(modified from Borkent and Wheeler 2013). Considerably large specimens, body elongate, mostly yellowish and/or brown. Two ocelli, in contact with eye margins. Laterotergite and mediotergite setose. Wing macrotrichia decumbent, some species without microtrichia. C clearly produced beyond apex of R₅, sc-r reaching R₁ almost always beyond origin of Rs, R₄ present, forming an elongated rectangular cell, M₁₊₂ short, barely twice length of r-m, M₁₊₂ forking slightly more distally than origin of M₄.

Holotype. 1♂, Colombia, Department of Cundinamarca, Chingaza National Natural Park (PNN), Alto de la Bandera locality; 4°34.351'N, 73°42.752'W; alt. 3660 m a.s.l.; forest; Malaise trap; L. Cifuentes leg.; (IAvH 2600, wing in Euparal on slide mounting, rest of the body in 96% ethanol, genitalia preserved in glycerine in microvial). Paratype. 1♂, Colombia, same data as holotype. (CEUA 11339, in alcohol).

General color yellow to light brown. Anepisternum bare. All coxae and hind femur with no dark markings. Wing darker along anterior margin, but without conspicuous maculae; sc-r beyond origin of Rs. Terminalia yellowish, wider than long. Lateral extension of gonocoxite long, slightly curved inwards, with an apical long dark spine. Gonostylus small and rounded, apical surface with scattered short spines. Parameres digitiform, apically bifurcated, with short dark spines.

Figure 2. 

A Habitus of Eudicrana silvaandina sp. nov., male (holotype) B Habitus of E. chingaza sp. nov., male (holotype), terminalia detached C Habitus of E. maculata sp. nov., male (holotype) D Male habitus of E. merizaldei sp. nov. (holotype) E Habitus of E. merizaldei sp. nov., female. Scale bar: 1mm

