. Haltere pale yellow.

Abdomen (Fig. 5) black, with sparse gray pruinescence. Male genitalia (Figs 6–10): syntergosternite circular. Epandrium broad, ventral apical angle with setae. Surstylus separated from epandrium and extended from interior of epandrium, tapering apically, curved in lateral view, with connective sclerite under cercus in posterior view; hypandrium narrow in the middle and wide on both sides, with small inner process. Postgonite with setae, slender and curved apically in lateral view; aedeagus tubular, ventral sclerite V-shaped concave apically in ventral view, with a single long spiny process on the bottom, forming an acute angle with aedeagus, aedeagus with membranous processes internally; aedeagal apodeme as long as aedeagus.

The new species is very similar to M. (Minettiella) sasakawai from China (Hainan) and Vietnam in the following characteristics: acrostichal setulae in four rows; mid and hind tarsomeres dark yellow, fore femur with eight posterior dorsal setae and five posterior ventral setae; wing slight yellow and hyaline, pale yellow at base; hypandrium narrow in the middle and wide on both sides, but it can be separated from the latter in the mesonotum with 1+3 dorsocentral setae, two pairs of strong acrostichal setulae present in front of prescutellar setae; the syntergosternite being circular; epandrium is broad. In M. (Minettiella) sasakawai, the mesonotum with 0 + 2 dorsocentral setae, a pair of strong acrostichal setulae present in front of prescutellar setae; the syntergosternite is semicircular and narrow under the spiracle; epandrium is slender.

China (Guizhou).

Figures 6–10. 

Minettia (Minettiella) dashahensis sp. nov. Male 6 syntergosternite and epandrium, lateral view 7 epandrial complex, posterior view 8 syntergosternite, anterior view 9 aedeagal complex, ventral view 10 aedeagal complex, lateral view. Scale bar: 0.5 mm.

Holotype : ♂ (CAUC), China, Guizhou: Fanjing mountain, 1300–1900 m, 29.vii.2001, Fang Zhao. Paratypes: 1♂, 5♀♀ (CAUC), China, Guizhou: Fanjing mountain, 1750–2200 m, 1.viii.2001, Fang Zhao; 1♂ (CAUC), China, Guizhou: Fanjing mountain, 950–1750 m, 2.viii.2001, Fang Zhao; 4♀♀ (CAUC), China, Guizhou: Fanjing mountain, 1300–1900 m, 29.vii.2001, Caixia Gao; 1♀ (CAUC), China, Guizhou: Fanjing mountain, 1900–2490 m, 30.vii.2001, Caixia Gao; 3♀♀ (CAUC), China, Guizhou: Fanjing mountain, 1750–2000 m, 1.viii.2001, Caixia Gao; 1♂6♀♀(CAUC), China, Guizhou: Fanjing mountain, 950–1750 m, 2.viii.2001, Caixia Gao; 1♀ (CAUC), China, Guizhou: Fanjing mountain, 1300 m, 27.vii.2001, Wanzhi Cai.

Figures 11–15. 

Minettia (Minettiella) longispina sp. nov. Male 11 head, anterior view 12 wing 13 habitus, lateral view 14 thorax, dorsal view 15 abdomen, dorsal view.

Latin, longispina, referring to the aedeagus with a long membranous spin-like processes internally.

Frons dark brown, anterior margin yellow, two black longitudinal stripes along fronto-orbital rows, extending from ocellar triangle to occiput. Arista blackish brown except paler basally, pubescent with longest setulae almost as long as 1/4 height of first flagellomere. Mesonotum with 0 + 2 dorsocentral setae, acrostichal setulae in six irregular rows. Surstylus separated from epandrium, distorted in lateral view and only one short curved claviform process, narrow apically and inwards curved in posterior view. Aedeagal apodeme long, nearly as long as aedeagus.

Male. Body length 3.6–3.8 mm, wing length 4.0–4.2 mm. Female. Body length 3.8–4.0 mm, wing length 4.0–4.5 mm.

