Checklist |
Corresponding author: Peter Huemer ( p.huemer@tiroler-landesmuseen.at ) Academic editor: Mark Metz
© 2020 Peter Huemer, Ole Karsholt.
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation:
Huemer P, Karsholt O (2020) Commented checklist of European Gelechiidae (Lepidoptera). ZooKeys 921: 65-140. https://doi.org/10.3897/zookeys.921.49197
|
The checklist of European Gelechiidae covers 865 species, belonging to 109 genera, with three species records which require confirmation. Further, it is the first checklist to include a complete coverage of proved synonyms of species and at generic level. The following taxonomic changes are introduced: Pseudosophronia constanti (Nel, 1998) syn. nov. of Pseudosophronia exustellus (Zeller, 1847), Metzneria expositoi Vives, 2001 syn. nov. of Metzneria aestivella (Zeller, 1839); Sophronia ascalis Gozmány, 1951 syn. nov. of Sophronia grandii Hering, 1933, Aproaerema incognitana (Gozmány, 1957) comb. nov., Aproaerema cinctelloides (Nel & Varenne, 2012) comb. nov., Aproaerema azosterella (Herrich-Schäffer, 1854) comb. nov., Aproaerema montanata (Gozmány, 1957) comb. nov., Aproaerema cincticulella (Bruand, 1851) comb. nov., Aproaerema buvati (Nel, 1995) comb. nov., Aproaerema linella (Chrétien, 1904) comb. nov., Aproaerema captivella (Herrich-Schäffer, 1854) comb. nov., Aproaerema semicostella (Staudinger, 1871) comb. nov., Aproaerema steppicola (Junnilainen, 2010) comb. nov., Aproaerema cottienella (Nel, 2012) comb. nov., Ptocheuusa cinerella (Chrétien, 1908) comb. nov., Pragmatodes melagonella (Constant, 1895) comb. nov., Pragmatodes albagonella (Varenne & Nel, 2010) comb. nov., Pragmatodes parvulata (Gozmány, 1953) comb. nov., Oxypteryx nigromaculella (Millière, 1872) comb. nov., Oxypteryx wilkella (Linnaeus, 1758) comb. nov., Oxypteryx ochricapilla (Rebel, 1903) comb. nov., Oxypteryx superbella (Zeller, 1839) comb. nov., Oxypteryx mirusella (Huemer & Karsholt, 2013) comb. nov., Oxypteryx baldizzonei (Karsholt & Huemer, 2013) comb. nov., Oxypteryx occidentella (Huemer & Karsholt, 2011) comb. nov., Oxypteryx libertinella (Zeller, 1872) comb. nov., Oxypteryx gemerensis (Elsner, 2013) comb. nov., Oxypteryx deserta (Piskunov, 1990) comb. nov., Oxypteryx unicolorella (Duponchel, 1843) comb. nov., Oxypteryx nigritella (Zeller, 1847) comb. nov., Oxypteryx plumbella (Heinemann, 1870) comb. nov., Oxypteryx isostacta (Meyrick, 1926) comb. nov., Oxypteryx helotella (Staudinger, 1859) comb. nov., Oxypteryx parahelotella (Nel, 1995) comb. nov., Oxypteryx graecatella (Šumpich & Skyva, 2012) comb. nov.; Aproaerema genistae (Walsingham, 1908) comb. rev., Aproaerema thaumalea (Walsingham, 1905) comb. rev.; Dichomeris neatodes Meyrick, 1923 sp. rev.; Caryocolum horoscopa (Meyrick, 1926) stat. rev.; Ivanauskiella occitanica (Nel & Varenne, 2013) sp. rev.; Apodia martinii Petry, 1911 sp. rev.; Caulastrocecis cryptoxena (Gozmány, 1952) sp. rev. Following Article 23.9.2 ICZN we propose Caryocolum blandella (Douglas, 1852) (Gelechia) nom. protectum and Caryocolum signatella (Eversmann, 1844) (Lita) nom. oblitum.
Europe, species diversity, cryptic diversity, DNA barcoding, synonymy, new combination
Lepidoptera, butterflies and moths, are among the best-known insects, and due to a long tradition of studying Lepidoptera in Europe our knowledge of European Lepidoptera is more comprehensive compared to other parts of the world. Even though Lepidoptera is a well-defined group they exhibit a huge diversity in size, colour and wing markings. Whereas everybody can recognize a butterfly the vast majority of Lepidoptera are small and often dull coloured insects. One such group is the family Gelechiidae. They have for a long time been rather neglected by most lepidopterists mainly due to their external similarity and lack of resources for their identification. Over the last couple of decades, the latter problem has partly been addressed, e.g.,
A checklist is the most basic taxonomic work on a group of organisms. It can be alphabetical or systematic, viz. trying to reflect the current knowledge of the relationship of the included taxa. This checklist is in systematic order, and it moreover includes synonyms and annotations. Its aim is to present an updated overview of the Gelechiidae known from Europe. This is highly appropriate as nearly a quarter of the currently known species have been described since 1990 (
This checklist of European Gelechiidae is the first one to include all known synonyms of genera and species of Europaean Gelechiidae. It is mainly based on data published in Fauna Europaea (
Taxonomically critical genera and species, especially possible cases of cryptic diversity (Fig.
For the purpose of the present checklist we define Europe in a broad sense, which includes the Ural Mountains, Russian parts of the Caucasus, the ‘European’ part of Kazakhstan, the Mediterranean islands and the Macaronesian Islands (except Cape Verde) (Fig.
Geographical boundaries of research area.World boundaries: https://www.arcgis.com/; SRTM-Data: https://dds.cr.usgs.gov/srtm/version2_1/SRTM3/.
The inclusion of the Russian parts of the Caucasus only added four species to the list (Acompsia caucasella Huemer & Karsholt, Neofriseria caucasicella Sattler, Chionodes caucasiella Huemer & Sattler and Scrobipalpa caucasica (Povolný)), which is surprising. One would expect a richer gelechiid fauna to occur in this vast and diverse mountain system. However, most likely the species inventory is simply underestimated as only few lepidopterists have done field research in this area so far.
The checklist is restricted to described nominal taxa. Potentially undescribed species (
The higher classification follows the molecular study of
Although our knowledge of European Gelechiidae has increased much over the last years, there are still available species-group names in the family which have not yet been associated with known species. Very few of these are likely to represent additional taxa, whereas most cases will be synonyms. Furthermore, several of the published synonyms need taxonomic re-assessment. We have not made special efforts to search for type specimens of such taxa for the purpose of the present checklist, and they should be considered in connection with taxonomic revisions within the Gelechiidae.
Many species-group names of European Gelechiidae have been combined in different genera since they were first made available. Following article 31.2 of the International Code of Zoological Nomenclature (
DNA barcodes have been sequenced for a significant number of the species included in the inventory (741 nominal species with sequences > 500 bp). These supported the compilation of the checklist and helped identify and fix yet unpublished synonyms and the systematic position of some species. Details to species and specimens are available on BOLD (
We tested the congruence of morphologically based species determinations and COI sequence data with the Barcode Index Number (BIN), a methodology recently proposed by Ratnasingham and Hebert (2013). This system clusters sequences into Operational Taxonomic Units (OTUs) regardless of their previous taxonomic assignment. It is based on a two-stage algorithm that groups the sequences in a cluster and automatically assigns new sequences. All high-quality sequences > 500 bp are recorded independently of the project origin and assigned to a BIN. Though BINs reflect classical Linnean taxonomy to a high level they were not used uncontested (
The checklist covers 865 nominal species of European Gelechiidae belonging to 109 genera, including 3 species with doubtful records (*). The majority belong to Gelechiinae (445 spp.), followed by Anomologinae (253 spp.), Anacampsinae (89 spp.), Dichomeridinae (47 spp.), Apatetrinae (29 spp.), and Thiotrichinae (5 spp.) (Table
Higher taxa | Species no. |
---|---|
Gelechiidae Stainton, 1854 | 865 |
Anacampsinae Bruand d’Uzelle, 1851 | 89 |
Anacampsini Bruand d’Uzelle, 1851 | 67 |
Chelariini Le Marchand, 1947 | 22 |
Dichomeridinae Hampson, 1918 | 47 |
Apatetrinae Le Marchand, 1947 | 29 |
Pexicopiini Hodges, 1986 | 6 |
Apatetrini Le Marchand, 1947 | 23 |
Thiotrichinae Karsholt, Mutanen, Lee & Kaila, 2013 | 5 |
Anomologinae Meyrick, 1926 | 253 |
Gelechiinae Stainton, 1854 | 445 |
Gelechiini Stainton, 1854 | 132 |
Gnorimoschemini Povolný, 1964 | 240 |
Litini |
73 |
A single species originally described in the Gelechiidae is excluded from the family, viz. Brachmia infuscatella Rebel, 1940, and is transferred to Autostichidae without generic assignation.
Numbers [1] – [202] refer to comments; * refers to doubtful records for the European fauna.
Gelechiidae Stainton, 1854
Anacampsinae Bruand d’Uzelle, 1851 [1]
Stomopteryginae Heslop, 1938, unavailable
Anacampsini Bruand d’Uzelle, 1851
Stomopteryx Heinemann, 1870 [2]
Inotica Meyrick, 1913
Acraeologa Meyrick, 1921
Kahelia Turati, 1922, unavailable
Stomopteryx detersella (Zeller, 1847)
egenella (Herrich-Schäffer, 1851), unavailable
palermitella (La Harpe, 1860)
tenuisignella Turati, 1924
obliterella Turati, 1924, unavailable
Stomopteryx bolschewickiella (Caradja, 1920)
Stomopteryx nugatricella Rebel, 1893 [3]
Stomopteryx mongolica Piskunov, 1975 [3]
Stomopteryx lineolella (Eversmann, 1844) [3]
Stomopteryx basalis (Staudinger, 1876)
oxychalca (Meyrick, 1937)
Stomopteryx deverrae (Walsingham, 1905) [4]
Stomopteryx flavoclavella Zerny, 1935 [5]
Stomopteryx remissella (Zeller, 1847) [6]
vetustella (Herrich-Schäffer, 1854)
tripunctigerella (Bruand d’Uzelle, 1859)
submissella (Frey, 1880), homonym
rufobasella (Rebel, 1916)
yunusemrei Koçak, 1986
Stomopteryx spathulella Nel, Varenne & Labonne, 2019 [6]
Stomopteryx orthogonella (Staudinger, 1871)
Stomopteryx flavipalpella Jäckh, 1959 [7]
Stomopteryx hungaricella Gozmány, 1957
Stomopteryx lusitaniella Corley & Karsholt, 2014
Stomoptery jeppeseni Karsholt & Šumpich, 2018
Stomopteryx alpinella Nel & Varenne, 2016
Stomopteryx schizogynae (Walsingham, 1908)
Aproaerema Durrant, 1897 [8]
Harpagus Stephens, 1834, homonym
Untomia Busck, 1906
Schuetzeia Spuler, 1910
Syncopacma Meyrick, 1925
Lixodessa Gozmány, 1957
Aproaerema patruella (Mann, 1857)
fulvistillella (Rebel, 1891)
Aproaerema coronillella (Treitschke, 1833)
fournieri (Nel, 1998)
Aproaerema incognitana (Gozmány, 1957) comb. nov. [8]
Aproaerema sangiella (Stainton, 1863)
Aproaerema cinctella (Clerck, 1759) [9]
vorticella (Scopoli, 1763)
ligulella ([Denis & Schiffermüller], 1775)
vittata (Fourcroy & Geoffroy, 1785)
vittatella (Villers, 1789)
albistrigella (Stephens, 1834)
ussuriella (Caradja, 1920)
finlandica (Gozmány, 1957)
Aproaerema cinctelloides (Nel & Varenne, 2012) comb. nov. [8]
Aproaerema larseniella (Gozmány, 1957)
ligulella auct.
Aproaerema wormiella (Wolff, 1958) [8]
parawormiella (Nel & Varenne, 2016)
Aproaerema azosterella (Herrich-Schäffer, 1854) comb. nov. [8]
Aproaerema ochrofasciella (Toll, 1936)
Aproaerema taeniolella (Zeller, 1839)
sircomella (Stainton, 1854)
Aproaerema montanata (Gozmány, 1957) comb. nov. [8]
Aproaerema albifrontella (Heinemann, 1870)
ignobilella (Heinemann, 1870)
Aproaerema cincticulella (Bruand, 1851) comb. nov.
Aproaerema vinella Bankes, 1898
fasciata Bankes, 1898, unavailable
biformella Schütze, 1902
Aproaerema buvati (Nel, 1995) comb. nov. [8]
Aproaerema linella (Chrétien, 1904) comb. nov. [8, 10]
schoenmanni (Gozmány, 1957)
Aproaerema albipalpella (Herrich-Schäffer, 1854)
leucopalpella (Herrich-Schäffer, 1854), unavailable
ruptella (Constant, 1865)
Aproaerema suecicella (Wolff, 1958) [11]
Aproaerema captivella (Herrich-Schäffer, 1854) comb. nov. [8]
sarothamnella (Zeller, 1868)
Aproaerema polychromella (Rebel, 1902)
argyrolobiella Caradja, 1920, unavailable
faceta (Meyrick, 1914)
Aproaerema karvoneni (Hackman, 1950) [12]
Aproaerema semicostella (Staudinger, 1871) comb. nov. [8]
albicapitella (Bidzilya, 1996)
Aproaerema steppicolella (Junnilainen, 2010) comb. nov. [8]
Aproaerema cottiennella (Nel, 2012) comb. nov. [8]
Aproaerema genistae (Walsingham, 1908) comb. rev. [8]
Aproaerema thaumalea (Walsingham, 1905) comb. rev. [8]
Aproaerema anthyllidella (Hübner, 1813) [13]
caliginosella (Duponchel, 1843)
elachistella (Stainton, 1859), subspecies
psoralella (Millière, 1865)
lachtensis (Erschoff, 1877)
sparsiciliella (Barrett, 1891)
infestella (Rebel, 1896)
natrixella (Weber, 1945)
brundini (Benander, 1945)
alfalfella Amsel, 1958
aureliana Căpuşe, 1964
Aproaerema lerauti Vives, 2001
Aproaerema mercedella Walsingham, 1908
Iwaruna Gozmány, 1957 [14]
Iwaruna heringi Gozmány, 1957
Iwaruna biguttella (Duponchel, 1843)
Iwaruna klimeschi Wolff, 1958
Iwaruna robineaui Nel, 2008
Anacampsis Curtis, 1827
Tachyptilia Heinemann, 1870
Agriastis Meyrick, 1914
Anacampsis populella (Clerck, 1759) [15]
tremella ([Denis & Schiffermüller], 1775)
boeberana (Fabricius, 1787)
populi (Haworth, 1828), emendation
laticinctella Stephens, 1834
tremulella Duponchel, 1839
atra (Strand, 1901), unavailable
lugens (Caradja, 1920)
sachalinensis (Matsumura, 1931)
fuscatella (Bentinck, 1934)
ambronella (Meder, 1934)
ceballosi Agenjo, 1959
Anacampsis blattariella (Hübner, 1796) [15]
thapsiella (Hübner, 1796)
blattariae (Haworth, 1828), emendation
atragriseella Bruand d’Uzelle, 1851
betulinella Vári, 1941
Anacampsis timidella (Wocke, 1887)
quercella (Chrétien, 1907)
disquei (Meess, 1907)
suberiella Caradja, 1920
Anacampsis scintillella (Fischer v. Röslerstamm, 1841) [16]
brunneella Herrich-Schäffer, 1854
contuberniella (Staudinger, 1859)
Anacampsis temerella (Lienig & Zeller, 1846)
pernigrella (Douglas, 1850)
Anacampsis trifoliella (Constant, 1890)
Anacampsis fuscella (Eversmann, 1844)
Anacampsis hirsutella (Constant, 1885)
Anacampsis obscurella ([Denis & Schiffermüller], 1775) [17]
subsequella (Hübner, 1796)
Anacampsis malella Amsel, 1959
Mesophleps Hübner, 1825 [18]
Brachyacma Meyrick, 1886
Lathontogenus Walsingham, 1897
Paraspistes Meyrick, 1905
Chretienia Spuler, 1910
Lipatia Busck, 1910
Stiphrostola Meyrick, 1923
Crossobela Meyrick, 1923
Xerometra Meyrick, 1925
Gnosimacha Meyrick, 1927
Bucolarcha Meyrick, 1929
Uncustriodonta Agenjo, 1952
Mesophleps corsicella (Herrich-Schäffer, 1856)
lala Agenjo, 1961
Mesophleps silacella (Hübner, 1796)
pyropella auct.
luteella (Hübner, 1896), unavailable
silacea (Haworth, 1828), emendation
apicellus Caradja, 1920
calaritanus Amsel, 1939
Mesophleps oxycedrella (Millière, 1871)
Mesophleps trinotella Herrich-Schäffer, 1856
aurantiella (Rebel, 1915)
subtilipennis (Turati, 1924)
Mesophleps ochracella (Turati, 1926)
orientella Nel & Nel, 2003
gallicella Varenne & Nel, 2011
Chelariini Le Marchand, 1947
Hypatimini Kloet & Hincks, 1945, unavailable
Anarsiini Amsel, 1977
Nothris Hübner, 1825 [19]
Nothris congressariella (Bruand, 1858)
declaratella Staudinger, 1859
Nothris lemniscellus (Zeller, 1839)
Nothris gregerseni Karsholt & Šumpich, 2015 [20]
Nothris verbascella ([Denis & Schiffermüller], 1775)
discretella Rebel, 1889
clarella Amsel, 1935
Nothris sulcella Staudinger, 1879
magna Nel & Peslier, 2007
Nothris radiata (Staudinger, 1879) [21]
Nothris skyvai Karsholt & Šumpich, 2015
Neofaculta Gozmány, 1955
Haplovalva Janse, 1958
Neofaculta ericetella (Geyer, 1832) [22]
gallinella (Treitschke, 1833)
lanceolella (Stephens, 1834)
fuscella (Duponchel, 1844)
subatrella (Duponchel, 1845)
quinquemaculella (Bruand d’Uzelle, 1859)
orcella (Zerny, 1927), subspecies
atlanticella (Amsel, 1938), subspecies
tenalella (Amsel, 1938)
amseli (Dufrane, 1955)
pyrenemontana (Dufrane, 1955)
betulea auct.