Male (Fig. 2A). Body length, 8.0–8.5 mm. Head (Fig. 3A). Width, 0.56 mm, height, 0.35 mm. Vertex and occiput yellowish, darkened around the ocelli, with abundant brownish-yellow short setae. Mid ocellus absent, lateral ocelli nearly touching eye margins. Eyes setose. Four long dark setae on occiput behind eye margin. Scape and pedicel yellow, cylindrical, scape slightly longer than pedicel, both with small brownish-yellow setae; 14 flagellomeres, mostly light brown; first flagellomere almost twice as long as second. Frons yellowish, setose; face yellowish, circular and setose; clypeus yellowish-brown, slightly elongate, sub-triangular, with abundant yellowish setae; palpus with palpifer plus four palpomeres, first parlpomere as long as second, light brown, distal flagellomeres gradually lighter, distal palpomere more than three times as long as penultimate. Labella well developed, cream-yellow. Thorax (Figs 3E, I). Scutum mostly yellowish, with a pair of elongated brownish stripes and a brownish line over acrostichals. Dorso-centrals slightly stronger than scattered setae over scutum, acrostichals undifferentiated from scutum setae, a number of stronger and longer black setae along lateral margins. Scutellum, yellowish, with scattered smaller setae over disc and two pairs of marginal setae slightly longer than remaining scutellar setae. Pleural sclerites yellowish brown, membrane pale-yellow. Antepronotum with four long darker setae, proespisternum with two stronger setae and some smaller ones. Proepimeron, anepisternum, katepisternum, mesepimeron, and metepisternum bare, laterotergite with 7–8 short setae on anterior half and 9–10 longer, darker setae on posterior half; mediotergite with longer dark setae along entire surface, dorsomedial setae shorter. Halter pedicel and knob yellowish, setose. Legs. Very elongate, yellowish brown, darker toward tip of femora and tibiae. Fore tibia with distal ventral oval depression with abundant and irregularly distributed trichia; first tarsomere 1.5 times tibia length. Mid tibia with short dark trichia irregularly arranged, with an apical ventral dark comb of setae, tarsi with dark, short, erect setae along entire length. Hind tibia with trichia as in mid tibia, but apical comb absent. Tibial spurs 1:2:2, light brown, spurs more than three times apical width of tibiae. Tarsal claws with a large apical tooth and a smaller basal tooth. Wing (Fig. 4A). Length 5.0–5.5 mm, width 2.0 mm. Membrane mostly hyaline, no defined maculae, but darkened along anterior margin, densely covered with decumbent macrotrichia on all cells and scarce microtrichia on anal lobe; wing veins light brown, anterior veins more strongly sclerotized. Sc complete, setose, reaching C slightly beyond level of R₄; sc-r present, bare, slightly more basal than mid of cell r1; first sector of Rs slightly oblique, R₁ long reaching C at about apical fifth of wing; C extending to slightly beyond tip of R₅, R₄ present, cell r1 rectangular, elongate, 8 times as long as wide; R₅ slightly curved posteriorly at apex; r-m setose, oblique. Medial and cubital veins complete, basally reaching wing margin, though not hardly sclerotized distally. M₁₊₂ stem almost twice the r-m length, M₁ slightly divergent from M₂ close to apex. Origin of M₄ more basal than level of medial fork; CuA curved towards posterior margin on apical third; pseudovein sclerotized to about third of CuA; CuP sclerotized to about mid of CuA. Abdomen. (Fig. 2A). Cylindrical with dark setae covering tergites and sternites; segments 1–6 yellowish brown, darker on distal half, 7–8 brownish. Terminalia (Fig. 5A, B). Light brown, wider than long. Gonocoxites almost fusing to each other mesally at ventral face, with a deep, slender incision between them; each gonocoxite with three rounded ventral lobes over the distal margin, inner surface covered with a set of homogeneous combs of dark setae. Dorsal lobe of gonocoxite, slightly extended inwards, inner surface covered with a set of combs of elongated setae and a long strong dark subapical spine; dorso-lateral projection of gonocoxite long, extending to about half the cercus length, slightly curved inwards, bearing a long dark spine at tip. Gonocoxal bridge strongly displaced towards base of terminalia. Gonostylus, apically round, surface with scattered dark short spines. Paramere wide, bifid with a pair of digitiform distal projections with short dark spines. Aedeagus elongate, weakly sclerotized. Cerci digitiform, very long, slender and setose.

Figure 3. 

Head, frontal view A Eudicrana silvaandina sp. nov. (holotype) B E. chingaza sp. nov. (holotype) C E. maculata sp. nov., (holotype) D E. merizaldei sp. nov. (holotype). Thorax, lateral view E Eudicrana silvaandina sp. nov. (holotype) F E. chingaza sp. nov. (holotype) G E. maculata sp. nov. (holotype) H E. merizaldei sp. nov. (holotype). Thorax, dorsal view I Eudicrana silvaandina sp. nov. (holotype) J E. chingaza sp. nov. (holotype) K E. maculata sp. nov. (holotype) L E. merizaldei sp. nov. (holotype). Scale bar: 0.25 mm.

Female. Unknown.

The specific epithet of this species combines the Latin word silva (nominative, noun, feminine) for “forest”, with andina (nominative, adjective feminine) for “Andean”, referring the presence of this species in the South American Andean ecosystem.

This species is easily discriminated from the other species described here by the faint darkened wing membrane along the entire anterior margin, the elongated, thin cerci, the long dorso-lateral projection of gonocoxite extending well beyond the gonostyle, bearing a distal spine, and the parameres with a pair of distal spinose projections.

Holotype. 1♂, Colombia, Department of Cundinamarca, Chingaza National Natural Park (PNN), Alto de la Bandera locality; 04°34.351'N, 73°42.752'W; alt. 3660 m a.l.s.; 15 Nov.–01 Dec. 2001; forest; Malaise trap; L. Cifuentes leg. (IAvH 2600, wing in Euparal on slide mounting, rest of the body in 96% ethanol, genitalia preserved in glycerine microvial).

