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New data on the hoverflies of Morocco (Diptera, Syrphidae) with faunistic and bibliographical inventories
expand article infoSouad Sahib, Ouafaa Driauach, Boutaïna Belqat
‡ University Abdelmalek Essaâdi, Tétouan, Morocco
Open Access

Abstract

All published records of 148 species of hoverflies from Morocco are reviewed and appropriate literature references, new locality records, and relevant comments are provided for each species. The list is supplemented with records from new field surveys. Two species, Eumerus obliquus (Fabricius, 1805) and Orthonevra brevicornis Loew, 1843 are recorded for the first time in Morocco. The new checklist comprises 150 nominal species from three subfamilies, 14 tribes, and 49 genera.

Keywords

Checklist, Diptera, hoverflies, Morocco, new records, North Africa, Palaearctic region, Syrphidae

Introduction

Syrphidae (Diptera: Brachycera), commonly named hoverflies, constitute, among the order Diptera, a taxonomically well-characterised family. Most adults feed on pollen and nectar and thus are potentially important pollinators and their conservation is essential to sustain the productivity of natural and agricultural landscapes (Moquet et al. 2018). The larvae of Syrphidae have various food habits; certain species are predators while others are phytophagous, mycophagous or saprophytophagous (Rotheray and Gilbert 1999, Rotheray 1993, Büchs 2003).

Hoverflies occur in all geographical regions except for Antarctica and some isolated islands. They are found from sea level to the highest altitudes in flower-rich habitats. The family contains more than 6,100 species Worldwide (Pape et al. 2011). Of the North African countries bordering the Mediterranean Sea, Morocco is the one best known faunistically. Since Becker and Stein (1913) published an initial list of 42 species, several authors have contributed to the knowledge of Moroccan Syrphidae (Gil Collado 1929, Séguy 1930, 1934, 1941, 1949, 1953, Kanervo 1939, Claussen 1989, Claussen and Hauser 1990, Hurkmans 1993, Kassebeer 1995, 1998, 1999, 2000, Părvu et al. 2006, Popović et al. 2015, Acanski et al. 2016, van Steenis et al. 2017, Vujić et al. 2018, Ebejer et al. 2019).

Morocco has the highest number of total Syrphidae species recorded in the North Africa Mediterranean area so far (N = 150). Ninety-one species are reported from Algeria (Peck 1988, Dirickx 1994, Haffaressas et al. 2017), while 69, 51 and 34 species are recorded from Tunisia (Peck 1988, Claussen and Hauser 1990, Dirickx 1994), Egypt (Peck 1988, Dirickx 1994, El-Hawagry and Gilbert 2019) and Libya (Peck 1988, Dirickx 1994), respectively. Below, we provide an annotated checklist of all syrphid species recorded from Morocco.

Altogether 150 species are recorded in this work, two species are newly recorded for Morocco, and 24 others are new records for at least one of the Moroccan regions: Rif (10), Eastern region (4), High Atlas (5), Middle Atlas (5), and Anti Atlas (6). Morocco has 12 endemic species.

Materials and methods

Syrphidae were collected from 79 localities across the main regions in Morocco, mostly in the Rif Mountains, including the Eastern region, the High Atlas, Middle Atlas and Anti Atlas. Additional material and databases indicated in the text are provided by other researchers.

Collecting was mainly carried out by sweep netting and Malaise trapping adults and/or rearing larvae and puparia in the laboratory from collected substrates in aquatic ecosystems following the technique used by Afzan and Belqat (2016). The majority of the Moroccan hoverflies were identified from external characteristics, except for some species for which male genital morphology was used to confirm their identity. Identifications were done with specific keys (Verlinden 1991, Kassebeer 1995a, Hauser and Kassebeer 1998, Kassebeer 1998b, Kassebeer 1999b, van Veen 2004, Marcos-García et al. 2007, Speight and Sarthou 2013). Fourty-six of the species were identified with the help of Bastiaan Wakkie (Brussels, Belgium).

A total of 690 Syrphidae specimens was collected from 79 Moroccan sites (Table 1, Fig. 1). Maps with collecting localities were created with ArcGIS (Geographic Information System). Literature references on the occurrence in Morocco are given for each species, together with the region of the country from which the species is known. Additional information is given in the section ’Comments’. The Distribution as it is given for individual species is based on Speight (2018) and Mengual et al. (2020), as the most up-to-date reference, although other published works were consulted for specific taxa in order to obtain a more accurate distribution.

Figure 1. 

Map of Morocco indicating the collecting sites for hoverflies in this study.

Table 1.

Sampling sites (in alphabetical order) with locality names, altitudes (meters), and geographical coordinates (latitude/longitude).

Station Locality Province Altitude (m) Geographical coordinates
Rif
1. Aïn Afersiw Mezine Chefchaouen 750 35°05.945'N, 05°20.439'W
2. Aïn Boughaba Jbel Bou Bessoui, Cedar Forest Chefchaouen 1530 34°58.779'N, 04°46.366'W
3. Aïn El Ma Bared Bouzthate Chefchaouen 1270 35°00.333'N, 05°12.105'W
4. Aïn El Maounzil National Park of Talassemtane Chefchaouen 1110 35°04.577'N, 05°10.406'W
5. Aïn Quanquben Jbel Bou Bessoui Chefchaouen 1600 34°57.750'N, 04°40.783'W
6. Aïn Takhninjoute Bab Rouida, National Park of Talassemtane Chefchaouen 1510 35°06.881'N, 05°08.270'W
7. Aïn Sidi Brahim Ben Arrif Road Moulay Abdessalam Larache 900 35°20.398'N, 05°32.712'W
8. Belyounech Jbel Moussa M’diq Fnideq 60 35°54.389'N, 05°23.630'W
9. Bni Maaden Bni Maaden Tétouan 20 35°34.420'N, 05°15.901'W
10. Dam Entrasol Ouedlaou Bni Ferten Chefchaouen 40 35°22.015'N, 05°10.626'W
11. Daya El Birdiyel Ânasar Chefchaouen 1300 35°01.089'N, 04°59.477'W
12. Dam Moulay Bouchta Larbaa Beni Hassan Tétouan 360 35°15.864'N, 05°21.221'W
13. Dam Nakhla Zinat Tétouan 240 35°26.954'N, 05°24.326'W
14. Dam Smir Bouzaghlale (M’diq) Tétouan 20 35°41.491'N, 05°22.673'W
15. Daya Amsemlil Jbel Bouhachem Tétouan 1060 35°15.596'N, 05°25.917'W
16. Daya El Ânassar Bab Berred Chefchaouen 1180 35°00.788"N/04°57.419'W
17. Daya Jbel Zemzem Jbel Zemzem Tétouan 220 35°45.457'N, 05°22.189'W
18. Daya Mtahen Jbel Bouhachem Larache 970 35°16.195'N, 05°26.158'W
19. Daya Rahrah Tanger Tanger-Assilah 60 35°46.063'N, 05°51.517'W
20. Daya Tazia Road Moulay Abdessalam Larache 720 35°20.814'N, 05°33.139'W
21. Douar Dacheryène Dacheryène Tétouan 130 35°33.863'N, 05°27.162'W
22. Douar Kitane Kitane Tétouan 50 35°32.412'N, 05°20.393'W
23. Forest Bab El Karne Tamakoute Chefchaouen 1250 34°58.510'N, 05°11.838'W
24. Forest El Mahfoura Zaouia Sidi Kacem, Amsa Tétouan 800 35°27.724'N, 05°25.807'W
25. Forest house National Park of Talassemtane Chefchaouen 1670 35°08.076'N, 05°08.262'W
26. Garden Ksar Al Rimal Kabila M’diq-Fnideq 10 35°43.806'N, 05°20.509'W
27. Jumb Kitane Kitane Tétouan 40 35°32.759'N, 05°20.420'W
28. Halouma Kitane Kitane Tétouan 140 35°31.912'N, 05°19.861'W
29. 1 km after Derdara Tanakoub Chefchaouen 510 35°06.419'N, 05°17.938'W
30. Ksar El Kébir Ksar El Kébir Larache 25 35°00.441'N, 05°53.352'W
31. Meadow Mizoghar El Khizana Chefchaouen 1000 35°02.725'N, 05°12.969'W
32. Meadow Fahs Lmhar Al Alyiene M’diq Fnideq 590 35°40.148'N, 05°26.537'W
33. Oued Aârkoub Aârkoub Chefchaouen 140 35°15.943'N, 04°50.631'W
34. Oued Achekrade Douar Aouzighen Tétouan 640 35°22.931'N, 05°20.364'W
35. Oued Ametrasse Chrafate Chefchaouen, 830 35°05.014'N, 05°05.130'W
36. Oued Azila Azila, Jbel Tidghine Al Hoceïma 1600 34°52.028'N, 04°32.609'W
37. Oued 15 km from Fifi Bouzthate Chefchaouen 1260 35°00.805'N, 05°12.365'W
38. Oued Boumarouil Larbaa Beni Hassen Tétouan 550 35°18.733'N, 05°21.271'W
39. Oued Bin El Ouidane Ksar Sghir Tanger-Tétouan 90 35°46.356'N, 05°31.343'W
40. Oued Derdara Derdara Chefchaouen 400 35°06.484'N, 05°17.147'W
41. Oued Mharhar Tanger Tanger-Assilah 5 35°35.040'N, 05°59.220'W
42. Oued El Koub Souk El Had Chefchaouen 120 35°01.298'N, 05°25.333'W
43. Oued Farda Akchour Chefchaouen 400 35°14.350'N, 05°10.460'W
44. Oued Jnane Niche Jnane Niche Chefchaouen 40 35°17.019'N, 04°51.233'W
45. Oued Majjou (Hafa meqlouba) Majjou village Chefchaouen 800 35°06.175'N, 05°10.836'W
46. Oued Martil Mhannech 2 Tétouan 9 35°33.693'N, 05°22.510'W
47. Oued Mezine Mezine Chefchaouen 780 35°06.104'N, 05°21.177'W
48. Oued Sahel Ben Karrich Tétouan 40 35°29.238'N, 05°26.352'W
49. Oued Sidi Ben Sâada Derdara Chefchaouen 220 35°03.921'N, 05°19.895'W
50. Oued Sidi Yahya Aârab Sidi Yahya Aârab Chefchaouen 60 35°17.545'N, 04°53.503'W
51. Oued Taïda Taïda Larache 500 35°22.250'N, 05°32.355'W
52. Stream at 1 km from Sidi Yahya Aârab Sidi Yahya Aârab Chefchaouen 2120 35°18.203'N, 04°53.965'W
53. Village Sebt Zinate Village Sebt Zinate Tanger-Assilah 60 35°39.272'N, 05°44.071'W
54. Tributary Oued Tazarine Mezine Chefchaouen 730 35°05.670'N, 05°21.991'W
55. Tributary Tarmast National Park of Al Hoceïma Al Hoceïma 170 35°10.666'N, 04°03.088'W
Eastern region
56. Farm Saf-Saf Douar Oueld Ben Amrou Aklim Berkane 60 34°54.122'N, 02°37.212'W
57. Oued El Khemis Road vers Aklim Berkane 80 34°54.330'N, 02°30.097'W
58. Oued Tafoughalt Tafoughalt, Beni Snassen Berkane 750 34°48.941'N, 02°24.471'W
Middle Atlas Mts
59. Aïn Ouilili Sidi Moulay Zarhoun Meknès 490 34°03.216'N, 05°31.050'W
60. Bridge Oued Oum Er-rabie (Douar Ahl Souss) El Had Labradia Fkih Bensalah 420 32°25.865'N, 06°31.883'W
61 Douar Ben Smim Ben Smim Ifrane 1690 33°28.200'N, 05°09.780'W
62. Douar Zaouiat Cheikh Zaouiat Cheikh Khénifra 700 32°39.246'N, 05°54.154'W
63. Oued d’Ifrane Gorge of Zad Ifrane 2140 33°01.740'N, 05°04.260'W
64. Vicinity of Ifrane Ifrane Ifrane 1660 33°32.580'N, 05°09.240'W
High Atlas Mts
65. Aïn Zarka of Meski Madkhal Meski Errachidia 970 31°51.388'N, 04°16.917'W
66. Agadir airport Ikhourbane Agadir 50 30°20.400'N, 09°26.820'W
67. Douar Akhlij Tnine Ourika Ourika Marrakech 870 31°22.385'N, 07°46.608'W
68. Douar Aourir Aouirir Agadir 200 30°28.980'N, 09°38.340'W
69. Lac Oukaïmeden Oukaïmeden Marrakech 2590 31°12.420'N, 07°51.120'W
70. Palm grove Igrane Merzouga village Hassilalbied Errachidia 710 31°08.520'N, 04°01.260'W
71. Vicinity of Asni Asni Marrakech 1400 31°14.880'N, 08°00.480'W
72. Vicinity of Ijoukak Ijoukak Marrakech 1790 30°56.100'N, 08°03.840'W
73. Tizi N’Test Tizi N’Test Taroudante 1850 30°51.180'N, 08°22.080'W
Anti Atlas Mts
74. Beach of Tamellalt Tamellalt Tiznit 70 29°43.080'N, 09°55.020'W
75. Douar Issafen Issafen Tata 1110 29°50.460'N, 08°32.100'W
76. Douar Zaouia Kelâa M’gouna Ouarzazate 1240 31°14.275'N, 06°08.169'W
77. 1 km before Douar Aïn Lahmar Jbel Ras Tarf Guelmim 190 29°01.320'N, 10°23.100'W
78. Msidira El Filaha Guelmim 260 28°56.865'N, 10°02.979'W
79. Oued Assa Assa Assa zag 310 28°36.507'N, 09°25.800'W