Neofaculta infernella (Herrich-Schäffer, 1854)
infernalis, unavailable
Neofaculta taigana Ponomarenko, 1998 [23]
Hypatima Hübner, 1825
Chelaria Haworth, 1828
Tituacia Walker, 1864
Stomylia Snellen, 1878
Allocota Meyrick, 1904, homonym
Cymatomorpha Meyrick, 1904
Deuteroptila Meyrick, 1904
Semodictis Meyrick, 1909
Allocotaniana Strand, 1913
Episacta Turner, 1919
Hypatima rhomboidella (Linnaeus, 1758) [24]
conscriptella (Hübner, 1805)
hubnerella (Donovan, 1806), incorrect original spelling
huebnerella (Donovan, 1806), justified emendation
conscripta Haworth, 1828, emendation
Anarsia Zeller, 1839 [25]
Ananarsia Amsel, 1959
Anarsia lineatella Zeller, 1839
pullatella (Hübner, 1796), nomen oblitum
pruniella Clemens, 1860
heratella Amsel, 1967, subspecies
tauricella Amsel, 1967, subspecies
Anarsia innoxiella Gregersen & Karsholt, 2017
Anarsia spartiella (Schrank, 1802)
robertsonella (Curtis, 1837)
genistae Stainton, 1854
genistella Doubleday, 1859, emendation
ragonotella Réal, 1994
krausei Réal, 1994
lhommella Réal, 1994
acutiloba Réal, 1994
pseudospartiella Réal, 1994
ungemachi Réal, 1994
Anarsia bilbainella (Rössler, 1877) [26]
burmanni Amsel, 1958
bizensis Réal, 1994
infundiblulella Réal, 1994
ovilella Réal, 1994
Anarsia eleagnella Kuznetsov, 1957
Anarsia dejoannisi Réal, 1994
Anarsia leberonella Réal, 1994
Anarsia sibirica Park & Ponomarenko, 1996
Anarsia stepposella Ponomarenko, 2002
psammobia Falkovitsh & Bidzilya, 2003
Anarsia acaciae Walsingham, 1896
Anarsia balioneura Meyrick, 1921
Dichomeridinae Hampson, 1918
Brachminae Omelko, 1999
Dichomerinae, misspelling
Dichomeris Hübner, 1818 [27]
Elasmion Hübner, 1808, unavailable
Oxybelia Hübner, 1825
Rhinosia Treitschke, 1833
Gaesa Walker, 1864
Uliaria Dumont, 1921
Cymotricha Meyrick, 1923
Acanthophila Heinemann, 1870
Mimomeris Povolný, 1978
Dichomeris acuminatus (Staudinger, 1876)
ianthes (Meyrick, 1887)
rusticus (Walsingham, 1892)
lotellus (Constant, 1893)
ammoxanthus (Meyrick, 1904)
ochrophanes (Meyrick, 1907)
sublotellus (Caradja, 1920)
Dichomeris cisti (Staudinger, 1859)
meridionella (Walsingham, 1891)
Dichomeris limbipunctellus (Staudinger, 1859) [28]
millierellus Stainton, 1873
Dichomeris neatodes Meyrick, 1923 sp. rev. [28]
Dichomeris helianthemi (Walsingham, 1903)
Dichomeris castellana (Schmidt, 1941)
Dichomeris juniperella (Linnaeus, 1761) [29]
juniperi Haworth, 1828, emendation
Dichomeris marginella (Fabricius, 1781)
fimbriella (Thunberg, 1788)
clarella (Treitschke, 1833)
Dichomeris ustalella (Fabricius, 1794)
capucinella (Hübner, 1796)
cornutus (Fabricius, 1798)
ustulatus (Fabricius, 1798), emendation
burgundiellus (Bruand d’Uzelle, 1859)
Dichomeris derasella ([Denis & Schiffermüller], 1775)
fasciella (Hübner, 1796)
unguiculatus (Fabricius, 1798)
coreanus Matsumura, 1931
paranthes Meyrick, 1936
Dichomeris limosellus (Schläger, 1849)
deflectivellus (Reutti, 1853)
Dichomeris nitiellus (Costantini, 1923)
Dichomeris rasilella (Herrich-Schäffer, 1854) [30]
lacrimella (Caradja, 1920)
insulella (Dumont, 1921)
occidentella (Zerny, 1927), subspecies
Dichomeris barbella ([Denis & Schiffermüller], 1775)
Dichomeris alacella (Zeller, 1839)
Dichomeris latipennella (Rebel, 1937)
scotosiella (Hackman, 1945)
piceana (Šulcs, 1968)
steueri Povolný, 1978
Anasphaltis Meyrick, 1925
Anasphaltis renigerellus (Zeller, 1839)
Acompsia Hübner, 1825 [31]
Brachycrossata Heinemann, 1870
Telephila Meyrick, 1923
Acompsia cinerella (Clerck, 1759)
murinella (Scopoli, 1763)
ardeliella (Hübner, 1817)
cinerea (Haworth, 1828), emendation
spodiella (Treitschke, 1833)
Acompsia pyrenaella Huemer & Karsholt, 2002 [32]
Acompsia antirrhinella Millière, 1866 [33]
Acompsia baldizzonei Pinzari, Nel & Pinzari, 2016
Acompsia maculosella (Stainton, 1851) [34]
Acompsia dimorpha Petry, 1904
Acompsia subpunctella Svensson, 1966
Acompsia delmastroella Huemer, 1998
Acompsia muellerrutzi Wehrli, 1925
Acompsia caucasella Huemer & Karsholt, 2002
Acompsia minorella Rebel, 1899
Acompsia tripunctella ([Denis & Schiffermüller], 1775) [35]
Acompsia ponomarenkoae Huemer & Karsholt, 2002
Acompsia schmidtiellus (Heyden, 1848)
durdhamellus (Stainton, 1849)
quadrinella (Herrich-Schäffer, 1854)
Brachmia Hübner, 1825 [36]
Claododes Heinemann, 1870, homonym
Eudodacles Snellen, 1889
Aulacomima Meyrick, 1904
Apethistis Meyrick, 1908
Brachmia dimidiella ([Denis & Schiffermüller], 1775) [37]
costiguttella (Lienig & Zeller, 1846)
kneri (Nowicki, 1864)
Brachmia blandella (Fabricius, 1798)
gerronella (Zeller, 1850)
Brachmia procursella Rebel, 1903
Brachmia inornatella (Douglas, 1850)
Helcystogramma Zeller, 1877
Ceratophora Heinemann, 1870, homonym
Dectobathra Meyrick, 1904
Teuchophanes Meyrick, 1914
Schemataspis Meyrick, 1918
Parelectra Meyrick, 1925, homonym
Psamathoscopa Meyrick, 1937
Anathyrsotis Meyrick, 1939
Parelectroides Clarke, 1952
Onebala auct.
Helcystogramma lineolella (Zeller, 1839)
Helcystogramma triannulella (Herrich-Schäffer, 1854)
sepiella (Steudel, 1866)
cinerea (Caradja, 1931)
macroscopa (Meyrick, 1932)
Helcystogramma lutatella (Herrich-Schäffer, 1854)
Helcystogramma rufescens (Haworth, 1828)
simplella (Eversmann, 1844)
diaphanella (Lienig & Zeller, 1846)
isabella (Stainton, 1849)
rufescentella (Doubleday, 1859), emendation
Helcystogramma albinervis (Gerasimov, 1929)
Helcystogramma arulensis (Rebel, 1929)
Helcystogramma klimeschi Ponomarenko & Huemer, 2001
Helcystogramma flavescens Junnilainen, 2010
Helcystogramma convolvuli (Walsingham, 1908)
chrypsilychna (Meyrick, 1914)
dryadopa (Meyrick, 1918)
effera (Meyrick, 1918)
emigrans (Meyrick, 1921)
Helcystogramma lamprostoma (Zeller, 1847) [38]
scutata (Meyrick, 1894)
Pseudosophronia Corley, 2001 [39]
Pseudosophronia exustellus (Zeller, 1847)
catharurga Meyrick, 1923
parahumerella Amsel, 1935
buvati Nel, 1998
constanti Nel, 1998, syn. nov.
Pseudosophronia cosmella (Constant, 1885)
Apatetrinae Le Marchand, 1947
Chrysoesthiinae Paclt, 1947, unavailable
Pexicopiini Hodges, 1986
Harpagidia Ragonot, 1895
Glaphyrerga Meyrick, 1925
Harpagidia magnetella (Staudinger, 1871)
pallidibasella Ragonot, 1895
melitophanes (Meyrick, 1931)
Pectinophora Busck, 1917
Pectinophora gossypiella (Saunders, 1844)
Pexicopia Common, 1958
Pexicopia malvella (Hübner, 1805) [40]
lutarea (Haworth, 1828), unavailable
umbrella auct.
Platyedra Meyrick, 1895
Aratrognathosia Gozmány, 1968, unavailable
Platyedra subcinerea (Haworth, 1828)
vilella (Zeller, 1847)
parviocellatella (Bruand d’Uzelle, 1851)
bathrosticta (Meyrick, 1937)
Sitotroga Heinemann, 1870
Nesolechia Meyrick, 1921
Syngenomictis Meyrick, 1927
Sitotroga psacasta Meyrick, 1908
celyphodes (Meyrick, 1909)
nea Walsingham, 1920
Sitotroga cerealella (Olivier, 1789)
hordei (Kirby, 1815)
arctella (Walker, 1864)
melanarthra (Lower, 1900)
palearis (Meyrick, 1913)
aenictopa (Meyrick, 1927)
ochrescens (Meyrick, 1938)
asemodes (Meyrick, 1938)
Apatetrini Le Marchand, 1947 [41]
Dactylotula Cockerell, 1888
Dactylota Snellen, 1876, homonym
Didactylota Walsingham, 1892
Rotundivalva Janse, 1951
Dactylotula altithermella (Walsingham, 1903)
Dactylotula kinkerella (Snellen, 1876) [42]
Apatetris Staudinger, 1879 [43]
Apatetris agenjoi Gozmány, 1954
Apatetris mediterranella Nel & Varenne, 2012 [44]
Catatinagma Rebel, 1903
Catatinagma trivittellum Rebel, 1903 [45]
Catatinagma kraterella Junnilainen & Nupponen, 2010 [46]
Coloptilia Fletcher, 1940
Colopteryx Hofmann, 1898, homonym
Coloptilia conchylidella (Hofmann, 1898)
Chrysoesthia Hübner, 1825 [47]
Microsetia Stephens, 1829
Chrysia Bruand d’Uzelle, 1851
Nomia Clemens, 1860, homonym
Chrysopora Clemens, 1860
Nannodia Heinemann, 1870
Anaphaula Walsingham, 1904
Chrysoesthia drurella (Fabricius, 1775) [48]
myllerella (Fabricius, 1794)
zinckenlla (Hübner, 1813)
druryella (Zeller, 1851), emendation
hermannella auct.
Chrysoesthia eppelsheimi (Staudinger, 1885)
Chrysoesthia verrucosa Tokár, 1999
Chrysoesthia sexguttella (Thunberg, 1794)
auropunctella (Thunberg, 1794)
aurofasciella (Stephens, 1834)
naeviferella (Duponchel, 1843)
stipella auct.
Chrysoesthia halimionella Bidzilya & Budashkin, 2015
Chrysoesthia atriplicella (Amsel, 1939) [49]
Chrysoesthia gaditella (Staudinger, 1859) [49]
Chrysoesthia aletris (Walsingham, 1919) [49]
Chrysoesthia boseae (Walsingham, 1908)
Chrysoesthia falkovitshi Lvovsky & Piskunov, 1989
Chrysoesthia hispanica Karsholt & Vives, 2014
Metanarsia Staudinger, 1871
Calyptrotis Meyrick, 1891
Epipararsia Rebel, 1914
Parametanarsia Gerasimov, 1930
Metanarsia modesta Staudinger, 1871 [50]
kurdistanella Amsel, 1959, subspecies
Metanarsia onzella Christoph, 1887
Metanarsia guberlica Nupponen, 2010
Metanarsia incertella (Herrich-Schäffer, 1861)
longivitella (Rebel, 1914)
halmyropis (Meyrick, 1926)
ramiferella (Lucas, 1940)
Oecocecis Guenée, 1870
Oecocecis guyonella Guenée, 1870 [51]
Thiotrichinae Karsholt, Mutanen, Lee & Kaila, 2013 [52]
Palumbininae Chapman, 1902, nomen nudum
Thiotricha Meyrick, 1886
Reuttia Hofmann, 1898
Mystax Caradja, 1920, homonym
Thiotricha majorella Rebel, 1910
Thiotricha subocellea (Stephens, 1834)
internella (Lienig & Zeller, 1846)
dissonella (Herrich-Schäffer, 1854)
subocellella (Doubleday, 1859), emendation
Thiotricha coleella (Constant, 1885)
Thiotricha wollastoni (Walsingham, 1884)
Palumbina Rondani, 1876
Thyrsostoma Meyrick, 1907
Palumbina guerinii (Stainton, 1858)
terebintella Rondani, 1876
pistaciae (Anagnostopoulos, 1935)
Anomologinae Meyrick, 1926
Aristoteliinae Le Marchand, 1947
Metzneriini Piskunov, 1975
Isophrictini Povolný, 1979
Bryotropha Heinemann, 1870 [53]
Mniophaga Pierce & Daltry, 1938
Adelphotropha Gozmány, 1955
Bryotropha sabulosella (Rebel, 1905)
Bryotropha domestica (Haworth, 1828)
domesticella (Doubleday, 1859), emendation
punctata (Staudinger, 1876)
salmonis (Walsingham, 1908)
algiricella Chrétien, 1917
Bryotropha vondermuhlli Nel & Brusseaux, 2003
Bryotropha rossica Anikin & Piskunov, 1996
tachengensis Li & Zheng, 1997
Bryotropha azovica Bidzilia, 1997
Bryotropha arabica Amsel, 1952
Bryotropha patockai Elsner & Karsholt, 2003
Bryotropha purpurella (Zetterstedt, 1839)
flavipalpella (Nylander, 1848)
Bryotropha tachyptilella (Rebel, 1916)
Bryotropha italica Karsholt & Rutten, 2005
Bryotropha politella (Stainton, 1851)
expolitella (Doubleday, 1859)
Bryotropha aliterrella (Rebel, 1935)
Bryotropha nupponeni Karsholt & Rutten, 2005
Bryotropha satschkovi Anikin & Piskunov, 2018
Bryotropha terrella ([Denis & Schiffermüller], 1775) [54]
inulella (Hübner, 1805)
pauperella (Hübner, 1825)
latella (Herrich-Schäffer, 1854)
lutescens (Constant, 1865)
suspectella (Heinemann, 1870)
alpicolella Heinemann, 1870
tenebrosella (Teich, 1886)
sardoterrella Schawerda, 1936
quignoni Dufrane, 1938, unavailable
joannisi Dufrane, 1938, unavailable
rufa Dufrane, 1938, unavailable
ochrea Dufrane, 1938, unavailable
Bryotropha sattleri Nel, 2003
Bryotropha desertella (Douglas, 1850) [55]
decrepidella (Herrich-Schäffer, 1854)
glabrella Heinemann, 1870
Bryotropha wolschrijni Karsholt & Rutten, 2005
Bryotropha heckfordi Karsholt & Rutten, 2005
Bryotropha figulella (Staudinger, 1859)
capnella (Constant, 1865)
cinnamomea Turati, 1934
Bryotropha plantariella (Tengström, 1848)
cinerosella (Tengström, 1848)
serrulatella (Tengström, 1848)
brevipalpella Rebel, 1893
Bryotropha galbanella (Zeller, 1839)
angustella (Heinemann, 1870)
ilmatariella (Hoffmann, 1893)
griseella (Caradja, 1920)
haareki (Strand, 1920)
fusconigratella (Palm, 1947)
Bryotropha boreella (Douglas, 1851)
Bryotropha sutteri Karsholt & Rutten, 2005
Bryotropha gallurella Amsel, 1952
Bryotropha hendrikseni Karsholt & Rutten, 2005
Bryotropha pallorella Amsel, 1952
mulinoides Amsel, 1952
zannonicola Hartig, 1953
Bryotropha hulli Karsholt & Rutten, 2005 [56]
Bryotropha plebejella (Zeller, 1847)
imperitella (Staudinger, 1859)
ancillula (Walsingham, 1908)
inexpectella Nel, 1999
Bryotropha dryadella (Zeller, 1850)
saralella Amsel, 1952
Bryotropha basaltinella (Zeller, 1839)
Bryotropha affinis (Haworth, 1828) [57]
tegulella (Herrich-Schäffer, 1854)
tectella (Herrich-Schäffer, 1854)
affinella (Doubleday, 1859), emendation
affinitella (Bruand d’Uzelle, 1859), emendation
Bryotropha umbrosella (Zeller, 1839) [58]
mundella (Douglas, 1850)
portlandicella (Richardson, 1890)
fulvipalpella Joannis, 1908
anacampsoidella (Hering, 1924)
oppositella auct.