Body light brown. Anepisternum bare. Coxae and hind femur without maculae. Wing membrane translucent; sc-r beyond of origin of Rs. Terminalia yellowish, wider than long. Dorso lateral-distal extension of gonocoxite short, with a distinctive apical spine. Gonostylus wide and lunular, inner surface with abundant spines. Cercus elongate, but wide on basal half. Paramere not bifid, elongate, with short spines.

Male (Fig. 2B). Body length, 7.0 mm. Head (Fig. 3B). Width, 0.60 mm, height, 0.37 mm. Vertex and occiput light brown, with abundant brownish-yellow short setae. Mid ocellus absent, lateral ocelli surrounded by dark brown, almost touching eye margin. Eyes setose. Four long dark setae on occiput behind eye margin. Scape and pedicel yellowish brown, cylindrical, scape slightly longer than pedicel, both with small brownish setae; 14 flagellomeres, first flagellomere almost twice as long as second. Frons yellowish brown, setose, face yellowish brown, elongate, setose; clypeus yellow, quite elongate, sub-triangular, with abundant brownish setae; palpus with palpifer plus four palpomeres, light brown, first palpomere as long as second, last palpomere more than three times as long as penultimate. Thorax (Fig. 3F, J). Scutum light brown, with a narrow brown stripe along acrostichal line and a pair of weak light brown slender band over dorsocentral lines; dorsocentrals present, slightly longer than other scattered setae on scutum, acrostichals not differentiated, a number of stronger and longer black setae along lateral margins. Scutellum yellowish brown, with scattered setae along distal margin. Pleural sclerites yellowish brown, ventral half of katepisternum slightly darker. Pleural membrane pale-yellow. Antepronotum with four strong, darker setae; proespisternum with a pair of darker setae. Proepimeron, anepisternum, katepisternum, mesepimeron, and metepisternum bare; laterotergite with shorter setae on anterior half and 9–10 long setae on posterior half; mediotergite with lateral longer dark setae along entire surface and dorsomedial setae shorter. Halter pedicel yellow, knob light brown, setose. Legs. Coxae brownish-yellow, femora, tibia, and tarsi light brown, darkened towards apex [femur, tibia and tarsus of front leg missing in the holotype]. Mid tibia with short setation irregularly arranged, with a distal comb ventrally and some dark, slightly longer setae laterally and ventrally [tarsi missing]; hind tibia trichia distributed as on mid tibia, with dark slightly longer setae laterally and dorsally, without apical ventral comb. Tibial spurs 1:2:2, light brown, hind spurs more than three times apical tibial width. Tarsal claws with large apical tooth, smaller basal tooth. Wing (Fig. 4B). Length 5.0 mm, width 2.0 mm. Membrane very light brown, no maculae, densely covered with decumbent macrotrichia on nearly all wing cells, and scarce microtrichia on anal lobe; veins brown. Sc complete, setose, reaching C well slightly beyond level of R₄; sc-r present, bare, just basal to mid of cell r1; first sector of Rs slightly oblique, R₁ long, reaching C at apical fifth of wing, C extending slightly beyond apex of R₅, R₄ present; cell r1 elongate, rectangular, setose, length about 8 times the width; R₅ gently curved at apex towards posterior margin. Medial and cubital veins complete basally, slightly less sclerotized close to margin. M₁₊₂ about twice r-m length, M₁ almost parallel to M₂ distally. Origin of M₄ more basal than level of medial fork. CuA curved towards wing margin at distal third; pseudovein sclerotized to distal third of CuA; CuP sclerotized to about mid of CuA. Abdomen (Fig. 2C) Segments light yellowish brown, distal two thirds darker, cylindrical, brownish setae covering tergites and sternites. Terminalia (Fig. 6A–D). Light brown, wider than longer. Gonocoxite almost fusing basally at ventral face, with a deep, slender incision between them; each gonocoxite with one distal ventral lobe at the distal margin, bearing a distal comb of straight dark setae. Dorsal lobe of each gonocoxite, slightly truncated at the apex, bearing at the inner surface scattered short spines and several set of combs of dark setae; dorso-lateral projection of gonocoxite short with a dark distal dark spine. Gonocoxal apodeme at about mid of terminalia. Gonostylus rounded and wider (lunular shape), inner surface bearing several and scattered short spines. Paramere digitiform, apical surface covered with short dark spines. Aedeagus elongate, weakly sclerotized and visible mesally. Cerci setose, digitiform and wider at basal half.