Nomenclature followed Speight (2018) and subfamily classification followed Mengual et al. (2015). Synonyms are given as far as they have been used in the literature dealing with the Moroccan fauna. All the specimens collected in this study are deposited in the collection of Diptera of the Department of Biology, Faculty of Sciences, University Abdelmalek Essaâdi, Tétouan.

Results

Subfamily SYRPHINAE Latreille, 1802

Tribe MELANOSTOMINI Williston, 1885

Genus MELANOSTOMA Schiner, 1860

Melanostoma mellinum (Linnaeus, 1758)

Literature records. Rif (Becker and Stein 1913: 88, Gil Collado 1929: 405, Kanervo 1939: 2–3). Central Plateau (Séguy 1934: 162). Middle Atlas (Kanervo 1939, Timon-David 1951: 145, Leclercq 1961: 241, Claussen and Hauser 1990: 435). High Atlas (Kanervo 1939, Timon-David 1951, Claussen 1989b: 359, 373, Claussen and Hauser 1990). Distribution map Dirickx (1994: 76, 218).

New sites. Rif: Daya Rahrah, 8/IV/2013, 4♂♂; Aïn El Ma Bared, 06/V/2013, 4♀♀, 06/V/2014, 2♂♂; Oued Derdara, 24/V/2013, 1♀; Daya El Ânassar, 24/V/2013, 1♂; Daya Mtahen, 30/V/2013, 1♂; Garden Ksar Al Rimal, 26/V/2013, 7♂♂, 17♀♀, 5/VI/2013, 1♂; Tributary Oued Tazarine, 11/VI/2013, 1♂; Oued Farda, 23/VI/2013, 1♂; Daya El Birdiyel, 27/VI/2013, 1♀; Forest house, 18/V/2014, 1♂, 1♀; Stream at 1 km from Sidi Yahya Aârab, 25/IV/2015, 1♂; Daya Amsemlil, 7/V/2015, 3♀♀, sweep net, leg. Sahib and Belqat. Eastern region: Farm Saf-Saf, 14/VI/2013, 3♂♂, 21♀♀, sweep net, leg. Sahib and Belqat. Middle Atlas: Oued d’Ifrane, 10/VI/2014, 4♂♂, sweep net, leg. Bot. High Atlas: Lac Oukaïmeden, 18/VI/2014, 1♂, 1♀, sweep net, leg. Bot.

Distribution. Holarctic.

Comment. New record for the Eastern Region.

Melanostoma scalare (Fabricius, 1794)

Literature records. Rif (Gil Collado 1929: 405). Listed (Claussen 1989b: 373). Distribution map (Dirickx 1994: 76, 219).

New sites. Rif: Aïn El Ma Bared, 06/V/2013, 11♀♀; Oued Mezine, 11/VI/2013, 1♀; Aïn Quanquben, 27/VI/2013, 6♀♀; Aïn Takhninjoute, 17/V/2014, 1♂; Oued El Koub, 30/V/2014, 1♀, sweep net, leg. Sahib and Belqat.

Distribution. Palaearctic, Afrotropical, and Indomalayan regions.

Genus PLATYCHEIRUS Le Peletier and Serville, 1828

Platycheirus ambiguus (Fallén, 1817)

Literature record. High Atlas (Kassebeer 1998b: 27).

Distribution. Palaearctic.

Platycheirus atlasi Kassebeer, 1998

Literature records. Middle Atlas (Kassebeer 1998b: 28). High Atlas (Kassebeer 1998b: 28).

Comment. Endemic to Morocco.

Distribution. Morocco.

Platycheirus fulviventris (Macquart, 1829)

Literature record. High Atlas (Ebejer et al. 2019: 150).

Distribution. Palaearctic.

Platycheirus manicatus (Meigen, 1822)

Literature records. High Atlas (Séguy 1961: 74). Listed (Claussen 1989b: 373). Distribution map (Dirickx 1994: 111, 268).

Distribution. Palaearctic, Greenland, and Alaska.

Platycheirus marokkanus Kassebeer, 1998

Literature record. High Atlas, Middle Atlas (Kassebeer 1998b: 31). Cited (Speight 2018: 209).

New site. Rif: Aïn Takhninjoute, 17/V/2014, 1♂. High Atlas: Douar Akhlij Tnine Ourika 18/IV/2015, 1♂, sweep net, leg. Sahib and Belqat.

Comment. New record for the Rif.

Distribution. Morocco and Portugal.

Genus XANTHANDRUS Verrall, 1901

Xanthandrus comtus (Harris, 1780)

Literature records. Rif (Gil Collado 1929: 405). Listed (Claussen 1989b: 374). Distribution map (Dirickx1994: 128, 290).

Distribution. Palaearctic.

Tribe PARAGINI Glumac, 1961

Genus PARAGUS Latreille, 1804

Paragus albifrons (Fallén, 1817)

Literature records. Middle Atlas (Kanervo1939: 2). Listed (Claussen 1989b: 373). Distribution map (Dirickx 1994: 98, 249).

Distribution. Palaearctic.

Paragus atlasi Claussen, 1989

Literature records. High Atlas (Claussen 1989b: 359–361). Cited (Dirickx 1994: 99, Speight 2018: 175).

Comment. Endemic to Morocco.

Distribution. Morocco.

Paragus bicolor (Fabricius, 1794)

Literature records. Rif (Becker and Stein 1913: 88). Central Plateau (Kanervo 1939: 2). Middle Atlas (Séguy 1930: 128, Kanervo 1939, Timon-David 1951: 144, Claussen 1989, Claussen and Hauser 1990). High Atlas (Kanervo 1939, Claussen 1989b: 361, 373, Claussen and Hauser 1990: 435). Anti Atlas (Séguy 1949: 156). Distribution map (Dirickx 1994: 98, 250).

New sites. Rif: Douar Dacheryène, 4/IV/2014, 1♀, sweep net, leg. Sahib and Belqat. High Atlas: Vicinity of Asni, 05/VI/2014, 1♀, sweep net, leg. Bot.

Distribution. Holarctic.

Paragus cinctus Schiner and Egger, 1853

Literature records. High Atlas (Claussen 1989b: 361, 373). Cited (Speight 2018: 176). Distribution map (Dirickx 1994: 99, 250).

New site. High Atlas: Vicinity of Asni, 05/VI/2014, 1♂, sweep net, leg. Bot.

Distribution. Western and Central Palaearctic.

Paragus coadunatus Rondani, 1847

Literature records. Middle Atlas (Claussen and Hauser 1990: 436). Cited (Speight 2018: 176). Distribution map (Dirickx 1994: 99, 251).

New site. Rif: Oued Mharhar, 16/VI/2009, 1♂, sweep net, leg. Bot.

Comment. New record from the Rif.

Distribution. Balearic Is, Canary Is, Sicily, Malta, Madeira, Turkey, and Morocco.

Paragus flammeus Goeldlin, 1971

Literature records. Middle Atlas (Claussen and Hauser 1990: 435). Cited (Speight 2018: 178). Distribution map (Dirickx 1994: 100, 252).

Distribution. Western and Central Palaearctic.

Paragus haemorrhous Meigen, 1822

Literature records. Recorded and listed from the Middle Atlas (Claussen 1989b: 359, 372). Distribution map (Dirickx 1994: 100, 252).

Distribution. Holarctic and Afrotropical regions.

Paragus hermonensis Kaplan, 1981

Literature record. Middle Atlas (Claussen 1989b: 361, 373). Distribution map (Dirickx 1994: 100, 101, 252).

Distribution. Mediterranean Basin.

Paragus pecchiolii Rondani, 1857

Literature records. Middle Atlas (Claussen and Hauser 1990: 435). Distribution map (Dirickx 1994: 101, 253).

Distribution. Western Palaearctic.

Paragus quadrifasciatus Meigen, 1822

Literature records. Central Plateau (Claussen 1989b: 361, 373). Middle Atlas (Claussen and Hauser 1990: 435, Timon-David 1951: 144, as Paragus pulcherrimus). Distribution map (Dirickx 1994: 101, 253).

New sites. Rif: Douar Kitane, 19/VI/2014, 1♀, sweep net, leg. Sahib and Belqat. High Atlas: Agadir airport, 7/IV/2015, 1♀, leg. Schmid-Egger.

Comment. New record from the Rif and the High Atlas.

Distribution. Palaearctic.

Paragus strigatus Meigen, 1822

Literature records. Central Plateau (Wiedemann 1824: 33, as Paragus bimaculatus). Middle Atlas (Claussen 1989b: 361, 373, Claussen and Hauser 1990: 435). High Atlas (Claussen 1989b: 361, 373). Cited (Speight 2018: 181). Distribution map (Dirickx 1994: 102, 254).

Distribution. Palaearctic.

Paragus tibialis (Fallén, 1817)

Literature records. Rif (Becker and Stein 1913: 88, Leclercq 1961: 241, as Paragus tibialis meridionalis). Middle Atlas (Kanervo 1939: 2, Leclercq 1961: 241). High Atlas (Gil Collado 1929: 403, 404 (as Paragus tibialis var. meridionalis), Kanervo 1939: 2). Anti Atlas (Séguy 1949: 156). Listed (Claussen 1989b: 359, 373). Distribution map (Dirickx 1994: 102, 254).