Bryotropha similis (Stainton, 1854)
thuleella (Zeller, 1857)
similella (Doubleday, 1859), emendation
pullifimbriella (Clemens, 1863)
confinis (Stainton, 1871)
obscurecinerea (Nolcken, 1871)
stolidella (Morris, 1872)
fuliginosella (Snellen, 1882)
tahavusella (Forbes, 1922)
clandestina (Meyrick, 1923)
dufraneella (Joannis, 1928)
novisimilis Li & Zheng, 1997
Bryotropha senectella (Zeller, 1839)
ciliatella (Herrich-Schäffer, 1854)
obscurella Heinemann, 1870
minorella Heinemann, 1870
phoebusella Millière, 1876
larseni Strand, 1927
Epidola Staudinger, 1859 [59]
Epidola stigma Staudinger, 1859
Epidola barcinonella Millière, 1867
Epidola semitica Amsel, 1942 [60]
Epidola nuraghella Hartig, 1939
Epidola melitensis Amsel, 1955
Aristotelia Hübner, 1825 [61]
Ergatis Heinemann, 1870, homonym
Eucatoptus Walsingham, 1897
Aristotelia decurtella (Hübner, 1813) [62]
turbatella (Treitschke, 1835)
amoenella (Joannis,1891)
Aristotelia decoratella (Staudinger, 1879)
Aristotelia leonhardi Krone, 1907
Aristotelia ericinella (Zeller, 1839) [63]
silendrella Caradja, 1920, unavailable
Aristotelia subdecurtella (Stainton, 1859) [64]
Aristotelia subericinella (Duponchel, 1843) [65]
prohaskaella (Rebel, 1907)
Aristotelia billii Varenne & Nel, 2013 [66]
Aristotelia montarcella Schmidt, 1941
Aristotelia heliacella (Herrich-Schäffer, 1854)
rogenhoferi (Staudinger, 1872)
Aristotelia pancaliella (Staudinger, 1871)
Aristotelia baltica Šulcs & Šulcs, 1983
coeruleopictella auct.
Aristotelia brizella (Treitschke, 1833)
Aristotelia brizelloidea Amsel, 1935
Aristotelia confusella Bidzilya & Budashkin, 2015
Aristotelia staticella Millière, 1876
Aristotelia mirandella Chrétien, 1908
Aristotelia frankeniae Walsingham, 1898
Aristotelia calastomella (Christoph, 1873)
Aristotelia mirabilis (Christoph, 1888)
Caulastrocecis Chrétien, 1931 [67]
Caulastrocecis pudicellus (Mann, 1861)
apicella (Caradja, 1920)
Caulastrocecis gypsella (Constant, 1893)
Caulastrocecis furfurella (Staudinger, 1871) [68]
Caulastrocecis cryptoxena (Gozmány, 1952) sp. rev. [68]
Caulastrocecis perexigella Junnilainen, 2010
Caulastrocecis interstratella (Christoph, 1873)
salinatrix (Meyrick, 1926)
Paranarsia Ragonot, 1895 [69]
Paranarsia joannisiella Ragonot, 1895
Megacraspedus Zeller, 1839 [70]
Chilopselaphus Mann, 1867
Chilopsephalus Rebel, 1901, misspelling
Toxoceras Chrétien, 1915, homonym
Toxidoceras Chrétien, 1923
Nevadia Caradja, 1920, homonym
Cauloecista Dumont, 1928
Reichardtiella Filipjev, 1931
Vadenia Caradja, 1933
Megacraspedus lanceolellus (Zeller, 1850) [71]
subdolellus Staudinger, 1859
hessleriellus Rössler, 1868
tutti Walsingham, 1897
grossisquammellus Chrétien, 1925
Megacraspedus bengtssoni Huemer & Karsholt, 2018
Megacraspedus junnilaineni Huemer & Karsholt, 2018
Megacraspedus uzunsyrtus Bidzilya & Budashkin, 2015
Megacraspedus similellus Huemer & Karsholt, 2018
Megacraspedus tokari Huemer & Karsholt, 2018
Megacraspedus dolosellus (Zeller, 1839) [72]
separatellus (Fischer von Röslerstamm, 1843)
incertellus Rebel, 1930
Megacraspedus neli Huemer & Karsholt, 2018
Megacraspedus faunierensis Huemer & Karsholt, 2018
Megacraspedus gredosensis Huemer & Karsholt, 2018
Megacraspedus cuencellus Caradja, 1920
Megacraspedus bidentatus Huemer & Karsholt, 2018
Megacraspedus fuscus Huemer & Karsholt, 2018
Megacraspedus trineae Huemer & Karsholt, 2018
Megacraspedus tristictus Walsingham, 1910
Megacraspedus alfacarellus Wehrli, 1926
Megacraspedus pusillus Walsingham, 1903
Megacraspedus skoui Huemer & Karsholt, 2018
Megacraspedus spinophallus Huemer & Karsholt, 2018 [73]
Megacraspedus occidentellus Huemer & Karsholt, 2018
Megacraspedus granadensis Huemer & Karsholt, 2018
Megacraspedus heckfordi Huemer & Karsholt, 2018
Megacraspedus tenuiuncus Huemer & Karsholt, 2018
Megacraspedus lativalvellus Amsel, 1954
Megacraspedus dejectella (Staudinger, 1859)
Megacraspedus devorator Huemer & Karsholt, 2018
Megacraspedus binotella (Duponchel, 1843) [74]
Megacraspedus brachypteris Huemer & Karsholt, 2018 [75]
Megacraspedus barcodiellus Huemer & Karsholt, 2018
Megacraspedus bilineatella Huemer & Karsholt, 1996
Megacraspedus andreneli Varenne & Nel, 2014 [76]
Megacraspedus sumpichi Huemer & Karsholt, 2018
Megacraspedus gallicus Huemer & Karsholt, 2018
Megacraspedus ribbeella (Caradja, 1920)
Megacraspedus numidellus (Chrétien, 1915)
mareotidellus Turati, 1924,
Megacraspedus albovenata Junnilainen, 2010
Megacraspedus longipalpella Junnilainen, 2010
Megacraspedus niphorrhoa (Meyrick, 1926)
Megacraspedus fallax (Mann, 1867)
Megacraspedus balneariellus (Chrétien, 1907)
Megacraspedus podolicus (Toll, 1942)
Megacraspedus knudlarseni Huemer & Karsholt, 2018
Megacraspedus imparellus (Fischer v. Röslerstamm, 1843) [77]
litovalvellus Junnilainen, 2010
Megacraspedus multispinella Junnilainen & Nupponen, 2010
Megacraspedus cerussatellus Rebel, 1930
Megacraspedus attritellus Staudinger, 1871
Megacraspedus lagopellus (Herrich-Schäffer, 1860)
Megacraspedus argyroneurellus Staudinger, 1871
Megacraspedus ibericus Huemer & Karsholt, 2018
Megacraspedus squalida Meyrick, 1926
escalerellus Schmidt, 1941
Megacraspedus pentheres Walsingham, 1920
Megacraspedus teriolensis Huemer & Karsholt, 2018 [78]
Megacraspedus korabicus Huemer & Karsholt, 2018
Megacraspedus quadristictus Lhomme, 1946
Megacraspedus eburnellus Huemer & Karsholt, 2001
Megacraspedus skulei Huemer & Karsholt, 2018
Megacraspedus peyerimhoffi Le Cerf, 1925
Megacraspedus peslieri Huemer & Karsholt, 2018
Dirhinosia Rebel, 1905 [79]
Dirhinosia cervinella (Eversmann, 1844)
trifasciella Rebel, 1905
Dirhinosia arnoldiella (Rebel, 1905)
Dirhinosia interposita Bidzilya & Budashkin, 2015
Psamathocrita Meyrick, 1925 [80]
Psamathocrita osseella (Stainton, 1860)
Psamathocrita argentella Pierce & Metcalfe, 1942
Psamathocrita dalmatinella Huemer & Tokár, 2000
Chimericorsa Varenne, Huemer & Nel, 2017
Chimericorsa nioloensis Varenne, Huemer & Nel, 2017
Spiniphallellus Bidzilya & Karsholt, 2008
Spiniphallellus desertus Bidzilya & Karsholt, 2008
Spiniphallellus chrysotosella Junnilainen, 2016
Deltophora Janse, 1950
Deltophora maculata (Staudinger, 1879)
Deltophora stictella (Rebel, 1927)
Deltophora gielisia Hull, 1995
Ivanauskiella Ivinskis & Piskunov, 1980 [81]
Spatuncusella Nel & Varenne, 2013
Ivanauskiella psamathias (Meyrick, 1891)
turkmenica auct.
Ivanauskiella occitanica (Nel & Varenne, 2013) sp. rev. [82]
Ptocheuusa Heinemann, 1870 [83]
Syneunetis Wallengren, 1881
Ptocheuusa paupella (Zeller, 1847) [84]
inulella (Curtis, 1850)
melanolepidella (Heydenreich, 1851)
perniveella (Bruand d’Uzelle, 1859)
Ptocheuusa inopella (Zeller, 1839) [85]
amesella Chrétien, 1908
Ptocheuusa abnormella (Herrich-Schäffer, 1854)
Ptocheuusa minimella (Rebel, 1936)
Ptocheuusa asterisci (Walsingham, 1903)
Ptocheuusa scholastica (Walsingham, 1903)
Ptocheuusa guimarensis (Walsingham, 1908)
Ptocheuusa sublutella Christoph, 1873
Ptocheuusa cinerella (Chrétien, 1908) comb. nov. [86]
Gladiovalva Sattler, 1960
Gladiovalva rumicivorella (Millière, 1881)
Gladiovalva aizpuruai Vives, 1990
Gladiovalva badidorsella (Rebel, 1935)
Ornativalva Gozmány, 1955
Pelostola Janse, 1960
Ornativalva heluanensis (Debski, 1913)
frankeniivorella (Chrétien, 1917)
oasicolella (Turati, 1924)
siculella (Mariani, 1937)
Ornativalva ornatella Sattler, 1967
Ornativalva tamariciella (Zeller, 1850)
Ornativalva pseudotamariciella Sattler, 1967
Ornativalva antipyramis (Meyrick, 1925)
Ornativalva plutelliformis (Staudinger, 1859)
olbiaella (Millière, 1861)
siewersiellus (Christoph, 1867)
sinuatella (Walsingham, 1904)
Ornativalva sieversi (Staudinger, 1871)
Ornativalva mixolitha (Meyrick, 1918)
bipunctella (Sattler, 1967), subspecies
Atremaea Staudinger, 1871
Calamotypa Meyrick, 1926
Atremaea lonchoptera Staudinger, 1871
exstans (Meyrick, 1926)
Amblypalpis Ragonot, 1886 [87]
Amblypalpis olivierella Ragonot, 1887
Parapodia Joannis, 1912 [88]
Cecidonostola Amsel, 1958
Parapodia sinaica (Frauenfeld, 1860)
tamaricicola Joannis, 1912
tamariciella (Amsel, 1958)
Isophrictis Meyrick, 1917 [89]
Isophrictis robinella (Chrétien, 1907)
microlina Meyrick, 1935
Isophrictis meridionella (Herrich-Schäffer, 1854)
Isophrictis constantina (Baker, 1888)
Isophrictis cerdanica Nel, 1995
Isophrictis lineatellus (Zeller, 1850)
albilineella (Bruand d´Uzelle, 1859)
Isophrictis kefersteiniellus (Zeller, 1850) [90]
senicula (Meyrick, 1913)
Isophrictis striatella ([Denis & Schiffermüller], 1775)
tanacetella (Schrank, 1802)
substriatella (Caradja, 1920), subspecies
Isophrictis corsicella Amsel, 1936
Isophrictis invisella (Constant, 1885)
Isophrictis anthemidella (Wocke, 1871) [91]
Isophrictis impugnata Gozmány, 1957
Pyncostola Meyrick, 1917
Pyncostola bohemiella (Nickerl, 1864)
tunesiella (Chrétien, 1915)
jablonkayi (Gozmány, 1954)
Metzneria Zeller, 1839 [92]
Cleodora Stephens, 1834, homonym
Parasia Duponchel, 1846
Archimetzneria Amsel, 1936
Metzneria paucipunctella (Zeller, 1839)
zimmermanni Hering, 1940
confusalis Lucas, 1956
luqueti Nel, 1995
Metzneria tenuiella (Mann, 1864)
seminivora (Walsingham, 1903)
infelix Walsingham, 1908
insignificans Walsingham, 1908
Metzneria neuropterella (Zeller, 1839)
nevropterella (Duponchel, 1843) [93]
gigantella Krulikowsky, 1909, unavailable
Metzneria aestivella (Zeller, 1839) [94]
carlinella (Stainton, 1851)
selaginella (Mann, 1855)
torridella (Mann, 1859)
dichroa Walsingham, 1908, subspecies.
expositoi Vives, 2001, syn. nov.