Figure 4. 

A Wing of Eudicrana silvaandina sp. nov. (holotype) B wing of E. chingaza sp. nov. (holotype) C wing of E. maculata sp. nov. (holotype) D, E wing of E. merizaldei sp. nov. D male (holotype) E female (paratype). Scale bar:1 mm.

Figure 5. 

Male terminalia of Eudicrana silvaandina sp. nov. (holotype), ventral view A photograph B drawing.

Female. Unknown.

The specific epithet of this species E. chingaza (Nominative, adjective feminine) refers to Natural National Park Chingaza, where the holotype was collected. This name comes from the muisca indigenous language of Colombia, meaning “mountain range of the gods of the night”. It is one of the largest paramo ecosystems of Colombia and is the type locality of E. chingaza.

This species can be clearly separated from E. silvaandina by the short latero-distal projection of the gonocoxite in E. chingaza, by the wider cercus, and the nearly translucent wing membrane, without a darkened anterior margin of the wing. E. maculata also has a short latero-distal projection of the gonocoxite, but has a clear maculation in the wing and sc-r is placed very close to the origin of Rs.

Figure 6. 

Male terminalia of Eudicrana chingaza sp. nov. (holotype) A, B ventral view C, D dorsal view A, C photograph B, D drawing.

Holotype. 1♂, Colombia, Department of Antioquia, San José de la Montaña municipality, El Congo rural settlement, paramo El Congo locality; 6°46.5652'N, 75°43.5702'W, alt. 3150 m a.s.l.; 25 Mar.–5 Apr. 2017; Proyecto Moscas de las flores. C .H–Sepúlveda and M. Wolff legs.; dwarf forests, sweeping net (CEUA 106590, wing in Euparal on slide mounting, rest of the body in 96% ethanol, genitalia preserved in glycerine microvial). Paratypes. 1♂, same data as holotype (CEUA 106591, in alcohol); 1♂, Colombia Department of Antioquia, Sonsón municipality, paramo of Sonsón locality; 5°4.1292'N, 75°14.8092'W; alt. 3000 m a.s.l.; 10–15 Sept. 2011; forest; Malaise trap; L. Rios leg.; (CEUA 113393, in alcohol).

Scutum with a wide dark brown band. Anepisternum with a set of short dorsal setae. Coxae darkened distally, hind femur with a blackish brown marking on basal third. Wing with a dark brown macula along margin at region of sc-r, base of Rs and R₄, r-m, bM, M₁₊₂ and base of medial fork and over entire apical third. Abdomen blackish brown except in the basal third of each segment. Terminalia as long as wide, with short dorso-lateral projection of gonocoxite bearing a strong distal spine. Cerci very long, wide basally and setose. Paramere bifid, apically rounded and wide, with long dark spines on the margin.