New sites. High Atlas: Vicinity of Asni, 05/VI/2014, 1♂, 07/VI/2014, 2♂♂; Vicinity of Ijoukak, 17/VI/2014, 1♂, sweep net, leg. Bot.

Distribution. Western Palaearctic.

Paragus vandergooti Marcos-Garcia, 1986

Literature records. Middle Atlas (Claussen and Hauser 1990: 435). High Atlas (Claussen 1989b: 362, 373). Cited (Speight 2018: 181). Distribution map (Dirickx 1994: 102, 255).

Distibution. France, Portugal, Spain, and Morocco.

Tribe CHRYSOTOXINI Newman, 1834

Genus CHRYSOTOXUM Meigen, 1803

Chrysotoxum intermedium (Meigen, 1822)

Literature records. Rif (Becker and Stein 1913: 86, Gil Collado 1929: 414). Central Plateau (Părvu et al. 2006: 275). Middle Atlas (Séguy 1930: 131, 1961: 120). High Atlas (Séguy 1930: 131, Kanervo 1939: 4, Gil Collado 1929: 414). Listed (Claussen 1989b: 372). Distribution map (Dirickx 1994: 47, 178).

New sites. Rif: 1 km after Derdara, 11/VI/2013, 1♀; Oued Azila, 27/VI/2013, 1♂; Daya Jbel Zemzem, 23/IV/2014, 1♂; Aïn El Maounzil, 17/V/2014, 1♀, sweep net, leg. Sahib and Belqat. Eastern region: Oued Tafoughalt, 25/XI/2014, 1♂, sweep net, leg. Sahib and Belqat. Middle Atlas: Douar Zaouiat Cheikh, 19/III/2008, 2♂♂, 1♀, sweep net, leg. A. van Eck. High Atlas: Vicinity of Asni, 6/VI/2014, 2♀♀, sweep net, leg. Bot. Anti Atlas: Douar Issafen, 14/VI/2015, 3♀♀, 1♂, sweep net, leg. Schmid-Egger.

Comment. New record from the Eastern region and the Anti Atlas.

Distribution. Europe and Morocco.

Chrysotoxum volaticum Séguy, 1961

Literature records. Middle Atlas (Timon-David 1951: 146, Leclercq 1961: 242, Séguy 1961: 123). High Atlas (Timon-David 1951, Claussen 1989b: 357, 372). Listed (Claussen 1989b: 372). Distribution map (Dirickx1994: 46, 49, 175, 181).

Distribution. Algeria and Morocco.

Tribe SYRPHINI Latreille, 1802

Genus DASYSYRPHUS Enderlein, 1838

Dasysyrphus albostriatus (Fallén, 1817)

Literature record. High Atlas (Kassebeer 1998a: 22).

Distribution. Palaearctic.

Genus EPISTROPHE Walker, 1852

Epistrophe eligans (Harris, 1780)

Literature records. Rif (Becker and Stein 1913: 88, as Syrphus ochrostoma). Middle Atlas (Kassebeer 1998a: 22). High Atlas (Kassebeer 1998a: 22). Listed (Claussen 1989b: 372, as Epistrophe ochrostoma).

New site. Rif: Daya Tazia, 25/VI/2014, 1♂, sweep net, leg. Sahib and Belqat.

Distribution. Western Palearctic.

Genus EPISYRPHUS> Matsumura and Adachi, 1917

Episyrphus balteatus (De Geer, 1776)

Literature records. Rif (Becker and Stein 1913: 88, as Syrphus balteatus, Gil Collado 1929: 406, Kanervo 1939: 3, as Epistrophe balteata). Central Plateau (Séguy 1930: 129, as Syrphus balteatus, Timon-David 1951: 144, as Epistrophe balteatus). High Atlas (Séguy 1930, Timon-David 1951). Listed (Claussen 1989b: 372). Distribution map (Dirickx 1994: 53, 189).

New sites. Rif: Village Sebt Zinate, 15/III/2008, 1♀, sweep net, leg. AvEck; Aïn Sidi Brahim Ben Arrif, 17/IV/2013, 1♂; Dam Nakhla, 24/V/2013, 1♂, 1♀; Aïn Boughaba, 24/V/2013, 1♀; Garden Ksar Al Rimal, 5/VI/2013, 1♀; Oued Aârkoub, 19/IV/2013, 4♀♀; Daya Rahrah, 2/XII/2013, 1♂, 1♀; Oued Sahel, 5/III/2014, 1♀; Dam Moulay Bouchta, 5/III/2014, 26♂♂; Forest house, 17/VI/2014, 1♂; Ksar El Kébir, 19/IV/2014, 1♂, 1♀; Daya Jbel Zemzem, 23/IV/2014, 13♂♂, 27/IV/2014, 1♀; Oued Taïda, 12/III/2015, 1♂; Stream at 1 km from Sidi Yahya Aârab, 25/IV/2015, 1♂; Aïn Quanquben, 25/IV/2015, 1♀; Oued Majjou (Hafa meqlouba), 25/XII/2015, 1♀; Forest Bab El Karne, 25/XII/2015, 1♂; Douar Kitane, 23/I/2016, 2♂♂, 1♀, sweep net, leg. Sahib and Belqat; Douar Kitane, 7/IV/2017, 54♂♂, 14♀♀, malaise trap, leg. Sahib and Belqat; Forest El Mahfoura, 13/V/2018, 18♂♂, 4♀♀, sweep net, leg. Sahib and Belqat. High Atlas: Aïn Zarka of Meski, 20/III/2008, 1♂, sweep net, leg. A. van Eck. Anti Atlas: Douar Zaouia, 11/VI/2015, 1♂, sweep net, leg. Sahib and Belqat.

Distribution. Palaearctic and Indomalayan regions.

Genus EUPEODES Osten Sacken, 1877

Eupeodes corollae (Fabricius, 1794)

Literature records. Rif (Becker and Stein 1913: 88, as Syrphus corollae, Kanervo 1939: 3, as Syrphus corollae, Gil Collado 1929: 406, as Syrphus corollae). Central Plateau (Timon David 1951: 144, as Syrphus corollae, Gil Collado 1929: 406). Middle Atlas (Leclercq 1961: 241, as Syrphus corollae). High Atlas (Séguy 1930, Timon-David 1951). Anti Atlas (Timon David 1951). Listed (Bigot 1884: 88, as Syrphus berber, Séguy 1930: 129, 1961: 103, Claussen 1989b: 372). Distribution map (Dirickx 1994: 89, 235 (as Metasyrphus corollae)).

New site. Rif: Village Sebt Zinate, 15/III/2008, 1♀, sweep net leg. AvEck; Aïn El Ma Bared, 27/IV/2013, 1♀; Garden Ksar Al Rimal, 20/V/2013, 1♀; Oued Bin EL Ouidane, 14/XI/2013, 1♀; Oued Sahel, 5/III/2014, 1♂; Aïn Takhninjoute, 17/V/2014, 31♂♂; Stream at 1 km from Sidi Yahya Aârab, 25/IV/2015, 2♂♂, 3♀♀; Oued Jnane Niche, 16/III/2014, 1♂, 25/IV/2015, 1♂, 1♀; Dam Smir, 27/IV/2014, 1♂; Oued Boumarouil, 10/V/2014, 1♂; Daya Jbel Zemzem, 23/IV/2014, 1♂, 3♀♀, 17/IV/2014, 6♂♂, 3♀♀; Oued Majjou (Hafa meqlouba), 10/V/2014, 1♂, 27/IV/2015, 1♂; Meadow Fahs Lmhar, 15/IV/2018, 1♀, sweep net, leg. Sahib and Belqat. Douar Kitane, 21/IV/2018, 1♂, reared by Sahib and Belqat; Forest El Mahfoura, 13/V/2018, 12♂♂, 7♀♀, sweep net, leg. Sahib and Belqat. Middle Atlas: Douar Zaouiat Cheikh, 19/III/2008, 1♀, sweep net, leg. A. van Eck. High Atlas: Vicinity of Asni, 6/VI/2014, 1♀; Vicinity of Ijoukak, 17/VI/2014, 1♀, sweep net, leg. Bot; Lac Oukaïmeden, 18/VI/2014, 1♂, sweep net, leg. Bot.

Distribution. Palaearctic, Afrotropical Region, and Taiwan.

Eupeodes latifasciatus (Macquart, 1829)

Literature records. Anti Atlas (Séguy 1949: 156, Séguy 1953: 84, as Syrphus latifasciatus). Listed (Claussen 1989b: 372). Distribution map (Dirickx 1994: 89, 236 (as Metasyrphus latifasciatus)).

New sites. Rif: Douar Kitane, 2/V/2014, 1♂, 2♀♀; Forest house, 18/V/2014, 1♂; Oued Ametrasse, 27/IV/2015, 1♂, sweep net, leg. Sahib and Belqat.

Comment. New record for the Rif.

Distribution. Holarctic and India.

Eupeodes luniger (Meigen, 1822)

Literature records. Central Plateau (Gil Collado 1929: 406, as Syrphus luniger. Listed (Claussen 1989b: 372). Distribution map (Dirickx 1994: 90, 237 (as Metasyrphus luniger)).

New sites. Rif: Oued Majjou (Hafa meqlouba), 10/V/2014, 3♂♂; Oued Martil, 13/XII/2013, 1♀; Belyounech, 9/VI/2013, 1♀, sweep net, leg. Sahib and Belqat; Douar Kitane, 24/IV/2017, 1♀, malaise trap, leg. Sahib and Belqat. Middle Atlas: Douar Ben Smim, 08/VI/2014, 1♂, sweep net, leg. Bot. High Atlas: Vicinity of Asni, 6/VI/2014, 1♀, sweep net, leg. Bot.

Comments. New records from the Rif, the Middle Atlas and the High Atlas.

Distribution. Palaearctic and northern India.

Eupeodes nuba (Wiedemann, 1830)

Literature records. High Atlas (Claussen 1989b: 359, 372). Cited (Bigot 1884: 88, as Syrphus rufinasutus, Séguy 1961: 107, as Syrphus rufinasutus, Speight 2018: 108). Distribution map (Dirickx 1994: 90, 238 (as Metasyrphus nuba)).

Distribution. Palaearctic.

Genus ISCHIODON Sack, 1913

Ischiodon aegyptius (Wiedemann, 1830)

Literature records. Central Plateau (Gil Collado 1929: 406, Timon-David 1951: 145). Anti Atlas (Timon-David 1951).

Distribution. Afrotropical Region, Mediterranean Basin, and Canary Islands.

Genus LAPPOSYRPHUS Dusek and Laska 1967

Lapposyrphus lapponicus (Zetterstedt, 1838)

Literature records. Rif (Becker and Stein 1913: 88, as Syrphus arcuatus). Listed (Claussen 1989b: 372, as Eupeodes lapponicus). Distribution map (Dirickx 1994: 89, 236 (as Metasyrphus lapponicus)).

Distribution. Palaearctic, northern America from Alaska to California.

Genus MELISCAEVA Frey, 1946

Meliscaeva auricollis (Meigen, 1822)

Literature records. Rif (Becker and Stein 1913: 89, as Syrphus auricollis, Gil Collado 1929: 406, as Syrphus auricollis). Central Plateau (Timon-David 1951: 145, as Epistrophe auricollis). Listed (Claussen 1989b: 373). Distribution map (Dirickx 1994: 76, 219).