Metzneria lappella (Linnaeus, 1758)
Metzneria castiliella (Möschler, 1866)
eatoni Walsingham, 1899
Metzneria littorella (Douglas, 1850)
quinquepunctella (Herrich-Schäffer, 1854)
Metzneria riadella Englert, 1974
Metzneria diffusella Englert, 1974
Metzneria fulva Labonne, Huemer, Thibault & Nel, 2019 [95]
Metzneria torosulella (Rebel, 1893) [95]
monochroa Walsingham, 1908
ignota Turati, 1922
Metzneria ehikeella Gozmány, 1954 [96]
Metzneria metzneriella (Stainton, 1851) [97]
falcatella (Bruand d´Uzelle, 1859)
Metzneria hilarella Caradja, 1920
Metzneria staehelinella Englert, 1974
Metzneria artificella (Herrich-Schäffer, 1861) [98]
litigiosella (Millière, 1879)
pannonicella Rebel, 1915
Metzneria agraphella (Ragonot, 1895)
incognita Walsingham, 1904
Metzneria aprilella (Herrich-Schäffer, 1854) [99]
igneella (Tengström, 1859)
sanguinolentella Joannis, 1910
Metzneria subflavella Englert, 1974 [100]
Metzneria filia Piskunov, 1979
Metzneria intestinella (Mann, 1864)
Metzneria santolinella (Amsel, 1936)
consimilella Hackman, 1946
Metzneria tristella Rebel, 1901
Metzneria campicolella (Mann, 1857) [101]
varennei Nel, 1997
Apodia Heinemann, 1870
Apodia bifractella (Duponchel, 1843)
inulella (Vallot, 1829), homonym
Apodia martinii Petry, 1911 sp. rev. [102]
Pragmatodes Walsingham, 1908 [103]
Pragmatodes fruticosella Walsingham, 1908
Pragmatodes melagonella (Constant, 1895) comb. nov. [103, 104]
Pragmatodes albagonella (Varenne & Nel, 2010) comb. nov. [103]
Pragmatodes cyrneogonella (Nel & Varenne, 2012) comb. nov. [103]
Pragmatodes parvulata (Gozmány, 1953) comb. nov. [103]
mediterranea (Nel & Luquet, 1997)
Argolamprotes Benander, 1945
Argolamprotes micella ([Denis & Schiffermüller], 1775)
asterella (Treitschke, 1833)
Monochroa Heinemann, 1870 [105]
Paltodora Meyrick, 1894
Catabrachmia Rebel, 1909
Monochroa rumicetella (Hofmann, 1868) [106]
acutangulella (Heinemann, 1870)
leptotechna (Meyrick, 1937)
Monochroa rebeli (Hering, 1927)
Monochroa sepicolella (Herrich-Schäffer, 1854) [107]
Monochroa rectifasciella (Fuchs, 1902) [107]
Monochroa tenebrella (Hübner, 1817) [108]
fuscocuprea (Haworth, 1828)
subcuprella (Stephens, 1834)
tenebrosella (Zeller, 1839)
parvella (Heydenreich, 1851)
fuscocuprella Doubleday, 1859, emendation
buffonella (Millière, 1876)
Monochroa scutatella (Müller-Rutz, 1920)
Monochroa dellabeffai (Rebel, 1932)
Monochroa servella (Zeller, 1839) [109]
farinosae (Stainton,1867)
Monochroa conspersella (Herrich-Schäffer, 1854)
questionella (Herrich-Schäffer, 1854)
morosa (Mühlig, 1864)
Monochroa tetragonella (Stainton, 1885)
gudmanni (Larsen, 1927)
Monochroa elongella (Heinemann, 1870)
micrometra (Meyrick, 1935)
Monochroa inflexella Svensson, 1992
Monochroa sperata Huemer & Karsholt, 2010
Monochroa lutulentella (Zeller, 1839)
brunickii (Rebel, 1913)
Monochroa aenigma Anikin & Piskunov, 2018
Monochroa saltenella (Benander, 1928)
Monochroa palustrellus (Douglas, 1850)
rozsikella (Rebel, 1909)
Monochroa divisella (Douglas, 1850)
csornensis Rebel, 1909
lepidolampra (Gozmány, 1952)
zarichella Piskunov, 1975
Monochroa lucidella (Stephens, 1834) [110]
scordiscella (Rebel, 1904)
unipunctella (Amsel, 1935)
immaculatella Huemer, 1996, subspecies
Monochroa simplicella (Lienig & Zeller, 1846)
impella (Piskunov, 1975)
Monochroa moyses Uffen, 1991
Monochroa arundinetella (Boyd, 1857) [111]
Monochroa suffusella (Douglas, 1850) [111]
oblitella (Doubleday, 1859)
peterseni (Teich, 1901)
Monochroa cytisella (Curtis, 1837)
fuscipennis (Humphreys & Westwood, 1845)
walkeriella (Douglas, 1850)
coenulentella (Herrich-Schäffer, 1854)
clinosema (Meyrick, 1935)
griseocapitella (Bentinck, 1949), unavailable
Monochroa ferrea (Frey, 1870)
latiuscula (Heinemann, 1870)
alfkeni (Amsel, 1938)
servella auct.
Monochroa nomadella (Zeller, 1868) [112]
Monochroa bronzella Karsholt, Nel, Fournier, Varenne & Huemer, 2013
Monochroa hornigi (Staudinger, 1883)
leptocrossa (Meyrick, 1926)
nordmanella Bruun, 1958
Monochroa niphognatha (Gozmány, 1953)
Oxypteryx Rebel, 1911 [113]
Eulamprotes Bradley, 1971
Lamprotes Heinemann, 1870, homonym
Argyritis Heinemann, 1870, homonym
Siderea Omelko, 1999
Oxypteryx nigromaculella (Millière, 1872) comb. nov. [113, 114]
punctatella (Staudinger, 1879)
morphochroma (Walsingham, 1900)
jactatrix (Meyrick, 1926)
angustipennis (Rebel, 1931)
craterotypa (Meyrick, 1939)
donskoffi (Nel & Luquet, 1997)
Oxypteryx wilkella (Linnaeus, 1758) comb. nov. [113, 115]
merianella (Linnaeus, 1758)
germarella (Geyer, 1832)
pictella (Zeller, 1839)
tarquiniella (Stainton, 1862)
Oxypteryx ochricapilla (Rebel, 1903) comb. nov. [113]
Oxypteryx superbella (Zeller, 1839) comb. nov. [113]
Oxypteryx mirusella (Huemer & Karsholt, 2013) comb. nov. [113]
Oxypteryx baldizzonei (Karsholt & Huemer, 2013) comb. nov. [113, 116]
Oxypteryx occidentella (Huemer & Karsholt, 2011) comb. nov. [113]
Oxypteryx libertinella (Zeller, 1872) comb. nov. [113, 117]
Oxypteryx gemerensis (Elsner, 2013) comb. nov. [113]
Oxypteryx deserta (Piskunov, 1990) comb. nov. [113]
Oxypteryx unicolorella (Duponchel, 1843) comb. nov. [113]
lucentella (Peyerimhoff, 1870)
Oxypteryx atrella ([Denis & Schiffermüller], 1775)
quadripunctella (Fabricius, 1781)
umbriferella (Herrich-Schäffer, 1854)
aurimaculella (Höfner, 1897)
ornata (Dufrane, 1942), unavailable
Oxypteryx nigritella (Zeller, 1847) comb. nov. [113]
Oxypteryx immaculatella (Douglas, 1850)
phaeella (Heckford & Langmaid, 1988)
Oxypteryx plumbella (Heinemann, 1870) comb. nov. [113]
Oxypteryx isostacta (Meyrick, 1926) comb. nov. [113]
Oxypteryx helotella (Staudinger, 1859) comb. nov. [113]
damonella (Millière, 1876)
algeriella (Baker, 1888)
doliodes (Meyrick, 1891)
striatopunctella (Rebel, 1891)
levisella (Chrétien, 1922)
Oxypteryx parahelotella (Nel, 1995) comb. nov. [113]
Oxypteryx graecatella (Šumpich & Skyva, 2012) comb. nov. [113]
Gelechiinae Stainton, 1954
Gelechiini Stainton, 1954
Xystophora Wocke, 1876
Doryphora Heinemann, 1870, homonym
Doryphorella Cockerell, 1888
Xystophora carchariella (Zeller, 1839)
Xystophora pulveratella (Herrich-Schäffer, 1854)
intaminatella (Stainton, 1860)
steudeliella (Frey, 1880)
Athrips Billberg, 1820 [118]
Rhynchopacha Staudinger, 1871
Epithectis Meyrick, 1895
Leobatus Walsingham, 1904
Ziminiola Gerasimov, 1930
Cremona Busck, 1934
Athrips spiraeae (Staudinger, 1871)
Athrips pruinosella (Lienig & Zeller, 1846)
Athrips rancidella (Herrich-Schäffer, 1854) [119]
triatomaea (Mühlig, 1864)
vepretella (Zeller, 1870)
superfetella (Peyerimhoff, 1877)
cotoneastri (Busck, 1934)
haifella Amsel, 1935
cerasivorella (Kuznetsov, 1960)
Athrips thymifoliella (Constant, 1893)
Athrips amoenella (Frey, 1882) [120]
allgunnensis Svensson, 1993, unavailable
Athrips nigricostella (Duponchel, 1842)
Athrips tetrapunctella (Thunberg, 1794)
lathyri (Stainton, 1865)
lathyrella (Doubleday, 1866), emendation
Athrips mouffetella (Linnaeus, 1758)
pedisequella ([Denis & Schiffermüller], 1775)
punctifera (Haworth, 1828)
Athrips asarinella (Chrétien, 1930)
Athrips medjella (Chrétien, 1900)
Athrips patockai (Povolný, 1979)
Athrips polymaculella Park, 1991
Athrips stepposa Bidzilya, 2005
Athrips aquila Junnilainen, 2010
Athrips bidzilyai Junnilainen, 2010
Athrips fagoniae (Walsingham, 1904)
Neofriseria Sattler, 1960
Neofriseria peliella (Treitschke, 1835) [121]
senecionella (Bruand d´Uzelle, 1859)
Neofriseria singula (Staudinger, 1876)
suppeliella (Walsingham, 1896)
amaurella (Rebel, 1927), homonym
ifranella (Lucas, 1956)
hispanicella (Amsel, 1953)
Neofriseria pseudoterrella (Rebel, 1928)
Neofriseria baungaardiella Huemer & Karsholt, 1999
Neofriseria hitadoella Karsholt & Vives, 2014 [122]
Neofriseria kuznetzovae Bidzilya, 2002 [123]
Neofriseria caucasicella Sattler, 1960
Neofriseria mongolinella Piskunov, 1987
Prolita Leraut, 1993
Lita Treitschke, 1833, homonym
Prolita sexpunctella (Fabricius, 1794)
virgella (Thunberg, 1794)
longicornis (Curtis, 1827)
longicornella (Doubleday, 1859), emendation
histrionella (Geyer, 1832)
zebrella (Treitschke, 1833)
alpicolo (Frey, 1867)
alternatella (Kearfott, 1908)
melanica (Strand, 1920), unavailable
petulans (Braun, 1925)
Prolita solutella (Zeller, 1839)
fumosella (Douglas, 1852)
cornubiae (Boyd, 1858)
pribitzeri (Rebel, 1889)
nigrobipunctatella (Lucas, 1932)
Sophronia Hübner, 1825 [124]
Sophronia semicostella (Hübner, 1813) [125]
marginella (Thunberg, 1794), homonym
Sophronia gelidella Nordman, 1941
Sophronia consanguinella Herrich-Schäffer, 1854 [126]
marginella Toll, 1936
Sophronia illustrella (Hübner, 1796)
Sophronia grandii Hering, 1933 [127]
ascalis Gozmány, 1951, syn. nov.
Sophronia chilonella (Treitschke, 1833) [128]
Sophronia finitimella Rebel, 1905
Sophronia acaudella Rebel, 1903
Sophronia curonella Standfuss, 1884
Sophronia humerella ([Denis & Schiffermüller], 1775)
Sophronia sicariellus (Zeller, 1839) [129]
Sophronia santolinae Staudinger, 1863
Mirificarma Gozmány, 1955 [130]
Helina Guenée, 1849, homonym
Mirificarma rhodoptera (Mann, 1866)
Mirificarma minimella Huemer & Karsholt, 2001
Mirificarma denotata Pitkin, 1984
Mirificarma maculatella (Hübner, 1796)
Mirificarma aflavella (Amsel, 1935)
Mirificarma flavella (Duponchel, 1844)
segetella (Zeller, 1847)
Mirificarma eburnella ([Denis & Schiffermüller], 1775)
formosella (Hübner, 1796), homonym
flammella (Hübner, 1825)
rufeoformosella (Bruand d´Uzelle, 1859)
Mirificarma fasciata Pitkin, 1984
Mirificarma lentiginosella (Zeller, 1839) [131]
Mirificarma pederskoui Huemer & Karsholt, 1999
Mirificarma cytisella (Treitschke, 1833) [132]
roseella (Hauder, 1918), unavailable
leonella Amsel, 1959, subspecies
Mirificarma monticolella (Rebel, 1931) [133]
Mirificarma interrupta (Curtis, 1827)
interuptella (Hübner, 1793), homonym
Mirificarma burdonella (Rebel, 1930) [134]
Mirificarma cabezella (Chrétien, 1925)
Mirificarma ulicinella (Staudinger, 1859) [135]
Mirificarma mulinella (Zeller, 1839)
caminariella (Fuchs, 1902)
nigraesilvae (Amsel, 1950)
Aroga Busck, 1914
Aroga velocella (Zeller, 1839) [136]
affiniella (Zetterstedt, 1839)
tesserella (Zetterstedt, 1839)
brunnea (Schöyen, 1882)
aterrimella (Rebel, 1889)
peperistis (Meyrick, 1926)
rupicolella (Müller-Rutz, 1934)
Aroga flavicomella (Zeller, 1839) [137]
aureodorsella (Bruand d´Uzelle, 1859)
Aroga eatoni Corley & Goodey, 2014
Aroga pascuicola (Staudinger, 1871)
eremella (Chrétien, 1915)
Aroga aristotelis (Millière, 1876)
astragali (Staudinger, 1879)
fulminella (Millière, 1882)
lacertella (Walsingham, 1904)
aplasticella (Rebel, 1913), unavailable
hyrcanella (Toll, 1948)
Aroga corsa Varenne & Nel, 2019
Aroga temporariella Sattler, 1960
Aroga balcanicola Huemer & Karsholt, 1999
Filatima Busck, 1939
Filatima angustipennis Sattler, 1961
albicosta auct.
Filatima pallipalpella (Snellen, 1884)
Filatima spurcella (Duponchel, 1843)
fuscantella (Heinemann, 1870)
Filatima transsilvanella Kovács & Kovács, 2002
Filatima algarbiella Corley, 2014
Filatima tephritidella (Duponchel, 1844)
tephriditella (Herrich-Schäffer, 1854)
Filatima textorella (Chrétien, 1908)
Filatima djakovica Anikin & Piskunov, 1996
Filatima incomptella (Herrich-Schäffer, 1854)
turbidella (Nolcken, 1871)
Filatima ukrainica Piskunov, 1971
Filatima zagulajevi Anikin & Piskunov, 1996
Chionodes Hübner, 1825 [138]
Chionodes lugubrella (Fabricius, 1794)
luctificella (Hübner, 1813)
lunatella (Zetterstedt, 1839)
Chionodes tragicella (Heyden, 1865)
libidinosa (Staudinger, 1871)
Chionodes soella Huemer & Sattler, 1995
Chionodes luctuella (Hübner, 1793) [139]
sauteriella (Zeller, 1868)
Chionodes aprilella Huemer & Sattler, 1995
Chionodes violacea (Tengström, 1848)
Chionodes mongolica Piskunov, 1979
ukrainica Piskunov, 1979
Chionodes holosericella (Herrich-Schäffer, 1854)
cognatella (Heinemann, 1870)
norvegiae (Strand, 1903)
dovrella (Grønlien, 1925)
meesi (Barca, 1932)
danieli (Osthelder, 1951)
Chionodes praeclarella (Herrich-Schäffer, 1854)
pergrandella (Rebel, 1917)
colorella (Caradja, 1920), unavailable
decolorella auct.