Head (Fig. 11) black. Face and parafacial with dense grayish white pruinescence. Frons dark brown, anterior margin yellow, as long as wide and parallel-sided, two black longitudinal stripes along fronto-orbital rows, extending from ocellar triangle to occiput; ocellar triangle black, ocellar setae developed, almost as long as posterior fronto-orbital setae and extending to anterior margin of frons; anterior fronto-orbital setae reclinate, shorter than posterior fronto-orbital setae. Gena ca. 1/6 height of eye. Antenna brownish yellow, first flagellomere blackish brown on 2/3 apex of dorsal margin, 1.7 times longer than high; arista blackish brown except paler basally, pubescent with longest setulae almost as long as 1/4 height of first flagellomere. Proboscis blackish brown, with pale yellow and black setulae; palpus black with black setulae.

Thorax (Fig. 14) black, with gray pruinescence. 0 + 2 dorsocentral setae, acrostichal setulae in six irregular rows, with pair of strong acrostichal setulae before prescutellar setae; a pair of prescutellar setae, almost as long as anteriormost dorsocentral setae. One anepisternal seta and one katepisternal seta. Legs black, tibia brownish yellow at base, mid and hind tarsomeres dark yellow (sometimes fore tarsomeres also dark yellow). Fore femur with eight posterior dorsal setae and five posterior ventral setae; fore tibia with one dorsal preapical seta and one short apical ventral seta. Mid femur with four anterior setae and one short apical posterior seta; mid tibia with one strong dorsal preapical seta and one strong apical ventral seta. Hind femur with one preapical anterior dorsal seta; hind tibia with one weak dorsal preapical seta and one short apical ventral seta. Wing slight yellow and hyaline, pale yellow at base; costa with 2^nd (between R₁ and R₂₊₃), 3^rd (between R₂₊₃ and R₄₊₅), and 4^th (between R₄₊₅ and M₁) sections in proportion of 7.1 : 2.0 : 1.5; r-m on middle of discal cell; ultimate and penultimate sections of M₁ in proportion of 4.9 : 2.8; ultimate section of CuA₁ ca. 1/6 of penultimate. Haltere pale yellow.

Abdomen (Figs 15) black, with sparse gray pruinescence. Male genitalia (Figs 16–20): syntergosternite circular, with one broad membranous ventral process. Epandrium narrow dorsally and broad ventrally, with one small ventral concave basally, ventral setae apically. Surstylus separated from epandrium, distorted in lateral view and only saw one short curved claviform process, narrow apically and inwards curved in posterior view, with connective broad sclerite under cercus at base; hypandrium narrow, U-shaped, with one distinct membranous inner process. Postgonite slender in ventral view, acute at tip, as long as half-length of aedeagus; aedeagus with long spiny processes internally, with distinct apical concave; aedeagal apodeme long, nearly as long as aedeagus.

The new species is very similar to Minettia (Minettiella) membranacea sp. nov. from Yunnan in the following characteristics: frons with two black longitudinal stripes along fronto-orbital rows, extending from ocellar triangle to occiput; mesonotum with 0 + 2 dorsocentral setae, acrostichal setulae in six irregular rows, and a pair of strong acrostichal setulae present in front of prescutellar setae; wing slight yellow and hyaline, pale yellow at base, but it can be separated from the latter in the fore femur with five posterior ventral setae, the mid femur with four anterior setae, the syntergosternite with one broad membranous process; the hypandrium narrow and U-shaped; the aedeagus with long spines ventrally. In M. (Minettiella) membranacea sp. nov., the fore femur with six posterior ventral setae, the mid femur with three anterior setae, the syntergosternite with one anterior median process in lateral view, one membranous band under ventral bridge; the hypandrium V-shaped; the aedeagus with membranous processes internally.

China (Guizhou).

Figures 16–20. 