New sites. Rif: Aïn El Ma Bared, 27/IV/2013, 1♀; Daya El Ânassar, 24/V/2013, 1♂; Belyounech, 9/VI/2013, 1♀; Oued Mezine, 11/VI/2013, 1♂; Dam Moulay Bouchta, 5/IV/2014, 1♀; Aïn Afersiw, 11/VI/2013, 3 ♂♂, 7♀♀; Dam Entrasol, 14/XI/2014, 1♀; Oued 15 km from Fifi, 6/V/2014, 1♀; Jumb Kitane, 10/III/2015, 1♀; Oued Majjou (Hafa meqlouba), 10/V/2014, 1♀; Douar Kitane, 2/II/2015, 1♂; Oued Sahel, 5/IV/2014, 1♂. Middle Atlas: Aïn Ouilili, 18/II/2016, 2♀♀, sweep net, leg. Sahib and Belqat.

Comment. New record for the Middle Atlas.

Distribution. Western Palaearctic.

Meliscaeva cinctella (Zetterstedt, 1843)

Literature records. Middle Atlas (Séguy 1934: 161, as Syrphus cinctellus). Listed (Claussen 1989b: 373). Distribution map (Dirickx 1994: 77, 219).

Distribution. Holarctic.

Genus SCAEVA Fabricius, 1805

Scaeva albomaculata (Macquart, 1842)

Literature records. Rif (Gil Collado 1929: 405, as Lasiopticus albomaculata). Central Plateau (Timon-David 1951: 145, as Lasiopticus albomaculata). Middle Atlas (Leclercq 1961: 241, Claussen 1989b: 359). Listed (Séguy 1961: 94, Claussen 1989b: 373). Cited (Speight 2018: 225). Distribution map (Dirickx 1994: 115, 275).

New sites. Rif: Belyounech, 9/VI/2013, 1♂, sweep net, leg. Sahib and Belqat. High Atlas: Aïn Zarka of Meski, 20/III/2008, 1♀, sweep net, leg. AvEck; Lac Oukaïmeden, 19/VI/2014, 1♀, sweep net, leg. Bot.

Comment. New records for the High Atlas.

Distribution. Palaearctic.

Scaeva dignota (Rondani, 1857)

Literature records. Middle Atlas (Claussen and Hauser 1990: 436). Distribution map (Dirickx 1994: 116, 275).

New sites. Rif: Oued Majjou (Hafa meqlouba), 27/IV/2015, 1♀; Daya Mtahen, 26/III/2016, 1♂, sweep net, leg. Sahib and Belqat.

Comment. New records for the Rif.

Distribution. Western Palaearctic.

Scaeva mecogramma (Bigot, 1860)

Literature records. Rif (Kassebeer1998a: 23). Middle Atlas (Kassebeer1998a: 23). High Atlas (Kassebeer1998a: 23).

Distribution. Spain, Southern France, Corsica, Italy, Switzerland, and Morocco.

Scaeva pyrastri (Linnaeus, 1758)

Literature records. Rif (Becker and Stein 1913: 88, as Catabomba pyrastri, Gil Collado 1929: 405). Central Plateau (Gil Collado 1929: 405, as Lasiopticus pyrastri, Timon-David 1951: 145, as Lasiopticus pyrastri). Middle Atlas (Séguy 1930: 128, as Lasiopticus pyrastri, Timon-David 1951: 145). Listed (Claussen 1989b: 373). Distribution map (Dirickx 1994: 116, 276).

New sites. Rif: Daya Jbel Zemzem, 23/IV/2014, 1♀; Stream at 1 km from Oued Sidi Yahya Aârab, 25/IV/2015, 1♀, sweep net, leg. Sahib and Belqat. Anti Atlas: 1 km before Douar Aïn Lahmar, 10/IV/2015, 1♀, sweep net, leg. Schmid-Egger.

Comment. New record for the Anti Atlas.

Distribution. Holarctic and India.

Scaeva selenitica (Meigen, 1822)

Literature records. High Atlas (Séguy 1930: 128, as Lasiopticus selenitica). Listed (Claussen 1989b: 373). Distribution map (Dirickx 1994: 116, 276).

Distribution. Palaearctic.

Genus SPHAEROPHORIA Lepeletier and Serville, 1828

Sphaerophoria interrupta (Fabricius, 1805)

Literature records. Rif (Becker and Stein 1913: 87, as Sphaerophoria menthastri, Kanervo 1939: 3). Central Plateau (Kanervo 1939: 3, as Sphaerophoria menthastri, Timon-David 1951: 145, as Sphaerophoria menthastri). Middle Atlas (Kanervo 1939, Timon-David 1951). High Atlas (Kanervo 1939). Anti Atlas (Timon-David 1951). Cited (Séguy 1961: 109, as Sphaerophoria menthastri). Listed (Claussen 1989b: 373). Distribution map (Dirickx 1994: 118).

Comment. S. interrupta records in Morocco need confirmation.

Distribution. Western Palaearctic and Siberia.

Sphaerophoria rueppelli (Wiedemann, 1830)

Literature records. Rif (Kanervo 1939: 4). Central Plateau (Timon-David 1951: 145, Kanervo 1939). High Atlas (Kanervo 1939, Timon-David 1951, Claussen 1989b). Anti Atlas (Timon-David 1951). Cited (Séguy 1961: 110). Listed (Claussen 1989b: 359, 373). Distribution map (Dirickx 1994: 119, 278).

New sites. Rif: Tributary Tarmast, 26/VI/2013, 1♂; Oued Sidi Ben Sâada, 6/V/2015, 2♂♂; Daya Amsemlil, 7/V/2015, 1♂. Eastern region: Farm Saf-Saf, 14/VI/2013, 4♂♂, sweep net, leg. Sahib and Belqat. Middle Atlas: Oued d’Ifrane: 10/VI/2014, 1♂, sweep net, leg. Bot.

Comment. New records for the Eastern region and the Middle Atlas.

Distribution. Palaearctic and Afrotropical regions.

Sphaerophoria scripta (Linnaeus, 1758)

Literature records. Rif (Becker and Stein 1913: 87, Gil Collado 1929: 406, Kanervo 1939: 4). Central Plateau (Kanervo 1939, Timon-David 1951: 145). Middle Atlas (Séguy 1930: 129, Kanervo 1939, Timon-David 1951, Leclercq 1961: 241, Claussen 1989). High Atlas (Kanervo 1939, Claussen 1989b: 359). Listed (Claussen 1989b: 359, 373). Distribution map (Dirickx 1994: 119, 278).

New sites. Rif: Oued Aârkoub, 19/IV/2013, 1♂; Dam Nakhla, 25/IV/2013, 1♀; Meadow Mizoghar, 27/IV/2013, 1♂, 1♀; Oued Derdara, 24/V/2013, 3♀♀; 1 km after Derdara, 11/VI/2013, 1♂, 1♀; Daya El Birdiyel, 27/VI/2013, 8♀♀, sweep net, leg. Sahib and Belqat; Palm grove Igrane, 12/VI/2014, 2♂♂, sweep net, leg. Bot; Aïn Quanquben, 27/VI/2013, 7♂♂, 7♀♀; Forest house, 18/V/2014, 1♂; Oued Sidi Ben Sâada, 6/V/2015, 1♂, 1♀; Daya Mtahen, 7/V/2015, 1♂; Daya Amsemlil, 7/V/2015, 1♂, 1♀; Forest El Mahfoura, 13/V/2018, 2♂♂, sweep net, leg. Sahib and Belqat. Eastern region: Farm Saf-Saf, 14/VI/2013, 47♂♂, 53♀♀, sweep net, leg. Sahib and Belqat. Middle Atlas: Oued d’Ifrane, 10/VI/2014, 2♂♂, sweep net, leg. Bot. High Atlas: Vicinity of Asni, 07/VI/2014, 2♂♂, sweep net, leg. Bot; Lac Oukaïmeden, 18/VI/2014, 1♂, sweep net, leg. Bot. Anti Atlas: Msidira, 18/V/2015, 1♂, 2♀♀, sweep net, leg. Sahib and Belqat.

Comment. New records for the Eastern region and the Anti Atlas.

Distribution. Palaearctic, Greenland, Nepal, and Kashmir.

Sphaerophoria taeniata (Meigen, 1822)

Literature records. Central Plateau (Timon-David 1951: 10, as Sphaerophoria menthastri var. taeniata). Listed (Claussen 1989b: 373). Distribution map (Dirickx 1994: 119, 279).

Distribution. Palaearctic.

Genus SYRPHUS Fabricius, 1775

Syrphus ribesii (Linnaeus, 1758)

Literature records. Rif (Kassebeer 1998a: 23).

New site. Rif: Douar Kitane, 30/I/2014, 1♀, sweep net, leg. Sahib and Belqat.

Distribution. Holarctic.

Syrphus vitripennis Meigen, 1822

Literature records. Rif (Gil Collado 1929: 406). Listed (Claussen 1989b: 374). Distribution map (Dirickx 1994: 125, 286).

Distribution. Palaearctic, western North America, and Taiwan.

Genus XANTHOGRAMMA Schiner, 1860

Xanthogramma dives (Rondani, 1857)

Literature record. High Atlas (Ebejer et al. 2019)

Distribution. Europe and Morocco.

Xanthogramma evanescens Becker, 1913

Literature records. Rif (Becker and Stein 1913: 87). Listed (Claussen 1989b: 374). Distribution map (Dirickx 1994: 128).

Comment. Endemic to Morocco.

Distribution. Morocco.

Xanthogramma marginale (Loew, 1854)

Literature records. Rif (Becker and Stein 1913: 86, as Xanthogramma marginale var. morenae), Gil Collado 1929: 406, as Xanthogramma marginale var. morenae), Kanervo 1939: 4). High Atlas (Kanervo 1939). Middle Atlas (Claussen and Hauser 1990: 436). Cited (Séguy 1961: 113, Speight 2018: 258). Listed (Claussen 1989b: 374). Distribution map (Dirickx 1994: 129, 291).

New sites. Rif: Village Sebt Zinate, 15/III/2008, 1♀, sweep net, leg. AvEck; Oued Majjou (Hafa meqlouba), 10/V/2014, 1♂; Oued Ametrasse, 18/IV/2015, 1♂, sweep net, leg. Sahib and Belqat; Douar Kitane, 7/IV/2017, 1♂, malaise trap, leg. Sahib and Belqat.

Distribution. Western part of the Mediterranean Basin.

Xanthogramma pedissequum (Harris, 1776)

Literature records. Rif (Gil Collado 1929: 406, as Xanthogramma ornatum). Listed (Claussen 1989b: 374). Distribution map (Dirickx 1994: 129, 292).

Comment. X. pedissequum records in Morocco need confirmation.

Distribution. Europe and Morocco.

Subfamily ERISTALINAE Rondani, 1845

Tribe CALLICERINI Brues and Melander, 1932

Genus CALLICERA Panzer, 1806

Callicera fagesii Guérin-Méneville, 1844

Literature record. High Atlas (Kassebber 1998a: 22).

Distribution. Europe and Morocco.

Callicera rufa Schummel, 1842

Literature records. Rif (Gil Collado 1929: 414). Listed (Claussen 1989b: 372). Distribution map (Dirickx 1994: 23, 139).

Distribution. Western Palaearctic.