Chionodes caucasicella Huemer & Sattler, 1995
Chionodes nubilella (Zetterstedt, 1839)
tarandella (Wocke, 1864)
Chionodes continuella (Zeller, 1839)
brumella (Clemens, 1864)
trimaculella (Packard, 1867)
albomaculella (Chambers, 1875)
Chionodes perpetuella (Herrich-Schäffer, 1854)
Chionodes apolectella (Walsingham, 1900)
Chionodes distinctella (Zeller, 1839)
striolatella (Heinemann, 1870)
tristella (Teich, 1889)
indistinctella (Rebel, 1901)
latiorella (Amsel, 1939)
unicolor (Toll, 1948)
deserticola Piskunov, 1979
Chionodes hayreddini Koçak, 1986
ochripalpella (Frey, 1880), homonym
Chionodes hinnella (Rebel, 1935)
Chionodes bastuliella (Rebel, 1931)
Chionodes electella (Zeller, 1839)
Chionodes viduella (Fabricius, 1794)
leucomella (Quenzel, 1802)
luctiferella (Herrich-Schäffer, 1856)
labradoriella (Clemens, 1863)
Chionodes nebulosella (Heinemann, 1870)
Chionodes fumatella (Douglas, 1850) [140]
celerella (Stainton, 1851)
oppletella (Herrich-Schäffer, 1854)
reuttiella (Heinemann, 1870)
nigricans (Heinemann, 1870)
syrticola (Staudinger, 1871)
brunnea (Teich, 1901), homonym
carpella Piskunov, 1971
Chionodes ignorantella (Herrich-Schäffer, 1854)
ochrisignella (Nolcken, 1871)
Gelechia Hübner, 1825 [141]
Guenea Bruand d´Uzelle, 1851
Cirrha Chambers, 1872
Oeseis Chambers, 1875
Mesogelechia Omelko, 1986
Gelechia rhombella ([Denis & Schiffermüller], 1775)
rhombea (Haworth, 1828), emendation
axilella (Thunberg, 1794)
Gelechia scotinella Herrich-Schäffer, 1854
conspurcatella Heinemann, 1870
confusella Heinemann, 1870
kiesenwetteri Heuäcker, 1873
lakatensis Rebel, 1904
baueri (Rebel, 1917)
Gelechia senticetella (Staudinger, 1859) [142]
limitanella Rebel, 1904
nigrostriella (Zerny, 1936)
Gelechia obscuripennis (Frey, 1880) [143]
melanotica (Burmann, 1950), unavailable
albicans (Burmann, 1950), unavailable
Gelechia sabinellus (Zeller, 1839)
hoffmanniella (Strand, 1902)
corsella (Rebel, 1930)
kalevalella (Kanerva, 1936)
Gelechia atlanticella (Amsel, 1955)
Gelechia nervosella (Zerny, 1927)
thuriferella (Cleu, 1936)
Gelechia sororculella (Hübner, 1817)
Gelechia jakovlevi Krulikovsky, 1905
nigrovittata Schantz, 1971
Gelechia muscosella Zeller, 1839
griseella Caradja, 1920
Gelechia cuneatella Douglas, 1852
Gelechia aspoecki Huemer, 1992
Gelechia asinella (Hübner, 1796)
aurorella Frey, 1882
Gelechia hippophaella (Schrank, 1802)
basalis Stainton, 1854
acupediella Frey, 1870
Gelechia basipunctella Herrich-Schäffer, 1854
basiguttella Heinemann, 1870
albicans Heinemann, 1870
Gelechia nigra (Haworth, 1828)
cautella Zeller, 1839
Gelechia turpella ([Denis & Schiffermüller], 1775)
populella (Hübner, 1796)
nebulea (Haworth, 1828), unavailable
pinguinella (Treitschke, 1832)
kochiella (Herrich-Schäffer, 1854)
Gelechia rhombelliformis Staudinger, 1871
Gelechia sirotina Omelko, 1986
Gelechia sestertiella Herrich-Schäffer, 1854
Gelechia mediterranea Huemer, 1991
Gelechia dujardini Huemer, 1991
Psoricoptera Stainton, 1854
Psoricoptera speciosella Teich, 1893
Psoricoptera gibbosella (Zeller, 1839)
triorthias (Meyrick, 1935)
lepigreella (Lucas, 1938)
Agnippe Chambers, 1872 [144]
Evippe Chambers, 1873
Phaetusa Chambers, 1875, homonym
Tholerostola Meyrick, 1917
Agnippe echinuloides Bidzilya & Li, 2010
Agnippe lunaki (Rebel, 1941)
penicillata (Amsel, 1961)
Agnippe pseudolella (Christoph, 1888)
cephalella (Caradja, 1920)
Holcophora Staudinger, 1871 [145]
Aponoaea Walsingham, 1905
Holcophora statices Staudinger, 1871
Holcophora inderskella (Caradja, 1920) [146]
Holcophora obtusipalpis (Walsingham, 1905)
cinerellus (Turati, 1930)
Gnorimoschemini Povolný, 1964
Gnorimoschema Busck, 1900
Lerupsia Riedl, 1965
Neoschema Povolný, 1967
Gnorimoschema soffneri (Riedl, 1965)
antiquum Povolný, 1966
Gnorimoschema herbichii (Nowicki, 1864) [147]
pusillella (Rebel, 1893)
tengstroemiella (Joannis, 1910)
pazsiczkyi (Rebel, 1913)
parentesella (Toll, 1936)
tengstroemi (Hackman, 1946)
mongoliae Povolný, 1973, subspecies
kamchaticum Povolný, 1977, subspecies
Gnorimoschema bodillum Karsholt & Nielsen, 1974
Gnorimoschema nupponeni Huemer & Karsholt, 2010
Gnorimoschema robustella (Staudinger, 1871)
syrphetopa (Meryick, 1926)
Gnorimoschema steueri Povolný, 1975
Gnorimoschema epithymella (Staudinger, 1859)
brunneomaculella (Hackman, 1946), subspecies
boerneri (Amsel, 1952), subspecies
kirgisicum Povolný, 1994, subspecies
Gnorimoschema nordlandicolella (Strand, 1902)
cyceonodes (Meyrick, 1924)
eucausta (Meyrick, 1929)
fennicella (Hackman, 1946)
Gnorimoschema nilsi Huemer, 1996
Gnorimoschema valesiella (Staudinger, 1877)
diabolicella (Hering, 1924)
charcoti (Meyrick, 1934)
hackmani (Schantz, 1952)
Gnorimoschema streliciella (Herrich-Schäffer, 1854)
Gnorimoschema hoefneri (Rebel, 1909)
Scrobipalpopsis Povolný, 1967
Scrobipalpopsis petasitis (Pfaffenzeller, 1867)
petasitella (Staudinger, 1867)
petasitae (Heinemann, 1870), emendation
Tecia Povolný, 1973
Tecia solanivora (Povolný, 1973)
Scrobipalpa Janse, 1951 [148]
Ilseopsis Povolný, 1965
Euscrobipalpa Povolný, 1967
Ergasiola Povolný, 1967
*Scrobipalpa aptatella (Walker, 1864) [149]
heliopa (Lower, 1900)
Scrobipalpa kasyi Povolný, 1968
Scrobipalpa notata (Povolný, 2001)
Scrobipalpa acuminatella (Sircom, 1850)
pulliginella (Sircom, 1850)
cirsiella (Stainton, 1851)
porcella (Heinemann, 1870)
ingloriella (Heinemann, 1870)
gracilella (Stainton, 1871)
Scrobipalpa skulei Huemer & Karsholt, 2010
Scrobipalpa hungariae (Staudinger, 1871)
Scrobipalpa adaptata (Povolný, 2001)
Scrobipalpa brahmiella (Heyden, 1862)
Scrobipalpa vasconiella (Rössler, 1877)
drahomirae Povolný, 1966
Scrobipalpa dorsolutea Huemer & Karsholt, 2010
Scrobipalpa amseli Povolný, 1966 [150]
Scrobipalpa hyssopi Nel, 2003 [150]
Scrobipalpa montanella (Chrétien, 1910)
Scrobipalpa corleyi Huemer & Karsholt, 2010
Scrobipalpa chrysanthemella (Hofmann, 1867)
opificella (Mann, 1878)
Scrobipalpa proclivella (Fuchs, 1886)
rancidella auct.
Scrobipalpa frugifera Povolný, 1969
hypothetica Povolný, 1973
Scrobipalpa oleksiyella Huemer & Karsholt, 2010
Scrobipalpa smithi Povolný & Bradley, 1964
Scrobipalpa occulta (Povolný, 2002)
sibirica Bidzilya, 2009
Scrobipalpa grisea Povolný, 1969
uralensis Povolný, 1973, unavailable
Scrobipalpa usingeri Povolný, 1969
Scrobipalpa clintoni Povolný, 1968
linella Piskunov, 1975
deleta Povolný, 1981
Scrobipalpa reiprichi Povolný, 1984 [151]
Scrobipalpa obsoletella (Fischer v. Röslerstamm, 1841)
miscitatella (Clarke, 1932)
bipunctella (Hartig, 1941)
calaritanella (Amsel, 1952)
hospes Povolný, 1964
Scrobipalpa feralella (Zeller, 1872)
rebeliella (Hauder, 1917)
Scrobipalpa halonella (Herrich-Schäffer, 1854)
Scrobipalpa perinii (Klimesch, 1951)
Scrobipalpa phagnalella (Constant, 1895)
staehelinella (Caradja, 1920), unavailable
Scrobipalpa tokari Huemer & Karsholt, 2010
Scrobipalpa karadaghi (Povolný, 2001)
Scrobipalpa heimi Huemer & Karsholt, 2010
Scrobipalpa acuta (Povolný, 2001)
Scrobipalpa soffneri Povolný, 1964
Scrobipalpa jariorum Huemer & Karsholt, 2010
Scrobipalpa murinella (Duponchel, 1843)
culminicolella (Staudinger, 1871)
excelsa (Frey, 1880)
Scrobipalpa wiltshirei Povolný, 1966
obrteliana Povolný, 1971, subspecies
Scrobipalpa caucasica (Povolný, 2001) [152]
bezengensis (Povolný, 2001)
Scrobipalpa pauperella (Heinemann, 1870) [153]
klimeschi Povolný, 1967
Scrobipalpa spumata (Povolný, 2001)
Scrobipalpa arenbergeri Povolný, 1973
Scrobipalpa mercantourica Varenne & Nel, 2018 [154]
Scrobipalpa nana Povolný, 1973
caroxyli (Falkovitsh & Bidzilya, 2006), subspecies
Scrobipalpa heretica Povolný, 1973
submagnificella Povolný, 1977
Scrobipalpa bigoti Povolný, 1973
tunesica Povolný, 1979, subspecies
Scrobipalpa dorsoflava (Povolný, 1996)
Scrobipalpa magnificella Povolný, 1967
Scrobipalpa abstrusa Huemer & Karsholt, 2010
Scrobipalpa superstes Povolný, 1977
Scrobipalpa remota Povolný, 1972
Scrobipalpa plesiopicta Povolný, 1969
Scrobipalpa bradleyi Povolný, 1971
glaserorum Povolný, 1977
meyricki auct.
Scrobipalpa selectella (Caradja, 1920)
fraterna Povolný, 1969
Scrobipalpa alterna (Falkovitsh & Bidzilya, 2006) [155]
Scrobipalpa lutea Povolný, 1977 [155]
Scrobipalpa griseoflava Bidzilya & Budashkin, 2011
Scrobipalpa niveifacies Povolný, 1977
milleri Povolný, 1977
Scrobipalpa indignella (Staudinger, 1879)
pseudobsoletellum (Povolný & Gregor, 1955)
hyoscyamivora (Gerasimov, 1940)
grossa Povolný, 1966
Scrobipalpa punctata (Povolný, 1996)
Scrobipalpa lagodes (Meyrick, 1926)
*Scrobipalpa deluccae Povolný, 1966
Scrobipalpa atriplicella (Fischer von Röslerstamm, 1841)
atrella (Thunberg, 1788), homonym
detersella (Clemens, 1860), homonym
infumatella (Fuchs, 1901)
brackenridgiella (Busck, 1903)
chenopodiella (Busck, 1916)
arogantella Povolný, 1967
altajica Povolný, 1969
Scrobipalpa suaedella (Richardson, 1893)
flavidorsella (Amsel, 1952)
hartigi Povolný, 1977
Scrobipalpa solitaria Povolný, 1969
*Scrobipalpa dagmaris Povolný, 1987
rezniki Piskunov, 1990
turkmenica Piskunov, 1990
Scrobipalpa suasella (Constant, 1895)
Scrobipalpa hendrikseni Huemer & Karsholt, 2010
Scrobipalpa halimifolia Bidzilya & Budashkin, 2011
Scrobipalpa traganella (Chrétien, 1915)
Scrobipalpa bazae Povolný, 1977
Scrobipalpa artemisiella (Treitschke, 1833) [156]
ancillella (Bruand d’Uzelle, 1851)
paniculatella (Novickij, 1924)
mongolensis Povolný, 1969
oreocyrniella (Petry, 1904), subspecies
syriaca Povolný, 1967, subspecies
Scrobipalpa stangei (Hering, 1889) [156]
saltenella (Meess, 1910)
Scrobipalpa suaedivorella (Chrétien, 1915)
detersipunctella (Toll, 1947)
Scrobipalpa bryophiloides Povolný, 1966 [157]
Scrobipalpa algeriensis Povolný & Bradley, 1964
Scrobipalpa deutschi Huemer & Karsholt, 2010
Scrobipalpa disjectella (Staudinger, 1859)
Scrobipalpa fontanensis Varenne & Nel, 2017
Scrobipalpa mixta Huemer & Karsholt, 2010
Scrobipalpa achtubica Anikin & Piskunov, 2018
Scrobipalpa rebeli (Preissecker, 1914)
fuscella (Klimesch, 1938)
japonica Povolný, 1977
Scrobipalpa gallicella (Constant, 1885)
Scrobipalpa ustulatella (Staudinger, 1871)
Scrobipalpa postulatella Huemer & Karsholt, 2010
Scrobipalpa filia Povolný, 1969
Scrobipalpa nitentella (Fuchs, 1902)
seminella (Pierce & Metcalfe, 1935)
Scrobipalpa costella (Humphreys & Westwood, 1845)
costimaculella (Bruand d´Uzelle, 1859)
Scrobipalpa hyoscyamella (Stainton, 1869)
Scrobipalpa portosanctana (Stainton, 1859)
eremaula (Meyrick, 1891)
lyciella (Walsingham, 1900)
desertella (Rebel, 1901)
bertramella (Lucas, 1940)
leroyella (Lucas 1950)
reisseri (Povolný & Gregor, 1955)
philolycii (Hering, 1957)
gallincolella auct.
Scrobipalpa vicaria (Meyrick, 1921)
tineiformis Povolný, 1967
Scrobipalpa ocellatella (Boyd, 1858) [158]
ocellatella (Stainton, 1859), homonym
submissella (Stainton, 1859)
horticolella (Rössler, 1866)
clarella (Caradja, 1920)
obscurior (Rebel, 1927)
orientale (Gregor & Povolný, 1954)
portosanctana auct.
Scrobipalpa pulchra Povolný, 1967
Scrobipalpa gecko (Walsingham, 1911)
Scrobipalpa hannemanni Povolný, 1966
furva Povolný, 1969, subspecies
gamanthi (Falkovitsh & Bidzilya, 2006), subspecies
Scrobipalpa erichi Povolný, 1964
Scrobipalpa divisella (Rebel, 1936)
Scrobipalpa voltinella (Chrétien, 1898)
Scrobipalpa corsicamontes Varenne & Nel, 2013
Scrobipalpa suaedicola (Mabille, 1906)
suaedicola (Amsel, 1939), homonym
mabillei Povolný, 1971
Scrobipalpa monochromella (Constant, 1895)
Scrobipalpa samadensis (Pfaffenzeller, 1870)
plantaginella (Stainton, 1883)
brunhildae (Schawerda, 1921)
zimmermanni (Zimmermann, 1923), unavailable
mariae (Zimmermann, 1926)
testacella (Rebel, 1935)
echo (Meyrick, 1937)
Scrobipalpa salinella (Zeller, 1847) [159]
omachella auct.
zernyella (Rebel, 1918)
corsicanum (Gregor & Povolný, 1954)
ignotum (Gregor & Povolný, 1954)
trebujenae Povolný, 1977
Scrobipalpa spergulariella (Chrétien, 1910) [159]
Scrobipalpa salicorniae (Hering, 1889) [159]
caliacrae (Caradja, 1932)
Scrobipalpa halimioniella Huemer & Karsholt, 2010
Scrobipalpa thymelaeae (Amsel, 1939)
Scrobipalpa halymella (Millière, 1864) [160]
Scrobipalpa camphorosmella Nel, 1999
Scrobipalpa stabilis Povolný, 1977 [160]
Scrobipalpa instabilella (Douglas, 1846)
lagunella (Chrétien, 1910)
strobilacella (Caradja, 1920), unavailable
salsolella (Amsel, 1935)
halymiphaga (Amsel, 1952)
Scrobipalpa peterseni (Povolný, 1965)
Scrobipalpa ergasima (Meyrick, 1916)
hyoscyamella (Rebel, 1912), homonym
mignatella (Caradja, 1920), unavailable
intestina (Meyrick, 1921)
mirabile (Gregor & Povolný, 1955)
pervada (Clarke, 1962)
Turcopalpa Povolný, 1973
Turcopalpa glaseri Povolný, 1973
Scrobipalpula Povolný, 1964 [161]
Scrobipalpula psilella (Herrich-Schäffer, 1854)
nocturnella (Staudinger, 1859)
pallidella (Heinemann, 1870)
killiasii (Frey, 1880)
astericolellum (Hering, 1957), unavailable
asiatica Povolný, 1968, subspecies
Scrobipalpula ramosella (Müller-Rutz, 1934)
Scrobipalpula seniorum Povolný, 2000
ptarmicae (Hering, 1957), unavailable
compositella (Povolný, 1964), unavailable
Scrobipalpula diffluella (Frey, 1870)
cacuminum (Frey, 1870)
diffluella (Heinemann, 1870)
bellidiastri (Klimesch, 1951)
uniflorellum (Hering, 1957), unavailable
Scrobipalpula tussilaginis (Stainton, 1867)
tussilaginella (Heinemann, 1870)
retusella (Rebel, 1891)
Phthorimaea Meyrick, 1902
Phthorimaea operculella (Zeller, 1873)
terrella (Walker, 1864)
solanella (Boisduval, 1874)
tabacella (Ragonot, 1879)
sedata (Butler, 1880)
argentinae Povolný, 1989
piscipellis auct.
epicentra auct.