Minettia (Minettiella) longispina sp. nov. Male 16 syntergosternite and epandrium, lateral view 17 epandrial complex, posterior view 18 syntergosternite, anterior view 19 aedeagal complex, ventral view 20 aedeagal complex, lateral view. Scale bar: 0.5 mm.

Holotype : ♂ (CAUC), China, Yunnan: Baoshan Baihualing hot spring, 1500 m, 29.v.2007, Xingyue Liu.

Latin, membranacea, referring to the aedeagus with membranous processes internally.

Figures 21–25. 

Minettia (Minettiella) membranacea sp. nov. Male 21 head, anterior view 22 wing 23 habitus, lateral view 24 thorax, dorsal view 25 abdomen, dorsal view.

Antenna brownish yellow, first flagellomere blackish brown on 2/3 apex of dorsal margin. Proboscis blackish brown, with pale yellow and black setulae. Mesonotum with two dorsocentral setae, acrostichal setulae in six irregular rows. Mid and hind tibia brownish yellow at base and tarsomeres dark yellow. Syntergosternite circular, broad dorsally and narrow ventrally, with one small anterior median process in lateral view, one membranous band under ventral bridge. Hypandrium median ventral side outward, V-shaped. Aedeagal apodeme long, slightly shorter than aedeagus.

Male. Body length 3.2 mm, wing length 4.3 mm. Female. Unknown.

Head (Fig. 21) black. Face and parafacial with dense grayish white pruinescence. Frons dark brown, as long as wide and parallel-sided, two wide black longitudinal stripes along fronto-orbital rows, extending from ocellar triangle to occiput; ocellar triangle black, ocellar setae developed, almost as long as posterior fronto-orbital setae and extending to anterior margin of frons; anterior fronto-orbital setae reclinate, shorter than posterior fronto-orbital setae. Gena ca. 1/7 height of eye. Antenna brownish yellow, first flagellomere blackish brown on 2/3 of the apex of dorsal margin, 1.7 times longer than high; arista blackish brown except paler basally, pubescent with longest setula almost as long as 1/3 height of first flagellomere. Proboscis blackish brown, with pale yellow and black setulae; palpus black with black setulae.

Thorax (Fig. 24) black, with brownish gray pruinescence. Two dorsocentral setae, acrostichal setulae in six irregular rows, with pair of strong acrostichal setulae before prescutellar setae; a pair of prescutellar setae, longer than anteriormost dorsocentral setae. One anepisternal seta and one katepisternal seta. Legs black, mid and hind tibia brownish yellow at base and tarsomeres dark yellow. Fore femur with eight posterior dorsal setae and six posterior ventral setae; fore tibia with one dorsal preapical seta and one short apical ventral seta. Mid femur with three anterior setae and one short apical posterior seta; mid tibia with one strong dorsal preapical seta and one strong apical ventral seta. Hind femur with one preapical anterior dorsal seta; hind tibia with one weak dorsal preapical seta and one short apical ventral seta. Wing slight yellow and hyaline, pale brown at base; costa with 2^nd (between R₁ and R₂₊₃), 3^rd (between R₂₊₃ and R₄₊₅), and 4^th (between R₄₊₅ and M₁) sections in proportion of 9.6 : 2.4 : 1.8; r-m before middle of discal cell; ultimate and penultimate sections of M₁ in proportion of 6.4 : 4.4; ultimate section of CuA₁ ca. 1/6 of penultimate. Haltere pale yellow.

Abdomen (Fig. 25) black, with sparse gray pruinescence. Male genitalia (Figs 26–30): syntergosternite circular, broad dorsally and narrow ventrally, with one small anterior median process in lateral view, one membranous band under ventral bridge. Epandrium slender. Surstylus separated from epandrium, with setae and curved in lateral view, with connective sclerite under cercus in posterior view; hypandrium median ventral side outward, V-shaped. Postgonite curved apically in ventral view, with long setae and nearly digitiform, aedeagus with membranous processes internally, with small apical concave; aedeagal apodeme long, slightly shorter than aedeagus.