Tribe CERIOIDINI Wahlgren, 1909

Genus CERIANA Rafinesque, 1815

Ceriana conopsoides (Linnaeus, 1758)

Literature records. Middle Atlas (Séguy 1930: 131, as Cerioides conopsoides). Listed (Claussen 1989b: 372). Distribution map (Dirickx 1994: 23, 140).

Distribution. Palaearctic.

Ceriana vespiformis (Latreille, 1804)

Literature records. Rif (Becker and Stein 1913: 88, as Cerioides vespiformis, Gil Collado 1929: 414). Central Plateau (Séguy 1930: 131, as Cerioides vespiformis, Leclercq 1961). Middle Atlas (Séguy 1930, Kanervo 1939: 5, Leclercq 1961: 242, as Cerioides vespiformis). High Atlas (Gil Collado 1929, as Cerioides vespiformis, Kanervo 1939, Claussen 1989b: 365, 372). Cited (Séguy 1961: 210, Speight 2018: 21). Distribution map (Dirickx 1994: 23, 140).

New sites. Rif: 1 km after Derdara, 11/VI/2013, 1♂; Meadow Mizoghar, 6/V/2014, 1♂; Oued Achekrade, 31/V/2014, 1♂, sweep net, leg. Sahib and Belqat.

Distribution. Western Palaearctic.

Tribe CHEILOSINI Williston, 1885

Genus CHEILOSIA Meigen, 1822

Cheilosia brunnipennis Becker, 1894

Literature records. High Atlas (Kanervo 1939: 2, as Chilosia flavipes, Kassebeer 1998c: 58, 59). Cited (Séguy 1961: 38, 39, Speight 2018: 29). Listed (Claussen 1989b: 372, as Cheilosia flavipes).

Distribution. Western Palaearctic.

Cheilosia grossa (Fallén, 1817)

Literature record. Hight Atlas (Kassebeer 1998c: 59). Cited (Speight 2018: 35).

Distribution. Palaearctic and India.

Cheilosia latifrons (Zetterstedt, 1843)

Literature records. Rif (Becker 1894: 395, as Chilosia marokkana, Becker and Stein 1913: 87, as Chilosia maroccana, Gil Collado 1929: 405, Kassebeer 1998c: 60). Middle Atlas, High Atlas (Kassebeer 1998c). Cental Plateau (Timon-David 1951: 144, as Chilosia intonsa). Cited (Séguy 1961: 43, as Chilosia maroccana). Listed (Claussen 1989b: 372, as Cheilosia maroccana). Distribution map (Dirickx 1994: 34, 36, 159).

New site. Rif: Oued Sidi Yahya Aârab, 25/IV/2015, 1♂, sweep net, leg. Sahib and Belqat.

Distribution. Palaearctic.

Cheilosia paralobi Malski, 1962

Literature records. Rif (Gil Collado 1929: 405, as Cheilosia longula). Middle Atlas (Claussen 1989b, Claussen and Hauser 1990: 436, Kassebeer 1998c: 61). High Atlas (Kassebeer 1998c: 61). Listed (Claussen 1989b: 372). Cited (Speight 2018: 44). Distribution map (Dirickx 1994: 38, 163).

Distribution. Portugal, Spain, Algeria and Morocco.

Cheilosia parva Kassebeer, 1998

Literature record. Middle Atlas (Kassebeer 1998c: 62).

Comment. Endemic to Morocco.

Distribution. Morocco.

Cheilosia rodgersi Wainwright, 1911

Literature records. Rif (Becker and Stein 1913: 88, Claussen 1989a: 283, Claussen 1989b: 372). Listed (Claussen 1989b: 372). Cited (Speight 2018: 48). Distribution map (Dirickx 1994: 40, 166).

Distribution. Portugal, southern Spain, and Morocco.

Cheilosia ruffipes (Zetterstedt, 1843)

Literature records. Middle Atlas (Claussen and Hauser 1990: 436 as Cheilosia soror, Kassebeer 1998c: 65, as Cheilosia rufipes). Distribution map (Dirickx 1994: 42, 169).

Distribution. Palaearctic.

Cheilosia scutellata (Fallén, 1817)

Literature records. Rif (Gil Collado 1929: 405, Kassebeer 1998c: 65). Middle Atlas (Kassebeer 1998c: 65). Listed (Claussen 1989b: 372). Distribution map (Dirickx 1994: 42, 169).

Distribution. Palaearctic.

Cheilosia variabilis (Panzer, 1798)

Literature record. Middle Atlas (Kassebeer 1998c: 65). Cited (Speight 2018: 52).

Distribution. Palaearctic.

Genus FERDINANDEA Rondani, 1844

Ferdinandea fumipennis Kassebeer, 1999

Literature records. Middle Atlas (Kassebeer 1999c: 153–162). High Atlas (Kassebeer 1999c). Cited (Speight 2018: 110).

Distribution. Portugal, Spain, France, Tunisia, and Morocco.

Tribe CHRYSOGASTERINI Shannon, 1922

Genus BRACHYOPA Meigen, 1822

Brachyopa atlantea Kassebeer, 2000

Literature record. High Atlas (Kassebeer 2000: 141–148). Cited (Speight 2018: 9).

Distribution. Southern Spain and Morocco.

Genus CHRYSOGASTER Meigen, 1803

Chrysogaster basalis Loew, 1857

Literature records. Middle Atlas (Claussen and Hauser 1990: 436, Kassebeer 1999b: 156). Distribution map (Dirickx 1994: 45, 173).

New site. High Atlas: Lac Oukaïmeden, 18/VI/2014, 1♂, sweep net, leg. Bot.

Comment. New record for the High Atlas.

Distribution. Western and Central Palaearctic.

Genus IGHBOULOMYIA Kassebeer, 1999

Ighboulomyia atlasi Kassebeer, 1999

Literature record. Middle Atlas (Kassebeer 1999a: 11–24).

New site. Middle Atlas: Oued d’Ifrane, 10/VI/2014, 8♂♂, 3♀♀, sweep net, leg. Bot.

Comments. Monotypic genus Ighboulomyia with Ighboulomyia atlasi, described by Kassebeer (1999a). The genus Ighboulomyia was placed in the tribe Brachyopini. The genus is endemic to Morocco.

Distribution. Morocco.

Genus MYOLEPTA Newman, 1838

Myolepta difformis Strobl, 1909

Literature records. High Atlas (Dirickx 1994: 93, as Myolepta philonis, Reemer et al. 2004: 558, 567). Cited (Speight 2018: 162).

Distribution. Portugal, Spain, Algeria, Tunisia, and Morocco.

Genus NEOASCIA Williston, 1886

Neoascia clausseni Hauser and Kassebeer 1998

Literature records. Rif (Gil Collado 1929: 40, as Neoascia podagrica). Central Plateau (Părvu et al. 2006: 275). High Atlas (Hauser and Kassebeer 1998: 38–42). Listed (Claussen 1989b: 373, as Neoascia podagrica). Distribution map (Dirickx 1994: 94, 244).

New sites. Rif: Oued Jnane Niche, 14/VI/2013, 1♀; Oued Majjou (Hafa meqlouba), 10/V/2014, 1♀, sweep net, leg. Sahib and Belqat.

Distribution. Algeria, Tunisia, and Morocco.

Genus ORTHONEVRA Macquart, 1829

Orthonevra bouazzai Kassebeer, 1999

Literature record. Middle Atlas (Kassebeer 1999a: 158–161).

Comment. Endemic to Morocco.

Distribution. Morocco.

Orthonevra brevicornis (Loew, 1843) (Fig. 2)

New record. Rif: Aïn Afersiw (Fig. 3), 11/VI/2013, 1♀, sweep net, leg. Sahib and Belqat.

Comment. New record for Morocco. In North Africa, the species was known only from Algeria (Djellab et al. 2013: 6). It prefers forests and wet woodland, standing-water bodies and flushes among Quercus forest, humid Pinus sylvestris forest and wet Salix woodland. It visits flowers such as Apiaceae, Cornus, Crateagus, Malus, Pyrus communis, Ranunculus, Rorippa, Salix (Speight 2018) in sunlit glades among woodland. The species is collected in the present study in a pond surrounded by Conifer reforestation, with the edges overgrown by grasses and herbaceous vegetation including fern, Rubus and Cistus monspeliensis.

Distribution. Western and Central Palaearctic.

Figure 2. 

Orthonevra brevicornis, female, lateral view.

Figure 3. 

Habitat of Orthonevra brevicornis: Aïn Afersiw environment.

Orthonevra elegans (Meigen, 1822)

Literature records. Rif (Becker and Stein 1913: 87). Listed (Claussen 1989b: 373). Distribution map (Dirickx 1994: 96, 246).

Distribution. Palaearctic.

Orthonevra schachti Claussen, 1989

Literature records. High Atlas (Claussen 1989b, Dirickx 1994: 97, Kassebeer 1999a: 160).

Comments. Endemic to Morocco.

Distribution. Morocco.

Genus RIPONNENSIA Maibach, Goeldin and Speight 1994

Riponnensia longicornis (Loew, 1843)

Literature records. Central Plateau (Séguy 1961: 23, as Orthonevra longicornis). High Atlas (Kanervo 1939: 2, as Orthonevra longicornis, Kassebeer 1999a: 162). Middle Atlas (Kassebeer 1999a: 162). Cited (Speight 2018: 224). Listed (Claussen 1989b: 373). Distribution map (Dirickx 1994: 97, 247).

Distribution. Mediterranean Basin.

Riponnensia splendens (Meigen, 1822)

Literature records. Rif (Gil Collado 1929: 405, as Chrysogaster splendens). Middle Atlas (Kassebeer 1999a). High Atlas (Claussen 1989b, as Orthonevra splendens, Kassebeer 1999a: 162). Recorded and listed (Claussen 1989b: 363, 373). Distribution map (Dirickx 1994: 97, 248 (as Orthonevra splendens)).

New site. Middle Atlas: Douar Zaouiat Cheikh, 19/III/2008, 2♂♂, sweep net, leg. A. van Eck.

Distribution. Western Palaearctic.

Tribe ERISTALINI Newman, 1834

Genus ANASIMYIA Schiner, 1864

Anasimyia contracta Claussen and Torp, 1980

Literature record. Middle Atlas (Kassebeer 1998a: 22).

Distribution. Western Palaearctic.

Genus ERISTALINUS Rondani, 1845

Eristalinus aeneus (Scopoli, 1763)

Literature records. Rif (Becker and Stein 1913: 86, as Lumpetia aenea, Gil Collado 1929: 406, 407 (as Eristalis aeneus), Leclercq 1961: 242, as Eristalis aeneus. Central Plateau (Timon-David 1951: 146, as Lathyrophtalmus aeneus, Gil Collado 1929, Séguy 1930: 129, as Lathyrophtalmus aeneus. High Atlas (Kanervo 1939: 5, as Lathyrophtalmus aeneus, Gill Collado 1929). Listed (Claussen 1989b: 372). Distribution map (Dirickx 1994: 53, 54, 190).

New sites. Rif: Jumb Kitane, 05/IV/2015, 1♂, sweep net, leg. Sahib and Belqat. High Atlas: Vicinity of Asni, 05/VI/2014, 1♀, sweep net, leg. Bot.

Distribution. Holarctic, Afrotropical, Oriental, and Australasian regions.

Eristalinus megacephalus (Rossi, 1794)

Literature records. Rif (Becker and Stein 1913: 85, as Eristalis quinquelineatus, Gil Collado 1929: 407, as Eristalis quinquelineatus). Central Plateau (Séguy 1930: 129, as Lathyrophthalmus quinquelineatus). Cited (Séguy 1961: 200). Listed (Claussen 1989b: 372). Distribution map (Dirickx 1994: 54, 190).