Tuta Kieffer & Jørgensen, 1910
Tuta absoluta (Meyrick, 1917)
Keiferia Busck, 1939 [162]
Keiferia lycopersicella (Walsingham, 1897)
Ephysteris Meyrick, 1908
Microcraspedus Janse, 1958
Opacopsis Povolný, 1964
Echinoglossa Clarke, 1965
Ephysteris promptella (Staudinger, 1859) [163]
despectella (Walker, 1863)
petiginella (Mann, 1867)
parvula (Staudinger, 1879)
cacomicra (Walsingham, 1908)
chersaea Meyrick, 1908
oschophora (Meyrick, 1910)
crystallista (Meyrick, 1911)
dispensata (Meyrick, 1921)
fanatica (Meyrick, 1921)
xanthorhabda (Gozmány, 1951)
australiae Povolný, 1977
Ephysteris tenuisaccus Nupponen, 2010
Ephysteris deserticolella (Staudinger, 1871)
albocapitella (Rebel, 1928)
buvati (Povolný, 1992)
Ephysteris insulella (Heinemann, 1870)
insularis (Staudinger, 1871)
praticolella (Christoph, 1872), subspecies
gallica (Povolný, 1992)
Ephysteris brachyptera Karsholt & Sattler, 1998
Ephysteris diminutella (Zeller, 1847) [164]
lunaki (Hartig, 1941)
treskensis Povolný, 1964
hispanica Povolný, 1981
foulonsensis Povolný, 1981
Ephysteris inustella (Zeller, 1839) [165]
delminiella (Rebel, 1904)
gredosensis (Rebel, 1935), subspecies
Ephysteris olympica Povolný, 1968
monticola Povolný, 1981
Ephysteris iberica Povolný, 1977
Ochrodia Povolný, 1966 [166]
Ochrodia subdiminutella (Stainton, 1867)
jamaicensis (Walsingham, 1897)
bucolica (Meyrick, 1904)
zygophyllella (Rebel, 1912)
ericnista (Meyrick, 1914)
ferritincta (Turner, 1919), subspecies
ochrodeta (Meyrick, 1923)
extorris (Meyrick, 1923)
crocoleuca (Meyrick, 1923)
unitella (Turati, 1930)
tribulivora (Dumont, 1931)
pulverea (Janse, 1950)
turgida (Janse, 1951)
pentamacula (Janse, 1958)
infallax (Gozmány, 1960)
tractatum (Gozmány, 1960)
Vladimirea Povolný, 1967
Distinxia Povolný, 1967
Vladimirea glebicolorella (Erschoff, 1874)
submaculata Povolný, 1967
Microlechia Turati, 1924
Hedma Dumont, 1932
Megalocypha Janse, 1960
Microlechia rhamnifoliae (Amsel & Hering, 1931)
rhamnifoliae (Amsel, 1935)
Microlechia chretieni Turati, 1924
microcasis (Meyrick, 1929)
micradelpha (Walsingham, 1900), homonym
hyoscyamella (Amsel & Hering, 1931), homonym
abzacella (Dumont, 1932)
hyoscyami (Amsel, 1935)
polioptera (Janse, 1960)
aellographa (Janse, 1960)
Microlechia klimeschi (Povolný, 1972)
Microlechia karsholti (Nupponen, 2010)
Cosmardia Povolný, 1965
Cosmardia moritzella (Treitschke, 1835)
morizella (Geyer, 1836)
roseella (Zetterstedt, 1839)
Lutilabria Povolný, 1965 [167]
Lutilabria lutilabrella (Mann, 1857)
robustella (Rebel, 1910)
olympica Huemer, 1993, subspecies
Lutilabria volgensis Anikin & Piskunov, 1996
Lutilabria prolata Junnilainen & Nupponen, 2010
Klimeschiopsis Povolný, 1967
Klimeschiopsis kiningerella (Duponchel, 1843) [168]
atralbella (Palm, 1947)
Klimeschiopsis discontinuella (Rebel, 1899)
Klimeschiopsis maritimaealpina Nel & Varenne, 2011
Klimeschiopsis terroris (Hartig, 1938)
Caryocolum Gregor & Povolný, 1954 [169]
Caryocolum fischerella (Treitschke, 1833)
Caryocolum tischeriella (Zeller, 1839) [170]
Caryocolum alsinella (Zeller, 1868) [171]
albifrontella (Heinemann, 1870)
tristella (Heinemann, 1870)
semidecandriella (Tutt, 1887)
semidecandrella (Threlfall & Stainton, 1887)
Caryocolum viscariella (Stainton, 1855)
crepusculella (Teich, 1889)
Caryocolum albifaciella (Heinemann, 1870)
behenella (Constant, 1890)
Caryocolum vicinella (Douglas, 1851) [172]
inflatella (Chrétien, 1901)
albescens (Bankes, 1909), unavailable
suffusa (Bankes, 1909), unavailable
Caryocolum bosalella (Rebel, 1936)
Caryocolum sciurella (Walsingham, 1908)
rubidella (Chrétien, 1908)
Caryocolum amaurella (Hering, 1924) [173]
viscariae (Schütze, 1926)
Caryocolum crypticum Huemer, Karsholt & Mutanen, 2014
Caryocolum tredosella Nel & Requena, 2017
Caryocolum oculatella (Thomann, 1930)
ochraceella (Thomann, 1929), homonym
Caryocolum leucofasciatum Huemer, 1989
Caryocolum petryi (Hofmann, 1899)
rougemonti (Rebel, 1907)
repentella (Chrétien, 1908)
benanderi (Hering, 1933)
Caryocolum baischi Huemer & Karsholt, 2010
Caryocolum repentis Huemer & Luquet, 1992
repentella auct.
Caryocolum siculum Bella, 2008
Caryocolum inflativorella (Klimesch, 1938)
xuthella (Rebel, 1941)
census (Gozmány, 1954)
Caryocolum saginella (Zeller, 1868) [174]
coussonella (Chrétien, 1908)
Caryocolum cauligenella (Schmid, 1863) [175]
Caryocolum trauniella (Zeller, 1868)
Caryocolum peregrinella (Herrich-Schäffer, 1854) [176]
melantypella (Mann, 1877)
Caryocolum delphinatella (Constant, 1890)
fiorii (Klimesch, 1953)
Caryocolum provinciella (Stainton, 1869)
Caryocolum mucronatella (Chrétien, 1900)
poschiavensis (Rebel, 1936)
Caryocolum leucomelanella (Zeller, 1839) [177]
gypsophilae (Stainton, 1869)
Caryocolum mazeli Huemer & Nel, 2005
Caryocolum leucothoracellum (Klimesch, 1953)
Caryocolum schleichi (Christoph, 1872) [178]
syriacum Povolný, 1977
dianthella (Chrétien, 1925), subspecies
hackeri Derra, 1985
improvisella (Rebel, 1936), subspecies
Caryocolum arenariella (Benander, 1937) [178]
Caryocolum marmorea (Haworth, 1828) [179]
manniella (Zeller, 1839)
marmorella (Doubleday, 1859), emendation
pulchra (Wollaston, 1858), subspecies
mediocorsa Varenne & Nel, 2013, subspecies
marmoreum auct.
Caryocolum pullatella (Tengström, 1848) [180]
pulla (Tengström, 1848)
subtractella (Walker, 1864)
livoniella (Teich, 1898)
agricolaris (Meyrick, 1933)
Caryocolum stramentella (Rebel, 1935)
emarginatum Huemer, 1988
Caryocolum hispanicum Huemer, 1988
Caryocolum confluens Huemer, 1988
Caryocolum srnkai Huemer & Karsholt, 2011
Caryocolum gallagenellum Huemer, 1989
Caryocolum fraternella (Douglas, 1851)
intermediella (Hodgkinson, 1897)
Caryocolum klosi (Rebel, 1917) [181]
Caryocolum interalbicella (Herrich-Schäffer, 1854)
quadrella (Fabricius, 1794), homonym
Caryocolum laceratella (Zeller, 1868)
thurneri (Pinker, 1953)
Caryocolum dauphini Grange & Nel, 2012
Caryocolum blandella (Douglas, 1852) nom. protectum [182]
signatella (Eversmann, 1844) nom. oblitum
maculea (Haworth, 1828), (nec Fabricius, 1794), emendation, misident.
Caryocolum blandelloides Karsholt, 1981
Caryocolum horoscopa (Meyrick, 1926) stat. rev. [183]
Caryocolum jaspidella (Chrétien, 1908)
Caryocolum proxima (Haworth, 1828)
maculiferella (Douglas, 1851)
maculivicinella (Bruand d´Uzelle, 1859)
horticolla (Peyerimhoff, 1871)
proximum auct.
Caryocolum blandulella (Tutt, 1887)
Caryocolum arenbergeri Huemer, 1989
Caryocolum tricolorella (Haworth, 1812)
contigua (Haworth, 1828)
acernella (Herrich-Schäffer, 1854)
Caryocolum fibigerium Huemer, 1988 [184]
Caryocolum junctella (Douglas, 1851) [185]
aganocarpa (Meyrick, 1935)
Caryocolum cassella (Walker, 1864)
melanotephrella (Erschoff, 1877)
albifasciella (Toll, 1936)
subvicinella (Hackman, 1946)
falellum Piskunov, 1975
Caryocolum moehringiae (Klimesch, 1954)
Caryocolum petrophila (Preissecker, 1914)
kemnerella (Palm, 1947)
Caryocolum huebneri (Haworth, 1828)
hubnerella (Doubleday, 1866)
knaggsiella (Stainton, 1866)
Caryocolum kroesmanniella (Herrich-Schäffer, 1854)
huebneri auct.
Tila Povolný, 1965
Tila capsophilella (Chrétien, 1900)
Pogochaetia Staudinger, 1879
Pogonochaetia Rye, 1881
Chaetopogon Rye, 1881
Pogochaetia solitaria Staudinger, 1879
ocymoidella (Walsingham, 1900), subspecies
cabreretsi Povolný, 1981
Agonochaetia Povolný, 1967
Sautereopsis Povolný, 1965
Agonochaetia terrestrella (Zeller, 1872) [186]
muestairella (Müller-Rutz, 1922)
Agonochaetia intermedia Sattler, 1968
Agonochaetia quartana Povolný, 1990
Canarischema Karsholt, 2017
Canarischema fuerteventura Karsholt, 2017
Sattleria Povolný, 1965 [187]
Sattleria melaleucella (Constant, 1865) [188]
mariae (Frey, 1867), unavailable
fusca (Burmann, 1954)
Sattleria arcuata Pitkin & Sattler, 1991
Sattleria pyrenaica (Petry, 1904) [189]
Sattleria taurandi Nel & Varenne, 2019
Sattleria karsholti Huemer & Hebert, 2011
Sattleria cottiella Huemer & Hebert, 2011
Sattleria marguareisi Huemer & Sattler, 1992
Sattleria izoardi Huemer & Sattler, 1992
Sattleria graiaeella Huemer & Hebert, 2011
Sattleria dolomitica Huemer, 2014
Sattleria basistrigella Huemer, 1997
Sattleria triglavica Povolný, 1987
Sattleria basistrigella Huemer, 1997
basistrigella (Müller-Rutz, 1934), unavailable
Sattleria dinarica Huemer, 2014
Sattleria haemusi Huemer, 2014
Sattleria dzieduszyckii (Nowicki, 1864)
tatrica (Gregor & Povolný, 1955)
Sattleria angustispina Pitkin & Sattler, 1991
Sattleria breviramus Pitkin & Sattler, 1991
Sattleria sophiae Timossi, 2014
Sattleria styriaca Pitkin & Sattler, 1991
Litini
Teleiodini Piskunov, 1973
Exoteleiini Omelko, 1999
Schneidereria Weber, 1957
Schneidereria pistaciella Weber, 1957 [191]
Teleiodes Sattler, 1960
Dubitationis Omelko & Omelko, 1998
Teleia Heinemann, 1870, homonym
Teleiodes vulgella ([Denis & Schiffermüller], 1775) [191]
aspera (Haworth, 1828)
Teleiodes italica Huemer, 1992 [192]
gallica Huemer, 1992
Teleiodes brevivalva Huemer, 1992 [192]
Teleiodes wagae (Nowicki, 1860)
marsata Piskunov, 1973
Teleiodes saltuum (Zeller, 1878) [193]
nigristrigella (Wocke, 1898)
Teleiodes kaitilai Junnilainen, 2010 [193]
Teleiodes luculella (Hübner, 1813) [194]
subrosea (Haworth, 1828)
Teleiodes flavimaculella (Herrich-Schäffer, 1854) [195]
rufipunctella (Steudel, 1882)
dealbella (Klemensiewicz, 1902), unavailable
herrichi (Dufrane, 1955), unavailable
Teleiodes albidorsella Huemer & Karsholt, 1999
Teleiodes albiluculella Huemer & Karsholt, 2001
Neotelphusa Janse, 1958
Neotelphusa sequax (Haworth, 1828)
apicistrigella (Duponchel, 1843)
sequaxella (Bruand d´Uzelle, 1859)
sequacella (Doubleday, 1859), emendation
Neotelphusa huemeri (Nel, 1998)
pseudocisti Leraut, 1997, unavailable
Neotelphusa traugotti (Huemer & Karsholt, 2001)
Neotelphusa cisti (Stainton, 1869)
Carpatolechia Căpuşe, 1964
Vicina Omelko, 1999
Carpatolechia decorella (Haworth, 1812)
humeralis (Zeller, 1839)
lyellella (Humphreys & Westwood, 1845)
incretella (Duponchel, 1845)
humeralella (Bruand d´Uzelle, 1851), emendation
marmoripennella (Bruand d´Uzelle, 1851)
pisticella (Nowicki, 1860)
scabra (Staudinger, 1870)
erschoffii (Frey, 1880)
subericolella (Caradja, 1920), unavailable
buckwelli (Lucas, 1956)
dumitrescui Căpuşe, 1964
Carpatolechia aenigma (Sattler, 1983)
Carpatolechia fugitivella (Zeller, 1839)
vovkella (Piskunov, 1973)
melanella (Romaniszyn, 1933), unavailable
Carpatolechia fugacella (Zeller, 1839)
nigrofasciella (Bruand d´Uzelle, 1851)
Carpatolechia minor (Kasy, 1978)
Carpatolechia filipjevi (Lvovsky & Piskunov, 1993)
Carpatolechia alburnella (Zeller, 1839)
seniculella (Eversmann, 1844)
radiella (Krulikowsky, 1909), unavailable
Carpatolechia notatella (Hübner, 1813)
euratella (Herrich-Schäffer, 1854)
oskella (Piskunov, 1973)
Carpatolechia proximella (Hübner, 1796)
peritella (Constant, 1885)
ochracella (Romaniszyn, 1933), unavailable
Carpatolechia intermediella Huemer & Karsholt, 1999
Carpatolechia epomidella (Tengström, 1869)
Pseudotelphusa Janse, 1958
Sattleria Căpuşe, 1968, homonym
Klaussattleria Căpuşe, 1968
Pseudotelphusa scalella (Scopoli, 1763) [196]
aleella (Fabricius, 1794)
bicolorella (Treitschke, 1832)
Pseudotelphusa istrella (Mann, 1866)
decuriella (Mann, 1872)
trifasciella (Rebel, 1916)
Pseudotelphusa occidentella Huemer & Karsholt, 1999
Pseudotelphusa paripunctella (Thunberg, 1794)
tigratella (Costa, 1834)
triparella (Zeller, 1839)
trijugella (Erschoff, 1877)
sultanella (Caradja, 1920)
griseella (Preissecker, 1931), unavailable
myricae (Gilles, 1936), unavailable
pseudowagae (Svensson, 1993), unavailable
Pseudotelphusa tessella (Linnaeus, 1758)
albinigrella ([Denis & Schiffermüller], 1775)
sturmella (Hübner, 1825)
berberidella (Hübner, 1825)
funestella (Geyer, 1832)
alboquadrella (Bruand d´Uzelle, 1859)
Istrianis Meyrick, 1918
Pseudoteleia Amsel, 1935
Istrianis myricariella (Frey, 1870)
Istrianis arenicolella (Caradja, 1920)
amilcarella (Lucas, 1933)
Istrianis pseudomyricariella Bidzilya & Karsholt, 2015
Istrianis nilssoni Bidzilya & Karsholt, 2015
Istrianis brucinella (Mann, 1872)
Istrianis femoralis (Staudinger, 1876)
comedonella (Staudinger, 1879)
gravosensis (Rebel, 1937)
angustipennis (Rebel, 1941)
funebrella (Rebel, 1941)
squamodorella auct.
Istrianis piskunovi Bidzilya & Karsholt, 2015
Streyella Janse, 1958
Streyella canariensis (Walsingham, 1908)
Streyella anguinella (Herrich-Schäffer, 1861)
ostentella (Zerny, 1934)
Teleiopsis Sattler, 1960
Teleiopsis terebinthinella (Herrich-Schäffer, 1856)
Teleiopsis latisacculus Pitkin, 1988
Teleiopsis diffinis (Haworth, 1828) [197]
dissimilella (Treitschke, 1833)
scabidella (Zeller, 1839)
friesella (Zetterstedt, 1839)
diffinella (Doubleday, 1859), emendation
groenliensis (Strand, 1920), unavailable
Teleiopsis lunariella (Walsingham, 1908)
Teleiopsis bagriotella (Duponchel, 1840) [197]
elatella (Herrich-Schäffer, 1854)
Teleiopsis laetitiae Schmid, 2011 [197]
Teleiopsis lindae Schmid, 2011
Teleiopsis albifemorella (Hofmann, 1867) [197]
Teleiopsis paulheberti Huemer & Mutanen, 2012 [197]
Teleiopsis rosalbella (Fologne, 1862) [197]
Xenolechia Meyrick, 1895 [198]
Xenolechia aethiops (Humphreys & Westwood, 1845)
aterrima (Edleston, 1844)
aethiopella (Doubleday, 1859), emendation
squamulella (Peyerimhoff, 1871)
tristis (Staudinger, 1879)
Xenolechia lindae Huemer & Karsholt, 1999
Xenolechia pseudovulgella Huemer & Karsholt, 1999
Altenia Sattler, 1960
Altenia perspersella (Wocke, 1862)
empetrella (Karvonen, 1932)
Altenia scriptella (Hübner, 1796) [199]
Altenia elsneriella Huemer & Karsholt, 1999
Altenia mersinella (Staudinger, 1879)
melanostictella (Ragonot, 1895)
sagittella (Caradja, 1920)
praedicta (Meyrick, 1923)
tribolopis (Meyrick, 1927)
Altenia wagneriella (Rebel, 1926)
danilevskyi (Piskunov, 1973)
Altenia modesta (Danilevsky, 1955)
Recurvaria Haworth, 1828
Lita Kollar, 1832
Telea Stephens, 1834, homonym
Aphanaula Meyrick, 1895
Hinnebergia Spuler, 1910
Recurvaria nanella ([Denis & Schiffermüller], 1775)
pumilella ([Denis & Schiffermüller], 1775)
nana Haworth, 1828, emendation
crataegella Busck, 1903
unicolor Rebel, 1927
pruniella auct.