Male (Fig. 2C). Body length, 7.5–8.0mm. Head (Fig. 3C). Width, 0.55–0.57 mm, height, 0.33–0.35 mm. Vertex brown, with abundant short brownish setae. Mid ocellus absent, lateral ocelli almost touching eye margin and surrounding by dark brown. Eyes setose. Occiput with three long dark setae behind eyes. Scape slightly longer than pedicel, ochre-yellow cylindrical, both with abundant dark setulae on distal half; 14 flagellomeres dark brown, first flagellomere with basal fourth whitish yellow, almost twice as long as second. Frons light brown, setose; face yellowish brown, semi-circular, setose; clypeus yellowish brown, elongate, with shorter setae on dorsal half and a set of dark longer setae on ventral half; palpus with palpifer plus four palpomeres, brown, first palpomere about as long as second, last palpomere light brown, more than three times as long as penultimate. Thorax (Fig. 3G, K). Scutum with yellowish area mesally, delimited by weak marks at anterior and lateral margins, with a pair of light brown lateral bands, blackish brown markings spread over scutum; scutum covered with scattered small setae, dorsocentrals differentiated, slightly longer than remaining setae, acrostichals not differentiated, a number of stronger and longer black setae along lateral margins. Scutellum yellowish-brown on disc, with brown distal margin, scattered homogeneous short setae on distal margin, with some longer setae. Pleural sclerites mostly ochre yellowish, with dark brown irregular spots. Pleural membrane pale yellow. Antepronotum with six longer setae, proespisternum with shorter irregularly dark setae. Anepisternum with a dorsal set of short setulae. Proepimeron, katepisternum, mesepimeron, and metepisternum bare, laterotergite with shorter setae on anterior half and 10–12 long setae on posterior half; mediotergite with long lateral setae and a number of smaller dorso-medial setae. Halter pedicel and knob light brown, setose. Legs. Fore coxae and mid femora cream-yellow, coxae darker at tips; all tibiae and tarsi dark brown, hind femur blackish-brown on basal third. Fore tibia with distal oval depression ventrally with abundant, irregularly distributed trichia, first tarsomere 1.5 times tibia length. Mid tibia with short irregularly arranged setation and some dark setae, a ventral comb apically, tarsi with dark, short erect setae along entire length. Hind tibia with blackish short setae laterally and dorsally along tibia length, ventral comb absent. Tibial spurs 1:2:2, dark brown, spurs more than three times tibia width at apex. Tarsal claws with large apical tooth, smaller basal tooth. Wing (Fig. 4C). Length 5.5–6.0 mm, width 2.0 mm. Membrane light brown, densely covered with decumbent macrotrichia, microtrichia present; a slightly darker macula along margin at region of sc-r, base of Rs and R₄, r-m, bM, M₁₊₂ and base of medial fork and over entire apical third. Sc complete, setose, reaching C well beyond base of Rs, near to level of R₄; sc-r present, bare, just slightly beyond level of origin of Rs; first sector of Rs only slightly oblique, setose; R₁ long, reaching C at apical fifth of wing; C extending shortly over apex of R₅; R₄ present, cell r4 elongate, setose, about 5.7 times as long as wide; R₅ slightly curved towards posterior margin at apex; r-m setose, oblique, shorter than M₁₊₂. Medial and cubital veins complete basally. M₁₊₂ twice length of r–m; origin of M₄ more basal than medial fork; CuA gently curved towards base at apical third; pseudovein sclerotized to distal third of CuA; CuP sclerotized to about basal third of CuA. Abdomen (Fig. 2B). Abdominal segments 2-6 elongate, cream yellow on basal fifth, blackish brown on distal four fifth, dark setae covering tergites and sternites. Terminalia (Fig. 7A–D). Dark brown, wider than long. Gonocoxites almost fusing at ventral face, with a thin incision between them; dorsal margin covered with several combs of elongated setae; inner surface of the ventral lobe of gonocoxite bearing scattered short spines and several combs of setae. Dorsally without a pronounced lobe, over the inner surface covered with short and scattered spines; dorsal latero-distal extensions short, with a strong external subdistal spine. Gonostylus short and rounded, apical surface with a set of dark combs. Paramere with two distal projection bearing dark spines distally. Aedeagus triangular distally. Cercus digitiform, long, extending well beyond distal margins of gonocoxite, slender, slightly wider at base, setose.

Figure 7. 

Male terminalia of Eudicrana maculata sp. nov. (holotype) A, B ventral view C, D dorsal view A, C photograph B, D drawing.

Female. Unknown.

The specific epithet maculata (nominative, adjective feminine) of this species refers to the evident dark brown maculae on the wing membrane.