The new species is very similar to Minettia (Minettiella) spinosa Shi & Yang from Hubei in the following characteristics: acrostichal setulae in six rows; the tarsomeres dark yellow, mid femur with three anterior setae and one short apical posterior seta; wing slight yellow and hyaline, but it can be separated from the latter in the face without spot; the mesonotum with pair of strong acrostichal setulae; the syntergosternite without setula near spiracle; the epandrium without subapical concavity; the hypandrium V-shaped. In M. (Minettiella) spinosa, the face with a yellow triangular median spot or only slightly yellow in center of face; the mesonotum without strong acrostichal setulae; the syntergosternite with one setula near spiracle; the epandrium with one small subapical concavity; the hypandrium H-shaped.

China (Yunnan).

Figures 26–30. 

Minettia (Minettiella) membranacea sp. nov. Male 26 syntergosternite and epandrium, lateral view 27 epandrial complex, posterior view 28 syntergosternite, anterior view 29 aedeagal complex, ventral view 30 aedeagal complex, lateral view. Scale bar: 0.5 mm.

Holotype : ♂(CAUC), China, Yunnan: Zhaotong Xiaocaoba, 1715 m, 15.ix.2009, Tingting Zhang. Paratypes: 1♀ (CAUC), China, Yunnan: as holotype; 1♂, 1♀ (CAUC), China, Yunnan: Zhaotong Xiaocaoba, 1715 m, 15.ix.2009, weina Cui; 1♀ (CAUC), China, Yunnan: Zhaotong Xiaocaoba, 1900 m, 15.ix.2009, weina Cui; 1♂, 1♀ (CAUC), China, Yunnan: Kunming Heilongtan, 2016 m, 23.vii.2006, Kuiyan Zhang; 1♀ (CAUC), China, Yunnan: Kunming Heilongtan, 2016 m, 23.vii.2006, Wenliang Li.

The specific epithet is named for the holotype locality in Yunnan, Zhaotong.

Face and parafacial with dense grayish white pruinescence. Anterior fronto-orbital setae reclinate, shorter than posterior fronto-orbital setae. Mesonotum with 0+2 dorsocentral setae, acrostichal setulae in six irregular rows. Mid and hind tarsomeres dark yellow (sometimes fore tarsomeres also dark yellow). Wing slight yellow and hyaline, brown at base. Dorsal of epandrium narrower than dorsal of syntergosternite. Surstylus separated from epandrium, claviform, with single long seta in the middle, expanded apicially in lateral view and cone-shaped in posterior view. Postgonite hairy. Aedeagal apodeme tubular, as long as half-length of aedeagus.

Figures 31–35. 

Minettia (Minettiella) zhaotongensis sp. nov. Male 31 head, anterior view 32 wing 33 habitus, lateral view 34 thorax, dorsal view 35 abdomen, dorsal view.

Male. Body length 3.9–4.2 mm, wing length 4.4–4.8 mm. Female. Body length 3.8–4.3 mm, wing length 4.3–4.9 mm.

Head (Fig. 31) black. Face and parafacial with dense grayish white pruinescence. Frons dark brown, as long as wide and parallel-sided, two black longitudinal stripes along fronto-orbital rows, extending from ocellar triangle to occiput; ocellar triangle black, ocellar setae developed, longer than anterior fronto-orbital setae and posterior fronto-orbital setae, extending to anterior margin of frons; anterior fronto-orbital setae reclinate, shorter than posterior fronto-orbital setae. Gena ca. 1/6 height of eye. Antenna brownish yellow, first flagellomere blackish brown on 2/3 apex of dorsal margin, 1.7 times longer than high; arista blackish brown except paler basally, pubescent with longest setulae almost as long as 1/4 height of first flagellomere. Proboscis blackish brown, with pale yellow and black setulae; palpus black with black setulae.