Recurvaria leucatella (Clerck, 1759)
leucatea Haworth, 1828, emendation
albocingulella (Duponchel, 1839)
Recurvaria thomeriella (Chrétien, 1901)
Recurvaria costimaculella Huemer & Karsholt, 2001
Coleotechnites Chambers, 1880
Evagora Clemens, 1860, homonym
Eidothea Chambers, 1873 (emendation and homonym)
Eucordylea Dietz, 1900
Pulicalvaria Freeman, 1963
Coleotechnites piceaella (Kearfott, 1903)
nigra (Kearfott, 1903), homonym
obscurella (Kearfott, 1907)
Exoteleia Wallengren, 1881
Paralechia Busck, 1903
Heringia Spuler, 1910, homonym
Heringiola Strand, 1917
Exoteleia dodecella (Linnaeus, 1758) [200]
duodecimcristata (Retzius, 1783), unavailable
punctulata (Fourcroy, 1785)
dodecea (Haworth, 1828), emendation
annulicornis (Stephens, 1834)
favillaticella (Zeller, 1839)
reussiella (Ratzeburg, 1840)
Exoteleia succinctella (Zeller, 1872)
oribatella (Rebel, 1918)
Stenolechia Meyrick, 1894
Poecilia Heinemann, 1870, homonym
Gibbosa Omelko, 1988
Stenolechia gemmella (Linnaeus, 1758)
nivella (Fabricius, 1794)
nivea (Haworth, 1828), emendation
lepidella (Zeller, 1839)
nigrovittella (Duponchel, 1839)
Parastenolechia Kanazawa, 1985
Origo Omelko, 1988
Tutor Omelko, 1988
Laris Omelko, 1988
Parastenolechia nigrinotella (Zeller, 1847)
nigralbella (Herrich-Schäffer, 1854), unavailable
Stenolechiodes Elsner, 1996
Stenolechiodes pseudogemmellus Elsner, 1996
Stenolechiodes macrolepiellus Huemer & Karsholt, 1999
Parachronistis Meyrick, 1925
Cochlevalva Omelko, 1986
Dentivalva Omelko, 1986
Parachronistis albiceps (Zeller, 1839) [201]
albicipitella (Herrich-Schäffer, 1854), emendation
albicapitella (Doubleday, 1859), emendation
Schistophila Chrétien, 1899
Schistophila laurocistella Chrétien, 1899
striatana (Lucas, 1937)
Unplaced genus
“Telphusa” cistiflorella (Constant, 1890) [202]
Approximately 200 comments on systematic problems, taxonomic changes and particularly potential cryptic diversity, are mainly derived from molecular data and are cross-referenced in the checklist: [1] – [202].
[1] Anacampsidae Bruand d’Uzelle, 1851 has priority over Gelechiidae Stainton 1854. The former name has hardly been used (
[2] Stomopteryx. This genus is in need of a taxonomic revision and includes several probable cases of cryptic diversity, and equally probably cases of over-splitting.
[3] Stomopteryx nugatricella / S. mongolica / S. lineolella. The taxonomy of these species is unresolved and should be checked in upcoming revisionary work.
[4] Stomopteryx deverrae. We have barcoded only North African specimens so far, including a syntype from Algeria, and the material from Spain should be sequenced in future to prove the occurrence in Europe.
[5] Stomopterayx flavoclavella. European samples from Spain slightly differ from a sequenced syntype from Morocco and cluster in a separate BIN. The suspected conspecificity will be addressed in an upcoming revision.
[6] Stomopteryx remissella. This species represents an unresolved species complex. DNA barcodes show an extraordinarily high and largely geographic variation, reflected by eight different BINs and differences in phenotype. The recently described Stomopteryx spathulella (
[7] Stomopteryx flavipalpella. A genetically variable species which clusters into three BINs without obvious geographic variation.
[8] Aproaerema. Recently A
[9] Aproaerema cinctella. This species clusters into two weakly separated DNA barcode clusters with max. distance of 1.86%, probably reflecting intraspecific variation.
[10] Aproaerema linella. A unicolorous, dissected male from Montenegro largely corresponds with the lectotype figured by
[11] Aproaerema suecicella. Two strongly divergent BINs (4.33% min. distance) show a geographic pattern and need to be tested for potential cryptic diversity.
[12] Aproaerema karvoneni. Three weakly separated BINs (1.61% min. distance) partially show geographic (probably intraspecific) variation.
[13] Aproaerema anthyllidella. The moderate DNA barcode variation with three BINs may reflect cryptic diversity, as e.g., suspected for the current synonym A. natrixella (Schmid pers. comm.) and some of the other five current synonyms, but has to be carefully checked with an integrative taxonomic approach.
[14] Iwaruna. Species in this genus share their BINs and partially overlap in DNA barcodes (I. biguttella and I. klimeschi) but differ in morphology. DNA barcodes of I. heringi, a species requiring taxonomic re-assessment, are unknown.
[15] Anacampsis populella / A. blattariella. A population from western Austria (Vorarlberg) shares its BIN with a unique specimen of A. populella from Finland though matching A. blattariella in morphology. This is most likely a case of a so far unrecognized introgression in these two species, though the weakly deviating DNA barcode may require further studies. All other sequenced specimens of both species group in separate BINs.
[16] Anacampsis scintillella. Two specimens from Spain cluster in a separate BIN.
[17] Anacampsis obscurella. Our limited data indicates geographically separated species with three BINs but requires additional revisionary work.
[18] Mesophleps. The genus was recently revised by
[19] Nothris. The sequence of species follows the revision by
[20] Nothris gregerseni. A specimen from Sweden clusters into a unique BIN (3.83% min. distance, but probably representing only an intraspecific split.
[21] Nothris radiata. The yet unpublished occurrence in Europe is based on a DNA barcoded specimen from Macedonia (Šumpich in prep.).
[22] Neofaculta ericetella. This species shows high intraspecific DNA barcode variation and clusters into three BINs without geographic variation.
[23] Neofaculta taigana. The occurrence of this Asian species in Europe will be dealt with separately by Aarvik, Berggren, Karsholt and Mutanen.
[24] Hypatima rhomboidella. Genetically variable species clustering into three BINs without geographic variation.
[25] Anarsia. The genus requires revisionary work and probably includes two undescribed species from Greece and Cyprus respectively.
[26] Anarsia bilbainella. A unique sequence from the type-area in Spain clusters into a separate BIN (1.26% min. distance).
[27] Dichomeris. As currently understood, Dichomeris is the largest genus within the Gelechiidae.
[28] Dichomeris limbipunctellus / D. neatodes. These two taxa, which have been regarded as conspecific, differ in phenotypy, show a different distribution pattern in the eastern (D. neatodes) and western Mediterranean (D. limbipunctellus), and cluster into two BINs. We accordingly list them as separate species and reinstate D. neatodes sp. rev. as a valid species.
[29] Dichomeris juniperella. The species splits into two strongly divergent BINs (5.26% min. distance), one widespread, and the other restricted to the southern Alps, reflecting possible cryptic diversity.
[30] Dichomeris rasilella. A single DNA barcode from Russia is highly divergent from other samples and clusters into a separate BIN (6.26% min. distance).
[31] Acompsia. The sequence of species follows the revision by
[32] Acompsia pyrenaeella. The species clusters into three BINs, one shared with phenotypically compared specimens of A. tripunctella and A. antirrhinella, indicating occasional introgression.
[33] Acompsia antirrhinella. Despite diagnostic morphological characters, this species shares the only known BIN with two genetically variable species, A. pyrenaella and A. tripunctella. See also comments under these species.
[34] Acompsia maculosella. Sequences of specimens from the southern Alps group into a separate BIN and are also separated by reduced forewing markings, but agree in genitalia morphology and are therefore tentatively considered as A. maculosella. [
[35] Acompsia tripunctella. A genetically highly variable species, which clusters into seven BINs, possibly reflecting cryptic diversity requiring revisionary work. One BIN is shared with A. pyrenaeella and A. antirrhinella. See also comments under these species.
[36] Brachmia. A species from Greece (Crete) is probably undescribed (Berggren in prep.).
[37] Brachmia dimidiella. A genetically variable species clustering into three different BINs.
[38] Helcystogramma lamprostoma. Male and female genitalia match Helcystogramma Zeller, 1877 and the species was placed in this genus in recent papers (
[39] Pseudosophronia. The identity of the three currently listed European species is somewhat doubtful and requires further analysis.
[40] Pexicopia malvella. The species splits into two BINs (4.33% min. distance) without geographic distinction and requires further analysis. The geographic variation in the forewing colour and pattern between specimens from Central Europe and South Europe is not reflected in the DNA barcode.
[41] Apatetrini. Genera and species of this tribe are in need of revision. Several of the included taxa do not cluster together in a barcode-based NJ tree and Apatetrini sensu auct. is likely not a monophyum.
[42] Dactylotula kinkerella. The species splits into two divergent clusters representing two BINs (4.49% min. distance).
[43] Apatetris. The two species listed here, A. agenjoi and A. mediterranella, are based on morphology not strictly congeneric with the type of the genus (A. mirabella Staudinger, 1879 from Turkey) and probably also not with each other, but are left here pending forthcoming revisionary work. Similarly, two yet unidentified species which are probably undescribed are not closely related and only tentatively assigned to Apatetris.
[44] Apatetris mediterranella. The species clusters into two geographically separate BINs (3.05% min. distance) and requires further revision.
[45] Catatinagma trivittellum. The species splits into two geographically separate and strongly divergent clusters, representing two BINs (5.11% min. distance). These should be tested for potential cryptic diversity with further sampling and a comprehensive morphological analysis.
[46] Catatinagma kraterella. The species does not cluster close to the type of the genus (C. trivittellum), instead appears closer to Apatetris mediterranella. It is, however, left in Catatinagma pending discovery of the unknown female and forthcoming revisionary work.
[47] Chrysoesthia. This genus lacks generic revision. Three yet unassigned, but sequenced species, may partly belong to the insufficiently revised taxa of the European fauna.
[48] Chrysoesthia drurella. This species splits into two strongly divergent BINs (3.69% min. distance) which partly overlap geographically and require careful re-assessment.
[49] Chrysoesthia atriplicella / C. gaditella / C. aletris. Morphological revisionary work and additional DNA barcoding efforts are required to determine if these three names represent one or more species. C. halymella (Amsel, 1935) also belongs to this complex (
[50] Metanarsia modesta. The species splits into two BINs, one only known from extra-European Armenia.
[51] Oecocecis guyonella. We were able to dissect both sexes from specimens provided by Christian Gibeaux. The female genitalia are rather similar to Metanarsia, but the male genitalia are strongly different. Therefore, and in the absence of molecular data, the systematic position is tentative and requires further revisionary work.
[52] For a discussion of the validity of Palumbininae, see
[53] Bryotropha. The sequence of species follows the revision by
[54] Bryotropha terrella. Two deviating DNA barcodes from Austria group into a separate BIN (2.94% min. distance) and the corresponding specimens require careful re-evaluation.
[55] Bryotropha desertella. A genetically variable species clustering into three BINs without geographical structure.
[56] Bryotropha hulli. The species clusters into two BINs without geographical separation.
[57] Bryotropha affinis. This species shares its BIN with one BIN of B. umbrosella.
[58] Bryotropha umbrosella. The species clusters into two BINs, one shared with B. affinis, which differs in phenotype and genitalia morphology (
[59] Epidola. Unrevised genus. The identity of Epidola grisea, described from a single male without an abdomen and collected in Sardinia (
[60] Epidola semitica. This species was described from a single male from Israel, but according to recently collected material it also occurs in Greece (new record for Europe, for detailed data see dataset in BOLD).
[61] Aristotelia. This genus is in strong need of a generic revision and includes several probably undescribed species.
[62] Aristotelia decurtella. This species genetically clusters into two BINs (2.25% min. distance) which are in need of morphological revision.
[63] Aristotelia ericinella. Specimens from Sardinia cluster separately into a different BIN (2.73% min. distance) and are considered as a separate species.
[64] Aristotelia subdecurtella. Two barcode clusters, grouping into different BINs that overlap in distribution.
[65] Aristotelia subericinella. The species identity is based on barcoded material from the type area (eastern Austria). Several additional clusters formerly identified as A. subericinella probably include cryptic diversity and are in strong need of taxonomic revision. These clusters are considered as unidentified taxa in our analysis.
[66] Aristotelia billii. DNA barcodes of this species are based upon the successfully sequenced holotype and prove a wide distribution from the Mediterranean to Kirgizia.
[67] Caulastrocecis. The genus is in need of revision.
[68] Caulastrocecis furfurella / C. cryptoxena. The former was considered as a senior synonym of C. cryptoxena but both are clearly divergent in DNA barcodes and represent different species (Bidzilya and Karsholt in prep.). We therefore reinstate C. cryptoxena sp. rev. as a valid species.
[69] Paranarsia. The systematic position of this genus is not fully resolved. The genitalia somewhat resemble those of Caulastrocesis but DNA barcodes are distant. Here we follow
[70] Megacraspedus. This genus was recently revised with 27 newly described species from Europe (
[71] Megacraspedus lanceolellus. Genetically extremely variable species, which clusters into 19 BINs of mainly geographic variation, with an intraspecific DNA barcode variation of 12.5% (
[72] Megacraspedus dolosellus. Genetically extremely variable species, which clusters into 23 BINs of mainly geographic variation, with an intraspecific DNA barcode variation of 13.8% (
[73] Megacraspedus spinophallus. Two barcode clusters, representing separate BINs with records from nearby localities.
[74] Megacraspedus binotella. Genetically variable species, which clusters into three BINs without clear geographic separation.
[75] Megacraspedus brachypteris. Genetically variable species, which clusters into four BINs without clear geographic separation.
[76] Megacraspedus andreneli. Two barcode clusters, representing separate BINs with records from nearby localities.
[77] Megacraspedus imparellus. Genetically variable species, which clusters into three BINs with probable geographic separation.
[78] Megacraspedus teriolensis. Genetically variable species, which clusters into two geographically distinct BINs.
[79] Dirhinosia. Species in this genus partly share DNA barcodes (D. cervinella and D. interposita) but differ in morphology (
[80] Psamathocrita. The genus is in need of revision. A probably undescribed species has been studied by Tokár and Junnilainen (in litt.) and Barton (in litt.).
[81] Ivanauskiella. This small genus seems to be more diverse than hitherto recognized, reflected an unidentified species from Russia and Spain. Some of the species are found in association with Limonium which is a likely host plant of the larvae (OK unpublished). Spatuncusella Nel & Varenne, 2013 was recently synonymized with Ivanauskiella (
[82] Ivanauskiella occitanica. This species was synonymized with I. psamathias by
[83] Ptocheuusa. The genus is in strong need of revision. Barcoding efforts for several validly described species failed to produce any sequences.
[84] Ptocheuusa paupella. The species clusters into three separate BINs without geographic separation.
[85] Ptocheuusa inopella. Two barcode clusters from Spain and Sweden represent three separate BINs and need to be re-examined.
[86] Ptocheuusa cinerella. We transfer Recurvaria cinerella Chrétien from Gelechiinae: Litini to Anomologinae as Ptocheuusa cinerella (Chrétien, 1908) comb. nov. The male genitalia are similar overall to other species of Ptocheuusa and seem sufficient for this new combination despite the lack of molecular data.
[87] Amblypalpis. The systematic position of this genus needs re-evaluation. It was recently published as new to Europe (
[88] Parapodia. Material from the western and eastern Mediterranean cluster into two strongly divergent BINs (5.43% min. distance). Although an initial morphological examination of females reveals no obvious diagnostic characters, these clusters should be tested for potential cryptic diversity by examining additional material and a widened morphological approach.
[89] Isophrictis. Unrevised genus, which includes cases of unresolved and apparently intraspecific DNA barcode divergence and probably some undescribed species, misidentified records or unrecognized synonymies for the European fauna. So far only six out of the twelve species in the checklist have been successfully barcoded.