This species is present in the transitional “dwarf forests” (Fig. 1B), between the high Andean forest and the paramos. This kind of environment is mainly dominated by the Melastomataceae species Tibuchina grossa (L.f) Cogn., the Cunoniaceae Wienmannia tomentosa L.f., the Clusiaceae Clusia multiflora Kunth, and the Winteraceae Drimys granadensis L.f.. This habitat is also rich in bryophyte mosses and the soil has a large layer of leaf litter. It is a very humid environment, usually with low temperature and low luminosity due to clouds.

Holotype. 1♂, Colombia, Department of Antioquia, Envigado municipality, private property Nubarrones; 6°8'45.626"N, 75°33'22.53"W; alt. 2200 m a.s.l.; 29 Jan.–11 Feb. 2017; M. Wolff and C. H-Sepúlveda; forest; Malaise trap (CEUA 108470, wing mounted in Euparal on microscope slide, rest of body in alcohol 96% ethanol). Paratypes. 2♂, 2♀, same location data as holotype, but differs on 26 Feb.–12 Mar. 2017 (CEUA 107038, in alcohol); 1♂, same location data as holotype, but differs on 23 Jun.–2 Jul. 2018 (CEUA 107040, in alcohol).

Body yellow to light brown. Anepisternum with a set of dorsal short setulae. Mid coxae and hind femur with dark markings. Wing with conspicuous maculae; sc-r reaching C beyond M₁₊₂ fork, at apical third of wing. Terminalia yellowish, with dorsal margin dark brown, lateral rounded apical extension with apical long dark spine.

Male (Fig. 2D). Body length, 7.5–8.5 mm. Head (Fig. 3D). Width, 0.54–0.56 mm, height, 0.30–0.35 mm. Vertex and occiput ochre-yellowish, light brown around ocelli, with abundant brownish-yellow short setae. Mid ocellus absent, lateral ocelli nearly touching eye margin. Eyes setose. Four long dark setae on occiput behind eye. Scape and pedicel ochre-yellowish, cylindrical, scape slightly longer than pedicel, both with small brownish setae; 14 flagellomeres, mostly light brown; first flagellomere almost twice as long as second. Frons and face ochre-yellowish, setose; clypeus yellowish-brown, slightly elongate, sub-triangular, with abundant light brown setae and some few darker setae; palpus with palpifer plus four palpomeres, first parlpomere as long as second, light brown, distal flagellomeres gradually lighter, distal palpomere more than three times as long as penultimate. Labella well-developed, cream-yellow. Thorax (Fig. 3H, L). Scutum with yellowish area between dorsocentrals, lateral margins, with a pair of light brown lateral bands; acrostichals indistinguishable from other setae, a number of stronger dark setae along lateral margins. Scutellum ochre-yellowish with light brown band along distal margin, scattered small setae over disc and two pairs of slightly longer marginal setae. Pleural sclerites ochre-yellowish, membrane cream-yellow. Antepronotum with three long dark setae, proespisternum with one stronger seta and some smaller ones. Anepisternum with a dorsal set of short setae, proepimeron, katepisternum, mesepimeron, and metepisternum bare, laterotergite with short 9–10 shorter setae on anterior half and 9–12 longer, darker setae on posterior half; mediotergite with longer dark setae along entire height, dorsomedial setae shorter. Halter pedicel light brown, knob yellowish, setose. Legs (Fig. 2D–E). Legs very elongate, ochre-yellowish, darker toward tip of femora and tibiae, base of hind femur darker. Fore tibia with distal ventral oval depression with abundant and irregularly distributed trichia; first tarsomere 1.5 times tibia length. Mid tibia with short dark trichia irregularly arranged and an apical ventral brown comb of setae, mid tarsus with dark, short, erect setae along entire length; hind tibia with trichia as in mid tibia, apex with row of dark setae. Tibial spurs 1:2:2, light brown, spurs more than three times apical width of tibiae. Tarsal claws with a large apical tooth and a smaller basal tooth. Wing (Fig. 4D). Length, 5.5–6.5 mm, width, 2.0 mm. Membrane hyaline with a pair of evident maculae, one more basally from anterior margin to the level of sc-r to base of medial fork, another one at distal third of wing; membrane densely covered with decumbent macrotrichia on all cells, microtrichia present; wing veins dark brown, anterior veins more strongly sclerotized. Sc complete, setose, reaching C slightly beyond level of R₄; sc-r present, bare, slightly more basal than mid of cell r1; first sector of Rs slightly oblique, R₁ long, reaching C at about apical fifth of wing; C extending to slightly beyond tip of R₅; R₄ present, cell rectangular, elongate, length almost 4 times width; R₅ slightly curved close to apex; r-m oblique, setose. Medial and cubital veins complete basally, reaching wing margin, though slightly less sclerotized close to apex. M₁₊₂ stem almost twice r-m length, M₁ slightly divergent from M₂ close to apex. Origin of M₄ more basal than level of medial fork, CuA curved towards posterior margin on apical third; pseudovein sclerotized to about mid of CuA; CuP sclerotized to less than half of CuA extension. Abdomen (Fig. 2D). Segments 1–7 ochre-yellowish on anterior third, brown on distal two thirds, segment 8 brownish; segments cylindrical, with dark setae covering tergites and sternites. Terminalia (Fig. 8A–D). Dark ochre-yellowish, slightly longer than wide. Gonocoxites close to each other mesally at ventral face, with a deep incision between them; each gonocoxite ventrally with one a wide short lobe on distal margin, inner surface with abundant scattered spines and set combs of setulae; dorso-lateral projection of gonocoxite slightly rounded, on the inner surface with several short dark spines and sets of combs of setulae, also at the distal margin a long strong dark spine. Gonostylus short, not conspicuous, close to apex of ventral lobe, rounded, inner surface with scattered dark short spines. Gonocoxal bridge strongly displaced towards base of terminalia. Paramere with two distal lobes, one wide apically with scattered spines and one slender, with long dark setae. Aedeagus elongate, weakly sclerotized. Cerci digitiform, densely covered with thin setae, extending well beyond lateral tip of gonocoxite.