Thorax (Fig. 34) black, with gray pruinescence. 0+2 dorsocentral setae, acrostichal setulae in six irregular rows, with pair of strong acrostichal setulae before prescutellar setae; a pair of prescutellar setae, almost as long as anteriormost dorsocentral setae. One anepisternal seta and one katepisternal seta. Legs black, tibia brownish yellow at base (sometimes tibia dark yellow), mid and hind tarsomeres dark yellow (sometimes fore tarsomeres also dark yellow). Fore femur with eight posterior dorsal setae and five posterior ventral setae; fore tibia with one dorsal preapical seta and one short apical ventral seta. Mid femur with three or four anterior setae and one short apical posterior seta; mid tibia with one strong dorsal preapical seta and one strong apical ventral seta. Hind femur with one preapical anterior dorsal seta; hind tibia with one weak dorsal preapical seta and one short apical ventral seta. Wing slight yellow and hyaline, brown at base; costa with 2^nd (between R₁ and R₂₊₃), 3^rd (between R₂₊₃ and R₄₊₅), and 4^th (between R₄₊₅ and M₁) sections in proportion of 9.6 : 2.5 : 1.7; r-m beyond middle of discal cell; ultimate and penultimate sections of M₁ in proportion of 6.5 : 3.7; ultimate section of CuA₁ ca. 1/5 of penultimate. Haltere pale yellow.

Abdomen (Fig. 35) black, with sparse gray pruinescence. Male genitalia (Figs 36–40): syntergosternite circular, broad dorsally and narrow ventrally. Dorsal of epandrium narrower than dorsal of syntergosternite. Surstylus separated from epandrium, claviform, with a long seta in the middle, expanded apically in lateral view and cone-shaped in posterior view, with connective weakly sclerotized claviform sclerite; hypandrium wide on both sides and concave in the middle, U-shaped. Postgonite hairy. Aedeagus with pair of ventral sclerites on basal 1/3 in ventral view, with membranous spiny processes apically, with small apical concave; aedeagal apodeme tubular, as long as half length of aedeagus.

The new species is very similar to Minettia (Minettiella) tianmushanensis Shi & Yang from Hainan in the following characteristics: mesonotum with 0 + 2 dorsocentral setae, a pair of strong acrostichal setulae present in front of prescutellar setae; the mid femur with three or four anterior setae; the surstylus is claviform, cone-shaped in posterior view, but it can be separated from the latter in the frons with two black longitudinal stripes along fronto-orbital rows; acrostichal setulae in six irregular rows; aedeagal apodeme as long as half length of aedeagus. In M. (Minettiella) tianmushanensis, the frons without black longitudinal stripe; acrostichal setulae in four irregular rows; aedeagal apodeme as long as aedeagus.

The new species is very similar to Minettia (Minettiella) membranacea sp. nov. from Yunnan in the following characteristics: frons with two black longitudinal stripes along fronto-orbital rows, extending from ocellar triangle to occiput; mesonotum with 0 + 2 dorsocentral setae, acrostichal setulae in six irregular rows, and a pair of strong acrostichal setulae present in front of prescutellar setae; wing slight yellow and hyaline, pale yellow at base, but it can be separated from the latter in the fore femur with five posterior ventral setae, the mid femur with four anterior setae, the syntergosternite with one broad membranous process; the hypandrium narrow and U-shaped; the aedeagus with long spines ventrally. In M. (Minettiella) membranacea sp. nov., the fore femur with six posterior ventral setae, the mid femur with three anterior setae, the syntergosternite with one anterior median process in lateral view, one membranous band under ventral bridge; the hypandrium V-shaped; the aedeagus with membranous processes internally.

China (Yunnan).

Figures 36–40. 

Minettia (Minettiella) zhaotongensis sp. nov. Male 36 syntergosternite and epandrium, lateral view 37 epandrial complex, posterior view 38 syntergosternite, anterior view 39 aedeagal complex, ventral view 40 aedeagal complex, lateral view. Scale bar: 0.5 mm.