[90] Isophrictis kefersteiniellus. Genetically highly variable species, which clusters into four BINs. A thorough evaluation of this problem is necessary.
[91] Isophrictis anthemidella. Genetically variable species, which clusters into three BINs. A thorough evaluation of this problem is necessary.
[92] Metzneria. The classic generic revision by
[93] Metzneria neuropterella. The species clusters into two BINs (2.89% min. distance) without geographic separation.
[94] Metzneria aestivella. The DNA barcode of a paratype of Metzneria expositoi Vives, 2001 from Spain fully agrees with that of M. aestivella. Also, the genitalia morphology of the two taxa is virtually identical, and we therefore consider M. expositoi to be a synonym of M. aestivella (syn. nov.).
[95] Metzneria fulva / Metzneria torosulella. Despite distinct diagnostic characters in phenotypic appearance and in the male genitalia, both species share barcodes.
[96] Metzneria ehikeella. The species clusters into two BINs (2.91% min. distance) without geographic separation.
[97] Metzneria metzneriella. This genetically variable species splits into four partly sympatric DNA barcode clusters, representing four BINs. A careful morphological examination of the problem is advisable.
[98] Metzneria artificella. Two weakly separated barcode clusters, representing geographically distinct BINs (1.46% min. distance), need to be re-examined.
[99] Metzneria aprilella. The species splits into three geographically separated DNA barcode clusters, representing three BINs. This possible case of cryptic diversity requires careful morphological re-examination.
[100] Metzneria subflavella. Two DNA barcodes referring to specimens from Spain and France respectively are strongly divergent and are considered separate species. These results are supported by genitalia morphology, with the Spanish specimen likely representing an undescribed species.
[101] Metzneria campicolella. Metzneria varennei Nel, 1997 was recently shown to be a synonym of M. campicolella (
[102] Apodia martinii. DNA barcodes of this species and A. bifractella with separate BINs (6.58% min. distance) fully support the species status for this long-disputed taxon. We therefore reinstate A. martinii sp. rev. as a valid species. Differences from A. bifractella in morphology, biology and distribution still need to be studied in detail.
[103] Pragmatodes. This genus, which has until now been placed in Gelechiini, has always been considered monotypic and endemic to the Canary Islands. However, a group of closely related species placed under Monochroa, i.e., Pragmatodes melagonella (Constant, 1895) comb. nov., Pragmatodes albagonella (Varenne & Nel, 2010) comb. nov., Pragmatodes cyrneogonlla (Nel & Varenne, 2012) comb. nov. and Pragmatodes parvulata (Gozmány, 1953) comb. nov., have similar genitalia which do not fit well with Monochroa, and their DNA barcodes cluster separately from that genus. Moreover, the known larvae of the above-mentioned species, as well as the type species of the genus (P. fruticosella) all feed on plants in the family Rubiaceae, an unusual feeding substrate for Gelechiidae. The genus includes additional, probably undescribed, species from South-East Europe and the Middle East.
[104] Pragmatodes melagonella. Specimens initially identified as this species from France and Bulgaria differ in the DNA barcode and also morphology and are considered as separate species. The type locality of P. melagonella is in France.
[105] Monochroa. This genus is in strong need of a generic revision and includes several probably undescribed species.
[106] Monochroa rumicetella. Two weakly separated BINs (2.12% min. distance) without geographic separation most probably reflect intraspecific variation.
[107] Monochroa sepicolella / M. rectifasciella.
[108] Monochroa tenebrella. Two weakly separated BINs (1.12% min. distance) without geographic separation most probably reflect intraspecific variation.
[109] Monochroa servella. Two BINs (2.89% min. distance) without geographic separation most probably reflect intraspecific variation.
[110] Monochroa lucidella. Despite a low intraspecific divergence, this species may include cryptic diversity as indicated by the morphologically and genetically (only short sequences available) weakly deviating subspecies immaculatella from Northern Italy.
[111] Monochroa arundinetella / M. suffusella. These two morphologically separate species represent one of the few cases of barcode sharing among European Gelechiidae. The author and year of description of M. arundinetella follow
[112] Monochroa nomadella. This genetically highly variable species clusters in four different and geographically separate BINs and is in strong need of revisionary work.
[113] Oxypteryx. Eulamprotes Bradley, 1971 with the type species E. atrella is shown to be a synonym of Oxypteryx Rebel, 1911 (
[114] Oxypteryx nigromaculella. A specimen from Greece clusters into a separate BIN and may represent a different species.
[115] Oxypteryx wilkella. Two specimens from Italy and Hungary respectively are strongly divergent from the large bulk of E. wilkella DNA barcodes and cluster into a separate BIN. The taxonomic status of this cluster requires careful evaluation.
[116] Oxypteryx baldizzonei. Two strongly divergent DNA barcode clusters, representing three BINs, have been considered as intraspecific variation by
[117] Oxypteryx libertinella. The geographic variation of DNA barcode clusters in this genetically highly variable species with eight BINs has been discussed by
[118] Athrips. The sequence of species follows the generic revision by
[119] Athrips rancidella. A specimen from Greece clusters separately into a second BIN (2.86% min. distance) and is in need of taxonomic re-evaluation.
[120] Athrips amoenella. A genetically highly variable species, which clusters into five BINs.
[121] Neofriseria peliella. Two weakly separated BINs (1.44% min. distance) without clear geographical separation most probably reflect intraspecific variation.
[122] Neofriseria hitadoella. A strongly divergent BIN from France with 3.85% min. distance to N. hitadoella from Morocco is considered as a probable cryptic species, but the problem needs to be carefully revised.
[123] Neofriseria kuznetzovae. This species was listed by
[124] Sophronia. Unrevised genus with some doubtful taxa lacking DNA barcodes.
[125] Sophronia semicostella. Two DNA barcode clusters, grouped into two BINs, show no clear geographic separation.
[126] Sophronia consanguinella. S. marginella was recently shown to be a junior synonym of this species (
[127] Sophronia grandii. The DNA barcode of a paratype of Sophronia ascalis Gozmány, 1951 fully agrees with that of S. grandii. The two taxa are virtually identical, and we therefore consider S. ascalis to be a synonym of S. grandii (syn. nov.).
[128] Sophronia chilonella. A single DNA barcode from Bulgaria of a specimen similar to S. chilonella strongly deviates and may represent the taxonomically disputed and unrevised S. acaudella.
[129] Sophronia sicariellus. A single DNA barcode sequence of 504bp from Germany strongly deviates, although it may represent intraspecific variation.
[130] Mirificarma. Several species show a high genetic variation which could indicate cryptic diversity. Therefore, despite available taxonomic revisions by
[131] Mirificarma lentiginosella. Two DNA barcode clusters, which separate into two BINs (1.7% min. distance) without geographic separation.
[132] Mirificarma cytisella. A genetically variable species, separated into four BINs without geographic separation.
[133] Mirificarma monticolella. Two DNA barcode clusters from Italy and Bulgaria are highly divergent and separate into two BINs (4.49% min. distance).
[134] Mirificarma burdonella. Two DNA barcodes from France show a deep split into two BINs (5.78% min. distance) and require taxonomic re-evaluation.
[135] Mirificarma ulicinella. Two DNA barcode clusters from France and Portugal are highly divergent and separate into two BINs (3.37% min. distance).
[136] Aroga velocella. The species splits into three BINs, which show no clear geographic separation. The attribution of authorship follows
[137] Aroga flavicomella. A genetically variable species, which splits into four BINs.
[138] Chionodes. The sequence of species follows the revision by
[139] Chionodes luctuella. DNA barcodes from central and northern Europe cluster into separate BINs (1.87% min. distance) which are currently not confirmed by morphology.
[140] Chionodes fumatella. DNA barcodes from central and northern Europe cluster into three geographically partially separated BINs and need taxonomic re-assessment.
[141] Gelechia. This genus includes at least one additional and probably undescribed species.
[142] Gelechia senticetella. DNA barcodes cluster into two geographically separate BINs with min. distances > 2% to the Nearest Neighbour, and need taxonomic re-assessment.
[143] Gelechia obscuripennis. This disputed taxon has recently been re-considered to be a separate species based on molecular data, morphology and biology (
[144] Agnippe. The genus (as Evippe Chambers, 1873) has traditionally been placed in the Litini. DNA barcodes of two species are not supportive of the systematic position of the genus in that tribe. We therefore follow
[145] Holcophora. The genera Holocophora and Aponoaea have been synonymized recently by
[146] Holcophora inderskella. The species was included in Holcophora by
[147] Gnorimoschema herbichii. Northern European populations of this species cluster into two BINs.
[148] Scrobipalpa. This extraordinary diverse genus still requires some taxonomic re-assessment, reflected by several yet unidentified barcode clusters which at least partly belong to undescribed species.
[149] Scrobipalpa aptatella. Records from Europe (France, Italy, former Yugoslavia) are unconfirmed (
[150] Scrobipalpa amseli / S. hyssopi. Both species clusters into the same BIN but differ in morphology of the male genitalia (
[151] Scrobipalpa reiprichi. Two geographically separate barcodes BINs (2.57% min distance) may reflect cryptic diversity, with altogether four potential species from preliminary morphological analysis (
[152] Scrobipalpa caucasica. Only known from the Caucasus. S. benzengensis (Povolný, 2001) is a junior synonym (
[153] Scrobipalpa pauperella. Some externally different specimens from northern Italy are slightly divergent in their DNA barcodes and may belong to a separate species.
[154] Scrobipalpa mercantourica. This species clusters together with Scrobipalpa arenbergeri but according to the original description differs in morphology. A taxonomic re-assessment seems advisable to fix the status of the taxon.
[155] Scrobipalpa alterna / S. lutea. Both species share barcodes and are virtually indistinguishable in genitalia characters, although the ground colour of the forewings is usually distinct with rare intermediates. A re-assessment of this group is in preparation (Bidzilya in litt.).
[156] Scrobipalpa artemisiella / Scrobipalpa stangei. These two species are clearly separated by their biology and female genitalia morphology, but share one barcode BIN. A second BIN of S. artemisiella based on a single sequence most probably reflects intraspecific variation.
[157] Scrobipalpa bryophiloides. A genetically variable species which clusters into two separate BINs and requires further evaluation.
[158] Scrobipalpa ocellatella. DNA barcodes of this species clusters into two weakly separated BINs (1.44% min distance), most probably reflecting intraspecific variation.
[159] Scrobipalpa salinella / S. salicorniae / S. spergulariella. Although these species show diagnostic morphology (
[160] Scrobipalpa halymella / S. stabilis. Both species cluster into the same BIN but differ weakly in morphology (
[161] Scrobipalpula. All five successfully sequenced species share BINs, but still show species-specific DNA barcode haplotypes.
[162] Keiferia lycopersicella. An American species introduced to Europe in 2008 which apparently has not established permanent populations (
[163] Ephysteris promptella. A genetically highly variable species clustering into four BINs. A taxonomic re-evaluation of this problem is necessary.
[164] Ephysteris diminutella. Two strongly divergent and geographically separate DNA barcode clusters reflected by two BINs (5.94% min. distance) require taxonomic revision.
[165] Ephysteris inustella. The year of description follows
[166] Ochrodia. An unidentified species from Greece (Crete) clusters with specimens from Saudia Arabia. The genus is in need of revision.
[167] Lutilabria lutilabrella. DNA barcodes from Slovenia and Slovakia cluster into separate BINs (3.41% min. distance) and need revisionary work.
[168] Klimeschiopsis kiningerella. Specimens from northern Italy cluster into a BIN separate from all other samples from various parts of Europe.
[169] Caryocolum. Despite extensive past revisionary work on this genus, it still includes a remarkable amount of unresolved taxonomic problems with several potential cryptic species.
[170] Caryocolum tischeriella. DNA barcodes cluster into three BINs without geographic separation.
[171] Caryocolum alsinella. A genetically highly variable species with strongly divergent DNA barcode clusters separated into three BINs. A thorough taxonomic re-assessment seems necessary.
[172] Caryocolum vicinella. DNA barcodes cluster into four BINs. A thorough taxonomic re-assessment seems necessary.
[173] Carayocolum amaurella. This genetically highly variable species clusters into five BINs, but alleged cryptic diversity is not supported by morphology (
[174] Caryocolum saginella. DNA barcode sequences with two BINs (5.46% min. distance). clearly support the existence of a separate species in the SW-Alps (Huemer in prep.).
[175] Caryocolum cauligenella. A single specimen from Spain strongly deviates in DNA barcode with a separate BIN and C. saginella instead of C. cauligenella as Nearest Neighbor (5.46% min. distance). However, the specimen clearly matches the latter in phenotypy and needs taxonomic re-assessment.
[176] Caryocolum peregrinella. This species splits into three highly divergent allopatric clusters which most probably represent different species (Huemer in prep.). One of the major problems in resolving the taxonomic mismatches is the status of the holotype of C. peregrinella, a female without an abdomen and unknown type-locality, stated as Europe (
[177] Caryocolum leucomelanella. Two DNA barcode clusters with separate BINs (2.73% min. distance) show no geographic pattern.
[178] Caryocolum schleichi / C. arenariella. Initially described as different species, the largely allopatric taxa of this group have been merged into a single species by
[179] Caryocolum marmorea spp. mediocorsa agrees in DNA barcode with the nominotypical subspecies.
[180] Caryocolum pullatella. This species shows an extraordinary genetic variation across its holarctic range (
[181] Caryocolum klosi. A single DNA barcode from the French Pyrenees is highly divergent with a separate BIN (4.17% min. distance) and may represent a different species.
[182] Caryocolum blandella. Lita signatella was described from an unstated number of specimens from Kazan in Russia (“provincia Casanensi”) (
Whereas the name Caryocolum blandella has been universally in use for a widespread European species since
[183] Caryocolum horoscopa. Initially described as a species, this taxon was recently considered to be a subspecies of Caryocolum blandella (
[184] Caryocolum fibigerium.
[185] Caryocolum junctella. Two barcode clusters with separate BINs show no clear geographic separation.
[186] Agonochaetia terrestrella. Specimens from Switzerland and Romania cluster into a separate BIN, but are considered as conspecific (
[187] Sattleria. The sequence of species follows
[188] Sattleria melaleucella. The species shares BINs with one cluster of the morphologically different S. pyrenaica, indicating occasional introgression.
[189] Sattleria pyrenaica. A genetically variable species with five different BINs, one shared with Sattleria melaleucella. The species requires taxonomic re-assessment.
[190] Litini.
[191] Schneidereria pistaciella Weber, 1957. The systematic placement of this genus and species follows
[192] Teleiodes vulgella / T. italica / T. brevivalva. These three species differ strongly in the male genitalia but share DNA barcodes.
[193] Teleiodes saltuum / T. kaitilai. Both species are closely related, mainly differing in the structures of the female genitalia. In DNA barcodes T. saltuum clusters into two BINs and T. kaitilai in a separate BIN.
[194] Teleiodes luculella. A genetically variable species, which clusters into three BINs. A thorough evaluation of this problem is necessary.
[195] Teleiodes flavimaculella. A genetically highly variable species, which clusters into three BINs. A re-evaluation of this problem is necessary.
[196] Pseudotelphusa tessella. Two weakly separated BINs (1.61% min. distance) without clear geographic separation are considered as intraspecific variation.
[197] Teleiopsis diffinis / T. bagriotella / T. albifemorella / T. paulheberti / T. rosalbella. These closely related species differ in morphology whereas barcodes give a more complex pattern. Genetic differences are generally weak with partial BIN sharing (i.e., T. rosalbella / T. albifemorella) and/or likely introgression in some taxa, while high intraspecific variation - with two BINs in three species - indicates possible further cryptic diversity.
[198] Xenolechia. Species in this genus share DNA barcodes but differ in morphology (
[199] Altenia scriptella. Two BINs without clear geographic separation are considered as intraspecific variation.
[200] Exoteleia dodecella. The taxonomy of dark specimens in this group, mainly observed in Central Europe, is disputed, though usually these are considered as infrasubspecific variation (
[201] Parachronistis albiceps. Genetically variable species, which clusters into four BINs without clear geographic separation.
[202] “Telphusa”. The placement of cistiflorella Constant, 1890 in the genus Telphusa follows
We are most grateful to Oleksiy Bidzilya (Kiev, Ukraine), Klaus Sattler (London, U.K.), and Jan Šumpich (Prague, Czech Republic) for various comments and support with unpublished observations. Jacques Nel (La Ciotat, France) and Thierry Varenne (Nice, France) supported our work with important material, Michel Billard (Saint Alban Leysse, France) with an excellent picture of Sattleria, and Roman Unterasinger helped with construction of the map. Tomas Pape (ZMUC) kindly commented on a relevant case, Vazrick Nazari (Ottawa, Canada) and particularly Mark Metz (Washington, U.S.A.) thoroughly reviewed the manuscript. Robert J. Heckford (Plympton, U.K.) is thanked for his careful language proofreading with valuable comments.
Selected references that have used the name Caryocolum blandella in the last 50 years
Data type: references