Distribution. Western Palaearctic and Afrotropical regions.

Eristalinus sepulchralis (Linnaeus, 1758)

Literature records. Rif (Becker and Stein 1913: 85, as Eristalis sepulchralis, Gil Collado 1929: 406, as Eristalis sepulchralis). Central Plateau (Gil Collado 1929). High Atlas (Claussen 1989b: 366, 372). Distribution map (Dirickx 1994: 54, 191).

Distribution. Palaearctic and India.

Eristalinus taeniops (Wiedemann, 1818)

Literature records. Rif (Becker and Stein 1913: 85, as Eristalis taeniops). Central Plateau, High Atlas (Séguy 1930: 130). Middle Atlas (Leclercq 1961: 243). Listed (Claussen 1989b: 372). Cited (Speight 2018: 79). Distribution map (Dirickx 1994: 55, 191).

New sites. Rif: Oued Martil, 13/XII/2013, 1♀; Halouma Kitane, 1/I/2015, 1♂, sweep net, leg. Sahib and Belqat; Oued Sidi Yahya Aârab, 3/II/2015, 2♂♂, 1♀. Middle Atlas: Bridge Oued Oum Er-rabie (Douar Ahl Souss), 17/V/2017, 1♂, reared by Sahib and Belqat.

Distribution. Palaearctic, Oriental, and Afrotropical regions.

Genus ERISTALIS Latreille, 1804

Eristalis arbustorum (Linnaeus, 1758)

Literature records. Rif (Becker and Stein 1913: 85, Gil Collado 1929: 407, Kanervo 1939: 4). Central Plateau (Timon-David 1951: 146). Middle Atlas (Claussen 1989b: 366, Timon-David 1951: 146, Séguy 1930: 129). Listed (Claussen 1989b: 372). Distribution map (Dirickx 1994: 55, 192).

New sites. Rif: Aïn Sidi Brahim Ben Arrif, 25/IV/2014, 1♂, sweep net, leg. Sahib and Belqat. Middle Atlas: Vicinity of Ifrane, 7/VI/2014, 1♀, sweep net, leg. Bot. High Atlas: Vicinity of Asni, 05/VI/2014, 2♂♂, 07/VI/2014, 1♀; Lac Oukaïmeden, 18/VI/2014, 3♂♂, 19/VI/2014, 1♂, 1♀, sweep net, leg. Bot. Anti Atlas: Douar Issafen, 12/IV/2015, 1♂; Douar Issafen, 14/IV/2015, 2♀♀, sweep net, leg. Schmid-Egger.

Comment. New records for the High Atlas and the Anti Atlas.

Distribution. Holarctic and Oriental regions.

Eristalis pertinax (Scopoli, 1763)

Literature records. Central Plateau, Middle Atlas, High Atlas (Séguy 1930: 130). Listed (Claussen 1989b: 372). Distribution map (Dirickx 1994: 56, 194).

Distribution. Western Palaearctic.

Eristalis similis (Fallén, 1817)

Literature records. Rif (Gil Collado 1929: 407, as Eristalis pratorum). High Atlas (Claussen 1989b: 366, 372). Cited (Séguy 1961: as Eristalis pratorum). Distribution map (Dirickx 1994: 56, 195).

New sites. Rif: Forest house, 18/V/2014, 1♂, 2♀♀, sweep net, leg. Sahib and Belqat. High Atlas: Douar Zaouiat Cheikh, 19/III/2008, 1♂, sweep net, leg. AvEck; Lac Oukaïmeden, 19/VI/2014, 1♀, sweep net, leg. Bot.

Distribution. Palaearctic.

Eristalis tenax (Linnaeus, 1758)

Literature records. Rif (Becker and Stein 1913: 85, as Erystalomyia tenax, Gil Collado 1929: 407, Kanervo 1939: 5, Leclercq 1961: 242). Central Plateau (Gil Collado 1929: 407, Timon-David 1951, as Erystalomyia tenax). Middle Atlas (Kanervo 1939, Timon-David 1951: 146, Leclercq 1961). High Atlas (Gil Collado 1929, Kanervo 1939, Timon-David 1951). Anti Atlas (Séguy 1949: 156, Timon-David 1951: 146). Cited (Séguy 1930: 130). Listed (Claussen 1989b: 372). Distribution map (Dirickx 1994: 57, 195).

New sites. Rif: Village Sebt Zinnat, 15/III/2008, 1♀; Belyounech, 9/V/2013, 1♂; Aïn Takhninjoute, 17/V/2014, 2♂♂; Forest house, 18/V/2014, 1♀; Jumb Kitane, 18/IV/2018, 1♀; Meadow Fahs Lmhar, 15/IV/2018, 2♂♂, sweep net, leg. Sahib and Belqat. High Atlas: Douar Zaouiat Cheikh, 19/III/2008, 1♂, sweep net, leg. AvEck; Vicinity of Asni, 05/VI/2014, 1♀, sweep net, leg. Bot; Lac Oukaïmeden, 18/VI/2014, 2♂♂, sweep net, leg. Bot.

Distribution. Cosmopolitan, except Antarctica.

Genus HELOPHILUS Meigen, 1822

Helophilus trivittatus (Fabricius, 1805)

Literature records. Rif (Becker and Stein 1913: 86, Gil Collado 1929: 407). Listed (Claussen 1989b: 372). Distribution map (Dirickx 1994: 70, 209).

Distribution. Palaearctic.

Genus MALLOTA Meigen, 1822

Mallota cimbiciformis (Fallen, 1817)

Literature records. Rif (Becker and Stein 1913: 85, as Mallota eristaloides). Listed (Claussen 1989b: 373). Distribution map (Dirickx 1994: 73, 213).

Distribution. Western and Central Palaearctic.

Mallota dusmeti Andreu, 1926

Literature records. High Atlas (Kassebeer 1998a: 23).

Distribution. Portugal, central Spain, Tunisia, and Morocco.

Genus MELANOGASTER Róndani, 1857

Melanogaster lindbergi Kassebeer, 1999

Literature records. Rif (Becker and Stein 1913: 87, as Chrysogaster macquarti). Middle Atlas (Kanervo 1939: 2, as Chrysogaster viduata, Kassebeer 1999a: 158). Cited (Séguy 1961: 27, 28 (as Chrysogaster viduata)). Listed (Claussen 1989b: 372, as Chrysogaster lucida). Distribution map (Dirickx 1994: 46, 175 (as Chrysogaster viduata)).

Comment. Endemic to Morocco.

Distribution. Morocco.

Genus MYATHROPA Rondani, 1845

Myathropa florea (Linnaeus, 1758)

Literature records. Rif (Becker and Stein 1913: 85, Gil Collado 1929: 407). Middle Atlas (Séguy 1930, Claussen 1989b: 366). Listed (Claussen 1989b: 373). Distribution map (Dirickx 1994: 92, 240).

New site. Rif: Oued 15 km from Fifi, 6/V/2014, 1♀, sweep net, leg. Sahib and Belqat.

Distribution. Palaearctic.

Genus PARHELOPHILUS Girschner, 1897

Parhelophilus versicolor (Fabricius, 1794)

Literature records. Central Plateau (Gil Collado 1929: 407, as Helophilus versicolor). Listed (Claussen 1989b: 373). Distribution map (Dirickx 1994: 104, 258).

Distribution. Palaearctic.

Tribe EUMERINI Smirnov, 1924

Genus EUMERUS Meigen, 1822

Eumerus amoenus loew, 1848

Literature records. Cited (Séguy 1961: 207, Speight 2018: 87). Listed (Claussen 1989b: 372). Distribution map (Dirickx 1994: 57, 196).

New sites. High Atlas: Douar Aourir, 8/IV/2015, 2♂♂. Anti Atlas: Beach of Tamelallt, 9/IV/2015, 1♂, sweep net, leg. Schmid-Egger.

Comment. New records for the Anti Atlas.

Distribution. Palaearctic.

Eumerus barbarus (Coquebert, 1804)

Literature records. Rif (Gil Collado 1929: 412, as Eumerus australis, Becker and Stein 1913: 86). Middle Atlas (Claussen 1989b: 363). High Atlas (Gil Collado 1929: 412). Cited (Séguy 1961: 208, Speight 2018: 88). Listed (Claussen 1989b: 372). Distribution map (Dirickx 1994: 58, 196).

Distribution. Mediterranean Basin.

Eumerus basalis Loew, 1848

Literature records. Middle Atlas (Séguy 1930: 130, as Eumerus angusticornis). Listed (Claussen 1989b: 372). Distribution map (Dirickx 1994: 58, 197).

Distribution. Western Palaearctic.

Eumerus caballeroi Gil-Collado, 1929

Literature records. Central Plateau (Gil Collado 1929: 412–414). Listed Claussen (1989b: 372). Cited (Speight 2018: 89). Distribution map (Dirickx 1994: 59, 197).

Distribution. Spain and Morocco.

Eumerus hungaricus Szilády, 1940

Literature record. Cited (Speight 2018: 92).

Distribution. Mediterranean Basin.

Eumerus lunatus (Fabricus, 1794)

Literature records. Rif (Becker and Stein 1913: 86, as Eumerus lunalatus, Leclercq 1961: 243). Listed (Claussen1989b: 372). Cited (Speight 2018: 94). Distribution map (Dirickx 1994: 62, 200).

Distibution. Afrotropical Region, Morocco. The Europe records require confirmation.

Eumerus melotus (Séguy, 1941)

Literature records. High Atlas (Séguy 1941: 13, as Lampetia melota). Listed (Claussen 1989b: 372). Distribution map (Dirickx 1994: 62).

Comments. Endemic to Morocco.

Distribution. Morocco.

Eumerus nudus Loew, 1848

Literature records. Rif (Becker and Stein 1913: 86). Cited (Speight 2018: 95). Listed (Claussen 1989b: 372). Distribution map (Dirickx 1994: 64, 201).

Distribution. Mediterranean Basin, Romania, and former Yugoslavia.

Eumerus obliquus (Fabricius, 1805) (Fig. 4)

New records. Rif: Oued Jnane Niche (Fig. 5), 25/IV/2015, 1♂, sweep net, leg. Sahib and Belqat. Eastern region: Oued El Khemis (Fig. 6), 11/IX/2013, 1♂, sweep net, leg. Sahib and Belqat.

Comments. New to Morocco. E. obliquus was recorded from Egypt (Steyskal and El-Bialy 1967, Peck 1988). The species is found in open ground, thinly vegetated, semi-arid dry grassland, often along the margins of seasonal rivers; it feeds from the juice of ripe, fallen fruits of Opuntia (Speight 2018). In Morocco, we collected adults by sweeping the vegetation around lotic habitats. As stated in the literature, we found the species in the margins of a temporary river. The wet section was reduced to a thin layer of water. Riparian vegetation consisted primarily of Opuntia ficus indica (Fig. 5), of Nerium oleander, and herbaceous vegetation. The second habitat was also a river flowing towards oued Moulouya with predominant vegetation consisted of Olea europea, Citrus sinensis, Citrus reticulata, Punica granatum, and Eucalyptus.

Distribution. Mediterranean Basin, the Canaries, Afrotropical, Malagasy, Australasian regions.

Figure 4. 

Eumerus obliquus, male, lateral view.

Figure 5. 