Figure 8. 

Male terminalia of Eudicrana merizaldei sp. nov. (holotype) A, B ventral view C, D dorsal view A, C photograph B, D drawing.

Female. (Fig. 2E). As a male, except for the following features. Body length, 7.5–8.0 mm. Wing (Fig. 4E). Length, 6.5–7 mm, width, 2.3–2.5 mm. Membrane maculae at same position but darker than in males. Terminalia (Fig. 9A–D). Short, ochre-yellow with brown apex of sternite 8. Sternite 8 as long as wide, gonapophyses well separated from each other medially at distal margin, each gonaphysis setose, with a long, stronger seta at distal margin close to tip, long setae at distal margin medially and three thin, long setae in between. Tergite 8 covered with small setae on distal three fourth, a row of stronger, longer setae along distal margin. Vaginal furca (sternite 9) long, slender, extending anteriorly beyond anterior margin of sternite 8, rounded at anterior end. Tergite 9+10 much wider than long, covered with setulae, but no longer setae. First segment of cercus about as long as, but wider than, second segment of cercus, both setose.

Figure 9. 

Female terminalia of Eudicrana merizaldei sp. nov. (paratype) A, B lateral view C, D ventral view A, C photograph B, D drawing.

The species epithet of E. merizaldei is dedicated to biologist Gabriel Merizalde, friend and nature lover, who for many years made possible our sampling in the wonderful forest of the private property called Nubarrones, in the Loma del Escobero neighborhood, to the south of the city of Medellín.

This species is very similar to E. maculata, e.g., in the color patterns on the body, likewise the setose anepisternum, and the wing membrane dark markings, although they differ in other features. The information on date of collection of the specimens of the type series suggest that this species may have its phenology related to the end of periods of great rainfall, from February to March and from June to early July. This species was found in an Andean forest dominated mainly by Clusia multiflora Kunth, Magolia spinalii (Lozano) Govaerts (Magnoliaceae), and Ceroxyilum vogelianum (Engel) H.Wendl. (Arecaceae).