Habitat of Eumerus obliquus: Oued Jnane Niche environment.

Figure 6. 

Habitat of Eumerus obliquus: Oued El Khemis environment.

Eumerus ornatus Meigen, 1822

Literature records. Middle Atlas (Séguy 1930: 130). Listed (Claussen 1989b: 372). Distribution map (Dirickx 1994: 64, 202).

Distribution. Western Palaearctic.

Eumerus pulchellus Loew, 1848

Literature records. Cited (Séguy 1961: 211). Listed (Claussen 1989b: 372). Distribution map (Dirickx 1994: 65, 203).

Distribution. Western Palaearctic.

Eumerus punctifrons Loew, 1857

Literature records. Rif (Leclercq 1961: 243). Listed (Claussen 1989: 372). Distribution map (Dirickx 1994: 65, 203).

Distribution. Afrotropical Region, southern part of the Mediterranean Basin.

Eumerus pusillus Loew, 1848

Literature records. High Atlas (Claussen 1989b: 363, 372). Distribution map (Dirickx 1994: 66, 203).

Distribution. Mediterranean Basin.

Eumerus sabulonum (Fallén, 1817)

Literature records. Cited (Séguy 1961: 212). Listed (Claussen 1989b: 372). Distribution map (Dirickx 1994: 66, 204).

Distribution. Western Palaearctic.

Eumerus schmideggeri Van Steenis, Hauser and Van Zuijen, 2017

Literature record. Anti Atlas (Van Steenis et al. 2017: 155–158).

Distribution. Algeria, Tunisia, and Morocco.

Eumerus strigatus (Fallén, 1817)

Literature records. Rif, High Atlas (Kanervo 1939: 5). Central Plateau (Timon-David 1951: 147). Listed (Claussen 1989b: 372). Distribution map (Dirickx 1994: 67, 205).

Distribution. Palaearctic, North America, Australia, and New Zealand.

Eumerus subornatus Claussen, 1989

Literature records. High Atlas (Claussen 1989b: 364, 365, 372). Cited (Speight 2018: 100). Distribution map (Dirickx 1994: 68).

Distribution. Portugal, France, Spain, and Morocco.

Eumerus truncatus Rondani, 1868

Literature records. Anti Atlas (Van Steenis et al. 2017: 160). Cited (Speight 2018: 102).

Distribution. Portugal, Spain, Greece, Italy, Tunisia, and Morocco.

Genus PLATYNOCHAETUS Wiedemann, 1830

Platynochaetus rufus Macquart, 1835

Literature records. Central Plateau (Gil Collado 1929: 407). Distribution map (Dirickx 1994: 113, 271).

Distribution. Maltese islands, Algeria, and Morocco.

Platynochaetus setosus (fabricius, 1794)

Literature records. High Atlas (Gil Collado 1929: 407). Listed (Claussen 1989b: 373). Cited (Speight 2018: 217). Distribution map (Dirickx 1994: 114, 272).

Distribution. Western part of the Mediterranean Basin.

Tribe MERODONTINI (Edwards, 1915)

Genus MERODON Meigen, 1803

Merodon aberrans Egger, 1860

Literature records. Cited (Séguy 1961: 174, as Lampetia aberrans, Speight 2018: 132). Listed (Claussen 1989b: 373). Distribution map (Dirickx 1994: 77, 220).

Distribution. Western Palaearctic.

Merodon arrasus Beck, 1921

Literature records. Rif (Becker 1921: 54, 55). Listed (Claussen 1989b: 373). Distribution map (Dirickx 1994: 78, 222).

Distribution. Tunisia, Algeria, and Morocco.

Merodon bequaerti Hurkmans, 1993

Literature records. Middle Atlas; Eastern region (Vujić et al. 2020: 98).

Comments. Merodon bequaerti belongs to the Merodon serrulatus species group (Vujić et al. 2020: 81); this species could be Merodon alexeji (Hurkmans 1993: 163).

Distribution. Algeria, Tunisia, Libya, and Morocco.

Merodon biarcuatus Curran, 1939

Literature records. Central Plateau (Curran 1939: 6, 7). Listed (Claussen 1989b: 373). Cited (Speight 2018: 137). Distribution map (Dirickx 1994: 79).

Distribution. France, Italy, Greece, Turkey, and Morocco.

Merodon cabanerensis Marcos-García, Vujić and Mengual, 2007

Literature records. High Atlas (Vujić et al. 2018: 113, 114, 135). Cited (Speight 2018: 137).

Distribution. Spain and Morocco.

Merodon chalybeus Wiedemann in Meigen, 1822

Literature records. Central plateau (Timon-David 1951: 146, as Lampetia spicata). Cited (Claussen 1989b: 365, 373 (as Merodon spicatus), Marcos-García et al. 2007: 546, Speight 2018: 138). Cited (Speight 2018: 138). Distribution map (Dirickx 1994: 87, 233).

Distribution. Mediterranean Basin.

Merodon clavipes (Fabricius, 1781)

Literature records. Cited (Hurkmans 1993: 178, 230, Marcos-García et al. 2007: 548, Speight 2018: 138).

Distribution. Western Palaearctic.

Merodon constans (Rossi, 1794)

Literature records. Rif (Becker and Stein 1913: 86, as Lampetia constans). Listed (Claussen 1989b: 373). Distribution map (Dirickx 1994: 81, 226).

Distribution. Western Palaearctic.

Merodon distinctus Palma, 1863

Literature records. Middle Atlas, Central Plateau (Timon-David 1951: 146, as Lampetia distincta). Listed (Claussen 1989b: 373). Distribution map (Dirickx 1994: 81, 226).

Distribution. Mediterranean Basin.

Merodon elegans Hurkmans, 1993

Literature records. Central Plateau (Hurkmans 1993: 195). Cited (Speight 2018: 141).

Distribution. Western part of Mediterranean Basin.

Merodon eques Fabricius, 1805

Literature records. Cited (Séguy 1961: 178, as lampetia eques). Listed (Claussen 1989b: 373). Distribution map (Dirickx 1994: 81, 227).

Distribution. Mediterranean Basin.

Merodon equestris (Fabricius, 1794)

Literature records. Central Plateau (Fabricius 1805: 240, as Eristalis ferrugineus). Cited (Séguy 1961: 178, 179 (as Lampetia equestris)). Listed (Claussen 1989: 373). Distribution map (Dirickx 1994: 81, 227).

Distribution. Holarctic.

Merodon escalerai Gil Collado, 1929

Literature records. Central Plateau (Gil Collado 1929: 408, 409). Listed (Claussen 1989b: 373). (Dirickx 1994: 82).

Distribution. Spain and Morocco.

Merodon geniculatus Strobl, 1909

Literature records. Rif (Gil Collado 1929: 408). Listed (Claussen 1989b: 365, 373). Cited (Speight 2018: 144). Distribution map (Dirickx 1994: 83, 229).

New Site. High Atlas: Lac Oukaïmeden, 18/VI/2014, 2♂♂, 1♀, 19/VI/2014, 6♂♂, sweep net, leg. Bot.

Comment. New record for the High Atlas.

Distribution. Mediterranean Basin.

Merodon ibericus Vujíc, 2015

Literature records. Rif (Becker and Stein 1913: 86, as Lampetia spinipes, Gil Collado 1929: 409, as Merodon spinipes var. avidus, Kanervo 1939: 5). Central Plateau (Séguy 1930: 130, Timon-David 1951: 146, as Lampetia spinipes). Middle Atlas (Séguy 1930: 130, Kanervo 1939: 5, as Lampetia spinipes var. avida, Popović et al. 2015: 796, Acanski et al. 2016: 3). High Atlas (Gil Collado 1929, Claussen 1989b). Cited (Séguy 1961: 176, as Lampetia avida, Speight 2018: 145). Listed (Claussen 1989: 365, 373). Distribution map (Dirickx 1994: 79, 223).

New site. High Atlas: Lac Oukaïmeden, 18/VI/2014, 1♂, 1♀, 19/VI/2014, 1♂, sweep net, leg. Bot.

Distribution. Portugal, Spain, and Morocco.

Merodon italicus Rondani, 1845

Literature records. Middle Atlas (Claussen and Hauser 1990: 437, as Merodon longicornis). Distribution map (Dirickx 1994: 84, 230).

Distribution. Mediterranean Europe, southern Russia, Lebanon, and Morocco.

Merodon maroccanus Gil Collado, 1929

Literature records. Central Plateau (Gil Collado 1929: 410, 411). Listed (Claussen 1989b: 373). Distribution map (Dirickx 1994: 84).

Comments. Endemic to Morocco.

Distribution. Morocco.

Merodon minutus Strobl, 1893

Literature records. Middle Atlas (Séguy 1961: 180, as lampetia minutus). Listed (Claussen 1989b: 373). Cited (Speight 2018: 149). Distribution map (Dirickx 1994: 84, 230).

Distribution. Southern France around the Mediterranean coast to the Balkan Peninsula.

Merodon murorum (Fabricius, 1794)

Literature records. Central Plateau (Fabricius 1794: 288, as Syrphus murorum, Meigen 1830: 354, as Merodon auripilus). High Atlas (Séguy 1941: 13, as Lampetia auripila). Cited (Séguy 1961: 176). Listed (Claussen 1989b: 373). Distribution map (Dirickx 1994: 79, 222).

Comments. Revised by Vujić et al. (2018: 123–125). Merodon auripilus is a junior synonym of M. murorum.

Distribution. Algeria, Tunisia, and Morocco.

Merodon pruni (Rossi, 1790)

Literature records. Rif (Becker and Stein 1913: 86, as Lampetia pruni, Gil Collado 1929: 407, 408 (as Merodon pruni var. obscurus)). Listed (Claussen 1989b: 373). Distribution map (Dirickx 1994: 85, 231).

Distribution. Western Palaearctic, including Turkmenistan and Iraq.

Merodon pumilus Macquart, 1849

Literature records. Rif (Becker and Stein 1913: 86, as Lampetia aenea, Kanervo 1939: 5, as Lampetia aenea, Ebejer et al. 2019: 150). Central Plateau (Gil Collado 1929: 407). Middle Atlas (Séguy 1930: 130). Cited (Séguy 1961: 174). Listed (Claussen 1989b: 373). Distribution map (Dirickx 1994: 77, 220).

Distribution. Spain, Algeria, and Morocco.

Merodon rufus Meigen, 1838

Literature records. Rif (Becker and Stein 1913: 86, as Lampetia rufa). Listed (Claussen 1989b: 373). Distribution (Dirickx 1994: 86, 232).

Distribution. Western and Central Palaearctic.

Merodon segetum (Fabricius, 1794)

Literature records. Cited (Peck 1988: 173). Listed (Claussen 1989b: 373). Distribution map (Dirickx 1994: 86, 232).

Distribution. Spain, Gibraltar, Macedonia, Crete, Algeria, and Morocco.

Merodon sophron Hurkmans, 1993

Literature records. Middle Atlas (Hurkmans 1993: 168, Vujic et al. 2020: 135–137).

Comment. Endemic to Morocco. Merodon sophron is one of the 13 species belonging to the Merodon serrulatus species group. It is redefined and redescribed by Vujic et al. (2020) on the material from the type locality of Azrou.

Distribution. Morocco.

Merodon tangerensis Hurkmans, 1993

Literature record. Rif (Hurkmans 1993: 172).

Comment. Endemic to Morocco.

Distribution. Morocco.

Merodon tricinctus Sack, 1913

Literature records. High Atlas, Anti Atlas (Kassebeer 1998a: 24). Middle Atlas (Timon-David 1951: 146, as Lampetia tricincta). Listed (Claussen 1989b: 373). Distribution map (Dirickx 1994: 88, 234).

Distribution. Western Palaearctic.

Merodon unguicornis Strobl 1909 (Fig. 7)

Literature record. Middle Atlas (Ebejer et al. 2019: 150).

New site. Rif: Forest house, 17/V/2014, 1♀, sweep net, leg. Sahib and Belqat.

Comment. New record for the Rif. M. unguicornis was cited for the first time in Morocco by Ebejer et al. (2019). In the present work, the female was collected at the locality of Forest house environment (Fig. 8), crossed by a spring and a brook and where the most predominant vegetation was formed by Abies marocana, Pinus negra, Pinus pinaster, Cedrus atlantica and Berberis hispanica. Collected on a misty day, as visualized in Figure 8.

Distribution. Spain and Morocco.

Figure 7. 

Merodon unguicornis, female, lateral view.

Figure 8. 

Habitat of Merodon unguicornis: Forest house environment.

Genus PSILOTA Meigen, 1822

Psilota atra (Fallén, 1817)

Literature records. High Atlas (Kassebeer 1995b: 395–400, as Psilota toubkalana). Cited (Smit and Vujić 2008: 350, Speight 2018: 220).

New Site. Middle Atlas: Douar Zaouiat Cheikh, 19/III/2008, 1♀, sweep net, leg. A. van Eck.

Comment. New record for the Middle Atlas.

Distribution. Europe and Morocco.

Tribe VOLUCELLINI Newman, 1834

Genus VOLUCELLA Geoffroy, 1762

Volucella inanis (Linnaeus, 1758)

Literature records. Middle Atlas (Claussen and Hauser 1990: 436). Distribution map (Dirickx 1994: 126, 289).

Distribution. Palaearctic.

Volucella liquida Erichson in Wagner, 1841

Literature records. Rif (Gil Collado 1929: 406, Leclercq 1961: 242). Central Plateau (Timon-David 1951: 146, Séguy 1930: 129). Middle Atlas (Kanervo 1939: 4, Timon-David 1951: 146, Séguy 1953: 84, Leclercq 1961). High Atlas (Séguy 1930: 129). Listed (Claussen 1989b: 374). Distribution (Dirickx 1994: 127, 289).

New site. Rif: Jumb Kitane, 19/IX/2017, 1♀, sweep net, leg. Sahib and Belqat.

Distribution. Algeria and Morocco.

Volucella zonaria (Poda, 1761)

Literature records. Central Plateau (Séguy 1930: 129). Listed (Claussen 1989b: 374). Distribution map (Dirickx 1994: 127, 290).

Distribution. Palaearctic.

Tribe XYLOTINI Bigot, 1883

Genus BRACHYPALPUS Macquart, 1834

Brachypalpus valgus (Panzer, 1798)

Literature records. High Atlas, Middle Atlas (Kassebeer 1998a: 22).

Distribution. Western Palaearctic.

Genus MILESIA Latreille, 1804

Milesia crabroniformis (Fabricius, 1775)

Literature records. Middle Atlas (Claussen and Hauser 1990: 437). Distribution map (Dirickx 1994: 92, 240).

Distribution. Europe, Turkey, Georgia, and Morocco.

Genus SPILOMYIA Meigen, 1803

Spilomyia maroccana Kuznetsov, 1997

Literature records. Rif (Becker and Stein 1913: 86, as Spilomyia digitate, Kuznetzov 1997: 203–205). High Atlas (Claussen 1989b: 366). Listed (Claussen 1989b: 373, as Spilomyia digitata). Distribution map (Dirickx 1994: 123, 283 (as Spilomyia digitata)).

Distribution. Algeria and Morocco.

Genus SYRITTA Lepeletier and Serville, 1828

Syritta flaviventris Macquart, 1842

Literature records. Central Plateau (Claussen 1989b: 366). Listed (Claussen 1989b: 373). Distribution map (Dirickx 1994: 123, 284).

New site. Eastern region: Farm Saf-Saf, 14/VI/2013, 1♂, 1♀, sweep net, leg. Sahib and Belqat.

Comment. New record for the Eastern region.

Distribution. Mediterranean Basin, Nearctic, Neotropical, Afrotropical regions, Madagascar and the Mascarene islands.

Syritta pipiens (Linnaeus, 1758)

Literature records. Rif (Becker and Stein 1913: 88, Gill Collado 1929: 412, Kanervo 1939: 5). Central Plateau (Séguy 1930: 130, Timon-David 1951, Pârvu et al. 2006: 275). Middle Atlas (Séguy 1930). High Atlas (Timon-David 1951: 146, Claussen 1989b: 366). Listed (Claussen 1989b: 373). Distribution map (Dirickx 994: 124, 285).

New sites. Rif: Bni Maaden, 5/XI/2013, 1♂; Dam Moulay Bouchta, 5/IV/2014, 1♀; Oued El Koub, 30/V/2014, 1♂, 1♀; Douar Kitane, 18/VI/2014, 2♀♀; Oued Majjou (Hafa meqlouba), 27/IV/2015, 1♂, sweep net, leg. Sahib and Belqat. Eastern region: Farm Saf-Saf, 14/VI/2013, 4♂♂, sweep net, leg. Sahib and Belqat. Middle Atlas: Vicinity of Ifrane, 09/VI/2014, 1♀, sweep net, leg. Bot. High Atlas: Vicinity of Asni, 07/VI/2014, 5♂♂; Tizi N’Test, 15/VI/2014, 1♂, sweep net, leg. Bot. Anti Atlas: Oued Assa, 21/V/2015, 1♂, sweep net, leg. Sahib and Belqat.

Comment. New records for the Eastern region and the Anti Atlas.

Distribution. Holarctic, South America, and Indomalayan Region.

Genus TEMNOSTOMA Lepeletier and Serville, 1828

Temnostoma bombylans (Fabricius, 1805)

Literature records. Middle Atlas (Séguy 1961: 155). Listed (Claussen 1989b: 374). Distribution map (Dirickx 1994: 125, 286).

Distribution. Palaearctic.

Genus XYLOTA Meigen, 1822

Xylota segnis (Linnaeus, 1758)

Literature records. Rif (Becker and Stein 1913: 86, as Zelima (Xylota) segnis, Gil Collado 1929: 412, Leclercq 1961: 243). High Atlas (Timon-David 1951: 147). Listed (Claussen 1989b: 372). Distribution map (Dirickx 1994: 130, 293).

New site. Rif: Aïn El Maounzil, 17/V/2014, 1♀; Oued El Koub, 30/V/2014, 1♀; Oued Sidi Ben Sâada, 6/V/2015, 1♂, sweep net, leg. Sahib and Belqat. High Atlas: Vicinity of Asni, 07/VI/2014, 1♂, sweep net, leg. Bot.

Distribution. Palaearctic and eastern North America.

Subfamily PIPIZINAE Mengual, Santos and Rojo 2015

Tribe PIPIZINI Williston, 1885

Genus HERINGIA Rondani, 1856

Heringia heringi (Zetterstedt, 1843)

Literature records. High Atlas, Middle Atlas (Kassebeer 1998a: 23).

Distribution. Palaearctic.

Genus PIPIZELLA Rondani, 1856

Pipizella thapsiana Kassebeer, 1995

Literature records. High Atlas (Kassebeer 1995a: 261–263). Cited (Speight 2018: 199).

New site. Middle Atlas: Douar Zaouiat Cheikh, 19/III/2008, 1♂, sweep net, leg. A. van Eck.

Comment. New record for the Middle Atlas.

Distribution. Portugal and Morocco.

Genus TRIGLYPHUS Loew

Triglyphus escalerai Gil Collado, 1929

Literature records. Rif (Gil Collado 1929: 404). Cited (Speight 2018: 253). Distribution map (Dirickx 1994: 125).

Distribution. Portugal, Croatia, Montenegro, and Morocco.

Discussion

The present work provides an important contribution to the Moroccan hoverfly fauna with two species recorded for the first time for Morocco: Eumerus obliquus (Fabricius, 1805) and Orthonevra brevicornis (Loew, 1843); nine new records are provided for the Rif, five for the Middle Atlas, and seven species are recorded for the first time for the Hight Atlas, the Anti Atlas, and the Eastern Region for which we have enlarged the distribution area. The 150 Moroccan species belong to three subfamilies, 14 tribes, and 49 genera (30 genera of Eristalinae, 16 of Syrphinae, and 3 of Pipizinae).

Knowledge of the hoverfly diversity of Morocco is unequal among regions. For instance, the Middle Atlas and the High Atlas have the same number of species (69), the Rif has 68 species, the Anti Atlas has 18 species, while the Eastern region has only seven species.

Four very common Palaearctic species were collected abundantly: Episyrphus balteatus (156 specimens), Sphaerophoria scripta (146 specimens), Eupeodes corollae (86 specimens) and Melanostoma mellinum (78 specimens).

Morocco is home to 12 endemic species: two are endemic to the Rif, four to the Middle Atlas (including both an endemic genus and species (Ighbouloumia atlasi)), three to the High Atlas, one to the central plateau, and two species, Platycheirus atlasi and Melanogaster lindbergi, to the Middle Atlas plus the High Atlas, and to the Rif plus the Middle Atlas, respectively.

Morocco with the greatest number of total species (150), has 75% of the North African syrphid fauna. Algeria, with 91 species, has 46% of the total fauna. Tunisia, with 69, has 35% of the fauna, whereas Egypt and Libya have 51 species (26%) and 34 species (17%), respectively.

Algeria, Tunisia, Libya, and Egypt share with Morocco 71, 55, 25, and 30 species, respectively, but these numbers are likely to change and a detailed comparison must await a systematic sampling of the whole region.

Of the two Iberian Peninsula countries, Spain has the best known Syrphiidae fauna with 417 species whereas Portugal has only 195 species (Marcos-Garcia et al. 2002, Speight 2015, van Eck 2011, 2016).

These new data on the Moroccan hoverfly fauna reflect the variety of suitable habitats and suggest that many more species can yet be found in Morocco, that provides a wide variety of geographical and climatic properties. New species are also likely to be discovered by genomic prospecting using molecular approaches.

Acknowledgements

The authors are grateful to the following colleagues for their help: Bastiaan Wakkie (Brussels, Belgium), John T. Smit (Leiden, Netherland), and Lair Xavier (France) for helping in the identification of some species; Dr Claus Claussen (Flensburg, Germany), Dr Gabriel Neve (Universite d’Aix-Marseille) for providing advice and bibliography, André van Eck (Amsterdam, Netherland), Sander Bot (Haren, Netherland), Dr Martin Hauser (California Department of Food and Agriculture), and Dr Schmid-Egger for providing additional material.

SS is most grateful to the programme CEBios for the opportunity they gave her to train at the Institute. She is also thankful to Dr Patrick Grootaert (Royal Belgian Institute of Natural Sciences, Brussels) and Mr Pol Limbourg (Royal Belgian Institute of Natural Sciences, Brussels) for their great help during her training in taxonomy at the collections and laboratory of the Royal Belgian Institute of Natural Sciences, Brussels. The authors warmly thank Dr. Martin Ebejer (Honorary Research Fellow, Department of Natural Sciences, National Museum of Wales, United Kingdom.) for his comments and corrections on the manuscript.

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